351:
903:
2488:
885:
2493:
138:
419:
965:
608:
1202:
1090:
942:
1155:
120:
2516:
794:
700:
670:
620:
1305:
561:
547:
1119:
1282:
849:
577:
is a nearly complete juvenile skeleton intertwined with three other individuals. Notably, three skulls and a good amount of arm material are known. The known manus has only two functional fingers, with the third being significantly reduced. Such a morphology is also seen in a wide variety of
438:
by professionals. Though the exact localities of these specimens are currently unknown due to their theft, although the colour of their sediments and geochemical analyses confirm their provenance from the Nemegt
Formation.
1715:
Owocki, K.; Kremer, B.; Cotte, M.; Bocherens, H. (2020). "Diet preferences and climate inferred from oxygen and carbon isotopes of tooth enamel of
Tarbosaurus bataar (Nemegt Formation, Upper Cretaceous, Mongolia)".
400:. Another specimen in this block is represented by MPC-D 102/110b, a large quadrate and quadratojugal. The third specimen in this assemblage is MPC-D 102/110c, a juvenile postcranial skeleton. Both specimens share
641:
represents a highly derived form in a series of oviraptorosaurs whose third digits have reduced over evolutionary time. Their cladogram is shown below (only relationships within
Oviraptoridae are shown):
2545:
1777:
427:
2555:
508:, is taken from the Mongolian word аварсан (avarsan, meaning rescued) given that specimens of the genus were redeemed from fossil poachers.
430:
along with a fourth specimen, MPC-D 102/11, representing a partial juvenile skeleton preserving the skull. They were later returned to the
415:
and the
Mongolian Palaeontological Centre. It was catalogued as MPC-D 102/12 and represents postcranial skeleton of an adult individual.
2509:
369:, MPC-D 100/33, was discovered in 1974 by the Soviet-Mongolian Palaeontological Expedition at the Bügiin Tsav locality in the
2487:
1770:
454:
381:
specimen is a block of an assemblage of three individuals buried in life positions that has been labelled as MPC-D 102/110.
412:
1637:
Chinzorig, T.; Kobayashi, Y.; Tsogtbaatar, K.; Currie, P. J.; Takasaki, R.; Tanaka, T.; Iijima, M.; Barsbold, R. (2018).
2550:
431:
2492:
1763:
497:
2519:
1874:
137:
411:
Another, isolated specimen was collected in 1998 at the
Guriliin Tsav locality by a joint expedition from the
533:
is unequivocally known from at least two localities of the Nemegt
Formation: Bügiin Tsav and Guriliin Tsav.
501:
469:
465:
350:
384:
The first individual, subcatalogued as MPC-D 102/110a, is a partial skeleton of a juvenile preserving the
2560:
2540:
2159:
902:
461:
408:
and preservation state, which may suggest that MPC-D 102/11 is part of the holotype block assemblage.
1928:
1725:
1689:
1650:
1639:"Ornithomimosaurs from the Nemegt Formation of Mongolia: manus morphological variation and diversity"
1591:
1532:
1519:
Funston, G. F.; Tsogtbaatar, C.; Tsogtbaatar, K.; Kobayashi, Y.; Sullivan, C.; Currie, P. J. (2020).
579:
477:
473:
259:
2050:
362:
884:
1901:
1741:
1580:"An Unusual New Theropod with a Didactyl Manus from the Upper Cretaceous of Patagonia, Argentina"
1195:
435:
389:
355:
132:
2078:
2043:
1619:
1560:
2263:
1996:
1974:
1940:
1786:
1733:
1697:
1658:
1609:
1599:
1550:
1540:
1468:
1436:
481:
418:
397:
370:
326:
964:
2188:
2101:
2085:
1859:
1460:
1456:
1201:
1089:
941:
607:
318:
30:
493:
306:
1729:
1693:
1654:
1595:
1536:
1154:
2442:
2249:
2226:
2111:
2036:
2021:
1614:
1579:
1555:
1520:
1464:
1113:
458:
793:
699:
669:
119:
2534:
2428:
2406:
2347:
2331:
2278:
2152:
2138:
2062:
1745:
1677:
1638:
1452:
1444:
1275:
959:
843:
649:
634:
629:
The describers' phylogenetic analysis places it as a derived member of the subfamily
596:
591:
586:
582:
221:
70:
2399:
2355:
2313:
2256:
2199:
2173:
1843:
1676:
Funston, G. F.; Mendonca, S. E.; Currie, P. J.; Barsbold, R.; Barsbold, R. (2018).
1084:
935:
810:
743:
447:
401:
334:
267:
1304:
619:
1737:
1701:
1662:
1604:
2449:
2382:
2320:
2242:
2235:
2071:
1913:
1472:
1443:
lived alongside other theropods, such as other oviraptorosaurs, tyrannosaurids,
1149:
1022:
723:
630:
311:
234:
45:
560:
2463:
2414:
2291:
2215:
2145:
2131:
2124:
1825:
1816:
1521:"A new two-fingered dinosaur sheds light on the radiation of Oviraptorosauria"
1484:
1476:
1448:
1036:
757:
526:
500:, in reference to the holotypic block assemblage preserving three skulls. The
195:
90:
55:
2421:
2370:
2363:
2298:
2166:
2028:
1886:
1834:
1480:
1178:
1118:
788:
694:
663:
517:
405:
208:
149:
95:
39:
1623:
1564:
1281:
848:
546:
2340:
2305:
2180:
2014:
1810:
879:
623:
574:
512:
378:
374:
330:
314:
182:
169:
85:
80:
65:
60:
50:
1545:
2208:
100:
75:
1804:
1488:
393:
159:
377:. It is represented by a partial subadult postcranial skeleton. The
1755:
1678:"Oviraptorosaur anatomy, diversity and ecology in the Nemegt Basin"
2435:
2391:
1251:
1060:
618:
606:
485:
443:
417:
385:
349:
434:
on
December 14 during the same year, where they were posteriorly
585:, however it can also be seen in other coelurosaurs such as the
322:
1797:
1759:
521:, this specimen shares several traits with other specimens of
127:
Holotype block assemblage (top) and line drawing (bottom)
426:
This block was in 2006 confiscated from poachers by the
1439:, one of the most productive fossil sites in Mongolia.
511:
Although MPC-D 100/33 has been largely referred as a
442:
All of the specimens were later assigned to the new
274:
2381:
2330:
2277:
2225:
2198:
2110:
2099:
2061:
1994:
1939:
1912:
1885:
1858:
310:, a mythical bird of Yakut folklore) is a genus of
496:, one of the multiple triple-headed eagles in the
1718:Palaeogeography, Palaeoclimatology, Palaeoecology
1682:Palaeogeography, Palaeoclimatology, Palaeoecology
1643:Palaeogeography, Palaeoclimatology, Palaeoecology
1479:are also known from the formation. Fossils of a
1771:
8:
2283:
2116:
2107:
2006:
2002:
1991:
1918:
1891:
1864:
1855:
1794:
1778:
1764:
1756:
118:
20:
1613:
1603:
1554:
1544:
1514:
1512:
1510:
1508:
1506:
1504:
1500:
428:General Intelligence Agency of Mongolia
615:, showing the two-fingered hand in (d)
413:Hayashibara Museum of Natural Sciences
7:
1267:
1243:
1236:
1170:
1141:
1134:
1105:
1076:
1052:
1028:
927:
871:
864:
835:
780:
773:
749:
739:
715:
686:
655:
645:
354:Holotype block assemblage prior to
14:
2546:Late Cretaceous dinosaurs of Asia
432:Mongolian Palaeontological Center
2515:
2514:
2491:
2486:
1303:
1280:
1200:
1153:
1117:
1088:
963:
940:
901:
883:
847:
792:
698:
668:
559:
545:
422:Skull of holotype MPC-D 102/110a
404:traits, such as the death pose,
325:, that lived in what is now the
136:
1578:Apesteguía, Sebastián (2016).
1:
2556:Fossil taxa described in 2020
637:. It has been suggested that
1738:10.1016/j.palaeo.2019.05.012
1702:10.1016/j.palaeo.2017.10.023
1663:10.1016/j.palaeo.2017.10.031
1605:10.1371/journal.pone.0157793
566:Body size of three specimens
552:Size comparison with a human
16:Extinct species of dinosaur
2577:
1525:Royal Society Open Science
525:including a reduced third
2504:
2484:
2286:
2119:
2009:
2005:
1990:
1971:
1921:
1894:
1867:
1854:
1793:
1296:
1272:
1265:
1248:
1241:
1234:
1192:
1175:
1168:
1146:
1139:
1132:
1110:
1103:
1081:
1074:
1057:
1050:
1033:
1026:
956:
932:
925:
900:Zamyn Khondt oviraptorid
898:
876:
869:
862:
840:
833:
807:
785:
778:
771:
754:
747:
737:
720:
713:
691:
684:
660:
653:
273:
266:
133:Scientific classification
131:
126:
117:
23:
578:theropods such as a few
626:
616:
470:Khishigjav Tsogtbaatar
423:
358:
622:
610:
466:Tsogtbaatar Chinzorig
421:
353:
580:carcharodontosaurian
474:Yoshitsugu Kobayashi
346:History of discovery
2551:Cretaceous Mongolia
2051:Scansoriopterygidae
1730:2020PPP...537j9190O
1694:2018PPP...494..101F
1655:2018PPP...494...91C
1596:2016PLoSO..1157793A
1546:10.1098/rsos.201184
1537:2020RSOS....701184F
1463:, and early birds.
1491:are also present.
1196:Heyuannia yanshini
627:
617:
611:Forelimb bones of
498:Altaic mythologies
492:, is derived from
462:Gregory F. Funston
424:
390:cervical vertebrae
359:
356:fossil preparation
333:. It includes the
2528:
2527:
2500:
2499:
2482:
2481:
2478:
2477:
2474:
2473:
2273:
2272:
2095:
2094:
2079:Similicaudipteryx
2044:Protarchaeopteryx
1986:
1985:
1982:
1981:
1967:
1966:
1963:
1962:
1959:
1958:
1469:pachycephalosaurs
1435:was found in the
1424:
1423:
1415:
1414:
1406:
1405:
1397:
1396:
1388:
1387:
1379:
1378:
1370:
1369:
1361:
1360:
1352:
1351:
1343:
1342:
1334:
1333:
1325:
1324:
1316:
1315:
1223:
1222:
1214:
1213:
1012:
1011:
1003:
1002:
994:
993:
985:
984:
976:
975:
914:
913:
822:
821:
599:oviraptorosaurs.
396:, hindlimbs, and
293:
292:
262:
2568:
2518:
2517:
2495:
2490:
2284:
2264:Epichirostenotes
2117:
2108:
2007:
2003:
1997:Oviraptorosauria
1992:
1975:Oviraptorosauria
1941:Oviraptorosauria
1919:
1892:
1865:
1856:
1849:
1848:
1795:
1787:Oviraptorosauria
1780:
1773:
1766:
1757:
1750:
1749:
1712:
1706:
1705:
1673:
1667:
1666:
1634:
1628:
1627:
1617:
1607:
1575:
1569:
1568:
1558:
1548:
1516:
1461:ornithomimosaurs
1457:therizinosaurids
1437:Nemegt Formation
1428:Paleoenvironment
1307:
1285:
1284:
1268:
1244:
1237:
1205:
1204:
1179:Heyuannia huangi
1171:
1157:
1142:
1135:
1121:
1106:
1092:
1077:
1053:
1029:
967:
945:
944:
928:
905:
887:
872:
865:
851:
836:
796:
781:
774:
750:
740:
716:
702:
687:
673:
672:
656:
646:
624:Life restoration
563:
549:
482:Philip J. Currie
398:caudal vertebrae
371:Nemegt Formation
327:Nemegt Formation
276:
253:
246:
233:
220:
141:
140:
122:
112:
42:
29:Temporal range:
21:
2576:
2575:
2571:
2570:
2569:
2567:
2566:
2565:
2531:
2530:
2529:
2524:
2496:
2470:
2377:
2326:
2269:
2221:
2194:
2189:Ojoraptorsaurus
2104:
2102:Caenagnathoidea
2091:
2086:Xingtianosaurus
2057:
1999:
1978:
1955:
1935:
1908:
1881:
1875:Avemetatarsalia
1860:Avemetatarsalia
1850:
1800:
1799:
1789:
1784:
1754:
1753:
1714:
1713:
1709:
1675:
1674:
1670:
1636:
1635:
1631:
1590:(7): e0157793.
1577:
1576:
1572:
1518:
1517:
1502:
1497:
1430:
1425:
1416:
1407:
1398:
1389:
1380:
1371:
1362:
1353:
1344:
1335:
1326:
1317:
1279:
1224:
1215:
1199:
1013:
1004:
995:
986:
977:
939:
915:
823:
667:
605:
571:
570:
569:
568:
567:
564:
555:
554:
553:
550:
539:
478:Corwin Sullivan
459:paleontologists
348:
319:Late Cretaceous
289:
282:
252:
244:
231:
218:
135:
113:
111:
110:
109:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
37:
36:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
2574:
2572:
2564:
2563:
2558:
2553:
2548:
2543:
2533:
2532:
2526:
2525:
2523:
2522:
2512:
2505:
2502:
2501:
2498:
2497:
2485:
2483:
2480:
2479:
2476:
2475:
2472:
2471:
2469:
2468:
2460:
2453:
2446:
2443:Machairasaurus
2439:
2432:
2425:
2418:
2411:
2403:
2396:
2387:
2385:
2379:
2378:
2376:
2375:
2367:
2360:
2352:
2344:
2336:
2334:
2328:
2327:
2325:
2324:
2317:
2310:
2302:
2295:
2287:
2281:
2275:
2274:
2271:
2270:
2268:
2267:
2260:
2253:
2250:Caenagnathasia
2246:
2239:
2231:
2229:
2227:Caenagnathinae
2223:
2222:
2220:
2219:
2212:
2204:
2202:
2196:
2195:
2193:
2192:
2185:
2177:
2170:
2163:
2156:
2149:
2142:
2135:
2128:
2120:
2114:
2112:Caenagnathidae
2105:
2100:
2097:
2096:
2093:
2092:
2090:
2089:
2082:
2075:
2067:
2065:
2059:
2058:
2056:
2055:
2047:
2040:
2037:Ningyuansaurus
2033:
2025:
2022:Incisivosaurus
2018:
2010:
2000:
1995:
1988:
1987:
1984:
1983:
1980:
1979:
1972:
1969:
1968:
1965:
1964:
1961:
1960:
1957:
1956:
1954:
1953:
1952:
1951:
1945:
1943:
1937:
1936:
1934:
1933:
1932:
1931:
1922:
1916:
1910:
1909:
1907:
1906:
1905:
1904:
1895:
1889:
1883:
1882:
1880:
1879:
1878:
1877:
1868:
1862:
1852:
1851:
1847:
1846:
1837:
1828:
1819:
1813:
1807:
1798:
1791:
1790:
1785:
1783:
1782:
1775:
1768:
1760:
1752:
1751:
1707:
1668:
1629:
1570:
1531:(10): 201184.
1499:
1498:
1496:
1493:
1453:alvarezsaurids
1445:dromaeosaurids
1429:
1426:
1422:
1421:
1418:
1417:
1413:
1412:
1409:
1408:
1404:
1403:
1400:
1399:
1395:
1394:
1391:
1390:
1386:
1385:
1382:
1381:
1377:
1376:
1373:
1372:
1368:
1367:
1364:
1363:
1359:
1358:
1355:
1354:
1350:
1349:
1346:
1345:
1341:
1340:
1337:
1336:
1332:
1331:
1328:
1327:
1323:
1322:
1319:
1318:
1314:
1313:
1310:
1309:
1295:
1292:
1291:
1288:
1287:
1271:
1266:
1264:
1261:
1260:
1257:
1256:
1247:
1242:
1240:
1235:
1233:
1230:
1229:
1226:
1225:
1221:
1220:
1217:
1216:
1212:
1211:
1208:
1207:
1191:
1188:
1187:
1184:
1183:
1174:
1169:
1167:
1164:
1163:
1160:
1159:
1145:
1140:
1138:
1133:
1131:
1128:
1127:
1124:
1123:
1114:Machairasaurus
1109:
1104:
1102:
1099:
1098:
1095:
1094:
1080:
1075:
1073:
1070:
1069:
1066:
1065:
1056:
1051:
1049:
1046:
1045:
1042:
1041:
1032:
1027:
1025:
1019:
1018:
1015:
1014:
1010:
1009:
1006:
1005:
1001:
1000:
997:
996:
992:
991:
988:
987:
983:
982:
979:
978:
974:
973:
970:
969:
955:
952:
951:
948:
947:
931:
926:
924:
921:
920:
917:
916:
912:
911:
908:
907:
897:
894:
893:
890:
889:
875:
870:
868:
863:
861:
858:
857:
854:
853:
839:
834:
832:
829:
828:
825:
824:
820:
819:
816:
815:
806:
803:
802:
799:
798:
784:
779:
777:
772:
770:
767:
766:
763:
762:
753:
748:
746:
738:
736:
733:
732:
729:
728:
719:
714:
712:
709:
708:
705:
704:
690:
685:
683:
680:
679:
676:
675:
659:
654:
652:
644:
633:in the family
604:
603:Classification
601:
589:, the avialan
587:tyrannosaurids
565:
558:
557:
556:
551:
544:
543:
542:
541:
540:
538:
535:
451:Oksoko avarsan
347:
344:
339:Oksoko avarsan
291:
290:
283:
279:Oksoko avarsan
271:
270:
264:
263:
242:
238:
237:
229:
225:
224:
216:
212:
211:
206:
199:
198:
193:
186:
185:
180:
173:
172:
167:
163:
162:
157:
153:
152:
147:
143:
142:
129:
128:
124:
123:
115:
114:
106:
105:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
43:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2573:
2562:
2559:
2557:
2554:
2552:
2549:
2547:
2544:
2542:
2539:
2538:
2536:
2521:
2513:
2511:
2507:
2506:
2503:
2494:
2489:
2466:
2465:
2461:
2459:
2458:
2454:
2452:
2451:
2447:
2445:
2444:
2440:
2438:
2437:
2433:
2431:
2430:
2429:Jiangxisaurus
2426:
2424:
2423:
2419:
2417:
2416:
2412:
2409:
2408:
2407:Ganzhousaurus
2404:
2402:
2401:
2397:
2394:
2393:
2389:
2388:
2386:
2384:
2380:
2373:
2372:
2368:
2366:
2365:
2361:
2358:
2357:
2353:
2350:
2349:
2348:Corythoraptor
2345:
2343:
2342:
2338:
2337:
2335:
2333:
2332:Oviraptorinae
2329:
2323:
2322:
2318:
2316:
2315:
2311:
2308:
2307:
2303:
2301:
2300:
2296:
2294:
2293:
2289:
2288:
2285:
2282:
2280:
2279:Oviraptoridae
2276:
2266:
2265:
2261:
2259:
2258:
2254:
2252:
2251:
2247:
2245:
2244:
2240:
2238:
2237:
2233:
2232:
2230:
2228:
2224:
2218:
2217:
2213:
2211:
2210:
2206:
2205:
2203:
2201:
2197:
2191:
2190:
2186:
2183:
2182:
2178:
2176:
2175:
2171:
2169:
2168:
2164:
2162:
2161:
2160:Leptorhynchos
2157:
2155:
2154:
2153:Gigantoraptor
2150:
2148:
2147:
2143:
2141:
2140:
2139:Chirostenotes
2136:
2134:
2133:
2129:
2127:
2126:
2122:
2121:
2118:
2115:
2113:
2109:
2106:
2103:
2098:
2088:
2087:
2083:
2081:
2080:
2076:
2074:
2073:
2069:
2068:
2066:
2064:
2063:Caudipteridae
2060:
2053:
2052:
2048:
2046:
2045:
2041:
2039:
2038:
2034:
2031:
2030:
2026:
2024:
2023:
2019:
2017:
2016:
2012:
2011:
2008:
2004:
2001:
1998:
1993:
1989:
1977:
1976:
1970:
1949:
1948:
1947:
1946:
1944:
1942:
1938:
1930:
1926:
1925:
1924:
1923:
1920:
1917:
1915:
1911:
1903:
1899:
1898:
1897:
1896:
1893:
1890:
1888:
1884:
1876:
1872:
1871:
1870:
1869:
1866:
1863:
1861:
1857:
1853:
1845:
1841:
1838:
1836:
1832:
1829:
1827:
1823:
1820:
1818:
1814:
1812:
1808:
1806:
1802:
1801:
1796:
1792:
1788:
1781:
1776:
1774:
1769:
1767:
1762:
1761:
1758:
1747:
1743:
1739:
1735:
1731:
1727:
1723:
1719:
1711:
1708:
1703:
1699:
1695:
1691:
1687:
1683:
1679:
1672:
1669:
1664:
1660:
1656:
1652:
1648:
1644:
1640:
1633:
1630:
1625:
1621:
1616:
1611:
1606:
1601:
1597:
1593:
1589:
1585:
1581:
1574:
1571:
1566:
1562:
1557:
1552:
1547:
1542:
1538:
1534:
1530:
1526:
1522:
1515:
1513:
1511:
1509:
1507:
1505:
1501:
1494:
1492:
1490:
1486:
1482:
1478:
1474:
1470:
1466:
1465:Hadrosauroids
1462:
1458:
1454:
1450:
1446:
1442:
1438:
1434:
1427:
1420:
1419:
1411:
1410:
1402:
1401:
1393:
1392:
1384:
1383:
1375:
1374:
1366:
1365:
1357:
1356:
1348:
1347:
1339:
1338:
1330:
1329:
1321:
1320:
1312:
1311:
1308:
1306:
1302:
1301:
1294:
1293:
1290:
1289:
1286:
1283:
1278:
1277:
1276:Jiangxisaurus
1270:
1269:
1263:
1262:
1259:
1258:
1255:
1254:
1253:
1246:
1245:
1239:
1238:
1232:
1231:
1228:
1227:
1219:
1218:
1210:
1209:
1206:
1203:
1198:
1197:
1190:
1189:
1186:
1185:
1182:
1181:
1180:
1173:
1172:
1166:
1165:
1162:
1161:
1158:
1156:
1152:
1151:
1144:
1143:
1137:
1136:
1130:
1129:
1126:
1125:
1122:
1120:
1116:
1115:
1108:
1107:
1101:
1100:
1097:
1096:
1093:
1091:
1087:
1086:
1079:
1078:
1072:
1071:
1068:
1067:
1064:
1063:
1062:
1055:
1054:
1048:
1047:
1044:
1043:
1040:
1039:
1038:
1031:
1030:
1024:
1021:
1020:
1017:
1016:
1008:
1007:
999:
998:
990:
989:
981:
980:
972:
971:
968:
966:
962:
961:
960:Corythoraptor
954:
953:
950:
949:
946:
943:
938:
937:
930:
929:
923:
922:
919:
918:
910:
909:
906:
904:
896:
895:
892:
891:
888:
886:
882:
881:
874:
873:
867:
866:
860:
859:
856:
855:
852:
850:
846:
845:
844:Ganzhousaurus
838:
837:
831:
830:
827:
826:
818:
817:
814:
813:
812:
805:
804:
801:
800:
797:
795:
791:
790:
783:
782:
776:
775:
769:
768:
765:
764:
761:
760:
759:
752:
751:
745:
742:
741:
735:
734:
731:
730:
727:
726:
725:
718:
717:
711:
710:
707:
706:
703:
701:
697:
696:
689:
688:
682:
681:
678:
677:
674:
671:
666:
665:
658:
657:
651:
650:Oviraptoridae
648:
647:
643:
640:
636:
635:Oviraptoridae
632:
625:
621:
614:
609:
602:
600:
598:
594:
593:
592:Balaur bondoc
588:
584:
581:
576:
562:
548:
536:
534:
532:
528:
524:
520:
519:
514:
509:
507:
503:
502:specific name
499:
495:
491:
487:
483:
479:
475:
471:
467:
463:
460:
456:
452:
449:
445:
440:
437:
433:
429:
420:
416:
414:
409:
407:
403:
399:
395:
391:
387:
382:
380:
376:
372:
368:
364:
357:
352:
345:
343:
341:
340:
336:
332:
328:
324:
320:
316:
313:
309:
308:
303:
299:
298:
287:
281:
280:
272:
269:
265:
261:
257:
251:
250:
243:
240:
239:
236:
230:
227:
226:
223:
222:Oviraptoridae
217:
214:
213:
210:
207:
204:
201:
200:
197:
194:
191:
188:
187:
184:
181:
178:
175:
174:
171:
168:
165:
164:
161:
158:
155:
154:
151:
148:
145:
144:
139:
134:
130:
125:
121:
116:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
41:
32:
26:
22:
19:
2561:Nemegt fauna
2541:Oviraptorids
2462:
2456:
2455:
2448:
2441:
2434:
2427:
2420:
2413:
2405:
2400:Conchoraptor
2398:
2390:
2369:
2362:
2356:Huanansaurus
2354:
2346:
2339:
2319:
2314:Tongtianlong
2312:
2304:
2297:
2290:
2262:
2257:Caenagnathus
2255:
2248:
2241:
2234:
2214:
2207:
2200:Elmisaurinae
2187:
2179:
2174:Microvenator
2172:
2165:
2158:
2151:
2144:
2137:
2130:
2123:
2084:
2077:
2070:
2049:
2042:
2035:
2027:
2020:
2013:
1973:
1844:Pennaraptora
1839:
1830:
1821:
1721:
1717:
1710:
1685:
1681:
1671:
1646:
1642:
1632:
1587:
1583:
1573:
1528:
1524:
1487:, and small
1485:crocodilians
1440:
1432:
1431:
1299:
1298:
1297:
1274:
1273:
1250:
1249:
1194:
1193:
1177:
1176:
1148:
1147:
1112:
1111:
1085:Conchoraptor
1083:
1082:
1059:
1058:
1035:
1034:
958:
957:
936:Huanansaurus
934:
933:
899:
878:
877:
842:
841:
811:Tongtianlong
809:
808:
787:
786:
756:
755:
744:Citipatiinae
722:
721:
693:
692:
662:
661:
638:
628:
612:
590:
583:allosauroids
572:
530:
522:
516:
510:
505:
489:
486:generic name
450:
448:type species
441:
425:
410:
402:taphonomical
383:
366:
360:
338:
337:
335:type species
305:
301:
296:
295:
294:
285:
278:
277:
268:Type species
255:
248:
247:
202:
189:
176:
24:
18:
2450:Nemegtomaia
2383:Heyuanninae
2243:Apatoraptor
2072:Caudipteryx
1929:Maniraptora
1914:Maniraptora
1688:: 101−120.
1477:titanosaurs
1473:ankylosaurs
1449:troodontids
1150:Nemegtomaia
1023:Heyuanninae
631:Heyuanninae
597:caudipterid
537:Description
529:. To date,
453:, named in
312:oviraptorid
302:Oak-soak-oh
235:Heyuanninae
228:Subfamily:
2535:Categories
2508:See also:
2464:Shixinggia
2415:Gobiraptor
2292:Luoyanggia
2216:Elmisaurus
2146:Eoneophron
2132:Beibeilong
2125:Anomalipes
1950:see below↓
1826:Dinosauria
1817:Sauropsida
1724:: 109190.
1649:: 91−100.
1495:References
1037:Shixinggia
758:Wulatelong
527:metacarpal
506:O. avarsan
361:The first
196:Saurischia
183:Dinosauria
2422:Heyuannia
2371:Rinchenia
2364:Oviraptor
2299:Nankangia
2167:Hagryphus
1902:Theropoda
1887:Theropoda
1835:Theropoda
1803:Kingdom:
1746:182937778
1481:pterosaur
789:Rinchenia
695:Oviraptor
664:Nankangia
518:Heyuannia
406:sediments
317:from the
209:Theropoda
156:Kingdom:
150:Eukaryota
2520:Category
2510:Timeline
2341:Citipati
2306:Nomingia
2181:Nomingia
2015:Avimimus
1811:Chordata
1809:Phylum:
1805:Animalia
1624:27410683
1584:PLOS ONE
1565:33204472
880:Citipati
575:holotype
513:paratype
436:prepared
379:holotype
375:Mongolia
363:specimen
331:Mongolia
315:dinosaur
284:Funston
254:Funston
215:Family:
170:Chordata
166:Phylum:
160:Animalia
146:Domain:
38:70
2209:Citipes
1815:Class:
1726:Bibcode
1690:Bibcode
1651:Bibcode
1615:4943716
1592:Bibcode
1556:7657903
1533:Bibcode
1489:mammals
457:by the
304:; from
241:Genus:
107:↓
2457:Oksoko
2321:Yulong
1744:
1622:
1612:
1563:
1553:
1475:, and
1441:Oksoko
1433:Oksoko
1300:Oksoko
724:Yulong
639:Oksoko
613:Oksoko
595:, and
531:Oksoko
523:Oksoko
494:Öksökö
490:Oksoko
484:. The
394:pelvis
367:Oksoko
307:Öksökö
297:Oksoko
288:, 2020
286:et al.
256:et al.
249:Oksoko
25:Oksoko
2436:Khaan
2392:Banji
1840:Clade
1831:Clade
1822:Clade
1742:S2CID
1252:Banji
1061:Khaan
444:genus
386:skull
203:Clade
190:Clade
177:Clade
2236:Anzu
1927:see
1900:see
1873:see
1620:PMID
1561:PMID
573:The
480:and
455:2020
446:and
323:Asia
260:2020
46:PreꞒ
2029:Kol
1734:doi
1722:537
1698:doi
1686:494
1659:doi
1647:494
1610:PMC
1600:doi
1551:PMC
1541:doi
515:of
373:of
365:of
329:in
321:of
2537::
1842::
1833::
1824::
1740:.
1732:.
1720:.
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1608:.
1598:.
1588:11
1586:.
1582:.
1559:.
1549:.
1539:.
1527:.
1523:.
1503:^
1483:,
1471:,
1467:,
1459:,
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476:,
472:,
468:,
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342:.
258:,
205::
192::
179::
96:Pg
40:Ma
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1692::
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1535::
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300:(
275:†
245:†
232:†
219:†
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
35:~
33:,
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