2609:
2615:
2622:
2602:
473:
in northwestern
Wyoming, and is composed of teeth, cranial, and postcranial bones. The bone fragments were broken and scattered suggesting a taphonomic pathway that includes trampling before burial. The holotype specimen (UM 83874) from Wyoming, USA consists of upper and lower dentition with dentary
497:
includes three incisors (I1–I3), one canine (C1), premolars (P2–P4), and molars (M1–M3), and the lower dentition includes two incisors (I1–I2), one canine (C1), premolars (P2–P4), and molars (M1–M3). A lower first incisor has not actually been recovered, but is believed to be present due to the
612:
M. C. McKenna. 1980. Late
Cretaceous and Early Tertiary vertebrate paleontological reconnaissance, Togwotee Pass area, northwestern Wyoming. Aspects of Vertebrate History: Essays in Honor of Edwin Harris Colbert, L. L. Jacobs (ed.), Museum of Northern Arizona Press
602:
W. C. Clyde. 1997. Stratigraphy and mammalian paleontology of the McCullough Peaks, northern
Bighorn Basin, Wyoming: Implications for biochronology, basin development, and community reorganization across the Paleocene-Eocene boundary. PhD Thesis, University of
579:
Rose, K. D., Chew, A. E., Dunn, R. H., Kraus, M. J., Fricke, H. C., & Zack, S. P. (2012). Earliest Eocene mammalian fauna from the
Paleocene-Eocene Thermal Maximum at Sand Creek Divide, southern Bighorn Basin, Wyoming. Museum of Paleontology, University of
502:
are tall and nearly vertical. The anterior teeth, the incisors and canines, were crowded. The second incisor has a massive oval crown. Lower premolars have smooth enamel set in a relatively deep jaw and the third premolar crown is large and bulbous.
616:
P. D. Gingerich and G. F. Gunnell. 1979. Systematics and
Evolution of the Genus Esthonyx (Mammalia, Tillodontia) in the Early Eocene of North America. Contributions from the Museum of Paleontology, University of Michigan
566:
Gingerich, P. D. (1989). New earliest
Wasatchian mammalian fauna from the Eocene of northwestern Wyoming: composition and diversity in a rarely sampled high-floodplain assemblage. Museum of Paleontology, University of
609:
K. D. Rose. 1981. The
Clarkforkian Land-Mammal Age and Mammalian Faunal Composition Across the Paleocene-Eocene Boundary. University of Michigan Papers on Paleontology 26:1-197
2686:
589:
Lucas S. G. and R. M. Schoch. 1998. Tillodontia. In C. M. Janis, K. M. Scott, and L. L. Jacobs (eds.), Evolution of
Tertiary Mammals of North America 1:268-273.
523:, or adapted for climbing. In contrast, bones of the foot suggest a somewhat cursorial or fossorial lifestyle. Worn enamel on the teeth additionally suggest
2750:
606:
D. W. Krause. 1982. Multituberculates from the
Wasatchian Land-Mammal Age, Early Eocene, of Western North America. Journal of Paleontology 56(2):271-294
2745:
2673:
2780:
490:
was relatively small, as indicated by an ulna length of about 50 millimetres (2.0 in) and a mandible about 60 millimetres (2.4 in).
486:
has been recovered, making the exact appearance and body size of the animal relatively difficult to determine. Compared to other tillodonts,
2608:
369:
2785:
642:
423:, an extinct group of mammals characterized by rodent-like incisors, clawed feet, and an elongated rostrum and mandibular symphysis.
2775:
388:
probably was a generalist terrestrial mammal that may have roamed around the ground, but was also capable of climbing trees.
2614:
2621:
515:. The large and laterally compressed claws and shallow trochlea of the astragalus suggests that the mode of life of
360:, about 56 to 50 million years ago. Fifty-six collections that have been recovered thus far include the remains of
2765:
498:
available space in the lower jaw. A lower third incisor was likely extremely reduced or absent. The incisors of
2770:
531:
likely spent a lot of time in trees, but also occasionally searched for food on the ground or dug for roots.
458:(Gingerich and Gunnell, 1979). The genus has a relatively problematic taxonomic history, as the type species
2641:
1497:
635:
262:
2755:
2712:
1238:
2601:
2341:
1527:
2740:
1751:
1440:
470:
345:
2496:
1920:
1781:
1652:
1575:
1431:
1062:
1027:
879:
2760:
2550:
2489:
2481:
2248:
2234:
1809:
1802:
1610:
1506:
1382:
1338:
1210:
1173:
973:
745:
520:
428:
195:
2465:
2400:
2355:
2220:
2178:
2082:
2073:
1941:
1410:
1345:
1295:
1180:
1016:
2436:
2429:
2422:
2393:
2362:
2320:
2313:
2206:
2185:
2062:
1997:
1976:
1892:
1871:
1837:
1700:
1686:
1679:
1638:
1624:
1596:
1561:
1538:
1417:
1352:
1331:
1281:
1152:
1091:
952:
931:
718:
341:
297:
121:
2518:
2327:
2306:
2283:
2227:
2213:
2199:
2055:
1983:
1934:
1913:
1864:
1767:
1742:
1728:
1714:
1645:
1631:
1568:
1547:
1403:
1389:
1324:
1317:
1245:
1231:
1145:
1098:
889:
2717:
2525:
2503:
2334:
2262:
2150:
1962:
1948:
1927:
1906:
1857:
1830:
1823:
1816:
1774:
1617:
1554:
1359:
1288:
1224:
1159:
1105:
2699:
2691:
2458:
2385:
2269:
2192:
2164:
1990:
1760:
1707:
1693:
1582:
1520:
1310:
1217:
772:
628:
2704:
2171:
1252:
2443:
2297:
1885:
1053:
1955:
2109:
2093:
1470:
1136:
2348:
1670:
921:
857:
474:
bone. Post cranial remains include the scapula, ulna, radius, ankle bones, and others.
27:
2734:
2578:
1848:
1603:
1373:
1267:
1083:
1005:
994:
349:
80:
328:
mammal, likely the size of a cat to raccoon, that lived in North
America during the
2510:
2450:
2241:
2048:
2039:
2028:
1969:
1721:
1513:
1479:
1447:
1201:
1187:
1113:
962:
512:
353:
222:
36:
31:
2664:
2562:
2376:
2255:
2141:
2015:
1878:
1396:
1166:
1121:
1038:
900:
420:
325:
182:
55:
431:, but differs from other esthonyids by having an unfused mandibular symphysis.
2571:
2539:
2116:
1899:
1461:
1274:
911:
868:
374:
357:
100:
65:
40:
2585:
2415:
2407:
2123:
2102:
1788:
1589:
1302:
703:
329:
169:
105:
49:
2658:
2473:
2276:
1795:
1486:
1260:
1076:
983:
840:
818:
784:
730:
675:
651:
380:
337:
144:
95:
90:
75:
70:
60:
2678:
1069:
110:
85:
941:
757:
687:
681:
669:
333:
156:
132:
2635:
511:
Tillodonts are considered to be generalists and some may have been
2639:
662:
624:
408:" meaning claw referencing the unjoined claw-like incisors.
620:
442:(Gingerich and Gunnell, 1979) and the additional species
340:. The only fossils that have been recovered are from the
462:
and other species of the genus are alternatively named
2536:
2373:
2294:
2138:
2012:
1739:
1667:
1458:
1428:
1370:
1198:
1133:
1050:
1034:
1023:
1012:
1001:
990:
979:
969:
958:
948:
937:
927:
917:
907:
896:
885:
875:
864:
853:
2648:
2561:
2535:
2372:
2293:
2137:
2092:
2072:
2038:
2011:
1847:
1750:
1738:
1666:
1537:
1496:
1469:
1457:
1427:
1369:
1197:
1132:
1049:
838:
783:
756:
729:
702:
438:are currently known and includes the type species
384:, both of which were ground-dwelling mammals.
636:
8:
352:of Wyoming, United States, and date to the
2636:
2020:
1747:
1466:
846:
835:
762:
735:
708:
699:
659:
643:
629:
621:
17:
540:
7:
575:
573:
562:
560:
558:
556:
554:
552:
550:
548:
546:
544:
2751:Paleocene mammals of North America
14:
2620:
2613:
2607:
2600:
370:Paleocene Eocene Thermal Maximum
2746:Eocene mammals of North America
446:(Gingerich and Gunnell, 1979),
396:The genus name comes from the "
372:along with other mammals like
289:(Gingerich & Gunnell 1979)
273:(Gingerich & Gunnell 1979)
253:(Gingerich & Gunnell 1979)
236:(Gingerich & Gunnell 1979)
1:
2781:Fossil taxa described in 1989
469:The type species comes from
466:by Lucas and Schoch (1998).
2802:
419:is placed in the suborder
2786:Prehistoric mammal genera
2597:
2023:
849:
834:
815:
765:
738:
711:
698:
658:
404:-" meaning joining, and "
336:in the early part of the
303:
296:
246:
241:
228:
221:
122:Scientific classification
120:
20:
482:No complete skeleton of
2776:Paleontology in Wyoming
493:The upper dentition of
310:Lucas & Schoch 1998
427:belongs to the family
2713:Paleobiology Database
527:stripped vegetation.
454:(Simpson, 1937), and
346:Fort Union Formations
1498:Pseudorhyncocyonidae
400:-" meaning absent, "
2728:
2727:
2700:Open Tree of Life
2642:Taxon identifiers
2633:
2632:
2629:
2628:
2595:
2594:
2342:Prozalambdalestes
2133:
2132:
2007:
2006:
1662:
1661:
830:
829:
826:
825:
811:
810:
807:
806:
803:
802:
452:Azygonyx latidens
450:(Simpson, 1937),
448:Azygonyx grangeri
444:Azygonyx ancylion
440:Azygonyx gunnelli
317:
316:
231:Azygonyx gunnelli
2793:
2766:Thanetian genera
2721:
2720:
2708:
2707:
2695:
2694:
2682:
2681:
2669:
2668:
2667:
2637:
2624:
2618:
2617:
2611:
2605:
2604:
2538:
2444:Kharmerungulatum
2375:
2298:Zalambdalestidae
2296:
2140:
2021:
2014:
1748:
1741:
1669:
1528:Pseudorhyncocyon
1467:
1460:
1430:
1372:
1200:
1135:
1054:Adapisoriculidae
1052:
1036:
1025:
1014:
1003:
992:
981:
971:
960:
950:
939:
929:
919:
909:
898:
887:
877:
866:
855:
847:
836:
763:
736:
709:
700:
693:
692:
660:
645:
638:
631:
622:
590:
587:
581:
577:
568:
564:
460:Azygonyx gunelli
456:Azygonyx xenicus
434:Five species of
311:
290:
282:
274:
266:
254:
115:
52:
26:Temporal range:
18:
2801:
2800:
2796:
2795:
2794:
2792:
2791:
2790:
2771:Ypresian genera
2731:
2730:
2729:
2724:
2716:
2711:
2703:
2698:
2690:
2685:
2677:
2672:
2663:
2662:
2657:
2644:
2634:
2625:
2612:
2599:
2591:
2557:
2531:
2368:
2289:
2129:
2110:Psittacotherium
2094:Stylinodontidae
2088:
2068:
2034:
2003:
1843:
1752:Paroxyclaenidae
1734:
1658:
1533:
1492:
1471:Gypsonictopidae
1453:
1441:Ferrequitherium
1423:
1365:
1193:
1137:Asioryctitheria
1128:
1045:
843:
822:
799:
779:
752:
725:
694:
665:
664:
654:
649:
599:
597:Further reading
594:
593:
588:
584:
578:
571:
565:
542:
537:
509:
480:
414:
394:
309:
288:
280:
272:
260:
252:
237:
234:
217:
214:
213:
198:
185:
172:
159:
147:
135:
116:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
47:
46:
44:
34:
24:
12:
11:
5:
2799:
2797:
2789:
2788:
2783:
2778:
2773:
2768:
2763:
2758:
2753:
2748:
2743:
2733:
2732:
2726:
2725:
2723:
2722:
2709:
2696:
2683:
2670:
2654:
2652:
2646:
2645:
2640:
2631:
2630:
2627:
2626:
2598:
2596:
2593:
2592:
2590:
2589:
2582:
2575:
2567:
2565:
2559:
2558:
2556:
2555:
2554:
2553:
2544:
2542:
2533:
2532:
2530:
2529:
2522:
2515:
2507:
2500:
2497:Sheikhdzheilia
2493:
2486:
2478:
2470:
2462:
2455:
2447:
2440:
2433:
2426:
2419:
2412:
2404:
2397:
2390:
2381:
2379:
2370:
2369:
2367:
2366:
2359:
2352:
2349:Zalambdalestes
2345:
2338:
2331:
2324:
2317:
2310:
2302:
2300:
2291:
2290:
2288:
2287:
2280:
2273:
2266:
2259:
2252:
2245:
2238:
2231:
2224:
2217:
2210:
2203:
2196:
2189:
2182:
2175:
2168:
2161:
2154:
2146:
2144:
2135:
2134:
2131:
2130:
2128:
2127:
2120:
2113:
2106:
2098:
2096:
2090:
2089:
2087:
2086:
2078:
2076:
2074:Onychodectidae
2070:
2069:
2067:
2066:
2059:
2052:
2044:
2042:
2036:
2035:
2033:
2032:
2024:
2018:
2009:
2008:
2005:
2004:
2002:
2001:
1994:
1987:
1980:
1973:
1966:
1959:
1952:
1945:
1938:
1931:
1924:
1921:Cryptopithecus
1917:
1910:
1903:
1896:
1889:
1882:
1875:
1868:
1861:
1853:
1851:
1845:
1844:
1842:
1841:
1834:
1827:
1820:
1813:
1806:
1799:
1792:
1785:
1782:Kiinkerishella
1778:
1771:
1764:
1756:
1754:
1745:
1736:
1735:
1733:
1732:
1725:
1718:
1711:
1704:
1697:
1690:
1683:
1675:
1673:
1671:Palaeoryctidae
1664:
1663:
1660:
1659:
1657:
1656:
1649:
1642:
1635:
1628:
1621:
1614:
1607:
1600:
1593:
1586:
1579:
1572:
1565:
1558:
1551:
1543:
1541:
1535:
1534:
1532:
1531:
1524:
1517:
1510:
1502:
1500:
1494:
1493:
1491:
1490:
1483:
1475:
1473:
1464:
1455:
1454:
1452:
1451:
1444:
1436:
1434:
1432:Horolodectidae
1425:
1424:
1422:
1421:
1414:
1407:
1400:
1393:
1386:
1378:
1376:
1367:
1366:
1364:
1363:
1356:
1349:
1342:
1335:
1328:
1321:
1314:
1307:
1299:
1292:
1285:
1278:
1271:
1264:
1257:
1249:
1242:
1235:
1228:
1221:
1214:
1206:
1204:
1195:
1194:
1192:
1191:
1184:
1177:
1170:
1163:
1156:
1149:
1141:
1139:
1130:
1129:
1127:
1126:
1118:
1110:
1102:
1095:
1088:
1080:
1073:
1066:
1063:Adapisoriculus
1058:
1056:
1047:
1046:
1044:
1043:
1032:
1028:Prokennalestes
1021:
1010:
999:
988:
977:
967:
956:
946:
935:
925:
922:Durlstotherium
915:
905:
894:
883:
880:Bayshinoryctes
873:
862:
858:Acristatherium
850:
844:
839:
832:
831:
828:
827:
824:
823:
816:
813:
812:
809:
808:
805:
804:
801:
800:
798:
797:
796:
795:
789:
787:
781:
780:
778:
777:
776:
775:
766:
760:
754:
753:
751:
750:
749:
748:
739:
733:
727:
726:
724:
723:
722:
721:
712:
706:
696:
695:
691:
690:
684:
678:
672:
663:
656:
655:
650:
648:
647:
640:
633:
625:
619:
618:
614:
610:
607:
604:
598:
595:
592:
591:
582:
569:
539:
538:
536:
533:
508:
505:
479:
476:
413:
410:
393:
390:
315:
314:
313:
312:
301:
300:
294:
293:
292:
291:
283:
281:(Simpson 1937)
275:
267:
255:
244:
243:
239:
238:
235:
226:
225:
219:
218:
216:Gingerich 1989
215:
206:
204:
200:
199:
193:
191:
187:
186:
180:
178:
174:
173:
167:
165:
161:
160:
155:
153:
149:
148:
143:
141:
137:
136:
131:
129:
125:
124:
118:
117:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
53:
28:Late Paleocene
25:
13:
10:
9:
6:
4:
3:
2:
2798:
2787:
2784:
2782:
2779:
2777:
2774:
2772:
2769:
2767:
2764:
2762:
2759:
2757:
2754:
2752:
2749:
2747:
2744:
2742:
2739:
2738:
2736:
2719:
2714:
2710:
2706:
2701:
2697:
2693:
2688:
2684:
2680:
2675:
2671:
2666:
2660:
2656:
2655:
2653:
2651:
2647:
2643:
2638:
2623:
2619:
2616:
2610:
2603:
2588:
2587:
2583:
2581:
2580:
2579:Boreoeutheria
2576:
2574:
2573:
2569:
2568:
2566:
2564:
2560:
2552:
2548:
2547:
2546:
2545:
2543:
2541:
2534:
2528:
2527:
2523:
2521:
2520:
2516:
2513:
2512:
2508:
2506:
2505:
2501:
2499:
2498:
2494:
2492:
2491:
2490:Parazhelestes
2487:
2484:
2483:
2482:Paranyctoides
2479:
2476:
2475:
2471:
2468:
2467:
2463:
2461:
2460:
2456:
2453:
2452:
2448:
2446:
2445:
2441:
2439:
2438:
2434:
2432:
2431:
2427:
2425:
2424:
2420:
2418:
2417:
2413:
2410:
2409:
2405:
2403:
2402:
2398:
2396:
2395:
2391:
2388:
2387:
2383:
2382:
2380:
2378:
2371:
2365:
2364:
2360:
2358:
2357:
2353:
2351:
2350:
2346:
2344:
2343:
2339:
2337:
2336:
2332:
2330:
2329:
2325:
2323:
2322:
2318:
2316:
2315:
2311:
2309:
2308:
2304:
2303:
2301:
2299:
2292:
2286:
2285:
2281:
2279:
2278:
2274:
2272:
2271:
2267:
2265:
2264:
2260:
2258:
2257:
2253:
2251:
2250:
2249:Plesiesthonyx
2246:
2244:
2243:
2239:
2237:
2236:
2235:Megalesthonyx
2232:
2230:
2229:
2225:
2223:
2222:
2218:
2216:
2215:
2211:
2209:
2208:
2204:
2202:
2201:
2197:
2195:
2194:
2190:
2188:
2187:
2183:
2181:
2180:
2176:
2174:
2173:
2169:
2167:
2166:
2162:
2160:
2159:
2155:
2153:
2152:
2148:
2147:
2145:
2143:
2136:
2126:
2125:
2121:
2119:
2118:
2114:
2112:
2111:
2107:
2105:
2104:
2100:
2099:
2097:
2095:
2091:
2085:
2084:
2080:
2079:
2077:
2075:
2071:
2065:
2064:
2060:
2058:
2057:
2053:
2051:
2050:
2046:
2045:
2043:
2041:
2037:
2031:
2030:
2026:
2025:
2022:
2019:
2017:
2010:
2000:
1999:
1995:
1993:
1992:
1988:
1986:
1985:
1981:
1979:
1978:
1974:
1972:
1971:
1967:
1965:
1964:
1960:
1958:
1957:
1953:
1951:
1950:
1946:
1944:
1943:
1939:
1937:
1936:
1932:
1930:
1929:
1925:
1923:
1922:
1918:
1916:
1915:
1911:
1909:
1908:
1904:
1902:
1901:
1897:
1895:
1894:
1890:
1888:
1887:
1883:
1881:
1880:
1876:
1874:
1873:
1869:
1867:
1866:
1862:
1860:
1859:
1855:
1854:
1852:
1850:
1849:Pantolestidae
1846:
1840:
1839:
1835:
1833:
1832:
1828:
1826:
1825:
1821:
1819:
1818:
1814:
1812:
1811:
1810:Paroxyclaenus
1807:
1805:
1804:
1803:Paraspaniella
1800:
1798:
1797:
1793:
1791:
1790:
1786:
1784:
1783:
1779:
1777:
1776:
1772:
1770:
1769:
1765:
1763:
1762:
1758:
1757:
1755:
1753:
1749:
1746:
1744:
1737:
1731:
1730:
1726:
1724:
1723:
1719:
1717:
1716:
1712:
1710:
1709:
1705:
1703:
1702:
1698:
1696:
1695:
1691:
1689:
1688:
1684:
1682:
1681:
1677:
1676:
1674:
1672:
1665:
1655:
1654:
1650:
1648:
1647:
1643:
1641:
1640:
1636:
1634:
1633:
1629:
1627:
1626:
1622:
1620:
1619:
1615:
1613:
1612:
1611:Myrmecoboides
1608:
1606:
1605:
1604:Megaleptictis
1601:
1599:
1598:
1594:
1592:
1591:
1587:
1585:
1584:
1580:
1578:
1577:
1573:
1571:
1570:
1566:
1564:
1563:
1559:
1557:
1556:
1552:
1550:
1549:
1545:
1544:
1542:
1540:
1536:
1530:
1529:
1525:
1523:
1522:
1518:
1516:
1515:
1511:
1509:
1508:
1507:Diaphyodectes
1504:
1503:
1501:
1499:
1495:
1489:
1488:
1484:
1482:
1481:
1477:
1476:
1474:
1472:
1468:
1465:
1463:
1456:
1450:
1449:
1445:
1443:
1442:
1438:
1437:
1435:
1433:
1426:
1420:
1419:
1415:
1413:
1412:
1408:
1406:
1405:
1401:
1399:
1398:
1394:
1392:
1391:
1387:
1385:
1384:
1383:Archaeoryctes
1380:
1379:
1377:
1375:
1374:Didymoconidae
1368:
1362:
1361:
1357:
1355:
1354:
1350:
1348:
1347:
1343:
1341:
1340:
1339:Protentomodon
1336:
1334:
1333:
1329:
1327:
1326:
1322:
1320:
1319:
1315:
1313:
1312:
1308:
1305:
1304:
1300:
1298:
1297:
1293:
1291:
1290:
1286:
1284:
1283:
1279:
1277:
1276:
1272:
1270:
1269:
1268:Chacopterygus
1265:
1263:
1262:
1258:
1255:
1254:
1250:
1248:
1247:
1243:
1241:
1240:
1236:
1234:
1233:
1229:
1227:
1226:
1222:
1220:
1219:
1215:
1213:
1212:
1211:Aboletylestes
1208:
1207:
1205:
1203:
1196:
1190:
1189:
1185:
1183:
1182:
1178:
1176:
1175:
1174:Sasayamamylos
1171:
1169:
1168:
1164:
1162:
1161:
1157:
1155:
1154:
1150:
1148:
1147:
1143:
1142:
1140:
1138:
1131:
1124:
1123:
1119:
1116:
1115:
1111:
1108:
1107:
1103:
1101:
1100:
1096:
1094:
1093:
1089:
1086:
1085:
1084:Deccanolestes
1081:
1079:
1078:
1074:
1072:
1071:
1067:
1065:
1064:
1060:
1059:
1057:
1055:
1048:
1041:
1040:
1033:
1030:
1029:
1022:
1019:
1018:
1011:
1008:
1007:
1006:Montanalestes
1000:
997:
996:
995:Microtherulum
989:
986:
985:
978:
976:
975:
974:Indoclemensia
968:
965:
964:
957:
955:
954:
947:
944:
943:
936:
934:
933:
926:
924:
923:
916:
914:
913:
906:
903:
902:
895:
892:
891:
884:
882:
881:
874:
871:
870:
863:
860:
859:
852:
851:
848:
845:
842:
837:
833:
821:
820:
814:
793:
792:
791:
790:
788:
786:
782:
774:
770:
769:
768:
767:
764:
761:
759:
755:
747:
743:
742:
741:
740:
737:
734:
732:
728:
720:
716:
715:
714:
713:
710:
707:
705:
701:
697:
689:
685:
683:
679:
677:
673:
671:
667:
666:
661:
657:
653:
646:
641:
639:
634:
632:
627:
626:
623:
617:25(7):125-153
615:
611:
608:
605:
601:
600:
596:
586:
583:
576:
574:
570:
563:
561:
559:
557:
555:
553:
551:
549:
547:
545:
541:
534:
532:
530:
526:
522:
518:
514:
506:
504:
501:
496:
491:
489:
485:
477:
475:
472:
467:
465:
461:
457:
453:
449:
445:
441:
437:
432:
430:
429:Esthonychidae
426:
422:
418:
411:
409:
407:
403:
399:
391:
389:
387:
383:
382:
377:
376:
371:
368:survived the
367:
363:
359:
355:
351:
350:Bighorn Basin
347:
343:
339:
335:
331:
327:
323:
322:
308:
305:
304:
302:
299:
295:
287:
284:
279:
276:
271:
268:
264:
259:
256:
251:
248:
247:
245:
240:
233:
232:
227:
224:
220:
212:
211:
205:
202:
201:
197:
196:Esthonychidae
192:
189:
188:
184:
179:
176:
175:
171:
166:
163:
162:
158:
154:
151:
150:
146:
142:
139:
138:
134:
130:
127:
126:
123:
119:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
51:
42:
38:
33:
29:
23:
19:
16:
2756:Clarkforkian
2649:
2606:
2584:
2577:
2570:
2524:
2517:
2511:Valentinella
2509:
2502:
2495:
2488:
2480:
2472:
2466:Mistralestes
2464:
2457:
2451:Khuduklestes
2449:
2442:
2435:
2428:
2421:
2414:
2406:
2401:Avitotherium
2399:
2392:
2384:
2361:
2356:Zhangolestes
2354:
2347:
2340:
2333:
2326:
2319:
2312:
2305:
2282:
2275:
2268:
2261:
2254:
2247:
2242:Meiostylodon
2240:
2233:
2226:
2221:Kuanchuanius
2219:
2212:
2205:
2198:
2191:
2184:
2179:Chungchienia
2177:
2170:
2163:
2157:
2156:
2149:
2122:
2115:
2108:
2101:
2083:Onychodectes
2081:
2061:
2054:
2049:Conoryctella
2047:
2040:Conoryctidae
2029:Schowalteria
2027:
1996:
1989:
1982:
1975:
1970:Palaeosinopa
1968:
1961:
1954:
1947:
1942:Gobipithecus
1940:
1933:
1926:
1919:
1912:
1905:
1898:
1891:
1884:
1877:
1870:
1863:
1856:
1836:
1829:
1822:
1815:
1808:
1801:
1794:
1787:
1780:
1773:
1766:
1759:
1727:
1722:Palaeoryctes
1720:
1713:
1706:
1699:
1692:
1685:
1678:
1651:
1644:
1637:
1630:
1623:
1616:
1609:
1602:
1595:
1588:
1581:
1574:
1567:
1560:
1553:
1546:
1526:
1519:
1514:Leptictidium
1512:
1505:
1485:
1480:Gypsonictops
1478:
1448:Horolodectes
1446:
1439:
1416:
1411:Kennatherium
1409:
1402:
1395:
1388:
1381:
1358:
1351:
1346:Puercolestes
1344:
1337:
1330:
1323:
1316:
1309:
1301:
1296:Ilerdoryctes
1294:
1287:
1280:
1273:
1266:
1259:
1251:
1244:
1237:
1230:
1223:
1216:
1209:
1202:Cimolestidae
1188:Ukhaatherium
1186:
1181:Uchkudukodon
1179:
1172:
1165:
1158:
1151:
1144:
1120:
1114:Sahnitherium
1112:
1104:
1097:
1090:
1082:
1075:
1068:
1061:
1037:
1026:
1017:Murtoilestes
1015:
1004:
993:
982:
972:
963:Hyotheridium
961:
951:
940:
930:
920:
910:
899:
888:
878:
867:
856:
817:
585:
528:
524:
516:
513:rhizophagous
510:
507:Paleoecology
499:
494:
492:
487:
483:
481:
468:
463:
459:
455:
451:
447:
443:
439:
435:
433:
424:
416:
415:
405:
401:
397:
395:
385:
379:
373:
365:
361:
354:Clarkforkian
338:Cenozoic Era
324:was a small
320:
319:
318:
306:
285:
277:
269:
257:
249:
230:
229:
223:Type species
209:
208:
37:Clarkforkian
32:Early Eocene
21:
15:
2741:Tillodontia
2563:Placentalia
2437:Gallolestes
2430:Eozhelestes
2423:Eoungulatum
2394:Aspanlestes
2377:Zhelestidae
2363:Zofialestes
2321:Barunlestes
2314:Anchilestes
2256:Plethorodon
2207:Higotherium
2186:Dysnoetodon
2142:Tillodontia
2063:Huerfanodon
2016:Taeniodonta
1998:Todralestes
1977:Pantolestes
1893:Bouffinomus
1879:Bisonalveus
1872:Bessoecetor
1838:Vulpavoides
1701:Lainoryctes
1687:Aceroryctes
1680:Aaptoryctes
1639:Prodiacodon
1625:Palaeictops
1597:Leptonysson
1562:Gallolestes
1539:Leptictidae
1418:Zeuctherium
1397:Didymoconus
1353:Scollardius
1332:Procerberus
1282:Didelphodus
1167:Kennalestes
1153:Bulaklestes
1122:Wyonycteris
1092:Garatherium
1039:Sinodelphys
953:Hovurlestes
932:Endotherium
901:Cokotherium
478:Description
471:Park County
421:Tillodontia
278:A. latidens
270:A. gunnelli
258:A. grangeri
250:A. ancylion
183:Tillodontia
48:56–50
2761:Wasatchian
2735:Categories
2572:Afrotheria
2551:Pantodonta
2540:Pantodonta
2519:Zhalmousia
2328:Beleutinus
2307:Alymlestes
2284:Yuesthonyx
2228:Lofochaius
2214:Interogale
2200:Franchaius
2117:Stylinodon
2056:Conoryctes
1984:Pentacodon
1935:Galethylax
1914:Coriphagus
1900:Buxolestes
1865:Aphronorus
1768:Euhookeria
1743:Pantolesta
1729:Pinoryctes
1715:Ottoryctes
1646:Protictops
1632:Praolestes
1569:Ictopidium
1548:Amphigyion
1462:Leptictida
1404:Hunanictis
1390:Ardynictis
1325:Pararyctes
1318:Paleotomus
1275:Cimolestes
1246:Bagalestes
1232:Ambilestes
1146:Asioryctes
1099:Namibiodon
912:Durlstodon
890:Bobolestes
869:Ambolestes
794:see below↓
746:Cynodontia
731:Cynodontia
686:Subclass:
535:References
521:scansorial
375:Phenacodus
358:Wasatchian
286:A. xenicus
177:Suborder:
41:Wasatchian
2665:Q21078353
2586:Xenarthra
2526:Zhelestes
2504:Sorlestes
2416:Borisodon
2408:Azilestes
2335:Kulbeckia
2263:Simplodon
2151:Adapidium
2124:Wortmania
2103:Ectoganus
1963:Pagonomus
1949:Kochictis
1928:Dyspterna
1907:Chadronia
1858:Amaramnis
1831:Spaniella
1824:Sororodon
1817:Pugiodens
1789:Kopidodon
1775:Fratrodon
1618:Ongghonia
1590:Leptictis
1555:Blacktops
1360:Tsaganius
1303:Maelestes
1289:Gelastops
1239:Avunculus
1225:Alveugena
1160:Daulestes
1106:Remiculus
719:Synapsida
704:Synapsida
668:Kingdom:
580:Michigan.
567:Michigan.
392:Etymology
330:Paleocene
326:tillodont
170:Cimolesta
128:Kingdom:
2659:Wikidata
2650:Azygonyx
2474:Oxlestes
2459:Lainodon
2386:Alostera
2277:Trogosus
2270:Tillodon
2193:Esthonyx
2165:Basalina
2158:Azygonyx
1991:Thelysia
1796:Merialus
1761:Dulcidon
1708:Nuryctes
1694:Eoryctes
1583:Lainodon
1521:Phakodon
1487:Sikuomys
1311:Naranius
1261:Betonnia
1218:Acmeodon
1077:Bustylus
984:Juramaia
841:Eutheria
819:Eutheria
785:Eutheria
773:Mammalia
758:Mammalia
682:Mammalia
676:Chordata
674:Phylum:
670:Animalia
652:Eutheria
603:Michigan
529:Azygonyx
525:Azygonyx
517:Azygonyx
500:Azygonyx
495:Azygonyx
488:Azygonyx
484:Azygonyx
464:Esthonyx
436:Azygonyx
425:Azygonyx
417:Azygonyx
412:Taxonomy
386:Azygonyx
381:Ectocion
366:Azygonyx
362:Azygonyx
342:Willwood
321:Azygonyx
307:Esthonyx
298:Synonyms
242:Species
210:Azygonyx
190:Family:
157:Mammalia
145:Chordata
140:Phylum:
133:Animalia
22:Azygonyx
2705:4945427
2692:1155184
2679:4831616
2172:Benaius
1253:Batodon
1070:Afrodon
680:Class:
613:323-343
348:in the
263:Simpson
203:Genus:
164:Order:
152:Class:
1886:Bogdia
942:Eomaia
688:Theria
334:Eocene
2718:40546
2687:IRMNG
1956:Oboia
1653:Wania
1576:Labes
402:zygos
265:1937)
2674:GBIF
2549:see
771:see
744:see
717:see
519:was
406:onyx
378:and
344:and
332:and
56:PreꞒ
356:to
2737::
2715::
2702::
2689::
2676::
2661::
572:^
543:^
364:.
106:Pg
50:Ma
2537:†
2514:?
2485:?
2477:?
2469:?
2454:?
2411:?
2389:?
2374:†
2295:†
2139:†
2013:†
1740:†
1668:†
1459:†
1429:†
1371:†
1306:?
1256:?
1199:†
1134:†
1125:?
1117:?
1109:?
1087:?
1051:†
1042:?
1035:†
1031:?
1024:†
1020:?
1013:†
1009:?
1002:†
998:?
991:†
987:?
980:†
970:†
966:?
959:†
949:†
945:?
938:†
928:†
918:†
908:†
904:?
897:†
893:?
886:†
876:†
872:?
865:†
861:?
854:†
644:e
637:t
630:v
398:a
261:(
207:†
194:†
181:†
168:†
111:N
101:K
96:J
91:T
86:P
81:C
76:D
71:S
66:O
61:Ꞓ
45:~
43:)
39:-
35:(
30:-
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