1453:
1459:
2009:
143:
450:
2039:
2015:
418:
119:
491:
differs markedly from the rostral morphology of other ceratopsians, possessing a sub-circular cross section, pitted texture instead of grooves, and a lack of lateral slicing edges. Furthermore, the proposed rostral bone appears to have fully fused with the premaxilla, a feature that is only seen in
436:
estimated a body length of 1 m (3.3 ft). Other material includes four predentary bones, cataloged as MTM V2009.193.1, V2009.194.1, V2009.195.1, and V2009.196.1; these are also believed to have belonged to
379:, cataloged as MTM V2009.192.1, consists only of a few skull fragments, including snout with proposed rostral bone, fused premaxillae, and maxillae fragments (beak and jaw fragments). These fossils are kept in the
500:
possesses a rostral bone in the first place, a defining feature of the ceratopsian group, and instead the hooked beak represents a highly derived, hypertrophied premaxilla. While there is the possibility
521:, which is interpreted as a floodplain and channel deposit formed by variegated clay, silt with interbedded grey and brown sand, and sandstone beds. This strata dates to the
2129:
673:
CzepiĹski, Ĺukasz; Madzia, Daniel (2024). "Osteology, phylogenetic affinities, and palaeobiogeographic significance of the bizarre ornithischian dinosaur
2144:
2139:
1452:
774:
622:Ĺsi, Attila; Butler, R.J.; Weishampel, David B. (2010-05-27). "A Late Cretaceous ceratopsian dinosaur from Europe with Asian affinities".
1458:
2032:
578:
380:
2149:
2008:
142:
2134:
505:
is a highly unique ceratopsian, it is currently referred to as an enigmatic ornithischian of uncertain affinities.
865:
767:
2124:
983:
701:
2059:
518:
2119:
2091:
2042:
892:
283:
2014:
631:
588:
260:
441:, although they are proportionately smaller, and probably came from other individuals of the genus.
1148:
760:
1868:
1818:
1250:
732:
655:
137:
2096:
702:"A Late Cretaceous ceratopsian dinosaur from Europe with Asian affinities (supplementary data)"
1951:
1889:
1852:
1326:
1213:
724:
647:
583:
458:
1981:
1779:
1604:
1306:
1001:
716:
682:
639:
421:
1875:
1827:
1786:
1722:
1625:
1576:
1278:
1202:
1157:
1031:
1017:
976:
850:
834:
547:
457:
When first described, the fossils were stated to most closely resemble those of the Asian
332:
593:
487:
However, more recent research has questioned this referral. The proposed rostral bone of
635:
1882:
1841:
1772:
1759:
1730:
1715:
1685:
1646:
1632:
1549:
1510:
1389:
1299:
1264:
1257:
1137:
1087:
1010:
990:
360:
219:
449:
2113:
1960:
1939:
1909:
1900:
1800:
1694:
1639:
1618:
1590:
1583:
1569:
1562:
1368:
1357:
1350:
1292:
1285:
1271:
1229:
1164:
1130:
1101:
1054:
736:
659:
562:
302:
69:
1974:
1925:
1918:
1834:
1807:
1793:
1701:
1674:
1667:
1660:
1611:
1597:
1489:
1476:
1407:
1396:
1343:
1317:
1222:
1188:
1171:
1123:
1116:
877:
825:
555:
536:
481:
476:
462:
344:
336:
325:
203:
399:
where the fossils were first discovered, combined with the given greek nomination
686:
1967:
1861:
1744:
1708:
1531:
1243:
1108:
493:
425:
352:
130:
44:
1737:
1653:
1503:
1236:
1024:
958:
945:
904:
783:
539:
471:
417:
235:
89:
54:
2082:
558:
531:
522:
154:
94:
38:
30:
728:
651:
118:
2076:
1538:
1524:
1517:
1080:
816:
807:
551:
543:
401:
384:
376:
328:
187:
174:
126:
84:
79:
64:
59:
49:
720:
643:
1932:
1094:
1038:
566:
396:
99:
74:
801:
453:
Scale bar 10 cm. Reconstruction based on initial ceratopsian identity
340:
164:
2053:
363:. Later research however has questioned this assignment, and treats
448:
416:
321:
392:
356:
2057:
1474:
943:
794:
756:
700:Ĺsi, Attila; Butler, R.J.; Weishampel, David B. (2010-05-27).
752:
474:
related to the protoceratopsids, but more primitive than the
432:
Although the fossils are fragmentary, the paper describing
529:
shared its environment with other dinosaurs such as
2066:
1950:
1908:
1899:
1851:
1817:
1758:
1684:
1548:
1487:
1406:
1367:
1325:
1316:
1212:
1186:
1147:
1052:
1000:
956:
903:
876:
849:
677:from the Late Cretaceous European archipelago".
359:, from where its ancestors may have migrated by
492:the ontogenetically oldest members of derived
405:, meaning "horned face". The specific name, "
768:
367:as an ornithischian of unresolved affinity.
8:
2054:
1905:
1764:
1554:
1495:
1484:
1471:
1373:
1322:
1194:
1064:
1060:
968:
964:
953:
940:
882:
855:
846:
791:
775:
761:
753:
525:stage, around 86 to 84 million years ago.
117:
20:
617:
615:
613:
611:
609:
605:
331:described in 2010. It lived during the
355:most closely related to forms in east
7:
496:. This has cast doubt as to whether
320:(pronounced "oi-ka-sera-tops") is a
2130:Late Cretaceous dinosaurs of Europe
14:
2145:Taxa named by David B. Weishampel
2038:
2037:
2013:
2007:
1457:
1451:
579:Timeline of ceratopsian research
381:Hungarian Natural History Museum
351:, was originally described as a
141:
466:. Those similarities indicate
1:
2140:Fossil taxa described in 2010
129:proposed rostral fused with
2166:
687:10.1093/zoolinnean/zlae048
16:Extinct genus of dinosaurs
2027:
2005:
1767:
1557:
1498:
1483:
1470:
1448:
1376:
1197:
1067:
1063:
971:
967:
952:
939:
885:
858:
845:
790:
289:
282:
138:Scientific classification
136:
125:
116:
23:
984:Micropachycephalosaurus
517:were discovered in the
409:, honors KĂĄroly Kozma.
454:
429:
2092:Paleobiology Database
452:
420:
2150:Ornithischian genera
589:2010 in paleontology
387:. The generic name,
2068:Ajkaceratops kozmai
1149:Archaeoceratopsidae
721:10.1038/nature09019
675:Ajkaceratops kozmai
644:10.1038/nature09019
636:2010Natur.465..466O
519:CsehbĂĄnya Formation
337:Tethyan archipelago
296:Ajkaceratops kozmai
2135:Fossils of Hungary
1869:Crittendenceratops
1819:Albertaceratopsini
679:Zoological Journal
550:birds, as well as
542:, other non-avian
455:
430:
2107:
2106:
2060:Taxon identifiers
2051:
2050:
2023:
2022:
2003:
2002:
1999:
1998:
1995:
1994:
1991:
1990:
1952:Pachyrhinosaurini
1890:Yehuecauhceratops
1853:Nasutoceratopsini
1754:
1753:
1466:
1465:
1446:
1445:
1442:
1441:
1438:
1437:
1434:
1433:
1430:
1429:
1426:
1425:
1327:Protoceratopsidae
1214:Leptoceratopsidae
1182:
1181:
1048:
1047:
935:
934:
931:
930:
927:
926:
923:
922:
715:(7297): 466â468.
630:(7297): 466â468.
584:List of dinosaurs
509:Palaeoenvironment
424:based on initial
339:, in what is now
313:
312:
263:
2157:
2100:
2099:
2087:
2086:
2085:
2055:
2041:
2040:
2018:
2017:
2011:
1982:Pachyrhinosaurus
1906:
1780:Machairoceratops
1765:
1605:Coahuilaceratops
1555:
1496:
1485:
1472:
1461:
1456:
1455:
1374:
1323:
1307:Zhuchengceratops
1195:
1065:
1061:
1002:Chaoyangsauridae
969:
965:
954:
941:
883:
856:
847:
840:
839:
792:
777:
770:
763:
754:
747:
746:
744:
743:
706:
697:
691:
690:
670:
664:
663:
619:
422:Life restoration
298:
294:
254:
247:
234:
218:
202:
146:
145:
121:
111:
41:
29:Temporal range:
21:
2165:
2164:
2160:
2159:
2158:
2156:
2155:
2154:
2110:
2109:
2108:
2103:
2095:
2090:
2081:
2080:
2075:
2062:
2052:
2047:
2019:
2012:
1987:
1946:
1895:
1876:Furcatoceratops
1847:
1828:Albertaceratops
1813:
1787:Menefeeceratops
1750:
1723:Tatankaceratops
1680:
1626:Mercuriceratops
1577:Arrhinoceratops
1544:
1492:
1479:
1462:
1450:
1422:
1402:
1363:
1312:
1279:Montanoceratops
1208:
1203:Graciliceratops
1191:
1178:
1158:Archaeoceratops
1143:
1057:
1044:
1032:Xuanhuaceratops
1018:Hualianceratops
996:
977:Albalophosaurus
961:
948:
919:
899:
872:
866:Avemetatarsalia
851:Avemetatarsalia
841:
835:Marginocephalia
797:
796:
786:
781:
751:
750:
741:
739:
704:
699:
698:
694:
672:
671:
667:
621:
620:
607:
602:
575:
548:enantiornithine
513:The fossils of
511:
459:protoceratopsid
447:
415:
373:
335:in the western
333:Late Cretaceous
309:
300:
292:
291:
278:
253:
245:
232:
216:
200:
140:
112:
110:
109:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
36:
35:
33:
27:
17:
12:
11:
5:
2163:
2161:
2153:
2152:
2147:
2142:
2137:
2132:
2127:
2125:Santonian life
2122:
2112:
2111:
2105:
2104:
2102:
2101:
2088:
2072:
2070:
2064:
2063:
2058:
2049:
2048:
2046:
2045:
2035:
2028:
2025:
2024:
2021:
2020:
2006:
2004:
2001:
2000:
1997:
1996:
1993:
1992:
1989:
1988:
1986:
1985:
1978:
1971:
1964:
1956:
1954:
1948:
1947:
1945:
1944:
1936:
1929:
1922:
1914:
1912:
1903:
1897:
1896:
1894:
1893:
1886:
1883:Nasutoceratops
1879:
1872:
1865:
1857:
1855:
1849:
1848:
1846:
1845:
1842:Medusaceratops
1838:
1831:
1823:
1821:
1815:
1814:
1812:
1811:
1804:
1797:
1790:
1783:
1776:
1773:Diabloceratops
1768:
1762:
1760:Centrosaurinae
1756:
1755:
1752:
1751:
1749:
1748:
1741:
1734:
1731:Titanoceratops
1727:
1719:
1716:Regaliceratops
1712:
1705:
1698:
1690:
1688:
1686:Triceratopsini
1682:
1681:
1679:
1678:
1671:
1664:
1657:
1650:
1647:Sierraceratops
1643:
1636:
1633:Navajoceratops
1629:
1622:
1615:
1608:
1601:
1594:
1587:
1580:
1573:
1566:
1558:
1552:
1550:Chasmosaurinae
1546:
1545:
1543:
1542:
1535:
1528:
1521:
1514:
1511:Brachyceratops
1507:
1499:
1493:
1488:
1481:
1480:
1475:
1468:
1467:
1464:
1463:
1449:
1447:
1444:
1443:
1440:
1439:
1436:
1435:
1432:
1431:
1428:
1427:
1424:
1423:
1421:
1420:
1419:
1418:
1412:
1410:
1404:
1403:
1401:
1400:
1393:
1390:Turanoceratops
1386:
1377:
1371:
1365:
1364:
1362:
1361:
1354:
1347:
1340:
1331:
1329:
1320:
1314:
1313:
1311:
1310:
1303:
1300:Unescoceratops
1296:
1289:
1282:
1275:
1268:
1265:Ischioceratops
1261:
1258:Gryphoceratops
1254:
1247:
1240:
1233:
1226:
1218:
1216:
1210:
1209:
1207:
1206:
1198:
1192:
1187:
1184:
1183:
1180:
1179:
1177:
1176:
1168:
1161:
1153:
1151:
1145:
1144:
1142:
1141:
1138:Sasayamagnomus
1134:
1127:
1120:
1113:
1105:
1098:
1091:
1088:Auroraceratops
1084:
1077:
1068:
1058:
1053:
1050:
1049:
1046:
1045:
1043:
1042:
1035:
1028:
1021:
1014:
1011:Chaoyangsaurus
1006:
1004:
998:
997:
995:
994:
991:Psittacosaurus
987:
980:
972:
962:
957:
950:
949:
944:
937:
936:
933:
932:
929:
928:
925:
924:
921:
920:
918:
917:
916:
915:
909:
907:
901:
900:
898:
897:
896:
895:
886:
880:
874:
873:
871:
870:
869:
868:
859:
853:
843:
842:
838:
837:
828:
819:
810:
804:
795:
788:
787:
782:
780:
779:
772:
765:
757:
749:
748:
692:
665:
604:
603:
601:
598:
597:
596:
591:
586:
581:
574:
571:
510:
507:
446:
445:Classification
443:
414:
411:
372:
369:
361:island-hopping
311:
310:
301:
287:
286:
280:
279:
275:A. kozmai
271:
269:
265:
264:
243:
239:
238:
230:
223:
222:
220:Neornithischia
214:
207:
206:
198:
191:
190:
185:
178:
177:
172:
168:
167:
162:
158:
157:
152:
148:
147:
134:
133:
123:
122:
114:
113:
105:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2162:
2151:
2148:
2146:
2143:
2141:
2138:
2136:
2133:
2131:
2128:
2126:
2123:
2121:
2118:
2117:
2115:
2098:
2093:
2089:
2084:
2078:
2074:
2073:
2071:
2069:
2065:
2061:
2056:
2044:
2036:
2034:
2030:
2029:
2026:
2016:
2010:
1984:
1983:
1979:
1977:
1976:
1972:
1970:
1969:
1965:
1963:
1962:
1961:Achelousaurus
1958:
1957:
1955:
1953:
1949:
1942:
1941:
1940:Styracosaurus
1937:
1935:
1934:
1930:
1928:
1927:
1923:
1921:
1920:
1916:
1915:
1913:
1911:
1910:Centrosaurini
1907:
1904:
1902:
1901:Eucentrosaura
1898:
1892:
1891:
1887:
1885:
1884:
1880:
1878:
1877:
1873:
1871:
1870:
1866:
1864:
1863:
1859:
1858:
1856:
1854:
1850:
1844:
1843:
1839:
1837:
1836:
1832:
1830:
1829:
1825:
1824:
1822:
1820:
1816:
1810:
1809:
1805:
1803:
1802:
1801:Wendiceratops
1798:
1796:
1795:
1791:
1789:
1788:
1784:
1782:
1781:
1777:
1775:
1774:
1770:
1769:
1766:
1763:
1761:
1757:
1747:
1746:
1742:
1740:
1739:
1735:
1733:
1732:
1728:
1725:
1724:
1720:
1718:
1717:
1713:
1711:
1710:
1706:
1704:
1703:
1699:
1697:
1696:
1695:Eotriceratops
1692:
1691:
1689:
1687:
1683:
1677:
1676:
1672:
1670:
1669:
1665:
1663:
1662:
1658:
1656:
1655:
1651:
1649:
1648:
1644:
1642:
1641:
1640:Pentaceratops
1637:
1635:
1634:
1630:
1628:
1627:
1623:
1621:
1620:
1619:Kosmoceratops
1616:
1614:
1613:
1609:
1607:
1606:
1602:
1600:
1599:
1595:
1593:
1592:
1591:Bravoceratops
1588:
1586:
1585:
1584:Bisticeratops
1581:
1579:
1578:
1574:
1572:
1571:
1570:Anchiceratops
1567:
1565:
1564:
1563:Agujaceratops
1560:
1559:
1556:
1553:
1551:
1547:
1541:
1540:
1536:
1534:
1533:
1529:
1527:
1526:
1522:
1520:
1519:
1515:
1513:
1512:
1508:
1506:
1505:
1501:
1500:
1497:
1494:
1491:
1486:
1482:
1478:
1473:
1469:
1460:
1454:
1416:
1415:
1414:
1413:
1411:
1409:
1405:
1399:
1398:
1394:
1392:
1391:
1387:
1384:
1383:
1379:
1378:
1375:
1372:
1370:
1369:Ceratopsoidea
1366:
1360:
1359:
1358:Protoceratops
1355:
1353:
1352:
1351:Breviceratops
1348:
1346:
1345:
1341:
1338:
1337:
1333:
1332:
1330:
1328:
1324:
1321:
1319:
1315:
1309:
1308:
1304:
1302:
1301:
1297:
1295:
1294:
1293:Udanoceratops
1290:
1288:
1287:
1286:Prenoceratops
1283:
1281:
1280:
1276:
1274:
1273:
1272:Leptoceratops
1269:
1267:
1266:
1262:
1260:
1259:
1255:
1253:
1252:
1248:
1246:
1245:
1241:
1239:
1238:
1234:
1232:
1231:
1230:Bainoceratops
1227:
1225:
1224:
1220:
1219:
1217:
1215:
1211:
1205:
1204:
1200:
1199:
1196:
1193:
1190:
1185:
1174:
1173:
1169:
1167:
1166:
1165:Helioceratops
1162:
1160:
1159:
1155:
1154:
1152:
1150:
1146:
1140:
1139:
1135:
1133:
1132:
1131:Mosaiceratops
1128:
1126:
1125:
1121:
1119:
1118:
1114:
1111:
1110:
1106:
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1102:Koreaceratops
1099:
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284:Binomial name
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40:
32:
26:
22:
19:
2120:Ceratopsians
2067:
1980:
1975:Stellasaurus
1973:
1966:
1959:
1938:
1931:
1926:Coronosaurus
1924:
1919:Centrosaurus
1917:
1888:
1881:
1874:
1867:
1860:
1840:
1835:Lokiceratops
1833:
1826:
1808:Xenoceratops
1806:
1799:
1794:Sinoceratops
1792:
1785:
1778:
1771:
1743:
1736:
1729:
1721:
1714:
1707:
1702:Nedoceratops
1700:
1693:
1675:Vagaceratops
1673:
1668:Utahceratops
1666:
1661:Terminocavus
1659:
1652:
1645:
1638:
1631:
1624:
1617:
1612:Judiceratops
1610:
1603:
1598:Chasmosaurus
1596:
1589:
1582:
1575:
1568:
1561:
1537:
1530:
1523:
1516:
1509:
1502:
1490:Ceratopsidae
1477:Ceratopsidae
1408:Ceratopsidae
1397:Zuniceratops
1395:
1388:
1382:Ajkaceratops
1381:
1380:
1356:
1349:
1344:Bagaceratops
1342:
1336:Ajkaceratops
1335:
1334:
1318:Coronosauria
1305:
1298:
1291:
1284:
1277:
1270:
1263:
1256:
1249:
1242:
1235:
1228:
1223:Asiaceratops
1221:
1201:
1189:Euceratopsia
1172:Yamaceratops
1170:
1163:
1156:
1136:
1129:
1124:Microceratus
1122:
1117:Liaoceratops
1115:
1107:
1100:
1093:
1086:
1079:
1073:Ajkaceratops
1072:
1071:
1037:
1030:
1023:
1016:
1009:
989:
982:
975:
893:Ornithischia
878:Ornithischia
830:
826:Ornithischia
821:
812:
740:. Retrieved
712:
708:
695:
678:
674:
668:
627:
623:
594:HaĹŁeg Island
565:turtles and
554:crocodiles,
530:
527:Ajkaceratops
526:
515:Ajkaceratops
514:
512:
503:Ajkaceratops
502:
498:Ajkaceratops
497:
489:Ajkaceratops
488:
486:
482:Ceratopsidae
477:Zuniceratops
475:
468:Ajkaceratops
467:
463:Bagaceratops
461:
456:
439:Ajkaceratops
438:
434:Ajkaceratops
433:
431:
406:
400:
395:, a town in
389:Ajkaceratops
388:
374:
365:Ajkaceratops
364:
348:
345:type species
317:Ajkaceratops
316:
315:
314:
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274:
273:
256:
250:Ajkaceratops
249:
248:
226:
210:
204:Ornithischia
194:
181:
25:Ajkaceratops
24:
18:
1968:Einiosaurus
1862:Avaceratops
1745:Triceratops
1709:Ojoceratops
1532:Monoclonius
1244:Ferrisaurus
1109:Kulceratops
563:bothremydid
540:ankylosaurs
494:ceratopsids
472:ceratopsian
426:ceratopsian
413:Description
353:ceratopsian
131:premaxillae
2114:Categories
2031:See also:
1738:Torosaurus
1654:Spiclypeus
1504:Agathaumas
1417:see belowâ
1237:Cerasinops
1025:Stenopelix
959:Ceratopsia
946:Ceratopsia
914:see belowâ
905:Ceratopsia
817:Dinosauria
784:Ceratopsia
742:2010-05-29
600:References
559:pterosaurs
556:azhdarchid
537:nodosaurid
236:Ceratopsia
188:Dinosauria
800:Kingdom:
737:205220451
660:205220451
569:lizards.
552:eusuchian
544:theropods
532:Mochlodon
523:Santonian
391:, honors
371:Discovery
349:A. kozmai
268:Species:
161:Kingdom:
155:Eukaryota
31:Santonian
2077:Wikidata
2043:Category
2033:Timeline
1539:Polyonax
1525:Dysganus
1518:Ceratops
1081:Aquilops
808:Chordata
806:Phylum:
802:Animalia
729:20505726
652:20505726
573:See also
480:and the
428:identity
402:ceratops
385:Budapest
377:holotype
329:dinosaur
175:Chordata
171:Phylum:
165:Animalia
151:Domain:
127:Holotype
37:85
2083:Q516014
1933:Spinops
1251:Gremlin
1039:Yinlong
632:Bibcode
407:kozmai"
397:Hungary
293:†
272:†
242:Genus:
106:↓
2097:167127
735:
727:
709:Nature
658:
650:
624:Nature
535:, the
343:. The
341:Europe
308:, 2010
306:et al.
257:et al.
831:Clade
822:Clade
813:Clade
733:S2CID
705:(PDF)
656:S2CID
567:teiid
470:is a
383:, in
322:genus
227:Clade
211:Clade
195:Clade
182:Clade
891:see
864:see
725:PMID
648:PMID
546:and
393:Ajka
375:The
357:Asia
261:2010
255:Ĺsi
45:Preę
1095:Beg
717:doi
713:465
683:doi
640:doi
628:465
324:of
303:Ĺsi
2116::
2094::
2079::
833::
824::
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731:.
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681:.
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638:.
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608:^
561:,
484:.
347:,
259:,
229::
213::
197::
184::
95:Pg
39:Ma
1943:?
1726:?
1385:?
1339:?
1175:?
1112:?
1076:?
776:e
769:t
762:v
745:.
719::
689:.
685::
662:.
642::
634::
246:â
233:â
217:â
201:â
100:N
90:K
85:J
80:T
75:P
70:C
65:D
60:S
55:O
50:ę
34:~
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