512:
128:
470:
146:
378:
in the same amber matrix. However, he refrained from describing this individual due to his inability to reliably determine its anatomical details. New imaging techniques developed since then allowed for later researchers to describe the smaller specimen in detail.
488:
legs. The size of the claws on the first, second, and third legs are generally similar, but the claws on the fourth leg are drastically different in size and shape.
1314:
1289:
1196:
503:
are visible. An elliptical organ is faintly visible on the side of the head, and the mouth hole at the front of the head is smooth.
1230:
1299:
374:, Kenneth W. Cooper noted the presence of a much smaller—seemingly poorly preserved—tardigrade potentially distinct from
345:
1158:
500:
1294:
1304:
145:
511:
1224:
Mapalo, Marc A.; Robin, Ninon; Boudinot, Brendon E.; Ortega-Hernández, Javier; Barden, Phillip (2021-10-13).
1309:
412:
349:
1197:"A New Eutardigrade (Tardigrada: Milnesiidae) in amber from the Upper Cretaceous (Turonian) of New Jersey"
1092:
197:
890:
357:
348:
PALE-4586 is embedded in a piece of amber collected in 1940 by
William M. Legg in Manitoba, Canada. The
322:
251:
127:
461:, the youngest known fossil tardigrade from the Miocene of the Dominican Republic, was named in 2021.
389:
as a new genus and species of tardigrade based on the smaller individual preserved in amber alongside
1046:
647:
495:
holotype specimen measures about 0.10 millimeters in length. There are no clear protuberances on the
441:
228:
873:
626:
611:
469:
931:
907:
383:
835:
757:
552:
290:
207:
1037:
362:
353:
140:
1207:
1265:
1247:
1177:
1127:
1109:
1053:
777:
457:
1255:
1239:
1167:
1117:
1101:
516:
435:
represents one of the only known fossil tardigrade species. The first named tardigrade was
411:", denoting "air", in reference to the holotype's apparent midair suspension in amber. The
312:
294:
48:
1122:
1087:
1260:
1225:
1204:
Studies on
Fossils in Amber, with Particular Reference to the Cretaceous of New Jersey
1283:
733:
536:
402:
85:
1149:
849:
685:
187:
425:
17:
951:
317:
60:
1105:
1088:"Cretaceous amber inclusions illuminate the evolutionary origin of tardigrades"
1059:
709:
661:
407:
286:
177:
105:
70:
1251:
1181:
1113:
581:
556:
447:
157:
110:
54:
1269:
1243:
1131:
1086:
Mapalo, Marc A.; Wolfe, Joanna M.; Ortega-Hernández, Javier (2024-08-06).
1172:
452:
341:
326:
100:
95:
80:
75:
65:
429:", meaning "finger", referencing the distinctive claws of the holotype.
496:
485:
283:
115:
90:
31:
167:
1206:. Netherlands: Backhuys Publishers. pp. 103–110. Archived from
455:) of New Jersey—in 2000, representing the oldest known tardigrade.
510:
468:
420:
394:
360:. The individual is curled and notably shriveled. The holotype of
304:
484:
is compact and barrel-like, with four short pairs of claw-tipped
540:
321:, another extinct tardigrade. The specimen was found near
1231:
Proceedings of the Royal
Society B: Biological Sciences
559:
below, with extinct species designated with a dagger (
480:
Like in other tardigrades, the general body shape of
967:
947:
773:
560:
551:
as closely related members of the tardigrade family
135:Holotype specimen under various imaging techniques
535:, Mapalo, Wolfe & Ortega-Hernández (2024)
382:In 2024, Mapalo, Wolfe & Ortega-Hernández
303:, known from a single individual preserved in
8:
1195:Bertolani, Roberto; Grimaldi, David (2000).
539:a combination of morphological features and
311:holotype is preserved in the same piece of
1081:
1079:
1077:
1075:
270:Mapalo, Wolfe & Ortega-Hernández, 2024
126:
38:
1259:
1171:
1121:
366:is preserved in the same amber piece as
1143:
1141:
1071:
227:Mapalo, Wolfe & Ortega-Hernández,
555:. These results are displayed in the
7:
1315:Monotypic prehistoric animal genera
923:
865:
841:
749:
725:
701:
677:
653:
604:
597:
573:
566:
289:("water bears") of the superfamily
531:To determine the relationships of
25:
1226:"A tardigrade in Dominican amber"
1148:Cooper, Kenneth W. (1964-01-01).
1060:Tardigrade (evolutionary history)
144:
1159:Psyche: A Journal of Entomology
445:—a species of the extant genus
401:, is derived from the modified
1154:Cooper, from Cretaceous amber"
1150:"The first fossil tardigrade:
1:
1290:Fossil taxa described in 2024
370:. In the 1964 description of
1202:. In Grimaldi, David (ed.).
491:In its shriveled state, the
439:in 1964. It was followed by
27:Genus of extinct tardigrades
1331:
1106:10.1038/s42003-024-06643-2
543:sequences. They recovered
451:from the Late Cretaceous (
29:
965:
945:
928:
921:
904:
887:
870:
863:
850:Ramazzottius oberhaeuseri
846:
839:
771:
754:
747:
730:
723:
706:
699:
682:
675:
658:
651:
624:
609:
602:
595:
578:
571:
537:phylogenetically analyzed
257:
250:
141:Scientific classification
139:
134:
125:
41:
874:Microhypsibius truncatus
686:Isohypsibius prosostomus
473:Details of the claws of
30:Not to be confused with
293:. The genus contains a
1300:Parachela (tardigrade)
1244:10.1098/rspb.2021.1760
1093:Communications Biology
734:Doryphoribius macrodon
528:
477:
419:, is derived from the
908:Calohypsibius ornatus
891:Acutuncus antarcticus
758:Hexapodibius micronyx
547:and the contemporary
514:
472:
1047:Milnesium swolenskyi
442:Milnesium swolenskyi
333:Discovery and naming
932:Hypsibius dujardini
662:Ramajendas frigidus
1173:10.1155/1964/48418
710:Halobiotus crispae
529:
478:
354:Saskatchewan River
352:is near where the
1295:Tardigrade genera
1054:Paradoryphoribius
1028:
1027:
1019:
1018:
1010:
1009:
1001:
1000:
992:
991:
983:
982:
825:
824:
816:
815:
807:
806:
798:
797:
789:
788:
778:Paradoryphoribius
637:
636:
458:Paradoryphoribius
387:Aerobius dactylus
275:
274:
264:Aerobius dactylus
231:
18:Aerobius dactylus
16:(Redirected from
1322:
1305:Campanian genera
1274:
1273:
1263:
1221:
1215:
1214:
1212:
1201:
1192:
1186:
1185:
1175:
1145:
1136:
1135:
1125:
1083:
969:
949:
924:
866:
842:
775:
750:
726:
702:
678:
654:
605:
598:
574:
567:
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517:life restoration
266:
262:
243:A. dactylus
226:
219:
149:
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130:
120:
57:
47:Temporal range:
39:
21:
1330:
1329:
1325:
1324:
1323:
1321:
1320:
1319:
1280:
1279:
1278:
1277:
1223:
1222:
1218:
1210:
1199:
1194:
1193:
1189:
1147:
1146:
1139:
1085:
1084:
1073:
1068:
1034:
1029:
1020:
1011:
1002:
993:
984:
826:
817:
808:
799:
790:
648:Isohypsibioidea
638:
509:
467:
335:
313:Late Cretaceous
271:
268:
260:
259:
246:
225:
217:
143:
121:
119:
118:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
53:
52:
49:Late Cretaceous
45:
35:
28:
23:
22:
15:
12:
11:
5:
1328:
1326:
1318:
1317:
1312:
1310:Canadian amber
1307:
1302:
1297:
1292:
1282:
1281:
1276:
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1216:
1213:on 2022-03-15.
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631:
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627:Macrobiotoidea
623:
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619:
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612:Eohypsibioidea
608:
603:
601:
596:
594:
591:
590:
587:
586:
577:
572:
570:
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519:of the coeval
508:
507:Classification
505:
466:
463:
334:
331:
295:single species
282:is a genus of
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569:
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515:Hypothetical
513:
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460:
459:
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422:
418:
414:
413:specific name
410:
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396:
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388:
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369:
365:
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359:
355:
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350:type locality
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252:Binomial name
249:
245:
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209:
206:
204:Superfamily:
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97:
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87:
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77:
72:
67:
62:
56:
50:
44:
40:
37:
33:
19:
1235:
1229:
1219:
1208:the original
1203:
1190:
1166:(2): 41–48.
1163:
1157:
1151:
1097:
1091:
1052:
1045:
1041:(tardigrade)
1038:
971:
970:
966:
950:
946:
930:
929:
906:
905:
889:
888:
872:
871:
848:
847:
836:Hypsibioidea
776:
772:
756:
755:
732:
731:
708:
707:
684:
683:
660:
659:
625:
610:
580:
579:
553:Hypsibioidea
548:
544:
532:
530:
524:
520:
492:
490:
481:
479:
474:
456:
446:
440:
436:
432:
431:
424:
416:
406:
398:
395:generic name
390:
386:
381:
375:
371:
367:
361:
338:
336:
316:
308:
299:
298:
291:Hypsibioidea
278:
277:
276:
263:
258:
242:
241:
221:
220:
208:Hypsibioidea
188:Eutardigrada
42:
36:
1152:Beorn leggi
465:Description
300:A. dactylus
287:tardigrades
1284:Categories
1100:(1): 953.
1066:References
523:(top) and
486:lobopodous
358:Cedar Lake
344:specimen,
329:, Canada.
323:Cedar Lake
178:Tardigrada
1252:0962-8452
1182:0033-2615
1114:2399-3642
582:Milnesium
557:cladogram
448:Milnesium
384:described
315:amber as
236:Species:
198:Parachela
164:Kingdom:
158:Eukaryota
55:Campanian
1270:34610770
1238:(1960).
1132:39107512
1123:11303527
1032:See also
972:Aerobius
545:Aerobius
533:Aerobius
527:(bottom)
525:Aerobius
501:eyespots
493:Aerobius
482:Aerobius
475:Aerobius
453:Turonian
433:Aerobius
426:dactylus
417:dactylus
405:prefix "
399:Aerobius
368:Aerobius
342:holotype
339:Aerobius
327:Manitoba
309:Aerobius
279:Aerobius
222:Aerobius
174:Phylum:
168:Animalia
154:Domain:
43:Aerobius
1261:8493197
497:cuticle
356:enters
284:extinct
261:†
240:†
214:Genus:
194:Order:
184:Class:
32:Aerobus
1268:
1258:
1250:
1180:
1130:
1120:
1112:
423:word "
393:. The
376:Beorne
307:. The
1211:(PDF)
1200:(PDF)
1039:Beorn
952:Beorn
549:Beorn
521:Beorn
499:. No
437:Beorn
421:Latin
408:aero-
403:Greek
391:Beorn
372:Beorn
363:Beorn
318:Beorn
305:amber
1266:PMID
1248:ISSN
1178:ISSN
1128:PMID
1110:ISSN
541:rRNA
337:The
229:2024
61:Preęž’
1256:PMC
1240:doi
1236:288
1168:doi
1118:PMC
1102:doi
563:):
346:MCZ
325:in
1286::
1264:.
1254:.
1246:.
1234:.
1228:.
1176:.
1164:71
1162:.
1156:.
1140:^
1126:.
1116:.
1108:.
1096:.
1090:.
1074:^
415:,
397:,
297:,
111:Pg
51:,
1272:.
1242::
1184:.
1170::
1134:.
1104::
1098:7
968:â€
948:â€
774:â€
561:â€
218:â€
116:N
106:K
101:J
96:T
91:P
86:C
81:D
76:S
71:O
66:ęž’
34:.
20:)
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