Ornithocheiridae - Knowledge (XXG)

2631:", which consists of travelling long distances without flapping using the vertical gradient of wind speed near the ocean surface as an advantage, at moderate flight speed. Several studies showed that most ornithocheirids sprawled their limbs to a large degree, similar to crocodiles, while other studies conclude that ornithocheirids were generally quadrupedal. Yet other studies concluded that ornithocheirids held their limbs more or less vertically extended, similar to an avian or mammalian configuration. Some studies in later genera show that ornithocheirids spend much of their time at sea, and as a result, they perhaps influenced the later pteranodontids with the same piscivorous diet, as well as their well-developed flight techniques. Analyses of limb proportions in the genus 1294: 2755: 10109: 997: 8876: 2610:. The skim-feeding hypothesis on ornithocheirids has been discounted in recent assessments of pterosaur skim-feeding, while dip-feeding is supported by a number of anatomical features. The elongated rostra of ornithocheirids are considered ideal for reaching into the water to grab swimming creatures; the rostral crests of ornithocheirids would have worked well as stabilizers for the jaws tips while being plunged into the water. Large, forward-facing eyes and well-developed 126: 7678: 153: 7684: 7061: 564: 8892: 10116: 5489: 7066: 1045: 6520: 1193: 10122: 2577: 6528: 6535: 8884: 1036:(MN 6594-V), a possible ornithocheirid (as it may be an anhanguerid), was calculated to have had a normal wingspan of 8.26 meters (27.1 ft), with another calculated maximum wingspan reached 8.70 meters (28.5 ft), indicating that the wingspans of toothed pterosaurs could exceed 7 meters (23 ft). 3908:

Eberhard D. Frey; Wolfgang Stinnesbeck; David M. Martill; Héctor E. Rivera-Sylva; Héctor Porras Múzquiz (2020). "The geologically youngest remains of an ornithocheirid pterosaur from the late Cenomanian (Late Cretaceous) of northeastern Mexico with implications on the paleogeography and extinction of

2781:

was found to contain a large number of pterosaur genera. The most diverse formation of the group is the Romualdo Formation, known for its wide variety of pterosaur remains. The formation dates back around 111 to 108 million years ago, also during the Albian stage of the Early Cretaceous. The Romualdo

1317:

The family Ornithocheiridae has had a controversial and very confusing taxonomic history; paleontologists who have studied this group seem to have had a different opinion on the composition of ornithocheirid taxonomy. A term called Anhangueridae was coined by Diogenes de Almeida Campos and Kellner in

584:

The family Ornithocheiridae is without a doubt, one of the most well-known pterosaur groups, mostly due to their very controversial and convoluted taxonomic history. Most of the ornithocheirid fossil record consists of isolated teeth, as well as fragmentary bones, reaching hundreds or even thousands

2597:

animals, mainly because they seem to have been suited for flight over marine settings; in fact, most ornithocheirids are known from lagoonal, coastal and marine deposits. Although the manner in which ornithocheirids gathered their food has not been researched in detail, it is generally thought that

1322:

priority. He therefore considered Anhangueridae a junior synonym of Ornithocheiridae in his study of pterosaur phylogeny in 2003. However, in many recent studies, Anhangueridae is recovered as a separate and distinct family from Ornithocheiridae, each containing different genera. The original term

958:, however, contradict this name synonymy, with Ornithocheiridae and Anhangueridae classified as different families, therefore following the 2014 analysis by Andres and colleagues instead. Other studies such as the ones by Kellner and colleagues in 2019 have used a different concept, classifying 1156:, though having similarities with other ornithocheirids, including the anterior end of the rostrum being transversely expanded, or having a low, bony sagittal crest that includes a smooth dorsal margin on the rostrum, still possesses some unique features. The most distinct characteristic of 1568:

is supported by several synapomorphies, such the teeth being short, straight, and relatively uniform in size. Several other recent studies such as the ones by Alexander Kellner and colleagues, or the one by Rodrigo Pêgas and colleagues, both in 2019, have recovered

3830:

Pêgas, Rodrigo V.; Holgado, Borja; Leal, Maria Eduarda C. (November 18, 2019). "On Targaryendraco wiedenrothi gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids".

2635:, however, show that some ornithocheirids were consistent with hopping, but the later genera were suggested that they most likely walked on four limbs, which consists on their wing-fingers as the front limbs, and using their hind limbs to balance. 546:. Ornithocheirids were excellent fish hunters, using various flight techniques to catch their prey, and were also capable of flying great distances without flapping constantly. Paleontologists suspect that ornithocheirids were the ancestors of the 1635:), altogether formed an unnamed clade within the Ornithocheiridae. However, as noted by Jacobs and colleagues, support for some of these arrangements is relatively weak due to the limited number of characters that can be scored, and the levels of 1372:

is another pterosaur that was initially classified within the Ornithocheiridae; later analysis have found it outside the Ornithocheiridae: Timothy Myers in 2015 and Nicholas Longrich and colleagues in 2018 for example, are two studies that found

5381:

Weishampel, David B; et al. (2004). "Dinosaur distribution (Early Cretaceous, North America)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 553-556.

536:, there is still no virtual consensus over the exact content and interrelationships of this group. Ornithocheirids were the most successful pterosaurs during their reign, and were also the largest pterosaurs before the appearance of the 3587: 2614:

are ideal for dip-feeding as well, which permits effective spotting of prey as well as judgement of distances when striking at them; as such, it seems likely that at least several ornithocheirids were efficient dip feeders. Sedate

1209:

The forelimbs of ornithocheirids were proportionally enormous, around five times longer than their legs. Substantial anchorage on the body is required given the mighty arms, and accordingly, ornithocheirids have robust

1281:(or tibiae) are similarly developed and of equal length to the femora. Although the feet in ornithocheirids are poorly known, they seem to be relatively small and gracile, with undeveloped claws and a hook-like fifth 4804:

Renan A. M. Bantim; Antônio A. F. Saraiva; Gustavo R. Oliveira; Juliana M. Sayão (2014). "A new toothed pterosaur (Pterodactyloidea: Anhangueridae) from the Early Cretaceous Romualdo Formation, NE Brazil".

3944:

Martill, David M.; Unwin, David M. (April 2012). "The world's largest toothed pterosaur, NHMUK R481, an incomplete rostrum of Coloborhynchus capito (Seeley, 1870) from the Cambridge Greensand of England".

1339:

In the past, many pterosaur genera were assigned to the Ornithocheiridae; however, following recent studies, these supposed ornithocheirids had been reclassified to other groups or families. The pterosaur

1526:. In the same year, Megan Jacobs and colleagues have recovered a different set of relationships for ornithocheirids in their analysis. Within the family, three clades emerge: the first one consists of 1020:, ornithocheirids were the largest; they were also among the most successful and widely distributed pterosaurs. Ornithocheirids were characterized by long jaws with spike-like teeth. Ornithocheirid 1124:

supported by these three species. Yet another feature shared by these three species is that the teeth are short, straight, and relatively uniform in size, something that is not present in other

528:

Ornithocheirids are generally infamous for having an enormously controversial and very confusing taxonomy. Although agreements that these animals were related, and therefore similar to

5230:

Lane, Jennifer A.; Maisey, John G. (September 2012). "The visceral skeleton and jaw suspension in the durophagous hybodontid shark Tribodus limae from the Lower Cretaceous of Brazil".

4549:"On Macrurosaurus semnus (Seeley), a Long tailed Animal with Proc lous Vertebrae from the Cambridge Upper Greensand, preserved in the Woodwardian Museum of the University of Cambridge" 893:

may belong to this family as well. However, their taxonomic status and precise relationships with other ornithocheirids are uncertain. In 2019, upon the description of the pterosaur

5427:

Cappetta H. 1987. Chondrichthyes II. Mesozoic and Cenozoic Elasmobranchii. Schultze H.-P. (ed.), Handbook of Paleoichthyology, Volume 3B. Gustav Fischer Verlag, Stuttgart, 193 p.

950:. However, back in 2001, Unwin considered the name Anhangueridae a junior synonym of Ornithocheiridae, a concept that was later followed by several paleontologists such as 1108:

margins of the premaxilla being convex in both anterior and lateral view, a feature that resulted in a bluntly rounded outline of the tip of the rostrum. The rostrum in

3638:

Pentland, Adele H.; Poropat, Stephen F.; Tischler, Travis R.; Sloan, Trish; Elliott, Robert A.; Elliott, Harry A.; Elliott, Judy A.; Elliott, David A. (December 2019).

550:; this is due to many shared aspects, such as unique flying techniques, capability of long-distance flights, and most of their diet, which mainly consisted of fish. 4891:

nov. gen., nov. sp., a long-necked, large pterosaurier from the Aptian of Brasil. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, B 83:145-172

3273:

Description of a new species of Anhangueridae (Pterodactyloidea) with comments on the pterosaur fauna from the Santana Formation (Aptian-Albian), northeastern Brazil

2619:

methods might have also been used when hunting -- examples of these methods are: reaching food while being alighted on the water surface, and shallow surface dives.

10214: 3977:

Kellner, A. W. A.; Campos, D. A.; Sayão, J. M.; Saraiva, A. N. A. F.; Rodrigues, T.; Oliveira, G.; Cruz, L. A.; Costa, F. R.; Silva, H. P.; Ferreira, J. S. (2013).

2643:

Ornithocheirids were a widespread type of pterosaur, with many fossil remains found across the world. The first true ornithocheirid specimens were uncovered in the

1318:

1985 to refer to pterosaurs that belong in this family. In 2001, however, Unwin argued that the name Ornithocheiridae refers to an identical group, and should have

4043:

First occurrence of the pterosaur Coloborhynchus (Pterosauria, Ornithocheiridae) from the Wessex Formation (Lower Cretaceous) of the Isle of Wight, England.

4684: 2655:, and dated back to the Albian stage of the Early Cretaceous. Within the fossil site, several other pterosaurs were also found, including the pterosaurs 4682:

Toothed pterosaurs from the Santana Formation (Cretaceous; Aptian-Albian) of northeastern Brazil. A reappraisal on the basis of newly described material

4701:

Fastnacht, M (2001). "First record of Coloborhynchus (Pterosauria) from the Santana Formation (Lower Cretaceous) of the Chapada do Araripe of Brazil".

3744:

Holgado, Borja; Pêgas, Rodrigo V.; Canudo, José Ignacio; Fortuny, Josep; Rodrigues, Taissa; Company, Julio; Kellner, Alexander W. A. (December 2019).

10201: 3640:"Ferrodraco lentoni gen. et sp. nov., a new ornithocheirid pterosaur from the Winton Formation (Cenomanian–lower Turonian) of Queensland, Australia" 1364:

in several recent studies such as the ones by Rodrigo Pêgas and colleagues, and Adele Pentland and colleagues, both studies of which are from 2019.

4590: 3446:

Kellner, Alexander W. A.; Caldwell, Michael W.; Holgado, Borja; Vecchia, Fabio M. Dalla; Nohra, Roy; Sayão, Juliana M.; Currie, Philip J. (2019).

3296:"LXI.— On the Ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge" 1032:, the largest toothed pterosaur, had a wingspan that may have reached 7 meters (23 ft). However, in 2013, a specimen referred to the genus 10273: 5008:(Thalassodromidae, Azhdarchoidea) from the Lower Cretaceous Santana Formation of Brazil, with a note on the nomenclature of Thalassodromidae". 4205:"An analysis of pterosaurian biogeography: implications for the evolutionary history and fossil record quality of the first flying vertebrates" 5521: 5471: 2602:, pushing their lower jaw through the water to snap up food upon impact, or fed by gleaning food from the water surface like some modern-day 10283: 4342:"On the Size and Flight Diversity of Giant Pterosaurs, the Use of Birds as Pterosaur Analogues and Comments on Pterosaur Flightlessness" 1498:. In 2019, a study performed by Adele Pentland and colleagues had found the Ornithocheiridae to comprise more genera; while the typical 10108: 1444:

was recovered as a stable sister taxon to the group Ornithocheiroidea. Some later analyses have also recovered this concept, with both

5493: 1585:

within the family Anhangueridae instead of the Ornithocheiridae. In 2020, a study by Borja Holgado and Pêgas had also recovered both

1346:

for example, was initially classified within the Ornithocheiridae; however, later analysis had found it in a different family called

5450: 5387: 5079: 3987:

Wellnhofer, 1987 (Pterodactyloidea, Anhangueridae) and other large pterosaurs from the Romualdo Formation, Lower Cretaceous, Brazil"

4130:

Myers, Timothy S. (November 2, 2015). "First North American occurrence of the toothed pteranodontoid pterosaur Cimoliopterus".

3746:"On a new crested pterodactyloid from the Early Cretaceous of the Iberian Peninsula and the radiation of the clade Anhangueria" 585:

of remains in some localities. The first uncovered ornithocheirid remains were described in 1861 by British paleontologist Sir

4965:

Pêgas, R. V.; Costa, F. R.; Kellner, A. W. A. (2018). "New Information on the osteology and a taxonomic revision of The genus

3700:"A new toothless pterosaur (Pterodactyloidea) from Southern Brazil with insights into the paleoecology of a Cretaceous desert" 5043:

Kellner, A.W.A.; Campos, D.A. (1996). "First Early Cretaceous theropod dinosaur from Brazil with comments on Spinosauridae".

4586: 1331:, therefore it is recovered as a larger group comprising the families Anhangueridae and Ornithocheiridae in recent analysis. 1180:, a pterosaur once assigned to the Ornithocheiridae, but some recent analysis have placed it within the more inclusive group 918:

In 2014, Andres and colleagues defined the Ornithocheiridae with a different definition: the most inclusive clade containing

5355:

Coombs, Walter P. (June 13, 1995). "A Nodosaurid Ankylosaur (Dinosauria: Ornithischia) from the Lower Cretaceous of Texas".

732:, he divided the family Ornithocheiridae into two subfamilies: Ornithocheirinae and Criorhynchinae; the former consisted of 152: 5266: 3877:"A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae" 2953:

Ornithocheirids were also partially distributed in North America, and several specimens are thought to belong to the genus

1293: 10278: 8875: 7683: 632:. However, in the same year, this was emended to Ornithocheiridae by Seeley himself following the article 11.7.1.3 of the 7677: 3520:"Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary" 1424:

had not been previously referred to the Ornithocheiridae besides a note added in proof to Unwin in 2001 that stated that

10288: 1727: 1464:

Different phylogenetic analysis have found Ornithocheiridae to comprise different genera, the most typical ones being

1028:

reached wingspans of up to 8 meters (26 ft) or more. Specimen NHMUK R481, a specimen that belongs to the species

844: 5443:

The Illustrated Encyclopedia of Pterosaurs: An illustrated natural history of the flying reptiles of the Mesozoic Era

1234:. Over 60 percent of the wing length is occupied by the wing fingers, making them among the longest possessed by any 4928:

Pinheiro, Felipe L.; Fortier, Daniel C.; Schultz, Cesar L.; De Andrade, José Artur F.G.; Bantim, Renan A.M. (2011).

2754: 4738:"On the presence of a giant pterosaur in the Lower Cretaceous (Aptian) of Chapada do Araripe (northeastern Brazil)" 1203: 7060: 5514: 5162: 1958: 1917: 1816: 1650:

recovered by Jacobs and colleagues, and the one to the right is a topology recovered by Pentland and colleagues.

1522:

to belong the Ornithocheiridae as well, specifically sister taxa within the Ornithocheirinae, closely related to

1092: 924: 140: 135: 5139: 1599:

has generally been recovered outside the Ornithocheiridae; subsequent analyses have found it as sister taxon to

10268: 5618: 4681: 2017: 1751: 1646:

showing the possible genera that are included within the ornithocheirid family. The cladogram to the left is a

4901:

Martill, David M. (2011). "A new pterodactyloid pterosaur from the Santana Formation (Cretaceous) of Brazil".

6519: 1488:, and altogether formed the family Ornithocheiridae. In 2018, Longrich and colleagues had included the genus 1448:

and the Ornithocheiroidea placed within the larger group Eupterodactyloidea. Another study in 2019 recovered

1420:

was used to define the Ornithocheiridae in Unwin's 2003 study; however, Andres and Myers in 2013 argued that

7096: 4424: 3698:

Kellner, Alexander W.A.; Weinschütz, Luiz C.; Holgado, Borja; Bantim, Renan A.M.; Sayão, Juliana M. (2019).

2000: 1253:. The tails of ornithocheirids are poorly known, though they appear to be composed of at least eleven short 1184:, outside the Ornithocheiridae. Other studies, however, have recovered it within the Anhangueridae instead. 1148:, which is similar to those seen in another ornithocheirid (though also recovered as an anhanguerid) called 7065: 6527: 4067:"A new pterodactyloid pterosaur from the Wessex Formation (Lower Cretaceous) of the Isle of Wight, England" 1494:

in their phylogenetic analysis, specifically as a member of the ornithocheirine subfamily, sister taxon to

10141: 10016: 8174: 4046: 3347: 2804: 2334: 1024:

varied in size, with smaller species having wingspans of approximately 4 meters (13 ft), while giant

988:

outside the Anhangueria due to being undiagnosable. Most recent studies have since followed this concept.

818: 768:

within the family Criorhynchidae (which is now considered synonymous to Ornithocheiridae), and recognized

725: 509:. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the 482: 10115: 3588:"A new species of Coloborhynchus (Pterosauria, Ornithocheiridae) from the mid-Cretaceous of North Africa" 3395: 10240: 10121: 4605: 4411: 4399: 3795: 2065: 2041: 1792: 955: 10179: 8891: 4765:

gen. et sp. nov. (Pterosauria, Pterodactyloidea) from the Santana Formation (Cretaceous) of NE Brazil"

3203:"An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England" 10188: 8269: 6708: 6618: 6534: 5507: 5309: 5196: 5088: 4635: 4353: 4051: 3757: 3651: 3459: 3339: 3242: 3079: 2847: 1941: 1775: 569: 8883: 3352: 996: 9773: 9462: 9149: 8448: 8375: 6143: 5863: 2644: 2082: 805: 785: 600: 280: 226: 125: 908: 772:

within the family Ornithocheiridae and subfamily Ornithocheirinae. He also designated the species

9302: 7776: 7619: 6762: 6565: 6552: 6466: 6308: 5939: 5914: 5247: 5212: 5143: 5104: 5025: 4986: 4786: 4718: 4568: 4529: 4490: 4451: 4185: 4147: 3926: 3848: 3607: 3448:"First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity" 3365: 3095: 1615:, the three formed the family Cimoliopteridae. In the analysis by Jacobs and colleagues, the two 1257:, and become relatively circular in cross section toward the end of the series. Like the related 835: 801: 469: 147: 10245: 4870:

nov. gen., nov. sp.: novo Pterosauria (Pterodactyloidea) da Chapada do Araripe, Ceara, Brasil."

2842:

were also present within the fossil site. Many other pterosaur were found within, including the

10227: 9728: 8543: 8472: 7714: 7701: 7644: 7407: 7305: 7207: 7125: 6903: 6880: 6846: 6776: 6317: 6233: 6185: 6159: 5872: 5777: 5467: 5446: 5383: 5337: 4822: 4663: 4381: 4298: 4234: 4008: 3783: 3721: 3677: 3551: 3487: 3419: 3276: 3176: 2962: 1406: 1274: 1087: 912: 701: 10232: 10219: 5072:"A new crested maniraptoran dinosaur from the Santana Formation (Lower Cretaceous) of Brazil" 9957: 9832: 9624: 9478: 9020: 8923: 8910: 8842: 8722: 8711: 7961: 7818: 7755: 7530: 7512: 7350: 7174: 6990: 6976: 6934: 6440: 6426: 6358: 6037: 5587: 5364: 5327: 5317: 5239: 5204: 5135: 5096: 5052: 5017: 4978: 4945: 4910: 4814: 4776: 4710: 4653: 4643: 4560: 4521: 4482: 4443: 4371: 4361: 4288: 4280: 4224: 4216: 4177: 4139: 4081: 3998: 3954: 3918: 3888: 3840: 3773: 3765: 3711: 3667: 3659: 3599: 3541: 3531: 3477: 3467: 3409: 3357: 3307: 3214: 3166: 3156: 3087: 2822: 2230: 1433: 904: 853: 685: 510: 506: 210: 10046: 9889: 9738: 9523: 9427: 9402: 9395: 9382: 9315: 9201: 9099: 9065: 8961: 8941: 8795: 8653: 8594: 8581: 8552: 8428: 8339: 8198: 8071: 8026: 7898: 7827: 7806: 7632: 7293: 7273: 7245: 7221: 7157: 7149: 7142: 7134: 6889: 6869: 6699: 6675: 6625: 6433: 6416: 6406: 6383: 6339: 6331: 6299: 6288: 6267: 6221: 6213: 6152: 6068: 6008: 5927: 5896: 5786: 5765: 5718: 5603: 5578: 4688: 3807: 3400: 2857: 2816: 2735: 2628: 2317: 1437: 1356: 1231: 1219: 1218:, which served the purpose of housing their substantial forelimb muscles. The shoulder or 1211: 1025: 889: 514: 5126:(Crocodylomorpha, Mesoeucrocodylia) from the Lower Cretaceous of Patagonia (Argentina)". 1112:

lacks a constriction that is posterior to the anterior rosette, a feature also shared by

1090:, a feature that is also seen in the ornithocheirid (or alternately anhanguerid) species 5313: 5200: 5122:

Ortega, F. J.; Z. B. Gasparini; A. D. Buscalioni; J. O. Calvo (2000). "A new species of

5092: 4639: 4357: 3761: 3655: 3463: 3343: 3083: 2782:

Formation is found to contain a variety of ornithocheirids (or anhanguerids), including

2627:

Similar to modern-day albatrosses, most ornithocheirids used a flight technique called "

1161: 10023: 9996: 9974: 9798: 9712: 9697: 9649: 9588: 9573: 9539: 9364: 9348: 9336: 9322: 9275: 9232: 9192: 9106: 9052: 8780: 8773: 8766: 8738: 8645: 8566: 8493: 8404: 8160: 8052: 7987: 7928: 7868: 7834: 7554: 7481: 7453: 7372: 7322: 7261: 7230: 6830: 6806: 6784: 6730: 6688: 6632: 6600: 6281: 6198: 6175: 6054: 5980: 5948: 5801: 5750: 5698: 5332: 5293: 4658: 4623: 4376: 4341: 4293: 4268: 4229: 4204: 4203:

Upchurch, Paul; Andres, Brian; Butler, Richard J.; Barrett, Paul M. (August 18, 2015).

3778: 3745: 3672: 3639: 3546: 3519: 3482: 3447: 3171: 3144: 3027: 2973:, which dated back to the Albian and Cenomanian stages. The formation includes several 2889: 2852: 2774: 2741: 2668: 2599: 2421: 2397: 2293: 1833: 1672: 1540: 1453: 1235: 1125: 1097: 1075: 1066: 1054: 1017: 1002: 972: 883: 677: 642: 625: 608: 574: 542: 456: 424: 391: 339: 317: 242: 5071: 4564: 4525: 4447: 4066: 1105: 10262: 10060: 10031: 10007: 9988: 9946: 9931: 9880: 9869: 9762: 9754: 9704: 9641: 9612: 9596: 9580: 9566: 9507: 9485: 9436: 9329: 9090: 9033: 8952: 8787: 8759: 8731: 8602: 8485: 8276: 8206: 8078: 8061: 7917: 7854: 7798: 7563: 7519: 7400: 7393: 7386: 7379: 7364: 7314: 7238: 7195: 7188: 7114: 6961: 6954: 6947: 6796: 6738: 6723: 6716: 6664: 6447: 6116: 6076: 6026: 5995: 5956: 5851: 5843: 5821: 5725: 5709: 5108: 5056: 5029: 4737: 4722: 4572: 4533: 4494: 3930: 3852: 3611: 3099: 2974: 2970: 2903: 2893: 2884: 2828: 2810: 2792: 2766: 2762: 2717: 2702: 2696: 2686: 2370: 2303: 2280: 1685: 1490: 1397: 1379: 1347: 1319: 1270: 1258: 1198: 1165: 1060: 1008: 947: 938: 877: 672:; no type species was designated, however. In 1876, however, Seeley pointed out that 613: 591: 547: 533: 529: 434: 372: 350: 328: 306: 83: 5251: 5216: 5147: 4990: 4790: 4455: 4189: 4151: 4003: 3978: 3369: 2901:

remains were found within the formation, with some specimens referred to the genera

2777:

called the Santana Group (sometimes known as the Santana Formation) in northeastern

1323:

Ornithocheirae by Seeley had been redefined as the least inclusive clade containing

724:

for the species with lanceolate jaw tips, a concept later favored by paleontologist

563: 10053: 9981: 9852: 9747: 9681: 9633: 9443: 9371: 9249: 9217: 8985: 8752: 8680: 8621: 8516: 8153: 8138: 8122: 8113: 8102: 7942: 7905: 7791: 7741: 7584: 7497: 7490: 7470: 7442: 7435: 7421: 7414: 7329: 7286: 7253: 6969: 6911: 6854: 6839: 6819: 6644: 6375: 6367: 6347: 6274: 6253: 6246: 6101: 5889: 5835: 5794: 5368: 3067: 2982: 2758: 2690: 2681: 2586: 2247: 1476:. In 2014, Andres and colleagues created the subfamily Ornithocheirinae to contain 1428:

appeared to be a small ornithocheirid. Phylogenetic analyses since then have found

1368: 1361: 1266: 1250: 1176: 1121: 1078:

on both tips, as well as robust fang-like teeth. The sagittal crest of the species

977: 895: 867: 738: 648: 586: 286: 269: 260: 5021: 4982: 4914: 4143: 4085: 4045:, 126, vol. 3, Proceedings of the Geologists' Association, pp. 377–380, 3958: 3844: 3716: 3699: 3603: 3586:

Jacobs, Megan L.; Martill, David M.; Ibrahim, Nizar; Longrich, Nick (March 2019).

3361: 1246: 1044: 5461: 5322: 4648: 4624:"High Diversity in Cretaceous Ichthyosaurs from Europe Prior to Their Extinction" 4622:

Valentin Fischer; Nathalie Bardet; Myette Guiomar & Pascal Godefroit (2014).

4366: 4220: 3536: 3114: 10173: 9939: 9899: 9818: 9782: 9516: 9291: 9224: 9167: 9113: 8970: 8638: 8509: 8501: 8461: 8420: 8412: 8287: 8254: 8222: 8214: 8185: 8146: 8130: 8089: 8018: 8010: 8003: 7949: 7890: 7882: 7543: 7505: 7428: 7083: 7016: 6983: 6919: 6260: 6129: 6084: 6061: 5736: 4267:

Bestwick, J.; Unwin, D.M.; Butler, R.J.; Henderson, D.M.; Purnell, M.A. (2018).

4065:

Steel, Lorna; Martill, David M.; Unwin, David M.; Winch, John D. (August 2005).

3330:

Unwin, D. M. (2003). "On the phylogeny and evolutionary history of pterosaurs".

3014: 2909: 2731: 2677: 2672: 2607: 2594: 2177: 1432:

as a rogue taxon either within the Pterodactyloidea, the Ornithocheiroidea, the

1342: 1309: 1192: 1181: 1049: 981: 951: 744: 709: 652:(meaning "ram beak") based on highly distinctive jaw fragments. Owen reassigned 518: 445: 58: 39: 17: 5488: 3769: 3663: 3472: 2576: 1510:

clade was included in their analysis, Pentland and colleagues found the genera

9907: 9862: 9790: 9665: 9657: 9492: 9471: 9416: 9284: 9259: 9209: 9174: 9120: 9008: 9000: 8993: 8745: 8668: 8660: 8559: 8523: 8361: 8262: 8241: 8230: 8167: 7784: 7577: 7570: 7005: 6896: 5903: 5560: 5070:

Martill, D.M.; Cruickshank, A.R.I.; Frey, E.; Small, P.G.; Clarke, M. (1996).

4872:

Resumos dos communicaçoes VIII Congresso bras. de Paleontologia e Stratigrafia

4818: 4781: 4760: 4486: 4181: 4170:

Earth and Environmental Science Transactions of the Royal Society of Edinburgh

3414: 3311: 3002: 2993: 2934: 2928: 2922: 2915: 2798: 2747: 2726: 2445: 1512: 1282: 1170: 1145: 1141: 1101: 761: 637: 537: 402: 103: 68: 10164: 5100: 3280: 3161: 9967: 9409: 9357: 9079: 9072: 9040: 8978: 8704: 8697: 8689: 8609: 8387: 8353: 8303: 8296: 8033: 7971: 7875: 7463: 6862: 6579: 6206: 6046: 5881: 5758: 5680: 5667: 5630: 5569: 5530: 4841: 3008: 2988: 2940: 2878: 2872: 2863: 2843: 2708: 2611: 1643: 1636: 1254: 712:

jaw tips. Therefore, to avoid confusion, Lydekker preferred to use the name

502: 197: 164: 108: 52: 5401: 5341: 4826: 4667: 4385: 4302: 4238: 4168:

Andres, Brian; Myers, Timothy S. (September 2012). "Lone Star Pterosaurs".

4012: 3893: 3876: 3787: 3725: 3681: 3555: 3491: 3423: 3180: 4950: 4929: 3518:

Longrich, Nicholas R.; Martill, David M.; Andres, Brian (March 13, 2018).

3219: 3202: 1222:

in ornithocheirids is set at a perpendicular angle to the spine, with the

10158: 10039: 9923: 9915: 9689: 9673: 9604: 9181: 8531: 8436: 8368: 8040: 7995: 7768: 6997: 6657: 5554: 3254:

Flugsaurier: Pterosaur Papers in Honour of Peter Wellnhofer. Zitteliana B

2997: 2977: 2946: 2868: 2838: 2713: 2616: 2462: 2192: 1647: 1518: 1410: 1303: 1278: 1223: 1021: 522: 413: 361: 184: 98: 93: 78: 73: 63: 43: 4469:

Nopcsa, Baron Francis (1923). "Notes on the British dinosaurs, Part IV:

3922: 10206: 4714: 2648: 1227: 903:, which was assigned to this family by Unwin, and recovered as a basal 113: 88: 4284: 3115:"The Ornithosauria: an Elementary Study of the Bones of Pterodactyles" 8630: 8397: 7979: 6587: 6454: 5548: 5243: 3091: 2898: 2778: 1262: 1242: 1215: 174: 10135: 4548: 4509: 4412:

http://h2vp.blogspot.com/2011/09/dinosaur-revolution-anhanguera.html

3295: 7728: 6324: 5208: 4934:, with comments on the relationships of Tapejaridae (Pterosauria)" 4591:"Wyleyia: a new bird humerus from the Lower Cretaceous of England" 2966: 2753: 2575: 1409:, more specifically placed within the family Cimoliopteridae in a 1292: 1191: 1043: 995: 604: 562: 10193: 4410:. Weblog entry, September 20, 2011. Accessed September 28, 2011: 1607:

is within the clade Targaryendraconia, again, closely related to

4269:"Pterosaur dietary hypotheses: a review of ideas and approaches" 2722: 2603: 633: 10139: 8908: 7699: 7081: 6550: 5665: 5541: 5503: 4842:"Description of Pterosaurian (Pterodactyloidea: Anhangueridae, 5499: 3138: 3136: 3134: 3132: 3130: 3128: 5294:"Endocranial Morphology of the Primitive Nodosaurid Dinosaur 3979:"The largest flying reptile from Gondwana: A new specimen of 5045:

Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen

2712:

were also found within the formation. Fossil remains of the

1653: 808:

pterosaurs, and divided Ornithocheiridae into three genera:

5187:

Hirayama, Ren (April 16, 1998). "Oldest known sea turtle".

5140:

10.1671/0272-4634(2000)020[0057:ANSOAC]2.0.CO;2

3068:"The early history of pterosaur discovery in Great Britain" 1100:

between these three species were also found, including the

2920:

Many fish remains were also found, assigned to the genera

2751:

were also found alongside the remains of ornithocheirids.

1230:. The shoulder girdle is also of typical construction for 5417:. United States Government Printing Office. pp. 3–7. 3396:"The Earliest Pterodactyloid and the Origin of the Group" 1360:; both pterosaurs were recovered as basal members of the 4250: 4248: 4024: 4022: 688:

in 1888. In the latter year, Lydekker acknowledged that

5292:

Paulina-Carabajal, A.; Lee, Y.N.; Jacobs, L.L. (2016).

3049: 3047: 3045: 3043: 1104:

having a tall and narrow shape in anterior aspect, the

976:

within the Anhangueridae, which, along with the family

796:

as its type species), and regarded it as distinct from

4840:

Veldmeijer, A.J.; Meijer, H.J.M.; Signore, M. (2009).

1603:, as mentioned earlier. Another possible position for 833:

In 2003, Unwin defined the family Ornithocheiridae as

589:, who assigned the fossil remains to a new species of 4163: 4161: 3149:

complex (Pterosauria) from the Cretaceous of England"

780:, but agreed with Hooley that it was synonymous with 5411:(DeKay) in the Western Interior of the United States 4969:(Pterodactyloidea, Tapejaridae, Thalassodrominae)". 3275:. National Science Museum Monographs. Vol. 17. 3271:

Kellner, Alexander W. A.; Tomida, Yukimitsu (2000).

3241:

Rodrigues, Taissa; Kellner, Alexander W. A. (2008).

1440:. In the phylogenetic analysis by Andres and Myers, 752:. In his review, Hooley also considered the species 636:. In 1874, Owen had proposed two new genera for the 452: 441: 430: 420: 409: 398: 387: 379: 368: 357: 346: 335: 324: 313: 302: 276: 10148: 10006: 9956: 9888: 9879: 9851: 9816: 9772: 9737: 9727: 9623: 9552: 9537: 9506: 9461: 9426: 9381: 9347: 9301: 9274: 9247: 9191: 9147: 9089: 9051: 9019: 8960: 8951: 8921: 8841: 8721: 8679: 8620: 8580: 8542: 8471: 8447: 8386: 8337: 8286: 8240: 8184: 8112: 8088: 8060: 8051: 7960: 7926: 7915: 7852: 7817: 7754: 7739: 7712: 7643: 7618: 7553: 7529: 7480: 7452: 7348: 7304: 7272: 7220: 7205: 7172: 7124: 7094: 7015: 6933: 6879: 6829: 6805: 6794: 6760: 6698: 6674: 6643: 6598: 6563: 6465: 6415: 6405: 6357: 6307: 6298: 6232: 6184: 6173: 6142: 6127: 6099: 6036: 6006: 5978: 5938: 5913: 5871: 5862: 5819: 5776: 5735: 5708: 5678: 5629: 5602: 3513: 3511: 3509: 3507: 3505: 3503: 3501: 1619:species had been found as sister taxa to the three 1297:Holotype skull and mandible of the ornithocheirids 1277:that anchor their hindlimb muscles. Ornithocheirid 966:, as well as several other close relatives such as 5004:Witton, Mark P. (October 2009). "A new species of 3972: 3970: 3968: 3072:Geological Society of London, Special Publications 2598:members of this family either fed like modern-day 915:and colleagues as the sister taxon of the former. 847:, and all its descendants. He included the genera 3825: 3823: 3821: 3819: 3817: 1593:within the Anhangueridae instead of this family. 668:, including the addition of a new species called 4742:Bollettino della Società Paleontologica Italiana 4125: 4123: 4121: 4119: 3332:Geological Society, London, Special Publications 3236: 3234: 3232: 3230: 1074:Ornithocheirids had elongated jaws with rounded 684:, a concept that was followed by paleontologist 5463:Pterosaurs: Natural History, Evolution, Anatomy 5267:"The Early Cretaceous Pterodactyloid Pterosaur 3633: 3631: 3629: 3627: 3625: 3623: 3621: 3441: 3439: 3437: 3435: 3433: 3389: 3387: 3385: 3383: 3381: 3379: 2996:. Within the fossil site, several specimens of 2855:(or thalassodromines, depending on the author) 4846:) remains from the Lower Cretaceous of Brazil" 4691:." Tekst. - Proefschrift Universiteit Utrecht. 4510:"On the Dinosauria of the Cambridge Greensand" 3870: 3868: 3866: 3864: 3862: 3739: 3737: 3735: 3693: 3691: 5515: 3012:, and along with these, remains of the shark 525:stages), around 140 to 90 million years ago. 8: 4866:Leonardi, G. & Borgomanero, G. (1985). " 1391:). In 2019, Pêgas and colleagues have found 5298:from the Early Cretaceous of North America" 4553:Quarterly Journal of the Geological Society 4514:Quarterly Journal of the Geological Society 4436:Proceedings of the Zoological Institute RAS 3325: 3323: 3321: 3196: 3194: 3192: 3190: 942:within the Ornithocheiridae, while placing 875:within the family, and also concluded that 10136: 9885: 9824: 9734: 9558: 9549: 9545: 9503: 9499: 9387: 9307: 9271: 9267: 9159: 9155: 9057: 9025: 8957: 8933: 8929: 8918: 8905: 8838: 8676: 8617: 8586: 8477: 8453: 8345: 8246: 8190: 8109: 8094: 8057: 7934: 7923: 7860: 7760: 7751: 7747: 7720: 7709: 7696: 7624: 7615: 7535: 7356: 7278: 7217: 7213: 7180: 7106: 7102: 7091: 7078: 6939: 6811: 6802: 6768: 6680: 6649: 6610: 6606: 6571: 6560: 6547: 6412: 6304: 6238: 6190: 6181: 6139: 6135: 6108: 6018: 6014: 5987: 5919: 5868: 5827: 5742: 5690: 5686: 5675: 5662: 5608: 5599: 5538: 5522: 5508: 5500: 4759:Ross A. Elgin & Eberhard Frey (2011). 624:. Later, in 1870, Seeley created the name 124: 31: 5331: 5321: 4949: 4780: 4657: 4647: 4375: 4365: 4292: 4228: 4050: 4002: 3892: 3777: 3715: 3671: 3545: 3535: 3481: 3471: 3413: 3351: 3218: 3170: 3160: 2593:Ornithocheirids are generally considered 4429:– valid taxon of azhdarchoid pterosaurs" 3991:Anais da Academia Brasileira de Ciências 3704:Anais da Academia Brasileira de Ciências 3581: 3579: 3577: 3575: 3573: 3571: 3569: 3567: 3565: 3243:"Review of the pterodactyloid pterosaur 620:as its type species, therefore creating 3039: 2961:). This pterosaur was uncovered in the 4327: 4315: 4254: 4110: 4098: 4028: 3803: 3793: 3394:Andres, B.; Clark, J.; Xu, X. (2014). 3300:Annals and Magazine of Natural History 3053: 1377:as sister taxon to the two species of 1164:crest, a feature that is only seen in 501:, meaning "bird hands") is a group of 3294:Holley, Reginald Walter (June 1914). 3000:were thought to belong to the genera 1548:species, and the third one comprises 1269:that project almost in line with the 804:revised the taxonomic history of the 7: 1140:, the teeth were found to have been 1086:extended to the anterior end of the 1048:Skull comparisons between different 1000:Comparison between the holotypes of 3143:Rodrigues, T.; Kellner, A. (2013). 2802:as well as close relatives such as 2437: 2413: 2389: 2362: 2309: 2299: 2272: 2222: 2215: 2208: 2184: 2172: 2057: 2033: 1992: 1933: 1909: 1902: 1895: 1808: 1767: 1743: 1719: 1712: 1705: 1678: 1668: 1395:, along with two other pterosaurs ( 616:meaning "bird hand"), and assigned 27:Family of ornithocheiran pterosaurs 5357:Journal of Vertebrate Paleontology 5128:Journal of Vertebrate Paleontology 4971:Journal of Vertebrate Paleontology 4340:Witton, M.P.; Habib, M.B. (2010). 4132:Journal of Vertebrate Paleontology 3909:Late Cretaceous ornithocheirids". 2765:of Brazil, with the extent of the 2651:, belonging to the infamous genus 911:and colleagues, was reassigned by 704:, while other species referred to 599:. In 1869, British paleontologist 25: 5080:Journal of the Geological Society 4736:Dalla, F.V.; Ligabue, G. (1993). 4565:10.1144/GSL.JGS.1876.032.01-04.50 4547:Seeley, H. G. (January 1, 1876). 4526:10.1144/GSL.JGS.1879.035.01-04.42 3875:Holgado, B.; Pêgas, R.V. (2020). 1556:. The close relationship between 800:. Later, in 2001, paleontologist 10120: 10114: 10107: 8890: 8882: 8874: 7682: 7676: 7064: 7059: 6533: 6526: 6518: 5487: 1452:within a different group called 826:as the type and only species of 742:, while the latter consisted of 151: 4448:10.31610/trudyzin/2012.316.1.40 4004:10.1590/S0001-37652013000100009 664:. He referred three species to 5466:. Princeton University Press. 5403:The Late Cretaceous Ammonites 5369:10.1080/02724634.1995.10011231 5163:"Turtle: Origin and evolution" 4769:Swiss Journal of Palaeontology 1307:(B), and holotype mandible of 1241:In adult ornithocheirids, the 1: 10274:Valanginian first appearances 5022:10.1016/j.cretres.2009.07.006 4983:10.1080/02724634.2018.1443273 4938:Acta Palaeontologica Polonica 4915:10.1016/j.cretres.2010.12.008 4144:10.1080/02724634.2015.1014904 4086:10.1016/j.cretres.2005.03.005 3959:10.1016/j.cretres.2011.09.003 3881:Acta Palaeontologica Polonica 3845:10.1080/08912963.2019.1690482 3717:10.1590/0001-3765201920190768 3604:10.1016/j.cretres.2018.10.018 3362:10.1144/GSL.SP.2003.217.01.11 1273:, but seem to lack prominent 1202:in flight, notice their high 1052:, notice the ornithocheirids 5323:10.1371/journal.pone.0150845 4703:Paläontologische Zeitschrift 4649:10.1371/journal.pone.0084709 4604:(4): 721–728. Archived from 4367:10.1371/journal.pone.0013982 4221:10.1080/08912963.2014.939077 3537:10.1371/journal.pbio.2001663 646:(meaning "maimed beak") and 4889:Santanadactylus brasilensis 1728:Coloborhynchus clavirostris 1538:, the second one comprises 1226:being much longer than the 1214:, and stout, deeply keeled 946:within the separate family 845:most recent common ancestor 774:Coloborhynchus clavirostris 754:Coloborhynchus clavirostris 10305: 10284:Taxa named by Harry Seeley 5441:Wellnhofer, Peter (1991). 5057:10.1127/njgpa/199/1996/151 4680:Veldmeijer, A.J. (2006). " 4041:Martill, David M. (2015), 3770:10.1038/s41598-019-41280-4 3664:10.1038/s41598-019-49789-4 3473:10.1038/s41598-019-54042-z 1354:being the sister taxon of 1144:, elongated, recurved and 760:. In 1967, paleontologist 728:in 1914. In his review of 10104: 9827: 9561: 9548: 9502: 9390: 9310: 9270: 9162: 9158: 9060: 9028: 8936: 8932: 8917: 8904: 8871: 8837: 8589: 8480: 8456: 8348: 8249: 8193: 8097: 7937: 7863: 7763: 7750: 7723: 7708: 7695: 7673: 7627: 7614: 7538: 7359: 7281: 7216: 7183: 7109: 7105: 7090: 7077: 7057: 6942: 6814: 6771: 6683: 6652: 6613: 6609: 6574: 6559: 6546: 6515: 6241: 6193: 6138: 6111: 6021: 6017: 5990: 5922: 5830: 5745: 5693: 5689: 5674: 5661: 5611: 5598: 5537: 4819:10.11646/zootaxa.3869.3.1 4782:10.1007/s13358-011-0017-4 4763:Barbosania gracilirostris 4487:10.1017/S0016756800085563 4182:10.1017/S1755691013000303 3415:10.1016/j.cub.2014.03.030 3312:10.1080/00222931408693521 3145:"Taxonomic review of the 2897:were also found. Several 2867:. Other animals such the 2796:, the targaryendraconian 2663:, the targaryendraconian 2459: 2442: 2435: 2418: 2411: 2394: 2387: 2367: 2360: 2331: 2314: 2307: 2297: 2277: 2270: 2244: 2227: 2220: 2213: 2206: 2189: 2182: 2079: 2062: 2055: 2038: 2031: 2014: 1997: 1990: 1959:Tropeognathus mesembrinus 1955: 1938: 1931: 1918:Siroccopteryx moroccensis 1914: 1907: 1900: 1893: 1830: 1817:Coloborhynchus fluviferox 1813: 1806: 1789: 1772: 1765: 1748: 1741: 1724: 1717: 1710: 1703: 1683: 1676: 1325:Anhanguera blittersdorffi 1093:Siroccopteryx moroccensis 1084:Tropeognathus mesembrinus 980:, forms the larger group 928:. They placed the genera 925:Anhanguera blittersdorffi 696:, and also distinguished 475: 468: 299: 294: 275: 268: 148:Scientific classification 146: 141:National Museum of Brazil 136:Tropeognathus mesembrinus 132: 123: 34: 5269:Coloborhynchus wadleighi 5101:10.1144/gsjgs.153.1.0005 3162:10.3897/zookeys.308.5559 2018:Camposipterus colorhinus 1752:Uktenadactylus wadleighi 1335:Formerly assigned genera 1265:of ornithocheirids have 822:. Unwin also designated 692:was the type species of 7097:Archaeopterodactyloidea 5232:Journal of Paleontology 5167:Encyclopædia Britannica 4932:Tupandactylus imperator 4761:"A new ornithocheirid, 4589:; Walker, C.A. (1973). 4423:Averianov A.O. (2012). 4408:H2VP: Paleobiomechanics 2959:Coloborhychus wadleighi 2721:were also present. The 2001:Camposipterus segwickii 1484:, as a sister taxon to 776:as the type species of 656:as the type species of 5296:Pawpawsaurus campbelli 4887:P. H. Buisonjé. 1980. 4508:Seeley, H. G. (1879). 4427:Ornithostoma sedgwicki 3894:10.4202/app.00751.2020 3710:(suppl 2): e20190768. 3066:Martill, D.M. (2010). 2770: 2590: 1314: 1206: 1071: 1013: 788:revalidated the genus 726:Reginald Walter Hooley 581: 567:Holotype specimens of 10241:Paleobiology Database 5460:Witton, Mark (2013). 5409:Scaphites hippocrepis 5400:William A.C. (1969). 4951:10.4202/app.2010.0057 4400:Dinosaur Revolution: 3220:10.5194/fr-4-189-2001 3201:Unwin, D. M. (2001). 3113:Seeley, H.G. (1870). 2757: 2623:Locomotion and flight 2584:stealing prey from a 2579: 2066:Cimoliopterus cuvieri 2042:Camposipterus nasutus 1793:Coloborhynchus capito 1296: 1196:Restoration of three 1195: 1047: 1030:Coloborhynchus capito 999: 956:phylogenetic analyses 566: 133:Restored skeleton of 10279:Turonian extinctions 6709:Archaeoistiodactylus 6619:Archaeoistiodactylus 4930:"New information on 4611:on February 6, 2012. 1942:Ornithocheirus simus 1776:Coloborhynchus reedi 1611:, and together with 1405:), within the clade 1329:Ornithocheirus simus 1188:Postcranial skeleton 1120:, therefore another 1080:Ornithocheirus simus 920:Ornithocheirus simus 841:Ornithocheirus simus 824:Ornithocheirus simus 784:. In 1994, however, 690:Ornithocheirus simus 640:British pterosaurs: 622:Ornithocheirus simus 570:Ornithocheirus simus 505:within the suborder 9774:Targaryendraconidae 9463:Ornithocheiriformes 9150:Ornithocheiromorpha 8449:Neopterodactyloidea 8376:Dsungaripteromorpha 6144:Campylognathoididae 5864:Eudimorphodontoidea 5314:2016PLoSO..1150845P 5271:from North America" 5265:Lee, Y.-N. (1994). 5201:1998Natur.392..705H 5093:1996JGSoc.153....5M 5010:Cretaceous Research 4903:Cretaceous Research 4868:Cearadactylus atrox 4640:2014PLoSO...984709F 4475:Geological Magazine 4398:Habib, M. (2011). " 4358:2010PLoSO...513982W 4330:, pp. 161–162. 4113:, pp. 159–160. 4074:Cretaceous Research 3947:Cretaceous Research 3923:10.18563/pv.43.1.e4 3762:2019NatSR...9.4940H 3656:2019NatSR...913454P 3592:Cretaceous Research 3464:2019NatSR...917875K 3344:2003GSLSP.217..139U 3084:2010GSLSP.343..287M 2992:and an indetermine 2645:Cambridge Greensand 2083:Cimoliopterus dunni 1160:is that it bores a 806:Cambridge Greensand 601:Harry Govier Seeley 281:Pterodactylus simus 227:Ornithocheiriformes 10289:Pterosaur families 9303:Istiodactyliformes 7777:Lonchognathosaurus 7620:Eupterodactyloidea 6763:Pterodactyliformes 6566:Pterodactylomorpha 6553:Pterodactylomorpha 6467:Pterodactylomorpha 6309:Angustinaripterini 5940:Eudimorphodontinae 5915:Eudimorphodontidae 5445:. Crescent Books. 4715:10.1007/bf03022595 4687:2012-03-17 at the 4273:Biological Reviews 4209:Historical Biology 3833:Historical Biology 3806:has generic name ( 3750:Scientific Reports 3644:Scientific Reports 3452:Scientific Reports 2771: 2769:shown in dark blue 2700:, and the dubious 2591: 1315: 1207: 1072: 1014: 836:Haopterus gracilis 798:Criorhynchus simus 782:Criorhynchus simus 758:Criorhynchus simus 662:Criorhynchus simus 582: 10256: 10255: 10228:Open Tree of Life 10142:Taxon identifiers 10133: 10132: 10129: 10128: 10102: 10101: 10098: 10097: 10094: 10093: 10090: 10089: 10086: 10085: 10082: 10081: 10078: 10077: 10074: 10073: 10070: 10069: 9812: 9811: 9808: 9807: 9729:Targaryendraconia 9723: 9722: 9533: 9532: 9457: 9456: 9453: 9452: 9243: 9242: 9143: 9142: 9139: 9138: 9135: 9134: 9131: 9130: 8900: 8899: 8869: 8868: 8865: 8864: 8861: 8860: 8833: 8832: 8829: 8828: 8825: 8824: 8821: 8820: 8817: 8816: 8813: 8812: 8809: 8808: 8805: 8804: 8576: 8575: 8544:Chaoyangopterinae 8473:Chaoyangopteridae 8333: 8332: 8329: 8328: 8325: 8324: 8321: 8320: 8317: 8316: 8313: 8312: 7848: 7847: 7844: 7843: 7715:Ornithocheiroidea 7702:Ornithocheiroidea 7691: 7690: 7671: 7670: 7667: 7666: 7663: 7662: 7645:Ornithocheiroidea 7610: 7609: 7606: 7605: 7602: 7601: 7598: 7597: 7594: 7593: 7408:Gladocephaloideus 7344: 7343: 7340: 7339: 7306:Aurorazhdarchidae 7208:Ctenochasmatoidea 7168: 7167: 7126:Germanodactylidae 7073: 7072: 7055: 7054: 7051: 7050: 7047: 7046: 7043: 7042: 7039: 7038: 7035: 7034: 6929: 6928: 6904:Dendrorhynchoides 6881:Batrachognathinae 6847:Dendrorhynchoides 6777:Changchengopterus 6756: 6755: 6752: 6751: 6748: 6747: 6542: 6541: 6513: 6512: 6509: 6508: 6505: 6504: 6501: 6500: 6497: 6496: 6493: 6492: 6489: 6488: 6485: 6484: 6401: 6400: 6397: 6396: 6393: 6392: 6318:Angustinaripterus 6234:Rhamphorhynchinae 6186:Rhamphorhynchidae 6169: 6168: 6160:Campylognathoides 6095: 6094: 5974: 5973: 5970: 5969: 5966: 5965: 5873:Raeticodactylidae 5815: 5814: 5811: 5810: 5778:Austriadraconidae 5657: 5656: 5653: 5652: 5649: 5648: 5473:978-0-691-15061-1 5195:(6677): 705–708. 4285:10.1111/brv.12431 3018:were also found. 2963:Paw Paw Formation 2888:, as well as the 2851:, as well as the 2730:, as well as the 2563: 2562: 2555: 2554: 2546: 2545: 2537: 2536: 2528: 2527: 2519: 2518: 2510: 2509: 2501: 2500: 2492: 2491: 2483: 2482: 2474: 2473: 2346: 2345: 2259: 2258: 2157: 2156: 2148: 2147: 2139: 2138: 2130: 2129: 2121: 2120: 2112: 2111: 2103: 2102: 2094: 2093: 1979: 1978: 1970: 1969: 1882: 1881: 1873: 1872: 1864: 1863: 1855: 1854: 1846: 1845: 1407:Targaryendraconia 913:Alexander Kellner 492: 491: 486: 264: 16:(Redirected from 10296: 10249: 10248: 10236: 10235: 10223: 10222: 10210: 10209: 10197: 10196: 10184: 10183: 10182: 10180:Ornithocheiridae 10169: 10168: 10167: 10150:Ornithocheiridae 10137: 10124: 10118: 10112: 10111: 9958:Coloborhynchinae 9886: 9841:Ornithocheiridae 9833:Brasileodactylus 9825: 9735: 9625:Ornithocheirinae 9559: 9554:Ornithocheiridae 9550: 9546: 9504: 9500: 9479:Brasileodactylus 9388: 9308: 9276:Lonchodraconidae 9272: 9268: 9160: 9156: 9058: 9026: 9021:Nyctosauromorpha 8958: 8934: 8930: 8924:Pteranodontoidea 8919: 8911:Pteranodontoidea 8906: 8895: 8894: 8887: 8886: 8879: 8878: 8843:Pteranodontoidea 8839: 8723:Quetzalcoatlinae 8712:Mistralazhdarcho 8677: 8618: 8587: 8478: 8454: 8346: 8247: 8191: 8110: 8095: 8058: 7962:Thalassodromidae 7935: 7924: 7861: 7819:Dsungaripterinae 7761: 7756:Dsungaripteridae 7752: 7748: 7721: 7710: 7697: 7686: 7681: 7680: 7625: 7616: 7536: 7531:Ctenochasmatinae 7513:Plataleorhynchus 7357: 7351:Ctenochasmatidae 7279: 7218: 7214: 7181: 7175:Euctenochasmatia 7107: 7103: 7092: 7079: 7068: 7063: 6991:Propterodactylus 6977:Ningchengopterus 6940: 6935:Pterodactyloidea 6812: 6803: 6769: 6681: 6650: 6611: 6607: 6572: 6561: 6548: 6537: 6531: 6530: 6523: 6522: 6441:Jianchangopterus 6427:Fenghuangopterus 6413: 6359:Rhamphorhynchini 6305: 6239: 6191: 6182: 6140: 6136: 6109: 6038:Dimorphodontidae 6019: 6015: 5988: 5920: 5869: 5828: 5743: 5691: 5687: 5676: 5663: 5609: 5600: 5593: 5592: 5588:Pterosauromorpha 5539: 5524: 5517: 5510: 5501: 5494:Ornithocheiridae 5492:Data related to 5491: 5477: 5456: 5428: 5425: 5419: 5418: 5416: 5397: 5391: 5379: 5373: 5372: 5352: 5346: 5345: 5335: 5325: 5289: 5283: 5282: 5262: 5256: 5255: 5244:10.1666/11-139.1 5227: 5221: 5220: 5184: 5178: 5177: 5175: 5173: 5158: 5152: 5151: 5119: 5113: 5112: 5076: 5067: 5061: 5060: 5040: 5034: 5033: 5016:(5): 1293–1300. 5001: 4995: 4994: 4962: 4956: 4955: 4953: 4925: 4919: 4918: 4898: 4892: 4885: 4879: 4864: 4858: 4857: 4844:Brasileodactylus 4837: 4831: 4830: 4801: 4795: 4794: 4784: 4756: 4750: 4749: 4733: 4727: 4726: 4698: 4692: 4678: 4672: 4671: 4661: 4651: 4619: 4613: 4612: 4610: 4595: 4587:Harrison, C.J.O. 4583: 4577: 4576: 4559:(1–4): 440–444. 4544: 4538: 4537: 4520:(1–4): 591–635. 4505: 4499: 4498: 4466: 4460: 4459: 4433: 4420: 4414: 4396: 4390: 4389: 4379: 4369: 4337: 4331: 4325: 4319: 4313: 4307: 4306: 4296: 4279:(4): 2021–2048. 4264: 4258: 4252: 4243: 4242: 4232: 4200: 4194: 4193: 4176:(3–4): 383–398. 4165: 4156: 4155: 4127: 4114: 4108: 4102: 4096: 4090: 4089: 4071: 4062: 4056: 4055: 4054: 4052:10.1.1.1064.6403 4038: 4032: 4026: 4017: 4016: 4006: 3974: 3963: 3962: 3941: 3935: 3934: 3905: 3899: 3898: 3896: 3872: 3857: 3856: 3839:(8): 1266–1280. 3827: 3812: 3811: 3805: 3801: 3799: 3791: 3781: 3741: 3730: 3729: 3719: 3695: 3686: 3685: 3675: 3635: 3616: 3615: 3583: 3560: 3559: 3549: 3539: 3515: 3496: 3495: 3485: 3475: 3443: 3428: 3427: 3417: 3391: 3374: 3373: 3355: 3327: 3316: 3315: 3291: 3285: 3284: 3268: 3262: 3261: 3251: 3238: 3225: 3224: 3222: 3198: 3185: 3184: 3174: 3164: 3140: 3123: 3122: 3110: 3104: 3103: 3092:10.1144/SP343.18 3063: 3057: 3051: 2823:Brasileodactylus 2572:Diet and feeding 2438: 2414: 2390: 2384:Ornithocheirinae 2363: 2357:Ornithocheiridae 2310: 2300: 2273: 2231:Brasileodactylus 2223: 2216: 2209: 2185: 2173: 2058: 2034: 1993: 1934: 1910: 1903: 1896: 1809: 1768: 1744: 1720: 1713: 1706: 1699:Ornithocheiridae 1679: 1669: 1654: 1434:Pteranodontoidea 1249:plate above its 1232:ornithocheiroids 1212:scapulocoracoids 984:. They assigned 905:eupterodactyloid 854:Brasileodactylus 756:as a synonym of 686:Richard Lydekker 628:to only contain 603:erected the new 507:Pterodactyloidea 495:Ornithocheiridae 481: 454: 443: 432: 422: 411: 400: 389: 383:Ornithocheirinae 381: 370: 359: 348: 337: 326: 315: 304: 278: 259: 256:Ornithocheiridae 254: 241: 225: 211:Pterodactyloidea 209: 196: 156: 155: 128: 118: 55: 38:Temporal range: 32: 21: 18:Ornithocheirinae 10304: 10303: 10299: 10298: 10297: 10295: 10294: 10293: 10269:Pteranodontoids 10259: 10258: 10257: 10252: 10244: 10239: 10231: 10226: 10218: 10213: 10205: 10200: 10192: 10187: 10178: 10177: 10172: 10163: 10162: 10157: 10144: 10134: 10125: 10106: 10066: 10047:Liaoningopterus 10002: 9952: 9890:Tropeognathinae 9875: 9847: 9821: 9804: 9768: 9739:Cimoliopteridae 9719: 9619: 9542: 9529: 9524:Zhenyuanopterus 9498: 9449: 9428:Istiodactylinae 9422: 9403:Longchengpterus 9396:Lingyuanopterus 9383:Istiodactylidae 9377: 9343: 9316:Hongshanopterus 9297: 9266: 9252: 9239: 9202:Hongshanopterus 9187: 9152: 9127: 9100:Barbaridactylus 9085: 9047: 9015: 8962:Pteranodontidae 8947: 8942:Santanadactylus 8926: 8913: 8896: 8889: 8881: 8873: 8857: 8801: 8796:Zhejiangopterus 8717: 8675: 8654:Palaeocursornis 8616: 8595:Montanazhdarcho 8582:Azhdarchiformes 8572: 8553:Chaoyangopterus 8538: 8467: 8443: 8429:Montanazhdarcho 8382: 8342: 8340:Azhdarchomorpha 8309: 8282: 8236: 8199:Aymberedactylus 8180: 8108: 8084: 8072:Aymberedactylus 8047: 8027:Thalassodromeus 7956: 7931: 7920: 7911: 7899:Montanazhdarcho 7857: 7840: 7828:Domeykodactylus 7813: 7807:Tendaguripterus 7744: 7735: 7717: 7704: 7687: 7675: 7659: 7639: 7633:Altmuehlopterus 7590: 7549: 7525: 7476: 7448: 7353: 7336: 7300: 7294:Gallodactylidae 7274:Aurorazhdarchia 7268: 7246:Normannognathus 7222:Gallodactylidae 7210: 7201: 7177: 7164: 7158:Tendaguripterus 7150:Normannognathus 7143:Germanodactylus 7135:Altmuehlopterus 7120: 7099: 7086: 7069: 7031: 7011: 6925: 6890:Batrachognathus 6875: 6870:Vesperopterylus 6825: 6799: 6790: 6765: 6744: 6700:Wukongopterinae 6694: 6676:Wukongopteridae 6670: 6639: 6626:Normannognathus 6603: 6594: 6568: 6555: 6538: 6525: 6517: 6481: 6461: 6434:Jianchangnathus 6417:Scaphognathidae 6407:Digibrevisauria 6389: 6384:Rhamphorhynchus 6353: 6340:Qinglongopterus 6332:Harpactognathus 6300:Rhamphorhynchae 6294: 6289:Qinglongopterus 6268:Harpactognathus 6228: 6222:Scaphognathinae 6214:Parapsicephalus 6178: 6165: 6153:Bergamodactylus 6132: 6123: 6105: 6091: 6069:Parapsicephalus 6032: 6011: 6009:Macronychoptera 6002: 5984: 5962: 5934: 5928:Arcticodactylus 5909: 5897:Raeticodactylus 5858: 5824: 5807: 5787:Arcticodactylus 5772: 5766:Raeticodactylus 5731: 5719:Austriadactylus 5704: 5683: 5670: 5645: 5625: 5619:Avemetatarsalia 5604:Avemetatarsalia 5594: 5579:Avemetatarsalia 5544: 5543: 5533: 5528: 5484: 5474: 5459: 5453: 5440: 5437: 5435:Further reading 5432: 5431: 5426: 5422: 5414: 5399: 5398: 5394: 5380: 5376: 5354: 5353: 5349: 5308:(3): e0150845. 5291: 5290: 5286: 5264: 5263: 5259: 5229: 5228: 5224: 5186: 5185: 5181: 5171: 5169: 5161:Zug, George R. 5160: 5159: 5155: 5121: 5120: 5116: 5074: 5069: 5068: 5064: 5042: 5041: 5037: 5003: 5002: 4998: 4977:(2): e1443273. 4967:Thalassodromeus 4964: 4963: 4959: 4927: 4926: 4922: 4900: 4899: 4895: 4886: 4882: 4865: 4861: 4839: 4838: 4834: 4803: 4802: 4798: 4758: 4757: 4753: 4735: 4734: 4730: 4700: 4699: 4695: 4689:Wayback Machine 4679: 4675: 4621: 4620: 4616: 4608: 4593: 4585: 4584: 4580: 4546: 4545: 4541: 4507: 4506: 4502: 4468: 4467: 4463: 4431: 4422: 4421: 4417: 4397: 4393: 4339: 4338: 4334: 4326: 4322: 4314: 4310: 4266: 4265: 4261: 4253: 4246: 4202: 4201: 4197: 4167: 4166: 4159: 4138:(6): e1014904. 4129: 4128: 4117: 4109: 4105: 4097: 4093: 4069: 4064: 4063: 4059: 4040: 4039: 4035: 4027: 4020: 3976: 3975: 3966: 3943: 3942: 3938: 3911:Palæovertebrata 3907: 3906: 3902: 3874: 3873: 3860: 3829: 3828: 3815: 3802: 3792: 3743: 3742: 3733: 3697: 3696: 3689: 3637: 3636: 3619: 3585: 3584: 3563: 3530:(3): e2001663. 3517: 3516: 3499: 3445: 3444: 3431: 3401:Current Biology 3393: 3392: 3377: 3353:10.1.1.924.5957 3329: 3328: 3319: 3306:(78): 529–557. 3293: 3292: 3288: 3270: 3269: 3265: 3249: 3240: 3239: 3228: 3200: 3199: 3188: 3142: 3141: 3126: 3112: 3111: 3107: 3065: 3064: 3060: 3052: 3041: 3036: 3024: 2858:Thalassodromeus 2853:thalassodromids 2834:Santanadactylus 2817:Araripedactylus 2736:Cetarthrosaurus 2641: 2629:dynamic soaring 2625: 2580:Restoration of 2574: 2569: 2564: 2556: 2547: 2538: 2529: 2520: 2511: 2502: 2493: 2484: 2475: 2347: 2318:Liaoningopterus 2260: 2158: 2149: 2140: 2131: 2122: 2113: 2104: 2095: 1980: 1971: 1883: 1874: 1865: 1856: 1847: 1639:are very high. 1462: 1438:Istiodactylidae 1357:Zhenyuanopterus 1337: 1291: 1236:pterodactyloids 1220:pectoral girdle 1190: 1166:pteranodontians 1126:ornithocheirans 1076:sagittal crests 1042: 1018:pterodactyloids 994: 890:Santanadactylus 794:C. clavirostris 670:C. clavirostris 561: 556: 548:pteranodontians 534:pteranodontians 515:Late Cretaceous 499:ornithocheirids 290: 284: 258: 252: 239: 223: 207: 194: 150: 119: 117: 116: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 50: 49: 47: 36: 35:Ornithocheirids 28: 23: 22: 15: 12: 11: 5: 10302: 10300: 10292: 10291: 10286: 10281: 10276: 10271: 10261: 10260: 10254: 10253: 10251: 10250: 10237: 10224: 10211: 10198: 10185: 10170: 10154: 10152: 10146: 10145: 10140: 10131: 10130: 10127: 10126: 10105: 10103: 10100: 10099: 10096: 10095: 10092: 10091: 10088: 10087: 10084: 10083: 10080: 10079: 10076: 10075: 10072: 10071: 10068: 10067: 10065: 10064: 10057: 10050: 10043: 10036: 10028: 10024:Caulkicephalus 10020: 10012: 10010: 10004: 10003: 10001: 10000: 9997:Uktenadactylus 9993: 9985: 9978: 9975:Coloborhynchus 9971: 9963: 9961: 9954: 9953: 9951: 9950: 9943: 9936: 9928: 9920: 9912: 9904: 9895: 9893: 9883: 9877: 9876: 9874: 9873: 9866: 9858: 9856: 9849: 9848: 9846: 9845: 9837: 9828: 9822: 9817: 9814: 9813: 9810: 9809: 9806: 9805: 9803: 9802: 9799:Targaryendraco 9795: 9787: 9778: 9776: 9770: 9769: 9767: 9766: 9759: 9751: 9743: 9741: 9732: 9725: 9724: 9721: 9720: 9718: 9717: 9713:Uktenadactylus 9709: 9701: 9698:Ornithocheirus 9694: 9686: 9678: 9670: 9662: 9654: 9650:Coloborhynchus 9646: 9638: 9629: 9627: 9621: 9620: 9618: 9617: 9609: 9601: 9593: 9589:Caulkicephalus 9585: 9577: 9574:Arthurdactylus 9570: 9562: 9556: 9543: 9540:Ornithocheirae 9538: 9535: 9534: 9531: 9530: 9528: 9527: 9520: 9512: 9510: 9497: 9496: 9489: 9482: 9475: 9467: 9465: 9459: 9458: 9455: 9454: 9451: 9450: 9448: 9447: 9440: 9432: 9430: 9424: 9423: 9421: 9420: 9413: 9406: 9399: 9391: 9385: 9379: 9378: 9376: 9375: 9368: 9365:Linlongopterus 9361: 9353: 9351: 9349:Mimodactylidae 9345: 9344: 9342: 9341: 9337:Lonchodectidae 9333: 9326: 9323:Linlongopterus 9319: 9311: 9305: 9299: 9298: 9296: 9295: 9288: 9280: 9278: 9265: 9264: 9255: 9253: 9248: 9245: 9244: 9241: 9240: 9238: 9237: 9233:Targaryendraco 9229: 9221: 9214: 9206: 9197: 9195: 9193:Lonchodectidae 9189: 9188: 9186: 9185: 9178: 9171: 9163: 9153: 9148: 9145: 9144: 9141: 9140: 9137: 9136: 9133: 9132: 9129: 9128: 9126: 9125: 9117: 9110: 9107:Muzquizopteryx 9103: 9095: 9093: 9087: 9086: 9084: 9083: 9076: 9069: 9061: 9055: 9053:Aponyctosauria 9049: 9048: 9046: 9045: 9037: 9029: 9023: 9017: 9016: 9014: 9013: 9005: 8997: 8990: 8982: 8975: 8966: 8964: 8955: 8949: 8948: 8946: 8945: 8937: 8927: 8922: 8915: 8914: 8909: 8902: 8901: 8898: 8897: 8872: 8870: 8867: 8866: 8863: 8862: 8859: 8858: 8856: 8855: 8854: 8853: 8847: 8845: 8835: 8834: 8831: 8830: 8827: 8826: 8823: 8822: 8819: 8818: 8815: 8814: 8811: 8810: 8807: 8806: 8803: 8802: 8800: 8799: 8792: 8784: 8781:Thanatosdrakon 8777: 8774:Quetzalcoatlus 8770: 8767:Phosphatodraco 8763: 8756: 8749: 8742: 8739:Arambourgiania 8735: 8727: 8725: 8719: 8718: 8716: 8715: 8708: 8701: 8694: 8685: 8683: 8674: 8673: 8665: 8657: 8650: 8646:Navajodactylus 8642: 8635: 8626: 8624: 8615: 8614: 8606: 8599: 8590: 8584: 8578: 8577: 8574: 8573: 8571: 8570: 8567:Shenzhoupterus 8563: 8556: 8548: 8546: 8540: 8539: 8537: 8536: 8528: 8520: 8513: 8506: 8498: 8494:Argentinadraco 8490: 8481: 8475: 8469: 8468: 8466: 8465: 8457: 8451: 8445: 8444: 8442: 8441: 8433: 8425: 8417: 8409: 8405:Argentinadraco 8401: 8393: 8391: 8384: 8383: 8381: 8380: 8372: 8365: 8358: 8349: 8343: 8338: 8335: 8334: 8331: 8330: 8327: 8326: 8323: 8322: 8319: 8318: 8315: 8314: 8311: 8310: 8308: 8307: 8300: 8292: 8290: 8284: 8283: 8281: 8280: 8273: 8266: 8259: 8250: 8244: 8238: 8237: 8235: 8234: 8227: 8219: 8211: 8203: 8194: 8188: 8182: 8181: 8179: 8178: 8171: 8164: 8161:Nemicolopterus 8157: 8150: 8143: 8135: 8127: 8118: 8116: 8107: 8106: 8098: 8092: 8086: 8085: 8083: 8082: 8075: 8067: 8065: 8055: 8053:Tapejariformes 8049: 8048: 8046: 8045: 8037: 8030: 8023: 8015: 8007: 8000: 7992: 7988:Argentinadraco 7984: 7976: 7967: 7965: 7958: 7957: 7955: 7954: 7946: 7938: 7932: 7929:Tapejaromorpha 7927: 7921: 7916: 7913: 7912: 7910: 7909: 7902: 7895: 7887: 7879: 7872: 7869:Argentinadraco 7864: 7858: 7853: 7850: 7849: 7846: 7845: 7842: 7841: 7839: 7838: 7835:Dsungaripterus 7831: 7823: 7821: 7815: 7814: 7812: 7811: 7803: 7795: 7788: 7781: 7773: 7764: 7758: 7745: 7740: 7737: 7736: 7734: 7733: 7724: 7718: 7713: 7706: 7705: 7700: 7693: 7692: 7689: 7688: 7674: 7672: 7669: 7668: 7665: 7664: 7661: 7660: 7658: 7657: 7656: 7655: 7649: 7647: 7641: 7640: 7638: 7637: 7628: 7622: 7612: 7611: 7608: 7607: 7604: 7603: 7600: 7599: 7596: 7595: 7592: 7591: 7589: 7588: 7581: 7574: 7567: 7559: 7557: 7555:Pterodaustrini 7551: 7550: 7548: 7547: 7539: 7533: 7527: 7526: 7524: 7523: 7516: 7509: 7502: 7494: 7486: 7484: 7482:Gnathosaurinae 7478: 7477: 7475: 7474: 7467: 7459: 7457: 7454:Moganopterinae 7450: 7449: 7447: 7446: 7439: 7432: 7425: 7418: 7411: 7404: 7397: 7390: 7383: 7376: 7373:Balaenognathus 7369: 7360: 7354: 7349: 7346: 7345: 7342: 7341: 7338: 7337: 7335: 7334: 7326: 7323:Aurorazhdarcho 7319: 7310: 7308: 7302: 7301: 7299: 7298: 7290: 7282: 7276: 7270: 7269: 7267: 7266: 7262:Moganopterinae 7258: 7250: 7242: 7235: 7231:Aurorazhdarcho 7226: 7224: 7211: 7206: 7203: 7202: 7200: 7199: 7192: 7184: 7178: 7173: 7170: 7169: 7166: 7165: 7163: 7162: 7154: 7146: 7139: 7130: 7128: 7122: 7121: 7119: 7118: 7110: 7100: 7095: 7088: 7087: 7082: 7075: 7074: 7071: 7070: 7058: 7056: 7053: 7052: 7049: 7048: 7045: 7044: 7041: 7040: 7037: 7036: 7033: 7032: 7030: 7029: 7028: 7027: 7021: 7019: 7013: 7012: 7010: 7009: 7002: 6994: 6987: 6980: 6973: 6966: 6958: 6951: 6943: 6937: 6931: 6930: 6927: 6926: 6924: 6923: 6916: 6908: 6900: 6893: 6885: 6883: 6877: 6876: 6874: 6873: 6866: 6859: 6851: 6843: 6835: 6833: 6831:Anurognathinae 6827: 6826: 6824: 6823: 6815: 6809: 6807:Anurognathidae 6800: 6795: 6792: 6791: 6789: 6788: 6785:Douzhanopterus 6781: 6772: 6766: 6761: 6758: 6757: 6754: 6753: 6750: 6749: 6746: 6745: 6743: 6742: 6735: 6731:Douzhanopterus 6727: 6720: 6713: 6704: 6702: 6696: 6695: 6693: 6692: 6689:Kunpengopterus 6684: 6678: 6672: 6671: 6669: 6668: 6661: 6653: 6647: 6641: 6640: 6638: 6637: 6633:Anurognathidae 6629: 6622: 6614: 6604: 6601:Monofenestrata 6599: 6596: 6595: 6593: 6592: 6584: 6575: 6569: 6564: 6557: 6556: 6551: 6544: 6543: 6540: 6539: 6516: 6514: 6511: 6510: 6507: 6506: 6503: 6502: 6499: 6498: 6495: 6494: 6491: 6490: 6487: 6486: 6483: 6482: 6480: 6479: 6478: 6477: 6471: 6469: 6463: 6462: 6460: 6459: 6451: 6444: 6437: 6430: 6422: 6420: 6410: 6403: 6402: 6399: 6398: 6395: 6394: 6391: 6390: 6388: 6387: 6380: 6372: 6363: 6361: 6355: 6354: 6352: 6351: 6344: 6336: 6328: 6321: 6313: 6311: 6302: 6296: 6295: 6293: 6292: 6285: 6282:Orientognathus 6278: 6271: 6264: 6257: 6250: 6242: 6236: 6230: 6229: 6227: 6226: 6218: 6210: 6203: 6199:Dolicorhamphus 6194: 6188: 6179: 6176:Breviquartossa 6174: 6171: 6170: 6167: 6166: 6164: 6163: 6156: 6148: 6146: 6133: 6128: 6125: 6124: 6122: 6121: 6112: 6106: 6100: 6097: 6096: 6093: 6092: 6090: 6089: 6081: 6073: 6065: 6058: 6055:Caelestiventus 6051: 6042: 6040: 6034: 6033: 6031: 6030: 6022: 6012: 6007: 6004: 6003: 6001: 6000: 5991: 5985: 5981:Zambellisauria 5979: 5976: 5975: 5972: 5971: 5968: 5967: 5964: 5963: 5961: 5960: 5953: 5949:Carniadactylus 5944: 5942: 5936: 5935: 5933: 5932: 5923: 5917: 5911: 5910: 5908: 5907: 5900: 5893: 5886: 5877: 5875: 5866: 5860: 5859: 5857: 5856: 5848: 5840: 5831: 5825: 5820: 5817: 5816: 5813: 5812: 5809: 5808: 5806: 5805: 5802:Seazzadactylus 5798: 5791: 5782: 5780: 5774: 5773: 5771: 5770: 5762: 5755: 5751:Carniadactylus 5746: 5740: 5733: 5732: 5730: 5729: 5722: 5714: 5712: 5706: 5705: 5703: 5702: 5699:Daohugoupterus 5694: 5684: 5679: 5672: 5671: 5666: 5659: 5658: 5655: 5654: 5651: 5650: 5647: 5646: 5644: 5643: 5642: 5641: 5635: 5633: 5627: 5626: 5624: 5623: 5622: 5621: 5612: 5606: 5596: 5595: 5591: 5590: 5581: 5572: 5563: 5557: 5551: 5542: 5535: 5534: 5529: 5527: 5526: 5519: 5512: 5504: 5498: 5497: 5496:at Wikispecies 5483: 5482:External links 5480: 5479: 5478: 5472: 5457: 5451: 5436: 5433: 5430: 5429: 5420: 5405:Scaphites leei 5392: 5374: 5363:(2): 298–312. 5347: 5284: 5257: 5238:(5): 886–905. 5222: 5179: 5153: 5114: 5062: 5051:(2): 151–166. 5035: 4996: 4957: 4944:(3): 567–580. 4920: 4909:(2): 236–243. 4893: 4880: 4859: 4832: 4813:(3): 201–223. 4796: 4775:(2): 259–275. 4751: 4728: 4693: 4673: 4614: 4578: 4539: 4500: 4481:(5): 193–199. 4461: 4415: 4391: 4352:(11): e13982. 4332: 4320: 4318:, p. 162. 4308: 4259: 4257:, p. 163. 4244: 4215:(6): 697–717. 4195: 4157: 4115: 4103: 4101:, p. 159. 4091: 4080:(4): 686–698. 4057: 4033: 4031:, p. 155. 4018: 3997:(1): 113–135. 3985:T. mesembrinus 3964: 3936: 3900: 3858: 3813: 3731: 3687: 3617: 3561: 3497: 3429: 3375: 3338:(1): 139–190. 3317: 3286: 3263: 3245:Coloborhynchus 3226: 3213:(1): 189–221. 3186: 3155:(308): 1–112. 3147:Ornithocheirus 3124: 3105: 3078:(1): 287–311. 3058: 3056:, p. 152. 3038: 3037: 3035: 3032: 3031: 3030: 3028:Pterosaur size 3023: 3020: 2955:Uktenadactylus 2890:crocodylomorph 2814:. The related 2788:Coloborhynchus 2759:Geological map 2742:Platypterygius 2687:ornithischians 2669:lonchodraconid 2661:Coloborhynchus 2653:Ornithocheirus 2640: 2637: 2624: 2621: 2573: 2570: 2568: 2565: 2561: 2560: 2553: 2552: 2549: 2548: 2544: 2543: 2540: 2539: 2535: 2534: 2531: 2530: 2526: 2525: 2522: 2521: 2517: 2516: 2513: 2512: 2508: 2507: 2504: 2503: 2499: 2498: 2495: 2494: 2490: 2489: 2486: 2485: 2481: 2480: 2477: 2476: 2472: 2471: 2468: 2467: 2458: 2455: 2454: 2451: 2450: 2441: 2436: 2434: 2431: 2430: 2427: 2426: 2422:Ornithocheirus 2417: 2412: 2410: 2407: 2406: 2403: 2402: 2398:Coloborhynchus 2393: 2388: 2386: 2380: 2379: 2376: 2375: 2366: 2361: 2359: 2353: 2352: 2349: 2348: 2344: 2343: 2340: 2339: 2330: 2327: 2326: 2323: 2322: 2313: 2308: 2306: 2298: 2296: 2294:Ornithocheirae 2290: 2289: 2286: 2285: 2276: 2271: 2269: 2266: 2265: 2262: 2261: 2257: 2256: 2253: 2252: 2243: 2240: 2239: 2236: 2235: 2226: 2221: 2219: 2214: 2212: 2207: 2205: 2202: 2201: 2198: 2197: 2188: 2183: 2181: 2171: 2159: 2155: 2154: 2151: 2150: 2146: 2145: 2142: 2141: 2137: 2136: 2133: 2132: 2128: 2127: 2124: 2123: 2119: 2118: 2115: 2114: 2110: 2109: 2106: 2105: 2101: 2100: 2097: 2096: 2092: 2091: 2088: 2087: 2078: 2075: 2074: 2071: 2070: 2061: 2056: 2054: 2051: 2050: 2047: 2046: 2037: 2032: 2030: 2027: 2026: 2023: 2022: 2013: 2010: 2009: 2006: 2005: 1996: 1991: 1989: 1986: 1985: 1982: 1981: 1977: 1976: 1973: 1972: 1968: 1967: 1964: 1963: 1954: 1951: 1950: 1947: 1946: 1937: 1932: 1930: 1927: 1926: 1923: 1922: 1913: 1908: 1906: 1901: 1899: 1894: 1892: 1889: 1888: 1885: 1884: 1880: 1879: 1876: 1875: 1871: 1870: 1867: 1866: 1862: 1861: 1858: 1857: 1853: 1852: 1849: 1848: 1844: 1843: 1840: 1839: 1834:Coloborhynchus 1829: 1826: 1825: 1822: 1821: 1812: 1807: 1805: 1802: 1801: 1798: 1797: 1788: 1785: 1784: 1781: 1780: 1771: 1766: 1764: 1761: 1760: 1757: 1756: 1747: 1742: 1740: 1737: 1736: 1733: 1732: 1723: 1718: 1716: 1711: 1709: 1704: 1702: 1694: 1693: 1690: 1689: 1682: 1677: 1675: 1673:Ornithocheirae 1667: 1652: 1642:Below are two 1583:Uktenadactylus 1571:Coloborhynchus 1562:Ornithocheirus 1546:Coloborhynchus 1541:Uktenadactylus 1528:Ornithocheirus 1524:Ornithocheirus 1508:Ornithocheirus 1504:Coloborhynchus 1496:Coloborhynchus 1482:Ornithocheirus 1478:Coloborhynchus 1474:Ornithocheirus 1470:Coloborhynchus 1461: 1458: 1454:Mimodactylidae 1336: 1333: 1290: 1289:Classification 1287: 1261:, the slender 1259:istiodactylids 1189: 1186: 1162:frontoparietal 1158:Caulkicephalus 1154:Caulkicephalus 1150:Caulkicephalus 1138:Coloborhynchus 1130:Coloborhynchus 1118:T. mesembrinus 1110:S. moroccensis 1098:synapomorphies 1067:Uktenadactylus 1064:(E and F) and 1055:Caulkicephalus 1041: 1038: 1016:Among toothed 1006:(A and C) and 1003:Ornithocheirus 993: 990: 986:Ornithocheirus 973:Caulkicephalus 960:Coloborhynchus 954:in 2013. Some 934:Ornithocheirus 930:Coloborhynchus 884:Arthurdactylus 873:Ornithocheirus 859:Coloborhynchus 828:Ornithocheirus 814:Coloborhynchus 810:Ornithocheirus 790:Coloborhynchus 778:Coloborhynchus 770:Ornithocheirus 734:Ornithocheirus 730:Ornithocheirus 722:Ornithocheirus 706:Ornithocheirus 694:Ornithocheirus 682:Ornithocheirus 678:junior synonym 666:Coloborhynchus 643:Coloborhynchus 630:Ornithocheirus 626:Ornithocheirae 609:Ornithocheirus 578:O. platyrhinus 575:junior synonym 560: 557: 555: 552: 543:Quetzalcoatlus 530:istiodactylids 490: 489: 488: 487: 479:Criorhynchidae 473: 472: 466: 465: 464: 463: 462: 461: 457:Uktenadactylus 450: 439: 428: 425:Ornithocheirus 418: 407: 396: 392:Coloborhynchus 377: 366: 355: 344: 340:Caulkicephalus 333: 322: 318:Arthurdactylus 311: 297: 296: 292: 291: 285: 273: 272: 266: 265: 250: 246: 245: 243:Ornithocheirae 237: 230: 229: 221: 214: 213: 205: 201: 200: 192: 188: 187: 182: 178: 177: 172: 168: 167: 162: 158: 157: 144: 143: 130: 129: 121: 120: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 56: 37: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 10301: 10290: 10287: 10285: 10282: 10280: 10277: 10275: 10272: 10270: 10267: 10266: 10264: 10247: 10242: 10238: 10234: 10229: 10225: 10221: 10216: 10212: 10208: 10203: 10199: 10195: 10190: 10186: 10181: 10175: 10171: 10166: 10160: 10156: 10155: 10153: 10151: 10147: 10143: 10138: 10123: 10119: 10117: 10110: 10063: 10062: 10061:Maaradactylus 10058: 10056: 10055: 10051: 10049: 10048: 10044: 10042: 10041: 10037: 10034: 10033: 10032:Cearadactylus 10029: 10026: 10025: 10021: 10019: 10018: 10014: 10013: 10011: 10009: 10008:Anhanguerinae 10005: 9999: 9998: 9994: 9991: 9990: 9989:Siroccopteryx 9986: 9984: 9983: 9979: 9977: 9976: 9972: 9970: 9969: 9965: 9964: 9962: 9959: 9955: 9949: 9948: 9947:Tropeognathus 9944: 9942: 9941: 9937: 9934: 9933: 9932:Siroccopteryx 9929: 9926: 9925: 9921: 9918: 9917: 9913: 9910: 9909: 9905: 9902: 9901: 9897: 9896: 9894: 9891: 9887: 9884: 9882: 9881:Anhangueridae 9878: 9872: 9871: 9870:Iberodactylus 9867: 9865: 9864: 9860: 9859: 9857: 9854: 9850: 9843: 9842: 9838: 9835: 9834: 9830: 9829: 9826: 9823: 9820: 9815: 9801: 9800: 9796: 9793: 9792: 9788: 9785: 9784: 9780: 9779: 9777: 9775: 9771: 9765: 9764: 9763:Cimoliopterus 9760: 9757: 9756: 9755:Camposipterus 9752: 9750: 9749: 9745: 9744: 9742: 9740: 9736: 9733: 9730: 9726: 9715: 9714: 9710: 9707: 9706: 9705:Siroccopteryx 9702: 9700: 9699: 9695: 9692: 9691: 9687: 9684: 9683: 9679: 9676: 9675: 9671: 9668: 9667: 9663: 9660: 9659: 9655: 9652: 9651: 9647: 9644: 9643: 9642:Camposipterus 9639: 9636: 9635: 9631: 9630: 9628: 9626: 9622: 9615: 9614: 9613:Tropeognathus 9610: 9607: 9606: 9602: 9599: 9598: 9597:Cimoliopterus 9594: 9591: 9590: 9586: 9583: 9582: 9581:Camposipterus 9578: 9576: 9575: 9571: 9569: 9568: 9567:Araripesaurus 9564: 9563: 9560: 9557: 9555: 9551: 9547: 9544: 9541: 9536: 9526: 9525: 9521: 9519: 9518: 9514: 9513: 9511: 9509: 9508:Boreopteridae 9505: 9501: 9495: 9494: 9490: 9488: 9487: 9486:Cearadactylus 9483: 9481: 9480: 9476: 9474: 9473: 9469: 9468: 9466: 9464: 9460: 9446: 9445: 9441: 9439: 9438: 9437:Istiodactylus 9434: 9433: 9431: 9429: 9425: 9419: 9418: 9414: 9412: 9411: 9407: 9405: 9404: 9400: 9398: 9397: 9393: 9392: 9389: 9386: 9384: 9380: 9374: 9373: 9369: 9367: 9366: 9362: 9360: 9359: 9355: 9354: 9352: 9350: 9346: 9339: 9338: 9334: 9332: 9331: 9330:Yixianopterus 9327: 9325: 9324: 9320: 9318: 9317: 9313: 9312: 9309: 9306: 9304: 9300: 9294: 9293: 9289: 9287: 9286: 9282: 9281: 9279: 9277: 9273: 9269: 9262: 9261: 9257: 9256: 9254: 9251: 9246: 9235: 9234: 9230: 9227: 9226: 9222: 9220: 9219: 9215: 9212: 9211: 9207: 9204: 9203: 9199: 9198: 9196: 9194: 9190: 9184: 9183: 9179: 9177: 9176: 9172: 9170: 9169: 9165: 9164: 9161: 9157: 9154: 9151: 9146: 9123: 9122: 9118: 9116: 9115: 9111: 9109: 9108: 9104: 9102: 9101: 9097: 9096: 9094: 9092: 9091:Nyctosauridae 9088: 9082: 9081: 9077: 9075: 9074: 9070: 9068: 9067: 9063: 9062: 9059: 9056: 9054: 9050: 9043: 9042: 9038: 9036: 9035: 9034:Alamodactylus 9031: 9030: 9027: 9024: 9022: 9018: 9011: 9010: 9006: 9003: 9002: 8998: 8996: 8995: 8991: 8988: 8987: 8983: 8981: 8980: 8976: 8973: 8972: 8968: 8967: 8965: 8963: 8959: 8956: 8954: 8953:Pteranodontia 8950: 8944: 8943: 8939: 8938: 8935: 8931: 8928: 8925: 8920: 8916: 8912: 8907: 8903: 8893: 8888: 8885: 8877: 8851: 8850: 8849: 8848: 8846: 8844: 8840: 8836: 8798: 8797: 8793: 8790: 8789: 8788:Wellnhopterus 8785: 8783: 8782: 8778: 8776: 8775: 8771: 8769: 8768: 8764: 8762: 8761: 8760:Hatzegopteryx 8757: 8755: 8754: 8750: 8748: 8747: 8743: 8741: 8740: 8736: 8734: 8733: 8732:Aralazhdarcho 8729: 8728: 8726: 8724: 8720: 8714: 8713: 8709: 8707: 8706: 8702: 8700: 8699: 8695: 8692: 8691: 8687: 8686: 8684: 8682: 8678: 8671: 8670: 8666: 8663: 8662: 8658: 8656: 8655: 8651: 8648: 8647: 8643: 8641: 8640: 8636: 8633: 8632: 8628: 8627: 8625: 8623: 8619: 8612: 8611: 8607: 8605: 8604: 8603:Radiodactylus 8600: 8597: 8596: 8592: 8591: 8588: 8585: 8583: 8579: 8569: 8568: 8564: 8562: 8561: 8557: 8555: 8554: 8550: 8549: 8547: 8545: 8541: 8534: 8533: 8529: 8526: 8525: 8521: 8519: 8518: 8514: 8512: 8511: 8507: 8504: 8503: 8499: 8496: 8495: 8491: 8488: 8487: 8486:Apatorhamphus 8483: 8482: 8479: 8476: 8474: 8470: 8464: 8463: 8459: 8458: 8455: 8452: 8450: 8446: 8439: 8438: 8434: 8431: 8430: 8426: 8423: 8422: 8418: 8415: 8414: 8410: 8407: 8406: 8402: 8400: 8399: 8395: 8394: 8392: 8389: 8385: 8378: 8377: 8373: 8371: 8370: 8366: 8364: 8363: 8359: 8356: 8355: 8351: 8350: 8347: 8344: 8341: 8336: 8306: 8305: 8301: 8299: 8298: 8294: 8293: 8291: 8289: 8285: 8279: 8278: 8277:Tupandactylus 8274: 8272: 8271: 8267: 8265: 8264: 8260: 8257: 8256: 8252: 8251: 8248: 8245: 8243: 8239: 8233: 8232: 8228: 8225: 8224: 8220: 8217: 8216: 8212: 8209: 8208: 8207:Caupedactylus 8204: 8201: 8200: 8196: 8195: 8192: 8189: 8187: 8183: 8177: 8176: 8172: 8170: 8169: 8165: 8163: 8162: 8158: 8156: 8155: 8151: 8149: 8148: 8144: 8141: 8140: 8136: 8133: 8132: 8128: 8125: 8124: 8120: 8119: 8117: 8115: 8111: 8105: 8104: 8100: 8099: 8096: 8093: 8091: 8087: 8081: 8080: 8079:Caupedactylus 8076: 8074: 8073: 8069: 8068: 8066: 8063: 8062:Caupedactylia 8059: 8056: 8054: 8050: 8043: 8042: 8038: 8036: 8035: 8031: 8029: 8028: 8024: 8021: 8020: 8016: 8013: 8012: 8008: 8006: 8005: 8001: 7998: 7997: 7993: 7990: 7989: 7985: 7982: 7981: 7977: 7974: 7973: 7969: 7968: 7966: 7963: 7959: 7952: 7951: 7947: 7945: 7944: 7940: 7939: 7936: 7933: 7930: 7925: 7922: 7919: 7918:Neoazhdarchia 7914: 7908: 7907: 7903: 7901: 7900: 7896: 7893: 7892: 7888: 7885: 7884: 7880: 7878: 7877: 7873: 7871: 7870: 7866: 7865: 7862: 7859: 7856: 7855:Azhdarchoidea 7851: 7837: 7836: 7832: 7830: 7829: 7825: 7824: 7822: 7820: 7816: 7809: 7808: 7804: 7801: 7800: 7799:Puntanipterus 7796: 7794: 7793: 7789: 7787: 7786: 7782: 7779: 7778: 7774: 7771: 7770: 7766: 7765: 7762: 7759: 7757: 7753: 7749: 7746: 7743: 7738: 7731: 7730: 7726: 7725: 7722: 7719: 7716: 7711: 7707: 7703: 7698: 7694: 7685: 7679: 7653: 7652: 7651: 7650: 7648: 7646: 7642: 7635: 7634: 7630: 7629: 7626: 7623: 7621: 7617: 7613: 7587: 7586: 7582: 7580: 7579: 7575: 7573: 7572: 7568: 7566: 7565: 7564:Beipiaopterus 7561: 7560: 7558: 7556: 7552: 7546: 7545: 7541: 7540: 7537: 7534: 7532: 7528: 7522: 7521: 7520:Tacuadactylus 7517: 7515: 7514: 7510: 7508: 7507: 7503: 7500: 7499: 7495: 7493: 7492: 7488: 7487: 7485: 7483: 7479: 7473: 7472: 7468: 7466: 7465: 7461: 7460: 7458: 7455: 7451: 7445: 7444: 7440: 7438: 7437: 7433: 7431: 7430: 7426: 7424: 7423: 7419: 7417: 7416: 7412: 7410: 7409: 7405: 7403: 7402: 7401:Forfexopterus 7398: 7396: 7395: 7394:Elanodactylus 7391: 7389: 7388: 7387:Cratonopterus 7384: 7382: 7381: 7380:Cathayopterus 7377: 7375: 7374: 7370: 7367: 7366: 7365:Ardeadactylus 7362: 7361: 7358: 7355: 7352: 7347: 7332: 7331: 7327: 7325: 7324: 7320: 7317: 7316: 7315:Ardeadactylus 7312: 7311: 7309: 7307: 7303: 7296: 7295: 7291: 7289: 7288: 7284: 7283: 7280: 7277: 7275: 7271: 7264: 7263: 7259: 7256: 7255: 7251: 7248: 7247: 7243: 7241: 7240: 7239:Cycnorhamphus 7236: 7233: 7232: 7228: 7227: 7225: 7223: 7219: 7215: 7212: 7209: 7204: 7198: 7197: 7196:Pterodactylus 7193: 7191: 7190: 7189:Diopecephalus 7186: 7185: 7182: 7179: 7176: 7171: 7160: 7159: 7155: 7152: 7151: 7147: 7145: 7144: 7140: 7137: 7136: 7132: 7131: 7129: 7127: 7123: 7117: 7116: 7115:Prejanopterus 7112: 7111: 7108: 7104: 7101: 7098: 7093: 7089: 7085: 7080: 7076: 7067: 7062: 7025: 7024: 7023: 7022: 7020: 7018: 7014: 7008: 7007: 7003: 7000: 6999: 6995: 6993: 6992: 6988: 6986: 6985: 6981: 6979: 6978: 6974: 6972: 6971: 6967: 6964: 6963: 6962:Herbstosaurus 6959: 6957: 6956: 6955:Eurolimnornis 6952: 6950: 6949: 6948:Dermodactylus 6945: 6944: 6941: 6938: 6936: 6932: 6922: 6921: 6917: 6914: 6913: 6909: 6906: 6905: 6901: 6899: 6898: 6894: 6892: 6891: 6887: 6886: 6884: 6882: 6878: 6872: 6871: 6867: 6865: 6864: 6860: 6857: 6856: 6852: 6849: 6848: 6844: 6842: 6841: 6837: 6836: 6834: 6832: 6828: 6822: 6821: 6817: 6816: 6813: 6810: 6808: 6804: 6801: 6798: 6797:Caelidracones 6793: 6787: 6786: 6782: 6779: 6778: 6774: 6773: 6770: 6767: 6764: 6759: 6741: 6740: 6739:Wukongopterus 6736: 6733: 6732: 6728: 6726: 6725: 6724:Darwinopterus 6721: 6719: 6718: 6717:Cuspicephalus 6714: 6711: 6710: 6706: 6705: 6703: 6701: 6697: 6691: 6690: 6686: 6685: 6682: 6679: 6677: 6673: 6667: 6666: 6665:Pterorhynchus 6662: 6660: 6659: 6655: 6654: 6651: 6648: 6646: 6642: 6635: 6634: 6630: 6628: 6627: 6623: 6621: 6620: 6616: 6615: 6612: 6608: 6605: 6602: 6597: 6590: 6589: 6585: 6582: 6581: 6577: 6576: 6573: 6570: 6567: 6562: 6558: 6554: 6549: 6545: 6536: 6532: 6529: 6521: 6475: 6474: 6473: 6472: 6470: 6468: 6464: 6457: 6456: 6452: 6450: 6449: 6448:Scaphognathus 6445: 6443: 6442: 6438: 6436: 6435: 6431: 6429: 6428: 6424: 6423: 6421: 6418: 6414: 6411: 6408: 6404: 6386: 6385: 6381: 6378: 6377: 6373: 6370: 6369: 6365: 6364: 6362: 6360: 6356: 6350: 6349: 6345: 6342: 6341: 6337: 6334: 6333: 6329: 6327: 6326: 6322: 6320: 6319: 6315: 6314: 6312: 6310: 6306: 6303: 6301: 6297: 6291: 6290: 6286: 6284: 6283: 6279: 6277: 6276: 6272: 6270: 6269: 6265: 6263: 6262: 6258: 6256: 6255: 6251: 6249: 6248: 6244: 6243: 6240: 6237: 6235: 6231: 6224: 6223: 6219: 6216: 6215: 6211: 6209: 6208: 6204: 6201: 6200: 6196: 6195: 6192: 6189: 6187: 6183: 6180: 6177: 6172: 6162: 6161: 6157: 6155: 6154: 6150: 6149: 6147: 6145: 6141: 6137: 6134: 6131: 6126: 6119: 6118: 6117:Eudimorphodon 6114: 6113: 6110: 6107: 6103: 6098: 6087: 6086: 6082: 6079: 6078: 6077:Peteinosaurus 6074: 6071: 6070: 6066: 6064: 6063: 6059: 6057: 6056: 6052: 6049: 6048: 6044: 6043: 6041: 6039: 6035: 6029: 6028: 6027:Herbstosaurus 6024: 6023: 6020: 6016: 6013: 6010: 6005: 5998: 5997: 5996:Peteinosaurus 5993: 5992: 5989: 5986: 5982: 5977: 5959: 5958: 5957:Eudimorphodon 5954: 5951: 5950: 5946: 5945: 5943: 5941: 5937: 5930: 5929: 5925: 5924: 5921: 5918: 5916: 5912: 5906: 5905: 5901: 5899: 5898: 5894: 5892: 5891: 5887: 5884: 5883: 5879: 5878: 5876: 5874: 5870: 5867: 5865: 5861: 5854: 5853: 5852:Preondactylia 5849: 5846: 5845: 5844:Peteinosaurus 5841: 5838: 5837: 5833: 5832: 5829: 5826: 5823: 5822:Eopterosauria 5818: 5804: 5803: 5799: 5797: 5796: 5792: 5789: 5788: 5784: 5783: 5781: 5779: 5775: 5768: 5767: 5763: 5761: 5760: 5756: 5753: 5752: 5748: 5747: 5744: 5741: 5738: 5734: 5728: 5727: 5726:Preondactylus 5723: 5721: 5720: 5716: 5715: 5713: 5711: 5710:Preondactylia 5707: 5701: 5700: 5696: 5695: 5692: 5688: 5685: 5682: 5677: 5673: 5669: 5664: 5660: 5639: 5638: 5637: 5636: 5634: 5632: 5628: 5620: 5616: 5615: 5614: 5613: 5610: 5607: 5605: 5601: 5597: 5589: 5585: 5582: 5580: 5576: 5573: 5571: 5567: 5564: 5562: 5558: 5556: 5552: 5550: 5546: 5545: 5540: 5536: 5532: 5525: 5520: 5518: 5513: 5511: 5506: 5505: 5502: 5495: 5490: 5486: 5485: 5481: 5475: 5469: 5465: 5464: 5458: 5454: 5452:0-517-03701-7 5448: 5444: 5439: 5438: 5434: 5424: 5421: 5413: 5412: 5408: 5404: 5396: 5393: 5389: 5388:0-520-24209-2 5385: 5378: 5375: 5370: 5366: 5362: 5358: 5351: 5348: 5343: 5339: 5334: 5329: 5324: 5319: 5315: 5311: 5307: 5303: 5299: 5297: 5288: 5285: 5281:(4): 755–763. 5280: 5276: 5275:Palaeontology 5272: 5270: 5261: 5258: 5253: 5249: 5245: 5241: 5237: 5233: 5226: 5223: 5218: 5214: 5210: 5209:10.1038/33669 5206: 5202: 5198: 5194: 5190: 5183: 5180: 5172:September 18, 5168: 5164: 5157: 5154: 5149: 5145: 5141: 5137: 5133: 5129: 5125: 5124:Araripesuchus 5118: 5115: 5110: 5106: 5102: 5098: 5094: 5090: 5086: 5082: 5081: 5073: 5066: 5063: 5058: 5054: 5050: 5046: 5039: 5036: 5031: 5027: 5023: 5019: 5015: 5011: 5007: 5000: 4997: 4992: 4988: 4984: 4980: 4976: 4972: 4968: 4961: 4958: 4952: 4947: 4943: 4939: 4935: 4933: 4924: 4921: 4916: 4912: 4908: 4904: 4897: 4894: 4890: 4884: 4881: 4877: 4873: 4869: 4863: 4860: 4855: 4851: 4847: 4845: 4836: 4833: 4828: 4824: 4820: 4816: 4812: 4808: 4800: 4797: 4792: 4788: 4783: 4778: 4774: 4770: 4766: 4764: 4755: 4752: 4748:(1): 131–136. 4747: 4743: 4739: 4732: 4729: 4724: 4720: 4716: 4712: 4708: 4704: 4697: 4694: 4690: 4686: 4683: 4677: 4674: 4669: 4665: 4660: 4655: 4650: 4645: 4641: 4637: 4634:(1): e84709. 4633: 4629: 4625: 4618: 4615: 4607: 4603: 4599: 4598:Palaeontology 4592: 4588: 4582: 4579: 4574: 4570: 4566: 4562: 4558: 4554: 4550: 4543: 4540: 4535: 4531: 4527: 4523: 4519: 4515: 4511: 4504: 4501: 4496: 4492: 4488: 4484: 4480: 4476: 4472: 4471:Acanthopholis 4465: 4462: 4457: 4453: 4449: 4445: 4441: 4437: 4430: 4428: 4419: 4416: 4413: 4409: 4405: 4403: 4395: 4392: 4387: 4383: 4378: 4373: 4368: 4363: 4359: 4355: 4351: 4347: 4343: 4336: 4333: 4329: 4324: 4321: 4317: 4312: 4309: 4304: 4300: 4295: 4290: 4286: 4282: 4278: 4274: 4270: 4263: 4260: 4256: 4251: 4249: 4245: 4240: 4236: 4231: 4226: 4222: 4218: 4214: 4210: 4206: 4199: 4196: 4191: 4187: 4183: 4179: 4175: 4171: 4164: 4162: 4158: 4153: 4149: 4145: 4141: 4137: 4133: 4126: 4124: 4122: 4120: 4116: 4112: 4107: 4104: 4100: 4095: 4092: 4087: 4083: 4079: 4075: 4068: 4061: 4058: 4053: 4048: 4044: 4037: 4034: 4030: 4025: 4023: 4019: 4014: 4010: 4005: 4000: 3996: 3992: 3988: 3986: 3982: 3981:Tropeognathus 3973: 3971: 3969: 3965: 3960: 3956: 3952: 3948: 3940: 3937: 3932: 3928: 3924: 3920: 3916: 3912: 3904: 3901: 3895: 3890: 3886: 3882: 3878: 3871: 3869: 3867: 3865: 3863: 3859: 3854: 3850: 3846: 3842: 3838: 3834: 3826: 3824: 3822: 3820: 3818: 3814: 3809: 3797: 3789: 3785: 3780: 3775: 3771: 3767: 3763: 3759: 3755: 3751: 3747: 3740: 3738: 3736: 3732: 3727: 3723: 3718: 3713: 3709: 3705: 3701: 3694: 3692: 3688: 3683: 3679: 3674: 3669: 3665: 3661: 3657: 3653: 3649: 3645: 3641: 3634: 3632: 3630: 3628: 3626: 3624: 3622: 3618: 3613: 3609: 3605: 3601: 3597: 3593: 3589: 3582: 3580: 3578: 3576: 3574: 3572: 3570: 3568: 3566: 3562: 3557: 3553: 3548: 3543: 3538: 3533: 3529: 3525: 3521: 3514: 3512: 3510: 3508: 3506: 3504: 3502: 3498: 3493: 3489: 3484: 3479: 3474: 3469: 3465: 3461: 3457: 3453: 3449: 3442: 3440: 3438: 3436: 3434: 3430: 3425: 3421: 3416: 3411: 3408:(9): 1011–6. 3407: 3403: 3402: 3397: 3390: 3388: 3386: 3384: 3382: 3380: 3376: 3371: 3367: 3363: 3359: 3354: 3349: 3345: 3341: 3337: 3333: 3326: 3324: 3322: 3318: 3313: 3309: 3305: 3301: 3297: 3290: 3287: 3282: 3278: 3274: 3267: 3264: 3259: 3255: 3248: 3246: 3237: 3235: 3233: 3231: 3227: 3221: 3216: 3212: 3208: 3207:Fossil Record 3204: 3197: 3195: 3193: 3191: 3187: 3182: 3178: 3173: 3168: 3163: 3158: 3154: 3150: 3148: 3139: 3137: 3135: 3133: 3131: 3129: 3125: 3120: 3116: 3109: 3106: 3101: 3097: 3093: 3089: 3085: 3081: 3077: 3073: 3069: 3062: 3059: 3055: 3050: 3048: 3046: 3044: 3040: 3033: 3029: 3026: 3025: 3021: 3019: 3017: 3016: 3011: 3010: 3005: 3004: 2999: 2995: 2991: 2990: 2985: 2984: 2979: 2976: 2975:ankylosaurian 2972: 2971:United States 2968: 2964: 2960: 2956: 2951: 2949: 2948: 2943: 2942: 2937: 2936: 2931: 2930: 2925: 2924: 2919: 2917: 2912: 2911: 2906: 2905: 2904:Santanachelys 2900: 2896: 2895: 2894:Araripesuchus 2891: 2887: 2886: 2885:Santanaraptor 2881: 2880: 2875: 2874: 2870: 2866: 2865: 2860: 2859: 2854: 2850: 2849: 2845: 2841: 2840: 2835: 2831: 2830: 2829:Cearadactylus 2825: 2824: 2819: 2818: 2813: 2812: 2811:Maaradactylus 2807: 2806: 2801: 2800: 2795: 2794: 2793:Araripesaurus 2789: 2785: 2784:Tropeognathus 2780: 2776: 2768: 2767:Santana Group 2764: 2763:Araripe Basin 2760: 2756: 2752: 2750: 2749: 2744: 2743: 2738: 2737: 2733: 2729: 2728: 2724: 2720: 2719: 2718:Macrurosaurus 2715: 2711: 2710: 2705: 2704: 2703:Eucercosaurus 2699: 2698: 2697:Acanthopholis 2693: 2692: 2688: 2684: 2683: 2679: 2675: 2674: 2670: 2666: 2665:Camposipterus 2662: 2658: 2654: 2650: 2646: 2638: 2636: 2634: 2630: 2622: 2620: 2618: 2613: 2609: 2605: 2601: 2596: 2589: 2588: 2583: 2582:Cimoliopterus 2578: 2571: 2566: 2559: 2551: 2550: 2542: 2541: 2533: 2532: 2524: 2523: 2515: 2514: 2506: 2505: 2497: 2496: 2488: 2487: 2479: 2478: 2470: 2469: 2466: 2465: 2464: 2457: 2456: 2453: 2452: 2449: 2448: 2447: 2440: 2439: 2433: 2432: 2429: 2428: 2425: 2424: 2423: 2416: 2415: 2409: 2408: 2405: 2404: 2401: 2400: 2399: 2392: 2391: 2385: 2382: 2381: 2378: 2377: 2374: 2373: 2372: 2371:Tropeognathus 2365: 2364: 2358: 2355: 2354: 2351: 2350: 2342: 2341: 2338: 2337: 2336: 2329: 2328: 2325: 2324: 2321: 2320: 2319: 2312: 2311: 2305: 2304:Anhangueridae 2302: 2301: 2295: 2292: 2291: 2288: 2287: 2284: 2283: 2282: 2281:Cearadactylus 2275: 2274: 2268: 2267: 2264: 2263: 2255: 2254: 2251: 2250: 2249: 2242: 2241: 2238: 2237: 2234: 2233: 2232: 2225: 2224: 2218: 2217: 2211: 2210: 2204: 2203: 2200: 2199: 2196: 2195: 2194: 2187: 2186: 2179: 2175: 2174: 2170: 2168: 2164: 2160: 2153: 2152: 2144: 2143: 2135: 2134: 2126: 2125: 2117: 2116: 2108: 2107: 2099: 2098: 2090: 2089: 2086: 2085: 2084: 2077: 2076: 2073: 2072: 2069: 2068: 2067: 2060: 2059: 2053: 2052: 2049: 2048: 2045: 2044: 2043: 2036: 2035: 2029: 2028: 2025: 2024: 2021: 2020: 2019: 2012: 2011: 2008: 2007: 2004: 2003: 2002: 1995: 1994: 1988: 1987: 1984: 1983: 1975: 1974: 1966: 1965: 1962: 1961: 1960: 1953: 1952: 1949: 1948: 1945: 1944: 1943: 1936: 1935: 1929: 1928: 1925: 1924: 1921: 1920: 1919: 1912: 1911: 1905: 1904: 1898: 1897: 1891: 1890: 1887: 1886: 1878: 1877: 1869: 1868: 1860: 1859: 1851: 1850: 1842: 1841: 1838: 1836: 1835: 1828: 1827: 1824: 1823: 1820: 1819: 1818: 1811: 1810: 1804: 1803: 1800: 1799: 1796: 1795: 1794: 1787: 1786: 1783: 1782: 1779: 1778: 1777: 1770: 1769: 1763: 1762: 1759: 1758: 1755: 1754: 1753: 1746: 1745: 1739: 1738: 1735: 1734: 1731: 1730: 1729: 1722: 1721: 1715: 1714: 1708: 1707: 1700: 1696: 1695: 1692: 1691: 1688: 1687: 1686:Anhangueridae 1681: 1680: 1674: 1671: 1670: 1666: 1664: 1660: 1656: 1655: 1651: 1649: 1645: 1640: 1638: 1634: 1630: 1629:C. colorhinus 1626: 1622: 1621:Camposipterus 1618: 1617:Cimoliopterus 1614: 1613:Camposipterus 1610: 1606: 1605:Cimoliopterus 1602: 1598: 1597:Cimoliopterus 1594: 1592: 1588: 1584: 1580: 1579:Tropeognathus 1576: 1575:Siroccopteryx 1572: 1567: 1566:Tropeognathus 1563: 1559: 1558:Siroccopteryx 1555: 1554:Camposipterus 1551: 1550:Cimoliopterus 1547: 1544:with several 1543: 1542: 1537: 1536:Siroccopteryx 1533: 1532:Tropeognathus 1529: 1525: 1521: 1520: 1515: 1514: 1509: 1505: 1501: 1500:Tropeognathus 1497: 1493: 1492: 1491:Siroccopteryx 1487: 1486:Tropeognathus 1483: 1479: 1475: 1471: 1467: 1466:Tropeognathus 1459: 1457: 1455: 1451: 1447: 1443: 1439: 1435: 1431: 1427: 1423: 1419: 1414: 1412: 1408: 1404: 1403:Cimoliopterus 1400: 1399: 1398:Camposipterus 1394: 1390: 1386: 1382: 1381: 1380:Cimoliopterus 1376: 1371: 1370: 1365: 1363: 1359: 1358: 1353: 1349: 1348:Boreopteridae 1345: 1344: 1334: 1332: 1330: 1326: 1321: 1320:nomenclatural 1312: 1311: 1306: 1305: 1300: 1295: 1288: 1286: 1284: 1280: 1276: 1272: 1271:femoral shaft 1268: 1267:femoral heads 1264: 1260: 1256: 1252: 1251:neural spines 1248: 1244: 1239: 1237: 1233: 1229: 1225: 1221: 1217: 1213: 1205: 1201: 1200: 1199:Tropeognathus 1194: 1187: 1185: 1183: 1179: 1178: 1173: 1172: 1167: 1163: 1159: 1155: 1151: 1147: 1143: 1139: 1135: 1131: 1127: 1123: 1119: 1115: 1111: 1107: 1106:anterolateral 1103: 1099: 1095: 1094: 1089: 1085: 1081: 1077: 1069: 1068: 1063: 1062: 1061:Tropeognathus 1057: 1056: 1051: 1050:anhanguerians 1046: 1039: 1037: 1035: 1034:Tropeognathus 1031: 1027: 1023: 1019: 1011: 1010: 1009:Tropeognathus 1005: 1004: 998: 991: 989: 987: 983: 979: 975: 974: 969: 965: 964:Tropeognathus 961: 957: 953: 949: 948:Anhangueridae 945: 941: 940: 939:Tropeognathus 935: 931: 927: 926: 921: 916: 914: 910: 906: 902: 898: 897: 892: 891: 886: 885: 880: 879: 878:Araripesaurus 874: 870: 869: 864: 860: 856: 855: 850: 846: 842: 838: 837: 831: 829: 825: 821: 820: 815: 811: 807: 803: 799: 795: 791: 787: 786:Yuong-Nam Lee 783: 779: 775: 771: 767: 763: 759: 755: 751: 747: 746: 741: 740: 735: 731: 727: 723: 719: 715: 711: 707: 703: 699: 695: 691: 687: 683: 679: 675: 671: 667: 663: 659: 655: 651: 650: 645: 644: 639: 635: 631: 627: 623: 619: 615: 614:Ancient Greek 611: 610: 606: 602: 598: 594: 593: 592:Pterodactylus 588: 579: 576: 573:(A to D) and 572: 571: 565: 558: 553: 551: 549: 545: 544: 539: 535: 531: 526: 524: 520: 516: 512: 508: 504: 500: 496: 484: 480: 477: 476: 474: 471: 467: 459: 458: 451: 448: 447: 440: 437: 436: 435:Siroccopteryx 429: 427: 426: 419: 416: 415: 408: 405: 404: 397: 394: 393: 386: 385: 384: 378: 375: 374: 373:Tropeognathus 367: 364: 363: 356: 353: 352: 351:Cimoliopterus 345: 342: 341: 334: 331: 330: 329:Camposipterus 323: 320: 319: 312: 309: 308: 307:Araripesaurus 301: 300: 298: 293: 288: 283: 282: 274: 271: 267: 262: 257: 251: 248: 247: 244: 238: 235: 232: 231: 228: 222: 219: 216: 215: 212: 206: 203: 202: 199: 193: 190: 189: 186: 183: 180: 179: 176: 173: 170: 169: 166: 163: 160: 159: 154: 149: 145: 142: 138: 137: 131: 127: 122: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 54: 45: 41: 33: 30: 19: 10149: 10113: 10059: 10054:Ludodactylus 10052: 10045: 10038: 10030: 10022: 10015: 9995: 9987: 9982:Nicorhynchus 9980: 9973: 9966: 9945: 9938: 9930: 9922: 9914: 9906: 9898: 9868: 9861: 9853:Hamipteridae 9840: 9839: 9831: 9797: 9789: 9781: 9761: 9753: 9748:Aetodactylus 9746: 9711: 9703: 9696: 9688: 9682:Ludodactylus 9680: 9672: 9664: 9656: 9648: 9640: 9634:Aetodactylus 9632: 9611: 9603: 9595: 9587: 9579: 9572: 9565: 9553: 9522: 9515: 9491: 9484: 9477: 9470: 9444:Liaoxipterus 9442: 9435: 9415: 9408: 9401: 9394: 9372:Mimodactylus 9370: 9363: 9356: 9335: 9328: 9321: 9314: 9290: 9283: 9258: 9250:Lanceodontia 9231: 9223: 9218:Lonchodectes 9216: 9208: 9200: 9180: 9173: 9166: 9119: 9112: 9105: 9098: 9078: 9071: 9064: 9039: 9032: 9007: 8999: 8992: 8986:Ornithostoma 8984: 8977: 8969: 8940: 8880: 8794: 8786: 8779: 8772: 8765: 8758: 8753:Eurazhdarcho 8751: 8744: 8737: 8730: 8710: 8703: 8696: 8688: 8681:Azhdarchinae 8667: 8659: 8652: 8644: 8637: 8629: 8622:Azhdarchidae 8608: 8601: 8593: 8565: 8558: 8551: 8530: 8522: 8517:Meilifeilong 8515: 8508: 8500: 8492: 8484: 8460: 8435: 8427: 8419: 8411: 8403: 8396: 8374: 8367: 8360: 8352: 8302: 8295: 8275: 8268: 8261: 8253: 8229: 8221: 8213: 8205: 8197: 8173: 8166: 8159: 8154:Huaxiapterus 8152: 8145: 8139:Eopteranodon 8137: 8129: 8123:Afrotapejara 8121: 8114:Sinopterinae 8103:Afrotapejara 8101: 8077: 8070: 8039: 8032: 8025: 8017: 8009: 8002: 7994: 7986: 7978: 7970: 7948: 7943:Bennettazhia 7941: 7906:Ornithostoma 7904: 7897: 7889: 7881: 7874: 7867: 7833: 7826: 7805: 7797: 7792:Ordosipterus 7790: 7783: 7775: 7767: 7742:Tapejaroidea 7727: 7631: 7585:Pterodaustro 7583: 7576: 7569: 7562: 7542: 7518: 7511: 7504: 7498:Huanhepterus 7496: 7491:Gnathosaurus 7489: 7471:Moganopterus 7469: 7462: 7443:Pterofiltrus 7441: 7436:Petrodactyle 7434: 7427: 7422:Liaodactylus 7420: 7415:Kepodactylus 7413: 7406: 7399: 7392: 7385: 7378: 7371: 7363: 7330:Huanhepterus 7328: 7321: 7313: 7292: 7287:Aerodactylus 7285: 7260: 7254:Petrodactyle 7252: 7244: 7237: 7229: 7194: 7187: 7156: 7148: 7141: 7133: 7113: 7004: 6996: 6989: 6982: 6975: 6970:Kryptodrakon 6968: 6960: 6953: 6946: 6918: 6912:Jeholopterus 6910: 6902: 6895: 6888: 6868: 6861: 6855:Jeholopterus 6853: 6845: 6840:Anurognathus 6838: 6820:Mesadactylus 6818: 6783: 6775: 6737: 6729: 6722: 6715: 6707: 6687: 6663: 6656: 6645:Darwinoptera 6631: 6624: 6617: 6586: 6578: 6524: 6453: 6446: 6439: 6432: 6425: 6382: 6376:Nesodactylus 6374: 6368:Cacibupteryx 6366: 6348:Sericipterus 6346: 6338: 6330: 6323: 6316: 6287: 6280: 6275:Nesodactylus 6273: 6266: 6259: 6254:Cacibupteryx 6252: 6247:Bellubrunnus 6245: 6220: 6212: 6205: 6197: 6158: 6151: 6115: 6102:Lonchognatha 6083: 6075: 6067: 6060: 6053: 6045: 6025: 5994: 5955: 5947: 5926: 5902: 5895: 5890:Pachagnathus 5888: 5880: 5850: 5842: 5836:Austriadraco 5834: 5800: 5795:Austriadraco 5793: 5785: 5764: 5757: 5749: 5724: 5717: 5697: 5583: 5574: 5565: 5462: 5442: 5423: 5410: 5407:Reeside and 5406: 5402: 5395: 5377: 5360: 5356: 5350: 5305: 5301: 5295: 5287: 5278: 5274: 5268: 5260: 5235: 5231: 5225: 5192: 5188: 5182: 5170:. Retrieved 5166: 5156: 5134:(1): 57–76. 5131: 5127: 5123: 5117: 5084: 5078: 5065: 5048: 5044: 5038: 5013: 5009: 5005: 4999: 4974: 4970: 4966: 4960: 4941: 4937: 4931: 4923: 4906: 4902: 4896: 4888: 4883: 4875: 4871: 4867: 4862: 4853: 4849: 4843: 4835: 4810: 4806: 4799: 4772: 4768: 4762: 4754: 4745: 4741: 4731: 4706: 4702: 4696: 4676: 4631: 4627: 4617: 4606:the original 4601: 4597: 4581: 4556: 4552: 4542: 4517: 4513: 4503: 4478: 4474: 4470: 4464: 4442:(1): 40–49. 4439: 4435: 4426: 4418: 4407: 4401: 4394: 4349: 4345: 4335: 4323: 4311: 4276: 4272: 4262: 4212: 4208: 4198: 4173: 4169: 4135: 4131: 4106: 4094: 4077: 4073: 4060: 4042: 4036: 3994: 3990: 3984: 3980: 3950: 3946: 3939: 3914: 3910: 3903: 3884: 3880: 3836: 3832: 3804:|last6= 3796:cite journal 3753: 3749: 3707: 3703: 3650:(1): 13454. 3647: 3643: 3595: 3591: 3527: 3524:PLOS Biology 3523: 3458:(1): 17875. 3455: 3451: 3405: 3399: 3335: 3331: 3303: 3299: 3289: 3281:110004312505 3272: 3266: 3257: 3253: 3244: 3210: 3206: 3152: 3146: 3118: 3108: 3075: 3071: 3061: 3013: 3007: 3001: 2987: 2983:Pawpawsaurus 2981: 2958: 2957:(originally 2954: 2952: 2945: 2939: 2933: 2927: 2921: 2914: 2908: 2902: 2892: 2883: 2877: 2871: 2862: 2856: 2846: 2837: 2833: 2827: 2821: 2815: 2809: 2803: 2797: 2791: 2787: 2783: 2772: 2746: 2740: 2734: 2732:ichthyosaurs 2725: 2716: 2707: 2701: 2695: 2691:Anoplosaurus 2689: 2682:Ornithostoma 2680: 2671: 2664: 2660: 2656: 2652: 2642: 2639:Paleoecology 2632: 2626: 2608:frigatebirds 2592: 2587:Lonchodectes 2585: 2581: 2567:Paleobiology 2557: 2461: 2460: 2444: 2443: 2420: 2419: 2396: 2395: 2383: 2369: 2368: 2356: 2333: 2332: 2316: 2315: 2279: 2278: 2248:Ludodactylus 2246: 2245: 2229: 2228: 2191: 2190: 2166: 2162: 2161: 2081: 2080: 2064: 2063: 2040: 2039: 2016: 2015: 1999: 1998: 1957: 1956: 1940: 1939: 1916: 1915: 1832: 1831: 1815: 1814: 1791: 1790: 1774: 1773: 1750: 1749: 1726: 1725: 1698: 1684: 1662: 1658: 1657: 1641: 1633:C. segwickii 1632: 1628: 1624: 1620: 1616: 1612: 1609:Aetodactylus 1608: 1604: 1601:Aetodactylus 1600: 1596: 1595: 1590: 1586: 1582: 1578: 1574: 1570: 1565: 1561: 1557: 1553: 1549: 1545: 1539: 1535: 1531: 1527: 1523: 1517: 1511: 1507: 1503: 1499: 1495: 1489: 1485: 1481: 1477: 1473: 1469: 1465: 1463: 1449: 1445: 1441: 1429: 1425: 1421: 1417: 1415: 1402: 1396: 1393:Aetodactylus 1392: 1388: 1384: 1378: 1375:Aetodactylus 1374: 1369:Aetodactylus 1367: 1366: 1362:Lanceodontia 1355: 1351: 1341: 1338: 1328: 1324: 1316: 1308: 1302: 1298: 1240: 1208: 1204:aspect ratio 1197: 1177:Ludodactylus 1175: 1169: 1157: 1153: 1152:. The genus 1149: 1137: 1133: 1129: 1122:synapomorphy 1117: 1113: 1109: 1091: 1083: 1079: 1073: 1065: 1059: 1053: 1033: 1029: 1015: 1007: 1001: 985: 978:Hamipteridae 971: 968:Ludodactylus 967: 963: 959: 943: 937: 933: 929: 923: 919: 917: 909:Brian Andres 900: 896:Mimodactylus 894: 888: 882: 876: 872: 868:Ludodactylus 866: 862: 858: 852: 848: 840: 834: 832: 827: 823: 817: 813: 809: 797: 793: 789: 781: 777: 773: 769: 766:Criorhynchus 765: 757: 753: 750:Criorhynchus 749: 743: 739:Lonchodectes 737: 733: 729: 721: 717: 714:Criorhynchus 713: 705: 700:by its tall 697: 693: 689: 681: 674:Criorhynchus 673: 669: 665: 661: 658:Criorhynchus 657: 653: 649:Criorhynchus 647: 641: 629: 621: 617: 607: 605:generic name 596: 590: 587:Richard Owen 583: 577: 568: 541: 527: 498: 494: 493: 478: 455: 444: 433: 423: 412: 401: 390: 382: 371: 360: 349: 338: 327: 316: 305: 279: 270:Type species 255: 233: 217: 134: 51:140–90 29: 10174:Wikispecies 9940:Thapunngaka 9900:Amblydectes 9819:Anhangueria 9783:Aussiedraco 9517:Boreopterus 9292:Lonchodraco 9225:Lonchodraco 9168:Aussiedraco 9114:Nyctosaurus 8971:Bogolubovia 8639:Bogolubovia 8510:Lacusovagus 8502:Eoazhdarcho 8462:Eoazhdarcho 8421:Leptostomia 8413:Keresdrakon 8288:Caiuajarina 8255:Bakonydraco 8223:Lacusovagus 8215:Keresdrakon 8186:Tapejarinae 8147:Huaxiadraco 8131:Bakonydraco 8090:Tapejaridae 8019:Leptostomia 8011:Lacusovagus 8004:Kariridraco 7950:Keresdrakon 7891:Leptostomia 7883:Keresdrakon 7544:Ctenochasma 7506:Lusognathus 7429:Otogopterus 7084:Lophocratia 7017:Lophocratia 6984:Pangupterus 6920:Sinomacrops 6261:Dorygnathus 6130:Novialoidea 6085:Rhamphinion 6062:Dimorphodon 5737:Caviramidae 5681:Pterosauria 5668:Pterosauria 5631:Pterosauria 5570:Archosauria 5531:Pterosauria 4328:Witton 2013 4316:Witton 2013 4255:Witton 2013 4111:Witton 2013 4099:Witton 2013 4029:Witton 2013 3756:(1): 4940. 3054:Witton 2013 3015:Leptostyrax 2910:Cearachelys 2775:Lagerstätte 2678:azhdarchoid 2673:Lonchodraco 2657:Amblydectes 2595:piscivorous 2178:Anhangueria 2163:Topology 2: 1837:NHMUK R481 1659:Topology 1: 1352:Boreopterus 1343:Boreopterus 1310:Aussiedraco 1247:supraneural 1245:develops a 1182:Anhangueria 992:Description 982:Anhangueria 952:Mark Witton 802:David Unwin 745:Amblydectes 660:, creating 538:azhdarchids 519:Valanginian 446:Thapunngaka 198:Pterosauria 40:Valanginian 10263:Categories 10017:Anhanguera 9908:Ferrodraco 9863:Hamipterus 9791:Barbosania 9666:Ferrodraco 9658:Draigwenia 9493:Hamipterus 9472:Barbosania 9417:Nurhachius 9285:Ikrandraco 9260:Draigwenia 9210:Ikrandraco 9175:Serradraco 9121:Volgadraco 9009:Volgadraco 9001:Tethydraco 8994:Pteranodon 8852:see below↓ 8746:Cryodrakon 8669:Volgadraco 8661:Tethydraco 8560:Jidapterus 8524:Microtuban 8362:Microtuban 8263:Europejara 8242:Tapejarini 8231:Vectidraco 8168:Sinopterus 7785:Noripterus 7654:see below↓ 7578:Gegepterus 7571:Eosipterus 7026:see below↓ 7006:Wenupteryx 6897:Cascocauda 6476:see below↓ 5904:Yelaphomte 5640:see below↓ 5561:Sauropsida 5087:(1): 5–8. 4402:Anhanguera 3260:: 219–228. 3034:References 3003:Turrilites 2994:nodosaurid 2935:Rhacolepis 2929:Rhinobatos 2923:Brannerion 2916:Araripemys 2805:Anhanguera 2799:Barbosania 2748:Sisteronia 2727:Enaliornis 2676:, and the 2633:Anhanguera 2446:Ferrodraco 2335:Anhanguera 1644:cladograms 1625:C. nasutus 1587:Ferrodraco 1513:Ferrodraco 1416:The genus 1385:C. cuvieri 1299:Ferrodraco 1283:metatarsal 1171:Pteranodon 1146:caniniform 1142:heterodont 1134:Anhanguera 1102:premaxilla 1096:; further 944:Anhanguera 849:Anhanguera 819:Anhanguera 762:Oskar Kuhn 710:lanceolate 638:Cretaceous 503:pterosaurs 403:Ferrodraco 204:Suborder: 9968:Aerodraco 9410:Luchibang 9358:Haopterus 9080:Simurghia 9073:Epapatelo 9041:Cretornis 8979:Dawndraco 8705:Azhdarcho 8698:Albadraco 8690:Aerotitan 8610:Alanqidae 8388:Alanqidae 8354:Cretornis 8304:Torukjara 8297:Caiuajara 8034:Tupuxuara 7972:Aerotitan 7876:Inabtanin 7464:Feilongus 6863:Luopterus 6580:Allkaruen 6207:Klobiodon 6047:Allkaruen 5882:Caviramus 5759:Caviramus 5547:Kingdom: 5109:131339386 5030:140174098 5006:Tupuxuara 4723:128410270 4709:: 23–36. 4573:129514056 4534:129277596 4495:129807044 4047:CiteSeerX 3931:225569843 3917:(1): e4. 3853:209595986 3612:134439172 3598:: 77–88. 3348:CiteSeerX 3119:Cambridge 3100:130116778 3009:Scaphites 2998:ammonoids 2989:Texasetes 2978:dinosaurs 2941:Tharrhias 2879:Mirischia 2873:Irritator 2869:theropods 2864:Tupuxuara 2844:tapejarid 2709:Trachodon 2165:Pentland 1637:homoplasy 1623:species ( 1460:Phylogeny 1450:Haopterus 1446:Haopterus 1442:Haopterus 1436:, or the 1430:Haopterus 1426:Haopterus 1422:Haopterus 1418:Haopterus 1279:shinbones 1275:processes 1255:vertebrae 1224:coracoids 1174:, and in 1070:(I and J) 1022:wingspans 1012:(B and D) 901:Haopterus 863:Haopterus 517:periods ( 171:Kingdom: 165:Eukaryota 10194:47449510 10159:Wikidata 10040:Guidraco 9924:Mythunga 9916:Haliskia 9690:Mythunga 9674:Guidraco 9605:Haliskia 9182:Unwindia 8532:Xericeps 8437:Xericeps 8369:Xericeps 8270:Tapejara 8041:Xericeps 7996:Banguela 7769:Banguela 6998:Samrukia 6658:Ceoptera 5555:Chordata 5553:Phylum: 5549:Animalia 5342:27007950 5302:PLOS ONE 5252:86285896 5217:45417065 5148:85946263 4991:90477315 4878:: 75–80. 4827:25283914 4791:89178816 4685:Archived 4668:24465427 4628:PLOS ONE 4456:67809186 4386:21085624 4346:PLOS ONE 4303:29877021 4239:26339122 4190:84617119 4152:86099117 4013:23538956 3788:30894614 3726:31432888 3682:31582757 3556:29534059 3492:31784545 3424:24768054 3370:86710955 3181:23794925 3022:See also 2980:such as 2947:Tribodus 2848:Tapejara 2839:Unwindia 2714:sauropod 2617:foraging 2612:flocculi 2600:skimmers 2463:Mythunga 2193:Guidraco 2169:(2019). 1665:(2019). 1648:topology 1591:Mythunga 1519:Mythunga 1411:polytomy 1389:C. dunni 1304:Mythunga 1301:(A) and 1228:scapulae 1168:such as 1128:such as 1114:O. simus 922:but not 843:, their 718:O. simus 698:O. simus 654:P. simus 618:P. simus 597:P. simus 580:(E to H) 540:such as 523:Turonian 470:Synonyms 414:Mythunga 362:Haliskia 249:Family: 185:Chordata 181:Phylum: 175:Animalia 161:Domain: 44:Turonian 10233:4947088 10207:3238833 10165:Q136890 9066:Alcione 8175:Wightia 5559:Class: 5333:4805287 5310:Bibcode 5197:Bibcode 5089:Bibcode 4856:: 9–40. 4850:Deinsea 4807:Zootaxa 4659:3897400 4636:Bibcode 4377:2981443 4354:Bibcode 4294:6849529 4230:4536946 3953:: 1–9. 3779:6426928 3758:Bibcode 3673:6776501 3652:Bibcode 3547:5849296 3483:6884559 3460:Bibcode 3340:Bibcode 3172:3689139 3153:ZooKeys 3080:Bibcode 2761:of the 2649:England 2180: 1701: 1661:Jacobs 1350:, with 1088:rostrum 764:placed 702:rostrum 554:History 295:Genera 191:Order: 139:in the 10246:156758 10220:117087 8631:Alanqa 8398:Alanqa 7980:Alanqa 6588:Sordes 6455:Sordes 5470: 5449: 5386: 5340: 5330: 5250: 5215: 5189:Nature 5146: 5107: 5028: 4989: 4825: 4789: 4721: 4666: 4656: 4571: 4532: 4493: 4454: 4384: 4374: 4301: 4291: 4237: 4227: 4188: 4150: 4049: 4011: 3929: 3851: 3786: 3776: 3724: 3680: 3670: 3610: 3554: 3544: 3490: 3480: 3422: 3368: 3350: 3279: 3179: 3169: 3121:: 113. 3098: 2899:turtle 2779:Brazil 2685:. The 2667:, the 2558: 2176: 2167:et al. 1697: 1663:et al. 1263:femora 1243:sacrum 1216:sterna 1026:morphs 792:(with 720:, and 676:was a 612:(from 559:Naming 485:, 1914 483:Hooley 289:, 1861 263:, 1870 261:Seeley 10215:IRMNG 7729:Piksi 6325:Dearc 5584:Clade 5575:Clade 5566:Clade 5415:(PDF) 5248:S2CID 5213:S2CID 5144:S2CID 5105:S2CID 5075:(PDF) 5026:S2CID 4987:S2CID 4787:S2CID 4719:S2CID 4609:(PDF) 4594:(PDF) 4569:S2CID 4530:S2CID 4491:S2CID 4452:S2CID 4432:(PDF) 4186:S2CID 4148:S2CID 4070:(PDF) 3927:S2CID 3849:S2CID 3608:S2CID 3366:S2CID 3250:(PDF) 3096:S2CID 2967:Texas 2604:terns 1136:. In 1058:(D), 1040:Skull 511:Early 234:Clade 218:Clade 10202:GBIF 5617:see 5468:ISBN 5447:ISBN 5384:ISBN 5338:PMID 5174:2015 4823:PMID 4811:3869 4664:PMID 4382:PMID 4299:PMID 4235:PMID 4009:PMID 3983:cf. 3808:help 3784:PMID 3722:PMID 3678:PMID 3552:PMID 3488:PMID 3420:PMID 3277:NAID 3177:PMID 3006:and 2944:and 2913:and 2882:and 2861:and 2836:and 2808:and 2790:and 2745:and 2723:bird 2706:and 2659:and 2606:and 1631:and 1589:and 1581:and 1564:and 1552:and 1534:and 1516:and 1506:and 1480:and 1472:and 1401:and 1387:and 1327:and 1132:and 1116:and 1082:and 970:and 936:and 887:and 871:and 816:and 748:and 736:and 716:for 708:had 634:ICZN 532:and 497:(or 287:Owen 59:PreꞒ 10189:EoL 5365:doi 5328:PMC 5318:doi 5240:doi 5205:doi 5193:392 5136:doi 5097:doi 5085:153 5053:doi 5049:199 5018:doi 4979:doi 4946:doi 4911:doi 4815:doi 4777:doi 4773:130 4711:doi 4654:PMC 4644:doi 4561:doi 4522:doi 4483:doi 4473:". 4444:doi 4440:316 4372:PMC 4362:doi 4289:PMC 4281:doi 4225:PMC 4217:doi 4178:doi 4174:103 4140:doi 4082:doi 3999:doi 3955:doi 3919:doi 3889:doi 3841:doi 3774:PMC 3766:doi 3712:doi 3668:PMC 3660:doi 3600:doi 3542:PMC 3532:doi 3478:PMC 3468:doi 3410:doi 3358:doi 3336:217 3308:doi 3215:doi 3167:PMC 3157:doi 3088:doi 3076:343 2965:of 2647:of 1313:(C) 907:by 680:of 521:to 513:to 10265:: 10243:: 10230:: 10217:: 10204:: 10191:: 10176:: 10161:: 5586:: 5577:: 5568:: 5361:15 5359:. 5336:. 5326:. 5316:. 5306:11 5304:. 5300:. 5279:37 5277:. 5273:. 5246:. 5236:86 5234:. 5211:. 5203:. 5191:. 5165:. 5142:. 5132:20 5130:. 5103:. 5095:. 5083:. 5077:. 5047:. 5024:. 5014:30 5012:. 4985:. 4975:38 4973:. 4942:56 4940:. 4936:. 4907:32 4905:. 4876:27 4874:, 4854:13 4852:. 4848:. 4821:. 4809:. 4785:. 4771:. 4767:. 4746:32 4744:. 4740:. 4717:. 4707:75 4705:. 4662:. 4652:. 4642:. 4630:. 4626:. 4602:16 4600:. 4596:. 4567:. 4557:32 4555:. 4551:. 4528:. 4518:35 4516:. 4512:. 4489:. 4479:60 4477:. 4450:. 4438:. 4434:. 4406:" 4380:. 4370:. 4360:. 4348:. 4344:. 4297:. 4287:. 4277:93 4275:. 4271:. 4247:^ 4233:. 4223:. 4213:27 4211:. 4207:. 4184:. 4172:. 4160:^ 4146:. 4136:35 4134:. 4118:^ 4078:26 4076:. 4072:. 4021:^ 4007:. 3995:85 3993:. 3989:. 3967:^ 3951:34 3949:. 3925:. 3915:43 3913:. 3887:. 3885:65 3883:. 3879:. 3861:^ 3847:. 3837:33 3835:. 3816:^ 3800:: 3798:}} 3794:{{ 3782:. 3772:. 3764:. 3752:. 3748:. 3734:^ 3720:. 3708:91 3706:. 3702:. 3690:^ 3676:. 3666:. 3658:. 3646:. 3642:. 3620:^ 3606:. 3596:95 3594:. 3590:. 3564:^ 3550:. 3540:. 3528:16 3526:. 3522:. 3500:^ 3486:. 3476:. 3466:. 3454:. 3450:. 3432:^ 3418:. 3406:24 3404:. 3398:. 3378:^ 3364:. 3356:. 3346:. 3334:. 3320:^ 3304:13 3302:. 3298:. 3258:28 3256:. 3252:. 3229:^ 3209:. 3205:. 3189:^ 3175:. 3165:. 3151:. 3127:^ 3117:. 3094:. 3086:. 3074:. 3070:. 3042:^ 2986:, 2969:, 2950:. 2938:, 2932:, 2926:, 2907:, 2876:, 2832:, 2826:, 2820:, 2786:, 2773:A 2739:, 2694:, 1627:, 1577:, 1573:, 1560:, 1530:, 1502:, 1468:, 1456:. 1413:. 1285:. 1238:. 962:, 932:, 899:, 881:, 865:, 861:, 857:, 851:, 839:, 830:. 812:, 595:: 236:: 220:: 109:Pg 53:Ma 10035:? 10027:? 9992:? 9960:? 9935:? 9927:? 9919:? 9911:? 9903:? 9892:? 9855:? 9844:? 9836:? 9794:? 9786:? 9758:? 9731:? 9716:? 9708:? 9693:? 9685:? 9677:? 9669:? 9661:? 9653:? 9645:? 9637:? 9616:? 9608:? 9600:? 9592:? 9584:? 9340:? 9263:? 9236:? 9228:? 9213:? 9205:? 9124:? 9044:? 9012:? 9004:? 8989:? 8974:? 8791:? 8693:? 8672:? 8664:? 8649:? 8634:? 8613:? 8598:? 8535:? 8527:? 8505:? 8497:? 8489:? 8440:? 8432:? 8424:? 8416:? 8408:? 8390:? 8379:? 8357:? 8258:? 8226:? 8218:? 8210:? 8202:? 8142:? 8134:? 8126:? 8064:? 8044:? 8022:? 8014:? 7999:? 7991:? 7983:? 7975:? 7964:? 7953:? 7894:? 7886:? 7810:? 7802:? 7780:? 7772:? 7732:? 7636:? 7501:? 7456:? 7368:? 7333:? 7318:? 7297:? 7265:? 7257:? 7249:? 7234:? 7161:? 7153:? 7138:? 7001:? 6965:? 6915:? 6907:? 6858:? 6850:? 6780:? 6734:? 6712:? 6636:? 6591:? 6583:? 6458:? 6419:? 6409:? 6379:? 6371:? 6343:? 6335:? 6225:? 6217:? 6202:? 6120:? 6104:? 6088:? 6080:? 6072:? 6050:? 5999:? 5983:? 5952:? 5931:? 5885:? 5855:? 5847:? 5839:? 5790:? 5769:? 5754:? 5739:? 5523:e 5516:t 5509:v 5476:. 5455:. 5390:. 5371:. 5367:: 5344:. 5320:: 5312:: 5254:. 5242:: 5219:. 5207:: 5199:: 5176:. 5150:. 5138:: 5111:. 5099:: 5091:: 5059:. 5055:: 5032:. 5020:: 4993:. 4981:: 4954:. 4948:: 4917:. 4913:: 4829:. 4817:: 4793:. 4779:: 4725:. 4713:: 4670:. 4646:: 4638:: 4632:9 4575:. 4563:: 4536:. 4524:: 4497:. 4485:: 4458:. 4446:: 4425:" 4404:. 4388:. 4364:: 4356:: 4350:5 4305:. 4283:: 4241:. 4219:: 4192:. 4180:: 4154:. 4142:: 4088:. 4084:: 4015:. 4001:: 3961:. 3957:: 3933:. 3921:: 3897:. 3891:: 3855:. 3843:: 3810:) 3790:. 3768:: 3760:: 3754:9 3728:. 3714:: 3684:. 3662:: 3654:: 3648:9 3614:. 3602:: 3558:. 3534:: 3494:. 3470:: 3462:: 3456:9 3426:. 3412:: 3372:. 3360:: 3342:: 3314:. 3310:: 3283:. 3247:" 3223:. 3217:: 3211:4 3183:. 3159:: 3102:. 3090:: 3082:: 2918:. 1383:( 460:? 453:† 449:? 442:† 438:? 431:† 421:† 417:? 410:† 406:? 399:† 395:? 388:† 380:† 376:? 369:† 365:? 358:† 354:? 347:† 343:? 336:† 332:? 325:† 321:? 314:† 310:? 303:† 277:† 253:† 240:† 224:† 208:† 195:† 114:N 104:K 99:J 94:T 89:P 84:C 79:D 74:S 69:O 64:Ꞓ 48:~ 46:, 42:- 20:)

Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.

Index