406:
422:
124:
442:
394:
Head without prominent posterior protuberances. Labrum with dentate anterior margin. Separate mentum absent. Anterior third of gulamentum not depressed. Pronotal disc with rounded lateral edges; dorsal surface without ridges or protuberances. Sternopleural suture absent. Prosternal process short.
405:
421:
1028:
366:
when disturbed. Its larval stage and many other life details are unknown due to its rarity. Males are typically 14–20 mm in length, while females are 14.4-27.5 mm.
1434:
1306:
1332:
441:
1280:
1319:
104:
1444:
1098:"Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea"
1429:
1404:
1025:
1047:"New fossil species of ommatids (Coleoptera: Archostemata) from the Middle Mesozoic of China illuminating the phylogeny of Ommatidae"
973:"The extant species of the genus Omma Newman and description of Beutelius gen. nov. (Coleoptera: Archostemata: Ommatidae: Ommatinae)"
343:
to
Australia. Three other extant species endemic to Australia that were formerly part of this genus were moved to the separate genus
1424:
1324:
1409:
395:
Punctured explanate elytral epipleura absent. CuA of hind wings forked; wedge cell present. Abdominal ventrites abutting.
1439:
202:
1419:
1241:
1337:
1414:
123:
784:
1194:
847:
658:
1358:
500:
1376:
1267:
1109:
801:
558:
504:
371:
921:"New species of Omma Newman from mid-Cretaceous Burmese amber (Coleoptera, Archostemata, Ommatidae)"
616:
479:
335:
522:(Callovian-Oxfordian), Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China
1399:
1174:
1146:
1008:
950:
876:
864:
813:
678:
642:
627:
600:
578:
515:
375:
231:
118:
1363:
1345:
1311:
1254:
1166:
1127:
1078:
1000:
992:
942:
550:
533:
1350:
1158:
1117:
1068:
1058:
984:
932:
1032:
1113:
1073:
1046:
893:
485:
363:
103:
1393:
1178:
1035:
at Joel Hallan’s
Biology Catalog. Texas A&M University. Retrieved on 11 May 2012.
1012:
972:
954:
825:
698:
432:
310:
61:
554:
469:
211:
1259:
1293:
1226:
1122:
1097:
889:
829:
36:
1232:
1217:
988:
868:
833:
821:
797:
694:
358:
330:
81:
46:
1170:
1131:
1063:
996:
946:
1246:
872:
748:
612:
345:
302:
274:
185:
155:
135:
86:
1082:
1004:
1162:
1211:
937:
920:
860:
390:
is distinguished from other ommatid beetles by the following characters:
326:
318:
76:
71:
56:
51:
41:
1285:
465:
340:
321:), over 200 million years ago, though the placement of this species in
91:
66:
1298:
971:
Escalona, Hermes E.; Lawrence, John F.; Ślipiński, Adam (2020-01-24).
325:
has been questioned. Numerous other fossil species are known from the
690:
654:
298:
175:
165:
145:
1188:
1371:
1145:
Jarzembowski, Edmund A.; Zheng, Daran; Zhao, Xianye (2021-11-12).
475:
Taxa labelled (?) are considered questionable by
Kirejtshuk, 2020
294:
1272:
415:
dating to the
Triassic-Jurassic boundary ~ 200 million years ago
1192:
1045:
Tan, Jingjing; Wang, Yongjie; Ren, Dong; Yang, Xingke (2012).
1147:"Is the beetle Omma (Insecta: Coleoptera) a living fossil?"
919:
Li, Yan-Da; Huang, Di-Ying; Cai, Chen-Yang (2021-11-30).
875:, Binton, Warwickshire, Lower Jurassic (Sinemurian),
356:
is strongly associated with wood, being found under
1201:
435:of Myanmar, dating to around 100 million years ago
1151:Geological Society, London, Special Publications
451:from the Mid-Cretaceous Burmese amber of Myanmar
317:, lived during the final stage of the Triassic (
411:Holotype (A,B) and paratype (C,D) specimens of
392:
333:of Europe and Asia. The only living species is
867:, Brown's wood, Warwickshire, Lower Jurassic (
8:
1189:
460:Species, temporal and spatial distribution
102:
20:
1121:
1072:
1062:
936:
370:occurs throughout eastern Australia from
386:According to Li, Huang & Cai, 2021,
1096:Kirejtshuk, Alexander G. (2020-02-17).
906:
401:
1435:Extant Late Triassic first appearances
7:
1377:DD1E8788-A60D-FFB2-FF23-C7C9425CF1A4
966:
964:
914:
912:
910:
767:Jarzembowski, Zheng & Zhao, 2022
736:Jarzembowski, Wang & Zheng, 2016
925:Deutsche Entomologische Zeitschrift
896:, Baissa, West Transbaikalia Russia
746:formerly placed in separate genus
514:(Tan, Wang, Ren & Yang, 2012)
14:
440:
420:
404:
122:
543:Tan, Wang, Ren & Yang, 2012
846:, Upper Jurassic (Tithonian),
1:
472:species have been described.
850:, Khoutiin-Khotgor, Mongolia
313:. The oldest species known,
1445:Taxa named by Edward Newman
1123:10.3390/geosciences10020073
783:, Middle to Late Jurassic,
1461:
1430:Rhaetian first appearances
1405:Endemic fauna of Australia
989:10.11646/zootaxa.4728.4.11
519:Pareuryomma ancistrodontum
720:Li, Huang & Cai, 2021
682:Tetraphalerus longicollis
507:, Inner Mongolia, China,
237:
230:
217:
210:
119:Scientific classification
117:
110:
101:
23:
1064:10.1186/1471-2148-12-113
1051:BMC Evolutionary Biology
804:, Issyk-Kul’, Kyrgyzstan
431:from the Mid-Cretaceous
661:, Bon-Tsagan, Mongolia
1026:Ommatidae Species List
561:, Karatau, Kazakhstan
499:Middle-Late Jurassic,
427:Holotype specimen of
397:
1425:Mesozoic life of Asia
1359:Paleobiology Database
1163:10.1144/SP521-2021-56
785:Togo-Khuduk Formation
626:(Deichmueller, 1886)
582:Tetraphalerus grandis
530:Cai & Huang, 2017
501:Jiulongshan Formation
1410:Insects of Australia
938:10.3897/dez.68.74174
888:, Lower Cretaceous (
879:, Charmouth, England
820:, Lower Cretaceous (
677:(Ponomarenko, 1997)
599:(Ponomarenko, 1964)
577:(Ponomarenko, 1964)
559:Karabastau Formation
505:Tiaojishan Formation
490:, recent, Australia
362:bark and exhibiting
1440:Archostemata genera
1114:2020Geosc..10...73K
848:Ulan-Ereg Formation
659:Dzun-Bain Formation
619:, Bavaria, Germany
617:Solnhofen Limestone
511:Omma ancistrodontum
309:is an example of a
1420:Cretaceous insects
1031:2012-10-23 at the
877:Charmouth Mudstone
865:Lilstock Formation
828:Lower Cretaceous (
796:, Lower Jurassic (
787:, Bakhar, Mongolia
744:(Kirejtshuk, 2020)
653:Lower Cretaceous (
646:Procarabus zitteli
641:(Oppenheim, 1888)
631:Pyrochroa brevipes
376:Central Queensland
1387:
1386:
1346:Open Tree of Life
1195:Taxon identifiers
1157:: SP521–2021–56.
886:Ponomarenko, 1966
859:, Late Triassic (
844:Ponomarenko, 1997
817:Carabus elongatus
794:Ponomarenko, 1969
781:Ponomarenko, 1997
768:
745:
721:
705:Omma davidbatteni
669:Ponomarenko, 1997
591:Ponomarenko, 1968
568:Ponomarenko, 1968
527:Omma daxishanense
286:
285:
268:Ponomarenko, 1964
206:
1452:
1415:Jurassic insects
1380:
1379:
1367:
1366:
1354:
1353:
1341:
1340:
1328:
1327:
1315:
1314:
1302:
1301:
1289:
1288:
1276:
1275:
1263:
1262:
1250:
1249:
1237:
1236:
1235:
1222:
1221:
1220:
1190:
1183:
1182:
1142:
1136:
1135:
1125:
1093:
1087:
1086:
1076:
1066:
1042:
1036:
1023:
1017:
1016:
968:
959:
958:
940:
916:
887:
858:
845:
836:, United Kingdom
819:
795:
782:
766:
743:
737:
729:
728:Kirejtshuk, 2020
719:
713:
689:Mid Cretaceous (
684:
670:
648:
633:
611:Upper Jurassic (
606:
604:Ommamima pilosum
592:
584:
569:
544:
532:Upper Jurassic (
531:
521:
489:
444:
424:
408:
280:
279:Kirejtshuk, 2020
269:
261:
260:Handlirsch, 1906
253:
252:Handlirsch, 1906
245:
201:
127:
126:
106:
96:
33:
29:Temporal range:
21:
16:Genus of beetles
1460:
1459:
1455:
1454:
1453:
1451:
1450:
1449:
1390:
1389:
1388:
1383:
1375:
1370:
1362:
1357:
1349:
1344:
1336:
1331:
1323:
1318:
1310:
1305:
1297:
1292:
1284:
1279:
1271:
1266:
1258:
1253:
1245:
1240:
1231:
1230:
1225:
1216:
1215:
1210:
1197:
1187:
1186:
1144:
1143:
1139:
1095:
1094:
1090:
1044:
1043:
1039:
1033:Wayback Machine
1024:
1020:
970:
969:
962:
918:
917:
908:
903:
885:
856:
843:
812:(Brodie, 1845)
811:
802:Dzhil Formation
793:
780:
735:
727:
707:
676:
674:Omma longicolle
668:
666:Omma antennatum
640:
625:
598:
590:
588:Omma jurassicum
576:
567:
549:Late Jurassic (
542:
529:
513:
496:
483:
462:
457:
456:
455:
452:
445:
436:
425:
416:
409:
384:
278:
267:
259:
251:
244:Oppenheim, 1888
243:
226:
223:
200:
121:
97:
95:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
32:Rhaetian–Recent
31:
30:
27:
17:
12:
11:
5:
1458:
1456:
1448:
1447:
1442:
1437:
1432:
1427:
1422:
1417:
1412:
1407:
1402:
1392:
1391:
1385:
1384:
1382:
1381:
1368:
1355:
1342:
1329:
1316:
1303:
1290:
1277:
1264:
1251:
1238:
1223:
1207:
1205:
1199:
1198:
1193:
1185:
1184:
1137:
1088:
1037:
1018:
983:(4): 547–574.
960:
931:(2): 341–348.
905:
904:
902:
899:
898:
897:
894:Zaza Formation
883:Omma sibiricum
880:
854:Omma liassicum
851:
837:
809:Omma elongatum
805:
788:
778:Omma altajense
774:
773:Other Species:
771:
770:
769:
753:
741:Omma manukyani
738:
730:
722:
714:
687:
686:
685:
671:
651:
650:
649:
634:
609:
608:
607:
593:
585:
570:
565:Omma aberratum
547:
546:
545:
540:Omma delicatum
537:
523:
495:
494:Fossil species
492:
484:Newman, 1839 (
464:The following
461:
458:
454:
453:
446:
439:
437:
426:
419:
417:
413:Omma liassicum
410:
403:
400:
399:
398:
383:
380:
301:in the family
284:
283:
282:
281:
270:
262:
257:Pyrochroophana
254:
249:Chalepocarabus
246:
235:
234:
228:
227:
224:
215:
214:
208:
207:
193:
189:
188:
183:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
143:
139:
138:
133:
129:
128:
115:
114:
108:
107:
99:
98:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
35:
34:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1457:
1446:
1443:
1441:
1438:
1436:
1433:
1431:
1428:
1426:
1423:
1421:
1418:
1416:
1413:
1411:
1408:
1406:
1403:
1401:
1398:
1397:
1395:
1378:
1373:
1369:
1365:
1360:
1356:
1352:
1347:
1343:
1339:
1334:
1330:
1326:
1321:
1317:
1313:
1308:
1304:
1300:
1295:
1291:
1287:
1282:
1278:
1274:
1269:
1265:
1261:
1256:
1252:
1248:
1243:
1239:
1234:
1228:
1224:
1219:
1213:
1209:
1208:
1206:
1204:
1200:
1196:
1191:
1180:
1176:
1172:
1168:
1164:
1160:
1156:
1152:
1148:
1141:
1138:
1133:
1129:
1124:
1119:
1115:
1111:
1107:
1103:
1099:
1092:
1089:
1084:
1080:
1075:
1070:
1065:
1060:
1056:
1052:
1048:
1041:
1038:
1034:
1030:
1027:
1022:
1019:
1014:
1010:
1006:
1002:
998:
994:
990:
986:
982:
978:
974:
967:
965:
961:
956:
952:
948:
944:
939:
934:
930:
926:
922:
915:
913:
911:
907:
900:
895:
891:
884:
881:
878:
874:
870:
866:
862:
857:Crowson, 1962
855:
852:
849:
842:
841:Omma gobiense
838:
835:
831:
827:
826:Purbeck Group
823:
818:
815:
810:
806:
803:
799:
792:
789:
786:
779:
775:
772:
765:
761:
757:
754:
751:
750:
742:
739:
734:
731:
726:
723:
718:
715:
711:
708:Jarzembowski
706:
703:
702:
700:
699:Burmese amber
696:
692:
688:
683:
680:
675:
672:
667:
663:
662:
660:
656:
652:
647:
644:
639:
635:
632:
629:
624:
623:Omma brevipes
621:
620:
618:
614:
610:
605:
602:
597:
594:
589:
586:
583:
580:
575:
571:
566:
563:
562:
560:
556:
552:
548:
541:
538:
535:
528:
524:
520:
517:
512:
509:
508:
506:
502:
498:
497:
493:
491:
487:
482:
481:
480:Omma stanleyi
476:
473:
471:
467:
459:
450:
443:
438:
434:
433:Burmese amber
430:
423:
418:
414:
407:
402:
396:
391:
389:
381:
379:
377:
373:
369:
368:Omma stanleyi
365:
361:
360:
355:
354:Omma stanleyi
352:
348:
347:
342:
338:
337:
336:Omma stanleyi
332:
328:
324:
320:
316:
312:
311:living fossil
308:
304:
300:
296:
292:
291:
277:
276:
271:
266:
263:
258:
255:
250:
247:
242:
239:
238:
236:
233:
229:
222:
221:
220:Omma stanleyi
216:
213:
209:
204:
199:
198:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
147:
144:
141:
140:
137:
134:
131:
130:
125:
120:
116:
113:
112:Omma stanleyi
109:
105:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
1202:
1154:
1150:
1140:
1105:
1101:
1091:
1054:
1050:
1040:
1021:
980:
976:
928:
924:
882:
853:
840:
816:
808:
790:
777:
763:
759:
755:
747:
740:
732:
725:Omma janetae
724:
716:
709:
704:
681:
673:
665:
645:
638:Omma zitteli
637:
630:
622:
603:
596:Omma pilosum
595:
587:
581:
573:
564:
555:Kimmeridgian
539:
526:
518:
510:
478:
477:
474:
463:
448:
447:Specimen of
429:Omma janetae
428:
412:
393:
387:
385:
367:
357:
353:
350:
344:
334:
322:
315:O. liassicum
314:
306:
289:
288:
287:
273:
264:
256:
248:
240:
225:Newman, 1839
219:
218:
212:Type species
196:
195:
111:
24:
18:
1294:iNaturalist
1227:Wikispecies
1102:Geosciences
890:Valanginian
830:Hauterivian
574:Omma grande
382:Description
339:, which is
1394:Categories
901:References
869:Hettangian
834:Weald Clay
822:Berriasian
798:Hettangian
717:Omma forte
701:, Myanmar
695:Cenomanian
449:Omma forte
364:thanatosis
359:Eucalyptus
331:Cretaceous
241:Procarabus
176:Coleoptera
156:Arthropoda
1400:Ommatidae
1179:244091355
1171:0305-8719
1132:2076-3263
1108:(2): 73.
1013:212766123
997:1175-5334
955:244783528
947:1860-1324
873:Blue Lias
791:Omma avus
760:Coronomma
749:Cionocups
613:Tithonian
551:Oxfordian
534:Oxfordian
346:Beutelius
303:Ommatidae
275:Cionocups
186:Ommatidae
142:Kingdom:
136:Eukaryota
1218:Q7090278
1212:Wikidata
1083:22776212
1029:Archived
1005:32229891
861:Rhaetian
764:axsmithi
733:Omma lii
536:), China
372:Victoria
327:Jurassic
319:Rhaetian
265:Ommamima
232:Synonyms
182:Family:
152:Phylum:
146:Animalia
132:Domain:
1351:3428713
1338:1903031
1312:1061320
1286:1096143
1273:2646454
1110:Bibcode
1074:3518168
1057:: 113.
977:Zootaxa
466:extinct
349:in 2020
341:endemic
299:beetles
192:Genus:
172:Order:
166:Insecta
162:Class:
1364:153451
1325:767667
1299:467298
1177:
1169:
1130:
1081:
1071:
1011:
1003:
995:
953:
945:
712:, 2020
710:et al.
691:Albian
655:Aptian
470:extant
205:, 1839
203:Newman
1372:Plazi
1307:IRMNG
1175:S2CID
1009:S2CID
951:S2CID
295:genus
293:is a
1333:NCBI
1320:ITIS
1281:GBIF
1260:67YK
1247:Omma
1233:Omma
1203:Omma
1167:ISSN
1128:ISSN
1079:PMID
1001:PMID
993:ISSN
981:4728
943:ISSN
756:Omma
553:and
503:and
486:type
468:and
388:Omma
329:and
323:Omma
307:Omma
290:Omma
197:Omma
37:PreꞒ
25:Omma
1268:EoL
1255:CoL
1242:AFD
1159:doi
1155:521
1118:doi
1069:PMC
1059:doi
985:doi
933:doi
892:),
871:),
863:),
839:(?)
832:),
824:),
807:(?)
800:),
776:(?)
664:(?)
657:),
636:(?)
615:),
572:(?)
525:(?)
374:to
297:of
1396::
1374::
1361::
1348::
1335::
1322::
1309::
1296::
1283::
1270::
1257::
1244::
1229::
1214::
1173:.
1165:.
1153:.
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