108:
506:
have distinct bifurcating ridges on the nutlet which are unique to section Torada. The overall shape of the samara is ovoid with an average length of up to 3.3 centimetres (1.3 in) and a wing width of 1.4 centimetres (0.55 in). The paired samaras for
568:
610:
Moss, P. T.; Greenwood, D. R.; Archibald, S. B. (2005). "Regional and local vegetation community dynamics of the Eocene
Okanagan Highlands (British Columbia β Washington State) from palynology".
584:
382:. The Allenby formation is similar with the birch and golden larch as strong pollen signals and traces of fir and spruce, however the cypress and palm are not distinct signals.
740:
679:
745:
735:
358:
in nature. The plant community preserved in the
Klondike Mountain formation is a mixed confer-broad leaf forest with large pollen elements of
750:
511:
have a 45Β° attachment angle and the attachment scar on the nutlet is 0.8 centimetres (0.31 in). While very similar in morphology to
397:
327:
730:
725:
424:
409:
330:
USNM 11018 in the Toroda Creek Graben of
Northeast Washington State. Location USNM 11018 is an outcropping of early Eocene,
107:
350:. Both the Allenby and Klondike Mountain Formations preserve upland temperate floras which were first interpreted as being
347:
335:
478:
were known only from the
Republic and Princeton fossil localities when first described. Of the three species, both
755:
413:
401:
647:
244:
515:, the two possibly related species can be separated by the coarse reticulum of veins which is found on
619:
310:
216:
474:
428:
301:
102:
692:
343:
697:
627:
464:
405:
305:
145:
623:
577:
Journal of the
Faculty of Science, Hokkaido University. Series 4, Geology and Mineralogy
720:
432:
293:
132:
714:
420:
417:
389:
263:
61:
363:
351:
297:
490:
has been described from both leaves and samaras. The three are among a number of
393:
355:
286:
191:
36:
670:
181:
81:
46:
392:, the holotype, number "USNM 396010 A,B" which is currently preserved in the
494:
species described from the
Republic and Princeton sites by Wolfe and Tanai.
444:
379:
86:
408:
and the paratype, "UBCB 2400", is in the paleobotanical collection of the
664:
375:
339:
331:
323:
158:
76:
71:
56:
51:
41:
322:
The species was described from a pair of complete fruit specimens. The
684:
448:
308:, Canada. It is one of three species belonging to the extinct section
282:
91:
66:
371:
290:
171:
641:
631:
454:
is in recognition of the type locality in the Toroda Creek Graben.
359:
279:
201:
119:
645:
367:
458:
is one of three species assigned to the extinct section
304:, United States, and the adjacent area of south central
441:
Journal of the
Faculty of Science, Hokkaido University
342:
specimen was recovered from sediments of the
Ypresian
654:
573:(maples) in the Cenozoic of Western North America"
354:, however further study has shown them to be more
326:fossil was recovered from Resner Canyon, location
425:United States Geological Survey, Denver office
569:"Systematics, Phylogeny, and Distribution of
486:are only known from the samara fossils, only
8:
562:
560:
558:
556:
554:
552:
550:
548:
642:
605:
603:
601:
20:
431:. Wolfe and Tanai published their 1987
296:. The species is solely known from the
544:
7:
535:by its notably larger overall size.
462:. The three members of the section
366:, but also having notable traces of
346:exposed in the Blue Flame mine near
612:Canadian Journal of Earth Sciences
398:National Museum of Natural History
328:National Museum of Natural History
14:
416:. The specimens were studied by
741:Ypresian plants of North America
106:
567:Wolfe, J.A.; Tanai, T. (1987).
300:sediments exposed in northeast
746:Extinct flora of North America
527:are distinguishable from both
410:University of British Columbia
1:
736:Fossil taxa described in 1987
388:was described from a pair of
751:Klondike Mountain Formation
348:Princeton, British Columbia
336:Klondike Mountain Formation
772:
583:(1): 1β246. Archived from
414:Victoria, British Columbia
396:collections housed at the
318:History and classification
334:, shale belonging to the
250:
243:
103:Scientific classification
101:
23:
731:Plants described in 1987
16:Extinct species of maple
726:Prehistoric angiosperms
427:and Toshimasa Tanai of
402:Smithsonian Institution
624:2005CaJES..42..187M
429:Hokkaido University
289:described from two
523:. The samaras of
266:& Tanai, 1987
756:Allenby Formation
708:
707:
693:Open Tree of Life
648:Taxon identifiers
513:A. washingtonense
488:A. washingtonense
475:A. washingtonense
344:Allenby Formation
271:
270:
236:A. toradense
763:
701:
700:
688:
687:
675:
674:
673:
643:
636:
635:
607:
596:
595:
593:
592:
564:
533:A.washingtonense
521:A.washingtonense
433:type description
406:Washington, D.C.
306:British Columbia
302:Washington state
259:
255:
221:
215:
111:
110:
96:
33:
29:Temporal range:
21:
771:
770:
766:
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632:10.1139/E04-095
609:
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599:
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566:
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541:
502:The samaras of
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320:
267:
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253:
252:
239:
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31:
30:
27:
17:
12:
11:
5:
769:
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738:
733:
728:
723:
713:
712:
706:
705:
703:
702:
689:
676:
660:
658:
656:Acer toradense
652:
651:
646:
638:
637:
618:(2): 187β204.
597:
543:
542:
540:
537:
525:A. stonebergae
504:Acer toradense
499:
496:
480:A. stonebergae
465:A. stonebergae
447:of the chosen
418:paleobotanists
400:, part of the
394:paleobotanical
386:Acer toradense
319:
316:
285:in the family
278:is an extinct
275:Acer toradense
269:
268:
262:
257:Acer toradense
248:
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60:
55:
50:
45:
40:
35:
34:
28:
25:Acer toradense
15:
13:
10:
9:
6:
4:
3:
2:
768:
757:
754:
752:
749:
747:
744:
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734:
732:
729:
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724:
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719:
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629:
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587:on 2011-10-04
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497:
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477:
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467:
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453:
450:
449:specific name
446:
442:
438:
434:
430:
426:
422:
421:Jack A. Wolfe
419:
415:
411:
407:
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249:
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245:Binomial name
242:
238:
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228:
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224:
223:
212:
209:
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205:
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167:
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138:
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134:
133:Tracheophytes
131:
128:
125:
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118:
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114:
109:
104:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
655:
615:
611:
589:. Retrieved
585:the original
580:
576:
570:
532:
529:A. toradense
528:
524:
520:
517:A. toradense
516:
512:
509:A. toradense
508:
503:
501:
491:
487:
484:A. toradense
483:
479:
473:
470:A. toradense
469:
463:
459:
456:A. toradense
455:
451:
440:
437:A. toradense
436:
385:
384:
364:golden larch
352:Microthermal
321:
309:
298:Early Eocene
274:
273:
272:
256:
251:
235:
234:
217:
202:
165:
152:
139:
126:
24:
18:
498:Description
356:mesothermal
287:Sapindaceae
192:Sapindaceae
146:Angiosperms
715:Categories
591:2011-11-16
539:References
182:Sapindales
452:toradense
445:etymology
390:specimens
229:Species:
210:Section:
116:Kingdom:
671:Q4673179
665:Wikidata
519:and not
340:paratype
332:Ypresian
324:holotype
188:Family:
159:Eudicots
32:Ypresian
698:7642123
685:9177769
620:Bibcode
443:. The
439:in the
423:of the
376:cypress
338:. The
294:samaras
283:species
254:†
233:†
198:Genus:
178:Order:
120:Plantae
472:, and
460:Torada
378:, and
372:spruce
311:Torada
291:fossil
222:Torada
172:Rosids
721:Maple
360:birch
280:maple
264:Wolfe
220:sect.
218:Acer
166:Clade
153:Clade
140:Clade
127:Clade
680:GBIF
571:Acer
531:and
492:Acer
482:and
435:for
380:palm
362:and
203:Acer
37:Preκ
628:doi
412:in
404:in
368:fir
717::
695::
682::
667::
626:.
616:42
614:.
600:^
581:22
579:.
575:.
547:^
468:,
374:,
370:,
314:.
168::
155::
142::
129::
87:Pg
634:.
630::
622::
594:.
214:β
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:κ
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