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Atretochoana

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1399: 77: 216: 1417: 52: 33: 1405: 1410: 299:. It has significant morphological differences from other caecilians, even the genera most closely related to it, even though those genera are aquatic. The skull is very different from those of other caecilians, giving the animal a broad, flat head. Its nostrils are sealed, and it has an enlarged mouth with a mobile cheek. Its body has a fleshy dorsal fin. 407:
that he designated the specimen a holotype, so it was not mentioned in the museum's catalogue of type specimens, and was placed beneath glass in a public display. There, it was noticed by the visiting English herpetologist Mark Wilkinson, who then borrowed the specimen to examine it with his American
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originally known only from two preserved specimens discovered by Sir Graham Hales in the Brazilian rainforest, while on an expedition with Sir Brian Doll in the late 1800s, but rediscovered in 2011 by engineers working on a hydroelectric dam project in Brazil. Until 1998, it was known only from the
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The specimen in the Vienna museum was known only to have originated from somewhere in South America, at least before 1945, most likely in the 19th century. Its lack of lungs was not known at this time, and the specimen was assigned to the species
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is a large caecilian at a length of 72.5 cm (28.5 in), while the Brasília specimen is larger still at 80.5 cm (31.7 in). By comparison, caecilians in general range in length from 11 to 160 cm (4.3 to 63.0 in).
510:. Neither have cold, fast-flowing water, as was originally thought, as there is less oxygen in warmer water. This makes its lack of lungs even more unusual, and the question of how it breathes has not yet been resolved. 437:, referring to a funnel or tube. Nussbaum and Wilkinson published further studies in 1997 describing in detail the caecilian's anatomy and morphology. In 1998, they discovered the second specimen in the 408:
colleague Ronald A. Nussbaum. Examination of the specimen showed it to have a number of unusual features, including the large number of splenial teeth observed by Taylor, but most unusually, closed
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Because of these and other distinctive features, Nussbaum and Wilkinson gave this species its own genus when they reported on the results of their research in a 1995 issue of the
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Nussbaum, Ronald A.; Wilkinson, Mark (1995). "A New Genus of Lungless Tetrapod: A Radically Divergent Caecilian (Amphibia: Gymnophiona)".
710: 2764: 786:"The largest lungless tetrapod: report on a second specimen of Atretochoana eiselti (Amphibia: Gymnophiona: Typhlonectidae) from Brazil" 741: 417: 941: 648: 2594: 2408: 1398: 2664: 2478: 2413: 487:. It is thought to be uncommon, with a limited distribution. It is likely a predator or scavenger, and is thought to be 76: 360: 818: 329:, allowing gas exchange. Its skull shows evidence of muscles not found in any other organism. The Vienna specimen of 2690: 1416: 1006: 2491: 1856: 1108: 317:, however, entirely lacks lungs, and has a number of other features associated with lunglessness, including sealed 476:, some of the few other lungless tetrapods, are aquatic as well. It was postulated to inhabit fast-flowing water. 438: 2559: 2344: 1206: 404: 239: 2754: 1264: 345:
in the media such as "penis snake", "man-aconda", and "floppy snake", owing to its visual similarity to the
2361: 1237: 369: 205: 1404: 1101: 571: 188: 1409: 2517: 2607: 2568: 2421: 2062: 1751: 968:(Taylor, 1968) (Amphibia: Gymnophiona: Typhlonectidae), in its natural habitat in Brazilian Amazonia" 750: 2140: 473: 326: 41: 381: 2716: 998: 766: 555: 71: 396:, and placed it in the former genus, taking much note only of its large size and high number of 2651: 2399: 2119: 933: 927: 679: 2698: 2499: 2465: 2149: 1490: 1427: 1338: 990: 937: 683: 2726: 2703: 2504: 1059: 2052: 1648: 1641: 1627: 982: 903: 800: 758: 550: 322: 215: 2677: 2278: 2248: 2177: 1955: 1522: 1479: 1469: 1383: 1191: 506:, but was not positively identified. In 2011, six individual organisms were found in the 754: 1948: 1923: 1821: 1716: 1580: 1249: 1218: 908: 672: 480: 450: 426: 151: 56: 1035:[Snake-shaped amphibian found in the Rio Madeira, RO] (in Portuguese). Globo. 2743: 2672: 2573: 2258: 2163: 2156: 2126: 1969: 1962: 1877: 1744: 1737: 1355: 1222: 1166: 541: 532: 507: 392: 260: 235: 61: 1002: 770: 32: 1934: 1888: 1758: 1698: 1601: 1459: 1348: 1310: 1276: 1157: 1124: 962:
Hoogmoed, Marinus Steven; Maciel, Adriano Oliveira; Coragem, Juliano Tupan (2011).
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In June 2011, an amphibian was photographed near Praia de Marahú on the island of
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are limbless amphibians with snake-like bodies, marked with rings like those of
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Himstedt, Werner (2000). "Lungless Tetradodes". In Hofrichter, Robert (ed.).
2288: 2224: 2010: 1976: 1902: 1870: 1768: 1617: 1566: 1503: 1362: 1083: 1060:"'Penis snake' discovered in Brazil is actually a rare species of amphibian" 833:"'Penis snake' discovered in Brazil is actually a rare species of amphibian" 785: 495: 384:. Taylor considered it to be similar to the aquatic caecilians of the genus 303: 296: 249: 230: 141: 128: 118: 88: 762: 884: 2538: 2378: 2170: 2112: 2105: 1916: 1803: 1175: 1148: 499: 397: 365: 284: 269: 108: 2625: 2586: 2439: 2028: 1796: 1705: 1673: 1536: 318: 2638: 2452: 2082: 1142: 409: 258:, in 1996. It was also found to be more closely related to the genus 98: 2355: 784:
Wilkinson, M.; Sebben, A.; Schwartz, E.N.F.; Schwartz, C.A. (1998).
2512: 338: 1033:"Anfíbio com formato de cobra é descoberto no Rio Madeira, em RO" 2612: 2299: 885:"Comparative morphology and evolution of the lungless caecilian 288: 2426: 2359: 1135: 1097: 1093: 526: 524: 522: 819:"Penis Snake is neither penis nor snake, but looks like both" 929:
Amphibians: The World of Frogs, Toads, Salamanders and Newts
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Boletim do Museu Paraense Emílio Goeldi - Ciências Naturais
412:, which showed it could not fill any lungs it might have. 291:, double the size of the next largest. Caecilians such as 325:. Its skin is filled with capillaries that penetrate the 302:
Most caecilians have a well-developed right lung and a
268:. The species is the largest of the few known lungless 2284: 2274: 2264: 2254: 2244: 2137: 2093: 2078: 2068: 2058: 2048: 1792: 1764: 1486: 1475: 1465: 1455: 1445: 1335: 1089:
2011 paper reporting the discovery of live specimens
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Due to the lack of information, it is classified as
2528: 2368: 2298: 2223: 2194: 2136: 2092: 2027: 2006: 1933: 1887: 1820: 1778: 1715: 1683: 1658: 1626: 1615: 1579: 1564: 1521: 1501: 1425: 1334: 1275: 1248: 1217: 1190: 531:Mark Wilkinson, John Measey, Marvalee Wake (2004). 461:Most caecilians are burrowers, but some, including 671: 1084:1998 paper reporting discovery of second specimen 889:(Taylor) (Amphibia: Gymnophiona: Typhlonectidae)" 855:Information on amphibian biology and conservation 472:is thought to be aquatic since its relatives and 368:for this species when it was first described by 485:International Union for Conservation of Nature 453:. Both of these specimens are mature females. 1109: 964:"Discovery of the largest lungless tetrapod, 883:Nussbaum, Ronald A.; Wilkinson, Mark (1997). 8: 576:: CS1 maint: multiple names: authors list ( 556:10.2305/IUCN.UK.2004.RLTS.T59493A11937533.en 433:, meaning "imperforate", and the Greek word 313:s relatives, have two well-developed lungs. 957: 955: 953: 921: 919: 2356: 2237: 2041: 2024: 2003: 1623: 1576: 1572: 1518: 1514: 1438: 1331: 1327: 1254: 1227: 1196: 1187: 1132: 1116: 1102: 1094: 403:Taylor did not inform the curators of the 248:in 1968, it was reclassified into its own 214: 50: 31: 20: 907: 896:Biological Journal of the Linnean Society 554: 272:, and the only known lungless caecilian. 242:, Vienna. Originally placed in the genus 844: 842: 705: 703: 701: 699: 616: 614: 612: 589: 587: 449:, and in 2011, grouped it in the family 878: 876: 874: 872: 518: 734: 732: 730: 728: 569: 380:, in honour of Viennese herpetologist 643: 641: 639: 7: 2750:IUCN Red List data deficient species 2518:EB2187D1-F00B-FFD2-FF08-FCD2FDEE3E5A 421:. The name they gave this genus was 849:Knapp, Michelle (3 February 2003). 542:IUCN Red List of Threatened Species 1039:from the original on 3 August 2012 909:10.1111/j.1095-8312.1997.tb01616.x 742:Proceedings of the Royal Society B 468:s relatives, are largely aquatic. 418:Proceedings of the Royal Society B 14: 1031:Ximenes, Marcela (31 July 2012). 1415: 1408: 1403: 1397: 75: 1058:Butler, Rhett (2 August 2012). 595:"Distribution and conservation" 502:, Brazil) that appeared to be 364:. The Vienna specimen was the 1: 1663:(Northeast Indian caecilians) 932:. New York: Firefly. p.  341:, it has been called various 2770:Amphibians described in 1968 2760:Endemic caecilians of Brazil 1526:(American tailed caecilians) 987:10.46357/bcnaturais.v6i3.608 678:. London: Belknap. pp.  168:Nussbaum and Wilkinson, 1995 1938:(South American caecilians) 361:Typhlonectes compressicauda 2786: 2765:Monotypic amphibian genera 793:Journal of Natural History 2339: 2240: 2199:(salamanders crown group) 2044: 2002: 1825:(Indo-African caecilians) 1584:(Asian tailed caecilians) 1575: 1517: 1441: 1395: 1330: 1326: 1307: 1257: 1230: 1199: 1186: 1131: 805:10.1080/00222939800770321 670:Naskrecki, Piotr (2005). 306:left lung. Some, such as 213: 194: 187: 72:Scientific classification 70: 48: 39: 30: 23: 2345:List of amphibian genera 1892:(Neotropical caecilians) 1631:(buried-eyed caecilians) 1507:(caecilians crown group) 1431:(caecilians total group) 713:. Natural History Museum 651:. Natural History Museum 624:. Natural History Museum 597:. Natural History Museum 405:Naturhistorisches Museum 240:Naturhistorisches Museum 374:Caecilians of the World 2314:several hundred genera 851:"Atretochoana eiselti" 763:10.1098/rspb.1995.0155 649:"Atretochoana eiselti" 445:was a sister taxon of 439:University of Brasília 372:in his 1968 monograph 370:Edward Harrison Taylor 549:: e.T59493A11937533. 337:Although it is not a 2600:Atretochoana-eiselti 2560:Atretochoana eiselti 2530:Atretochoana eiselti 2015:(salamanders + frogs 1720:(aquatic caecilians) 1688:(African caecilians) 966:Atretochoana eiselti 887:Atretochoana eiselti 674:The Smaller Majority 535:Atretochoana eiselti 474:lungless salamanders 378:Typhlonectes eiselti 321:, and an absence of 226:Atretochoana eiselti 198:Atretochoana eiselti 2303:(frogs crown group) 2141:Batrachosauroididae 1783:(common caecilians) 755:1995RSPSB.261..331N 42:Conservation status 16:Genus of amphibians 1428:Gymnophionomorpha 323:pulmonary arteries 2737: 2736: 2699:Open Tree of Life 2500:Open Tree of Life 2362:Taxon identifiers 2353: 2352: 2335: 2334: 2331: 2330: 2327: 2326: 2323: 2322: 2305: 2233: 2219: 2218: 2201: 2150:Batrachosauroides 2037: 2019: 1998: 1997: 1994: 1993: 1990: 1989: 1986: 1985: 1940: 1894: 1827: 1785: 1722: 1690: 1665: 1633: 1628:Scolecomorphidae 1611: 1610: 1586: 1560: 1559: 1528: 1509: 1491:Albanerpetontidae 1433: 1393: 1392: 1339:Albanerpetontidae 1322: 1321: 1318: 1317: 1303: 1302: 1299: 1298: 1295: 1294: 1012:on 13 August 2012 749:(1362): 331–339. 689:978-0-647-02562-8 222: 221: 169: 65: 2777: 2730: 2729: 2720: 2719: 2707: 2706: 2694: 2693: 2681: 2680: 2668: 2667: 2655: 2654: 2642: 2641: 2629: 2628: 2616: 2615: 2603: 2602: 2590: 2589: 2577: 2576: 2564: 2563: 2562: 2549: 2548: 2547: 2521: 2520: 2508: 2507: 2495: 2494: 2482: 2481: 2469: 2468: 2456: 2455: 2443: 2442: 2430: 2429: 2417: 2416: 2404: 2403: 2402: 2389: 2388: 2387: 2357: 2304: 2301: 2286: 2276: 2266: 2256: 2246: 2238: 2232: 2227: 2200: 2197: 2139: 2095: 2080: 2070: 2060: 2050: 2042: 2036: 2031: 2025: 2018: 2013: 2004: 1939: 1936: 1893: 1890: 1826: 1823: 1794: 1784: 1781: 1766: 1721: 1718: 1689: 1686: 1664: 1661: 1632: 1629: 1624: 1585: 1582: 1577: 1573: 1527: 1524: 1523:Rhinatrematidae 1519: 1515: 1508: 1505: 1488: 1477: 1467: 1457: 1447: 1439: 1432: 1429: 1420: 1419: 1412: 1407: 1401: 1337: 1332: 1328: 1255: 1228: 1197: 1188: 1181: 1180: 1133: 1118: 1111: 1104: 1095: 1072: 1071: 1069: 1067: 1055: 1049: 1048: 1046: 1044: 1028: 1022: 1021: 1019: 1017: 1011: 1005:. Archived from 972: 959: 948: 947: 923: 914: 913: 911: 893: 880: 867: 866: 864: 862: 846: 837: 836: 829: 823: 822: 821:. 4 August 2012. 815: 809: 808: 790: 781: 775: 774: 736: 723: 722: 720: 718: 707: 694: 693: 677: 667: 661: 660: 658: 656: 645: 634: 633: 631: 629: 618: 607: 606: 604: 602: 591: 582: 581: 575: 567: 565: 563: 558: 528: 467: 312: 229:is a species of 218: 200: 167: 80: 79: 59: 54: 53: 35: 21: 2785: 2784: 2780: 2779: 2778: 2776: 2775: 2774: 2740: 2739: 2738: 2733: 2725: 2723: 2715: 2710: 2702: 2697: 2689: 2684: 2676: 2671: 2663: 2658: 2650: 2645: 2637: 2632: 2624: 2619: 2611: 2606: 2598: 2593: 2585: 2580: 2572: 2567: 2558: 2557: 2552: 2543: 2542: 2537: 2524: 2516: 2511: 2503: 2498: 2490: 2485: 2477: 2472: 2464: 2459: 2451: 2446: 2438: 2433: 2425: 2420: 2412: 2407: 2398: 2397: 2392: 2383: 2382: 2377: 2364: 2354: 2349: 2319: 2302: 2294: 2279:Triadobatrachus 2249:Czatkobatrachus 2230: 2228: 2226: 2215: 2198: 2190: 2178:Peratosauroides 2132: 2088: 2063:Kulgeriherpeton 2034: 2032: 2030: 2021: 2016: 2014: 2012: 2009: 2008: 1982: 1956:Luetkenotyphlus 1937: 1929: 1891: 1883: 1824: 1822:Grandisoniidae 1816: 1782: 1774: 1719: 1717:Typhlonectidae 1711: 1687: 1679: 1662: 1654: 1630: 1620: 1607: 1583: 1581:Ichthyophiidae 1569: 1556: 1525: 1511: 1506: 1497: 1480:Chinlestegophis 1470:Rubricacaecilia 1435: 1430: 1421: 1414: 1389: 1314: 1291: 1271: 1244: 1213: 1207:Tetrapodomorpha 1192:Tetrapodomorpha 1182: 1138: 1137: 1127: 1122: 1080: 1075: 1065: 1063: 1057: 1056: 1052: 1042: 1040: 1030: 1029: 1025: 1015: 1013: 1009: 970: 961: 960: 951: 944: 925: 924: 917: 891: 882: 881: 870: 860: 858: 848: 847: 840: 831: 830: 826: 817: 816: 812: 788: 783: 782: 778: 738: 737: 726: 716: 714: 709: 708: 697: 690: 669: 668: 664: 654: 652: 647: 646: 637: 627: 625: 620: 619: 610: 600: 598: 593: 592: 585: 568: 561: 559: 530: 529: 520: 516: 465: 459: 355: 310: 283:is the largest 278: 209: 202: 196: 183: 180:A. eiselti 166: 74: 66: 55: 51: 44: 17: 12: 11: 5: 2783: 2781: 2773: 2772: 2767: 2762: 2757: 2755:Typhlonectidae 2752: 2742: 2741: 2735: 2734: 2732: 2731: 2721: 2708: 2695: 2682: 2669: 2656: 2643: 2630: 2617: 2604: 2591: 2578: 2565: 2550: 2534: 2532: 2526: 2525: 2523: 2522: 2509: 2496: 2483: 2470: 2457: 2444: 2431: 2418: 2405: 2390: 2374: 2372: 2366: 2365: 2360: 2351: 2350: 2348: 2347: 2340: 2337: 2336: 2333: 2332: 2329: 2328: 2325: 2324: 2321: 2320: 2318: 2317: 2316: 2315: 2309: 2307: 2296: 2295: 2293: 2292: 2282: 2272: 2262: 2252: 2241: 2235: 2221: 2220: 2217: 2216: 2214: 2213: 2212: 2211: 2205: 2203: 2192: 2191: 2189: 2188: 2181: 2174: 2167: 2160: 2153: 2145: 2143: 2134: 2133: 2131: 2130: 2123: 2116: 2109: 2101: 2099: 2090: 2089: 2087: 2086: 2076: 2066: 2056: 2045: 2039: 2022: 2007: 2000: 1999: 1996: 1995: 1992: 1991: 1988: 1987: 1984: 1983: 1981: 1980: 1973: 1966: 1959: 1952: 1949:Brasilotyphlus 1944: 1942: 1931: 1930: 1928: 1927: 1924:Schistometopum 1920: 1913: 1906: 1898: 1896: 1885: 1884: 1882: 1881: 1874: 1867: 1860: 1853: 1846: 1839: 1831: 1829: 1818: 1817: 1815: 1814: 1807: 1800: 1789: 1787: 1776: 1775: 1773: 1772: 1762: 1755: 1748: 1741: 1734: 1726: 1724: 1713: 1712: 1710: 1709: 1702: 1694: 1692: 1681: 1680: 1678: 1677: 1669: 1667: 1656: 1655: 1653: 1652: 1649:Scolecomorphus 1645: 1637: 1635: 1621: 1616: 1613: 1612: 1609: 1608: 1606: 1605: 1598: 1590: 1588: 1570: 1567:Stegokrotaphia 1565: 1562: 1561: 1558: 1557: 1555: 1554: 1547: 1540: 1532: 1530: 1512: 1502: 1499: 1498: 1496: 1495: 1484: 1473: 1463: 1453: 1442: 1436: 1426: 1423: 1422: 1396: 1394: 1391: 1390: 1388: 1387: 1380: 1373: 1366: 1359: 1352: 1344: 1342: 1324: 1323: 1320: 1319: 1316: 1315: 1308: 1305: 1304: 1301: 1300: 1297: 1296: 1293: 1292: 1290: 1289: 1288: 1287: 1281: 1279: 1273: 1272: 1270: 1269: 1268: 1267: 1265:Dissorophoidea 1258: 1252: 1250:Dissorophoidea 1246: 1245: 1243: 1242: 1241: 1240: 1231: 1225: 1219:Batrachomorpha 1215: 1214: 1212: 1211: 1210: 1209: 1200: 1194: 1184: 1183: 1179: 1178: 1169: 1160: 1151: 1145: 1136: 1129: 1128: 1123: 1121: 1120: 1113: 1106: 1098: 1092: 1091: 1086: 1079: 1078:External links 1076: 1074: 1073: 1050: 1023: 981:(3): 241–262. 949: 942: 915: 868: 838: 835:. August 2012. 824: 810: 799:(4): 617–627. 776: 724: 695: 688: 662: 635: 608: 583: 517: 515: 512: 481:Data Deficient 458: 455: 451:Typhlonectidae 376:. He named it 354: 351: 277: 274: 220: 219: 211: 210: 203: 192: 191: 185: 184: 177: 175: 171: 170: 159: 155: 154: 152:Typhlonectidae 149: 145: 144: 139: 132: 131: 126: 122: 121: 116: 112: 111: 106: 102: 101: 96: 92: 91: 86: 82: 81: 68: 67: 57:Data Deficient 49: 46: 45: 40: 37: 36: 28: 27: 15: 13: 10: 9: 6: 4: 3: 2: 2782: 2771: 2768: 2766: 2763: 2761: 2758: 2756: 2753: 2751: 2748: 2747: 2745: 2728: 2722: 2718: 2713: 2709: 2705: 2700: 2696: 2692: 2687: 2683: 2679: 2674: 2670: 2666: 2661: 2657: 2653: 2648: 2644: 2640: 2635: 2631: 2627: 2622: 2618: 2614: 2609: 2605: 2601: 2596: 2592: 2588: 2583: 2579: 2575: 2570: 2566: 2561: 2555: 2551: 2546: 2540: 2536: 2535: 2533: 2531: 2527: 2519: 2514: 2510: 2506: 2501: 2497: 2493: 2488: 2484: 2480: 2475: 2471: 2467: 2462: 2458: 2454: 2449: 2445: 2441: 2436: 2432: 2428: 2423: 2419: 2415: 2410: 2406: 2401: 2395: 2391: 2386: 2380: 2376: 2375: 2373: 2371: 2367: 2363: 2358: 2346: 2342: 2341: 2338: 2313: 2312: 2311: 2310: 2308: 2306: 2297: 2291: 2290: 2283: 2281: 2280: 2273: 2271: 2270: 2263: 2261: 2260: 2259:Notobatrachus 2253: 2251: 2250: 2243: 2242: 2239: 2236: 2234: 2222: 2209: 2208: 2207: 2206: 2204: 2202: 2193: 2187: 2186: 2182: 2180: 2179: 2175: 2173: 2172: 2168: 2166: 2165: 2164:Palaeoproteus 2161: 2159: 2158: 2157:Opisthotriton 2154: 2152: 2151: 2147: 2146: 2144: 2142: 2135: 2129: 2128: 2127:Marmorerpeton 2124: 2122: 2121: 2117: 2115: 2114: 2110: 2108: 2107: 2103: 2102: 2100: 2098: 2091: 2085: 2084: 2077: 2075: 2074: 2067: 2065: 2064: 2057: 2055: 2054: 2047: 2046: 2043: 2040: 2038: 2026: 2023: 2020: 2005: 2001: 1979: 1978: 1974: 1972: 1971: 1970:Mimosiphonops 1967: 1965: 1964: 1963:Microcaecilia 1960: 1958: 1957: 1953: 1951: 1950: 1946: 1945: 1943: 1941: 1935:Siphonopidae 1932: 1926: 1925: 1921: 1919: 1918: 1914: 1912: 1911: 1907: 1905: 1904: 1900: 1899: 1897: 1895: 1889:Dermophiidae 1886: 1880: 1879: 1878:Sylvacaecilia 1875: 1873: 1872: 1868: 1866: 1865: 1861: 1859: 1858: 1854: 1852: 1851: 1847: 1845: 1844: 1840: 1838: 1837: 1833: 1832: 1830: 1828: 1819: 1813: 1812: 1808: 1806: 1805: 1801: 1799: 1798: 1791: 1790: 1788: 1786: 1777: 1771: 1770: 1763: 1761: 1760: 1756: 1754: 1753: 1752:Potamotyphlus 1749: 1747: 1746: 1745:Nectocaecilia 1742: 1740: 1739: 1738:Chthonerpeton 1735: 1733: 1732: 1728: 1727: 1725: 1723: 1714: 1708: 1707: 1703: 1701: 1700: 1696: 1695: 1693: 1691: 1682: 1676: 1675: 1671: 1670: 1668: 1666: 1657: 1651: 1650: 1646: 1644: 1643: 1642:Crotaphatrema 1639: 1638: 1636: 1634: 1625: 1622: 1619: 1614: 1604: 1603: 1599: 1597: 1596: 1592: 1591: 1589: 1587: 1578: 1574: 1571: 1568: 1563: 1553: 1552: 1548: 1546: 1545: 1541: 1539: 1538: 1534: 1533: 1531: 1529: 1520: 1516: 1513: 1510: 1500: 1493: 1492: 1485: 1482: 1481: 1474: 1472: 1471: 1464: 1462: 1461: 1454: 1452: 1451: 1444: 1443: 1440: 1437: 1434: 1424: 1418: 1413: 1411: 1406: 1400: 1386: 1385: 1381: 1379: 1378: 1374: 1372: 1371: 1367: 1365: 1364: 1360: 1358: 1357: 1356:Anoualerpeton 1353: 1351: 1350: 1346: 1345: 1343: 1340: 1333: 1329: 1325: 1313: 1312: 1306: 1285: 1284: 1283: 1282: 1280: 1278: 1274: 1266: 1262: 1261: 1260: 1259: 1256: 1253: 1251: 1247: 1239: 1238:Temnospondyli 1235: 1234: 1233: 1232: 1229: 1226: 1224: 1223:Temnospondyli 1220: 1216: 1208: 1204: 1203: 1202: 1201: 1198: 1195: 1193: 1189: 1185: 1177: 1173: 1170: 1168: 1167:Sarcopterygii 1164: 1161: 1159: 1155: 1152: 1150: 1146: 1144: 1140: 1139: 1134: 1130: 1126: 1119: 1114: 1112: 1107: 1105: 1100: 1099: 1096: 1090: 1087: 1085: 1082: 1081: 1077: 1061: 1054: 1051: 1038: 1034: 1027: 1024: 1008: 1004: 1000: 996: 992: 988: 984: 980: 976: 969: 967: 958: 956: 954: 950: 945: 943:1-55209-541-X 939: 935: 931: 930: 922: 920: 916: 910: 905: 902:(1): 39–109. 901: 897: 890: 888: 879: 877: 875: 873: 869: 857:. AmphibiaWeb 856: 852: 845: 843: 839: 834: 828: 825: 820: 814: 811: 806: 802: 798: 794: 787: 780: 777: 772: 768: 764: 760: 756: 752: 748: 744: 743: 735: 733: 731: 729: 725: 712: 706: 704: 702: 700: 696: 691: 685: 681: 676: 675: 666: 663: 650: 644: 642: 640: 636: 623: 617: 615: 613: 609: 596: 590: 588: 584: 579: 573: 557: 552: 548: 544: 543: 538: 536: 527: 525: 523: 519: 513: 511: 509: 508:Madeira River 505: 501: 497: 492: 490: 486: 482: 477: 475: 471: 464: 456: 454: 452: 448: 447:Potamotyphlus 444: 440: 436: 432: 428: 424: 420: 419: 413: 411: 406: 401: 399: 395: 394: 393:Potamotyphlus 389: 388: 383: 379: 375: 371: 367: 363: 362: 352: 350: 348: 344: 340: 335: 332: 328: 324: 320: 316: 309: 305: 300: 298: 294: 290: 286: 282: 275: 273: 271: 267: 266:Typholonectes 263: 262: 261:Potamotyphlus 257: 256: 251: 247: 246: 241: 237: 236:type specimen 232: 228: 227: 217: 212: 207: 201: 199: 193: 190: 189:Binomial name 186: 182: 181: 176: 173: 172: 165: 164: 160: 157: 156: 153: 150: 147: 146: 143: 140: 137: 134: 133: 130: 127: 124: 123: 120: 117: 114: 113: 110: 107: 104: 103: 100: 97: 94: 93: 90: 87: 84: 83: 78: 73: 69: 63: 58: 47: 43: 38: 34: 29: 26: 22: 19: 2574:Atretochoana 2529: 2414:Atretochoana 2400:Atretochoana 2370:Atretochoana 2369: 2287: 2277: 2267: 2257: 2247: 2231:total group) 2183: 2176: 2169: 2162: 2155: 2148: 2125: 2118: 2111: 2104: 2081: 2071: 2061: 2051: 2035:total group) 2033:(salamanders 2017:crown group) 1975: 1968: 1961: 1954: 1947: 1922: 1915: 1908: 1901: 1876: 1869: 1862: 1855: 1848: 1841: 1834: 1809: 1802: 1795: 1780:Caeciliidae 1767: 1759:Typhlonectes 1757: 1750: 1743: 1736: 1731:Atretochoana 1730: 1729: 1704: 1699:Boulengerula 1697: 1672: 1647: 1640: 1602:Uraeotyphlus 1600: 1593: 1549: 1542: 1535: 1504:Gymnophiona 1489: 1478: 1468: 1460:Funcusvermis 1458: 1448: 1402: 1382: 1375: 1368: 1361: 1354: 1349:Albanerpeton 1347: 1311:Lissamphibia 1309: 1277:Lissamphibia 1171: 1162: 1158:Osteichthyes 1153: 1125:Lissamphibia 1064:. Retrieved 1053: 1041:. Retrieved 1026: 1014:. Retrieved 1007:the original 978: 974: 965: 928: 899: 895: 886: 859:. Retrieved 854: 827: 813: 796: 792: 779: 746: 740: 715:. Retrieved 673: 665: 653:. Retrieved 626:. Retrieved 599:. Retrieved 572:cite journal 560:. Retrieved 546: 540: 534: 503: 493: 478: 470:Atretochoana 469: 463:Atretochoana 462: 460: 446: 443:Atretochoana 442: 434: 430: 423:Atretochoana 422: 416: 414: 402: 391: 387:Typhlonectes 385: 382:Josef Eiselt 377: 373: 359: 356: 343:common names 336: 331:Atretochoana 330: 315:Atretochoana 314: 308:Atretochoana 307: 301: 293:Atretochoana 292: 280: 279: 265: 259: 255:Atretochoana 254: 253: 245:Typhlonectes 243: 225: 224: 223: 197: 195: 179: 178: 163:Atretochoana 162: 161: 135: 25:Atretochoana 24: 18: 2712:SeaLifeBase 2634:iNaturalist 2582:AmphibiaWeb 2554:Wikispecies 2448:iNaturalist 2394:Wikispecies 2185:Prodesmodon 1910:Geotrypetes 1864:Indotyphlus 1857:Idiocranium 1850:Hypogeophis 1843:Grandisonia 1836:Gegeneophis 1685:Herpelidae 1660:Chikilidae 1595:Ichthyophis 1544:Epicrionops 1377:Wesserpeton 1370:Shirerpeton 1016:22 February 717:22 February 655:22 February 628:22 February 601:22 February 562:17 November 425:, from the 347:human penis 276:Description 129:Gymnophiona 2744:Categories 2343:See also: 2269:Prosalirus 2210:115 genera 2097:Karauridae 2073:Triassurus 1811:Oscaecilia 1618:Teresomata 1551:Rhinatrema 1450:Eocaecilia 1286:see below↓ 1062:. Mongabay 861:21 January 622:"Taxonomy" 514:References 504:A. eiselti 489:viviparous 297:earthworms 281:A. eiselti 2385:Q15727328 2289:Vieraella 2225:Salientia 2120:Kuzbassia 2011:Batrachia 1977:Siphonops 1903:Dermophis 1871:Praslinia 1769:Ymboirana 1363:Celtedens 1176:Tetrapoda 1141:Kingdom: 1066:11 August 995:1981-8114 711:"Biology" 496:Mosqueiro 327:epidermis 304:vestigial 270:tetrapods 250:monotypic 231:caecilian 174:Species: 95:Kingdom: 89:Eukaryota 2652:10939991 2545:Q1916885 2539:Wikidata 2379:Wikidata 2196:Urodela 2171:Parrisia 2113:Kokartus 2106:Karaurus 1917:Gymnopis 1804:Caecilia 1149:Chordata 1147:Phylum: 1143:Animalia 1043:1 August 1037:Archived 1003:83940114 771:84351569 398:splenial 366:holotype 287:to lack 285:tetrapod 148:Family: 119:Amphibia 109:Chordata 105:Phylum: 99:Animalia 85:Domain: 62:IUCN 3.1 2727:4810107 2704:3617943 2691:1938463 2626:2430995 2613:1025251 2505:3617929 2466:1197800 2440:2430994 2029:Caudata 1797:Apodops 1706:Herpele 1674:Chikila 1537:Amazops 751:Bibcode 483:by the 457:Biology 431:atretos 410:choanae 400:teeth. 353:History 319:choanae 252:genus, 238:in the 208:, 1968) 158:Genus: 125:Order: 115:Class: 60: ( 2724:uBio: 2665:668511 2492:264005 2479:668191 2427:102438 2300:Anura 2229:(frogs 2083:Urupia 2053:Egoria 1384:Yaksha 1001:  993:  940:  769:  686:  498:(near 435:choana 206:Taylor 2717:66922 2678:59493 2647:IRMNG 2639:28097 2513:Plazi 2461:IRMNG 2453:28096 1172:Clade 1163:Clade 1154:Clade 1010:(PDF) 999:S2CID 971:(PDF) 892:(PDF) 789:(PDF) 767:S2CID 680:46–47 500:Belém 466:' 429:word 427:Greek 339:snake 311:' 289:lungs 264:than 142:Apoda 136:Clade 2686:NCBI 2673:IUCN 2660:ITIS 2621:GBIF 2587:1948 2487:NCBI 2474:ITIS 2435:GBIF 1263:see 1236:see 1205:see 1068:2013 1045:2012 1018:2011 991:ISSN 938:ISBN 863:2010 719:2012 684:ISBN 657:2012 630:2012 603:2012 578:link 564:2021 547:2004 390:and 2608:EoL 2595:ASW 2569:ADW 2422:EoL 2409:ASW 983:doi 904:doi 801:doi 759:doi 747:261 551:doi 2746:: 2714:: 2701:: 2688:: 2675:: 2662:: 2649:: 2636:: 2623:: 2610:: 2597:: 2584:: 2571:: 2556:: 2541:: 2515:: 2502:: 2489:: 2476:: 2463:: 2450:: 2437:: 2424:: 2411:: 2396:: 2381:: 1221:/ 1174:: 1165:: 1156:: 997:. 989:. 977:. 973:. 952:^ 936:. 934:81 918:^ 900:62 898:. 894:. 871:^ 853:. 841:^ 797:32 795:. 791:. 765:. 757:. 745:. 727:^ 698:^ 682:. 638:^ 611:^ 586:^ 574:}} 570:{{ 545:. 539:. 521:^ 491:. 349:. 138:: 2285:† 2275:† 2265:† 2255:† 2245:† 2138:† 2094:† 2079:† 2069:† 2059:† 2049:† 1793:† 1765:† 1494:? 1487:† 1483:? 1476:† 1466:† 1456:† 1446:† 1341:? 1336:† 1117:e 1110:t 1103:v 1070:. 1047:. 1020:. 985:: 979:6 946:. 912:. 906:: 865:. 807:. 803:: 773:. 761:: 753:: 721:. 692:. 659:. 632:. 605:. 580:) 566:. 553:: 537:" 533:" 204:( 64:)

Index


Conservation status
Data Deficient
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Amphibia
Gymnophiona
Apoda
Typhlonectidae
Atretochoana
Binomial name
Taylor

caecilian
type specimen
Naturhistorisches Museum
Typhlonectes
monotypic
Potamotyphlus
tetrapods
tetrapod
lungs
earthworms
vestigial
choanae
pulmonary arteries

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