2700:
2695:
1855:
525:
136:
1860:
115:
606:
Rodolfo Salas-Gismondi; Jorge W. Moreno-Bernal; Torsten M. Scheyer; Marcelo R. Sánchez-Villagra; Carlos
Jaramillo (2018). "New Miocene Caribbean gavialoids and patterns of longirostry in crocodylians".
671:
396:. Based on the fragmentary remains recovered, the species was differentiated from other members of its family by unique positioning and geometry of its skull elements.
2796:
388:
body terminated in a long, laterally flattened tail at one end, and a specialized, narrow snout at the other. As with the other members of its family, the snout of
768:
679:
3012:
3007:
2956:
2876:
2783:
482:. This has led to conclusions that gavialoids were partially or primarily saltwater reptiles prior to the evolution of the group's one or two
3002:
2694:
761:
2699:
2027:
1854:
310:
2073:
1942:
432:
642:
135:
754:
1934:
1101:
2155:
2127:
1773:
1094:
1859:
2148:
706:"A gharial from the Oligocene of Puerto Rico: transoceanic dispersal in the history of a non-marine reptile"
559:"A gharial from the Oligocene of Puerto Rico: transoceanic dispersal in the history of a non-marine reptile"
2634:
2408:
2738:
2658:
2432:
2141:
2969:
2902:
2822:
2618:
2611:
2424:
2416:
2300:
937:
857:
2642:
2943:
2863:
2770:
2134:
1823:
1816:
1766:
1564:
261:
466:. This is in contrast with the living gavialoids, which reside entirely in the freshwater rivers of
2997:
1802:
1529:
530:
2666:
2575:
2567:
2293:
1571:
1550:
1543:
1204:
1080:
884:
624:
130:
2974:
2907:
2827:
2889:
2809:
2801:
2582:
2081:
2048:
1809:
1536:
1341:
735:
588:
2894:
2814:
462:
form part of the deposits where the specimen was found, indicating a marine distribution for
2560:
1795:
1658:
1621:
1557:
1452:
1318:
725:
717:
616:
578:
570:
436:
408:
203:
190:
2650:
2626:
2589:
2509:
2257:
1984:
1743:
1728:
1693:
1636:
1506:
1362:
1307:
1124:
1108:
393:
2523:
2501:
2243:
2235:
2193:
1991:
1926:
1912:
1785:
1498:
1482:
1411:
1369:
1327:
1299:
1277:
1234:
1169:
1156:
1131:
1116:
1087:
1018:
974:
959:
730:
705:
583:
558:
494:
2991:
2721:
2553:
2546:
2531:
2516:
2479:
2377:
2362:
2354:
2271:
2250:
2221:
2214:
2207:
2097:
2011:
1999:
1949:
1919:
1904:
1890:
1672:
1643:
1613:
1475:
1444:
1431:
1419:
1403:
1385:
1348:
1248:
1072:
1050:
236:
72:
628:
2716:
2539:
2472:
2385:
2370:
2337:
2326:
2278:
2060:
1970:
1957:
1877:
1756:
1707:
1700:
1628:
1490:
1467:
1292:
1284:
1255:
1197:
1190:
1065:
950:
842:
817:
483:
459:
397:
289:
620:
2934:
2761:
2494:
2487:
2285:
2264:
2228:
2200:
2179:
1721:
1714:
1686:
1679:
1665:
1606:
1459:
1395:
1355:
1334:
1262:
1241:
1183:
1057:
1025:
982:
471:
440:
424:
402:
362:
354:
47:
2854:
2448:
2312:
2117:
2041:
2034:
2020:
1599:
1216:
996:
896:
808:
778:
520:
467:
381:
373:
334:
223:
213:
92:
57:
650:
505:(gavial). This roughly translates to "shore gavial", referring to its rather
2398:
1736:
1519:
1003:
869:
513:
means "from Puerto Rico", in reference to the specimen's country of origin.
506:
474:
sediments possibly have been washed onto the delta from inland sources, the
452:
428:
385:
366:
346:
338:
147:
114:
97:
41:
30:
739:
721:
592:
574:
557:
Vélez-Juarbe, Jorge; Brochu, Christopher A.; Santos, Hernán (2007-03-06).
447:
remains recovered were extremely fragmentary, consisting of an incomplete
2928:
2848:
2755:
2601:
2089:
1650:
1270:
1010:
826:
802:
420:
167:
87:
82:
67:
62:
52:
2961:
2881:
2788:
989:
746:
704:
Vélez-Juarbe, Jorge; Christopher A Brochu; Hernán Santos (2007-03-06).
392:
was extremely long and narrow, tapering into a thin structure past the
342:
327:
177:
102:
77:
34:
1586:
796:
479:
157:
2732:
478:
specimen found so far was from a location that had essentially been
1032:
448:
400:
analysis indicates its closest relatives were the extinct gavials
330:
2948:
2868:
2775:
2105:
967:
427:
deposits dating around 28 million years old, from the Middle to
122:
An image (A) and a drawing (B) of the underside of the skull of
2736:
1875:
1154:
935:
789:
750:
678:. World Wide Fund for Nature. 2008-01-25. Archived from
649:. World Wide Fund for Nature. 2008-07-22. Archived from
285:
439:
in Puerto Rico, have been an adequate supply of other
296:
276:
2918:
2838:
2745:
2599:
2447:
2396:
2336:
2310:
2177:
2115:
2059:
2010:
1968:
1888:
1783:
1754:
1585:
1517:
1430:
1383:
1317:
1214:
1167:
1042:
949:
895:
868:
841:
762:
8:
552:
550:
548:
546:
544:
542:
423:, designated UPRMP 3094, was discovered in
2733:
2457:
2346:
2318:
2185:
2065:
1976:
1896:
1885:
1872:
1591:
1436:
1427:
1226:
1222:
1175:
1164:
1151:
946:
932:
874:
847:
838:
786:
769:
755:
747:
113:
20:
729:
582:
538:
672:"Gharial - Habitat & Distribution"
455:and other scattered cranial elements.
286:Vélez-Juarbe, Brochu & Santos 2007
470:. While fossils found in prehistoric
7:
716:(1615). The Royal Society: 1245–54.
569:(1615). The Royal Society: 1245–54.
609:Journal of Systematic Palaeontology
349:. Two species have been described:
14:
509:distribution. The specific name
3013:Extinct animals of the Caribbean
3008:Prehistoric pseudosuchian genera
2698:
2693:
1858:
1853:
710:Proceedings of the Royal Society
563:Proceedings of the Royal Society
523:
134:
415:The type species of the genus,
384:body plan. An elongated, squat
431:. The deposits, part of the
1:
621:10.1080/14772019.2018.1495275
419:, was described in 2007. The
353:from the Middle Oligocene of
2840:Aktiogavialis puertoricensis
351:Aktiogavialis puertoricensis
124:Aktiogavialis puertoricensis
16:Species of reptile (fossil)
3029:
451:featuring elements of the
3003:Oligocene crocodylomorphs
2711:
2691:
2460:
2349:
2321:
2188:
2068:
1979:
1899:
1884:
1871:
1851:
1594:
1439:
1229:
1225:
1178:
1163:
1150:
945:
931:
877:
850:
837:
785:
273:
268:
131:Scientific classification
129:
121:
112:
23:
2028:"Crocodylus" gariepensis
2301:"Tomistoma" lusitanicum
2074:"Crocodylus" megarhinus
1943:"Crocodylus" megarhinus
365:of the Late Miocene of
2920:Aktiogavialis caribesi
722:10.1098/rspb.2006.0455
575:10.1098/rspb.2006.0455
493:, is derived from the
380:followed the standard
359:Aktiogavialis caribesi
2970:Paleobiology Database
2903:Paleobiology Database
2823:Paleobiology Database
458:Marine sediments and
2576:"Tomistoma" coppensi
2568:"Tomistoma" cairense
2294:"Tomistoma" cairense
1935:"Crocodylus" affinis
1102:"Crocodylus" affinis
435:Formation along the
337:that lived from the
2583:"Tomistoma" dowsoni
531:Paleontology portal
345:in what is now the
2156:C. thorbjarnarsoni
1205:Listrognathosuchus
1081:Brachyuranochampsa
682:on October 5, 2008
2985:
2984:
2890:Open Tree of Life
2810:Open Tree of Life
2739:Taxon identifiers
2730:
2729:
2707:
2706:
2689:
2688:
2685:
2684:
2681:
2680:
2677:
2676:
2443:
2442:
2173:
2172:
2169:
2168:
2165:
2164:
2128:C. anthropophagus
1867:
1866:
1849:
1848:
1845:
1844:
1841:
1840:
1837:
1836:
1833:
1832:
1581:
1580:
1379:
1378:
1342:Eurycephalosuchus
1146:
1145:
1142:
1141:
1095:"Crocodylus" acer
927:
926:
923:
922:
919:
918:
915:
914:
615:(12): 1049–1075.
489:The genus' name,
464:A. puertoricensis
417:A. puertoricensis
319:
318:
313:
293:
281:A. puertoricensis
264:
3020:
2978:
2977:
2965:
2964:
2952:
2951:
2939:
2938:
2937:
2911:
2910:
2898:
2897:
2885:
2884:
2872:
2871:
2859:
2858:
2857:
2831:
2830:
2818:
2817:
2805:
2804:
2792:
2791:
2779:
2778:
2766:
2765:
2764:
2734:
2702:
2697:
2659:G. pachyrhynchus
2561:Siquisiquesuchus
2458:
2347:
2319:
2186:
2066:
1977:
1897:
1886:
1873:
1862:
1857:
1824:C. wannlangstoni
1817:C. venezuelensis
1659:Globidentosuchus
1622:Centenariosuchus
1592:
1453:Allognathosuchus
1437:
1428:
1319:Orientalosuchina
1227:
1223:
1176:
1165:
1152:
947:
933:
875:
848:
839:
832:
831:
787:
771:
764:
757:
748:
743:
733:
691:
690:
688:
687:
668:
662:
661:
659:
658:
639:
633:
632:
603:
597:
596:
586:
554:
533:
528:
527:
526:
409:Siquisiquesuchus
304:
298:
284:
278:
255:
248:
235:
204:Archosauriformes
191:Archosauromorpha
139:
138:
117:
107:
44:
29:Temporal range:
21:
3028:
3027:
3023:
3022:
3021:
3019:
3018:
3017:
2988:
2987:
2986:
2981:
2973:
2968:
2960:
2955:
2947:
2942:
2933:
2932:
2927:
2914:
2906:
2901:
2893:
2888:
2880:
2875:
2867:
2862:
2853:
2852:
2847:
2834:
2826:
2821:
2813:
2808:
2800:
2795:
2787:
2782:
2774:
2769:
2760:
2759:
2754:
2741:
2731:
2726:
2703:
2673:
2635:G. curvirostris
2612:G. bengawanicus
2595:
2590:Toyotamaphimeia
2510:Hesperogavialis
2451:
2439:
2392:
2340:
2332:
2315:
2306:
2258:Megadontosuchus
2182:
2161:
2149:C. palaeindicus
2111:
2055:
2006:
1985:Aldabrachampsus
1973:
1964:
1893:
1880:
1863:
1829:
1803:C. brevirostris
1779:
1750:
1744:Wannaganosuchus
1729:Stangerochampsa
1694:Orthogenysuchus
1637:Chinatichampsus
1577:
1513:
1507:Wannaganosuchus
1426:
1388:
1375:
1363:Orientalosuchus
1313:
1308:Stangerochampsa
1219:
1210:
1172:
1159:
1138:
1125:Prodiplocynodon
1109:Portugalosuchus
1038:
941:
911:
891:
864:
833:
792:
791:
781:
775:
703:
700:
695:
694:
685:
683:
670:
669:
665:
656:
654:
641:
640:
636:
605:
604:
600:
556:
555:
540:
529:
524:
522:
519:
305:Salas-Gismondi
254:
246:
233:
133:
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
39:
38:
27:
17:
12:
11:
5:
3026:
3024:
3016:
3015:
3010:
3005:
3000:
2990:
2989:
2983:
2982:
2980:
2979:
2966:
2953:
2940:
2924:
2922:
2916:
2915:
2913:
2912:
2899:
2886:
2873:
2860:
2844:
2842:
2836:
2835:
2833:
2832:
2819:
2806:
2793:
2780:
2767:
2751:
2749:
2743:
2742:
2737:
2728:
2727:
2725:
2724:
2719:
2712:
2709:
2708:
2705:
2704:
2692:
2690:
2687:
2686:
2683:
2682:
2679:
2678:
2675:
2674:
2672:
2671:
2663:
2655:
2647:
2639:
2631:
2623:
2615:
2607:
2605:
2597:
2596:
2594:
2593:
2586:
2579:
2572:
2564:
2557:
2550:
2543:
2536:
2528:
2524:Maomingosuchus
2520:
2513:
2506:
2502:Harpacochampsa
2498:
2491:
2484:
2476:
2469:
2461:
2455:
2445:
2444:
2441:
2440:
2438:
2437:
2429:
2425:T. lusitanicum
2421:
2413:
2409:T. calaritanum
2404:
2402:
2394:
2393:
2391:
2390:
2382:
2374:
2367:
2359:
2350:
2344:
2334:
2333:
2331:
2330:
2322:
2316:
2311:
2308:
2307:
2305:
2304:
2297:
2290:
2282:
2275:
2268:
2261:
2254:
2247:
2244:Maomingosuchus
2240:
2236:Leptorrhamphus
2232:
2225:
2218:
2211:
2204:
2197:
2194:Dollosuchoides
2189:
2183:
2178:
2175:
2174:
2171:
2170:
2167:
2166:
2163:
2162:
2160:
2159:
2152:
2145:
2142:C. falconensis
2138:
2131:
2123:
2121:
2113:
2112:
2110:
2109:
2102:
2094:
2086:
2078:
2069:
2063:
2057:
2056:
2054:
2053:
2045:
2038:
2031:
2024:
2016:
2014:
2008:
2007:
2005:
2004:
1996:
1992:Dzungarisuchus
1988:
1980:
1974:
1969:
1966:
1965:
1963:
1962:
1954:
1946:
1939:
1931:
1927:Australosuchus
1923:
1916:
1913:Antecrocodylus
1909:
1900:
1894:
1889:
1882:
1881:
1876:
1869:
1868:
1865:
1864:
1852:
1850:
1847:
1846:
1843:
1842:
1839:
1838:
1835:
1834:
1831:
1830:
1828:
1827:
1820:
1813:
1806:
1799:
1791:
1789:
1781:
1780:
1778:
1777:
1774:M. latrubessei
1770:
1762:
1760:
1752:
1751:
1749:
1748:
1740:
1733:
1725:
1718:
1711:
1704:
1697:
1690:
1683:
1676:
1669:
1662:
1655:
1647:
1640:
1633:
1625:
1618:
1610:
1603:
1595:
1589:
1583:
1582:
1579:
1578:
1576:
1575:
1568:
1561:
1554:
1547:
1540:
1533:
1525:
1523:
1515:
1514:
1512:
1511:
1503:
1499:Procaimanoidea
1495:
1487:
1483:Hassiacosuchus
1479:
1472:
1464:
1456:
1449:
1440:
1434:
1425:
1424:
1416:
1412:Menatalligator
1408:
1400:
1391:
1389:
1384:
1381:
1380:
1377:
1376:
1374:
1373:
1370:Protoalligator
1366:
1359:
1352:
1345:
1338:
1331:
1328:Dongnanosuchus
1323:
1321:
1315:
1314:
1312:
1311:
1304:
1300:Procaimanoidea
1296:
1289:
1281:
1278:Hassiacosuchus
1274:
1267:
1259:
1252:
1245:
1238:
1235:Albertochampsa
1230:
1220:
1215:
1212:
1211:
1209:
1208:
1201:
1194:
1187:
1179:
1173:
1170:Alligatoroidea
1168:
1161:
1160:
1157:Alligatoroidea
1155:
1148:
1147:
1144:
1143:
1140:
1139:
1137:
1136:
1132:Planocraniidae
1128:
1121:
1117:Pristichampsus
1113:
1105:
1098:
1091:
1088:Charactosuchus
1084:
1077:
1069:
1062:
1054:
1046:
1044:
1040:
1039:
1037:
1036:
1029:
1022:
1019:Trilophosuchus
1015:
1007:
1000:
993:
986:
979:
975:Harpacochampsa
971:
964:
960:Australosuchus
955:
953:
943:
942:
936:
929:
928:
925:
924:
921:
920:
917:
916:
913:
912:
910:
909:
908:
907:
901:
899:
893:
892:
890:
889:
888:
887:
878:
872:
866:
865:
863:
862:
861:
860:
851:
845:
835:
834:
830:
829:
820:
811:
805:
799:
790:
783:
782:
776:
774:
773:
766:
759:
751:
745:
744:
699:
696:
693:
692:
663:
634:
598:
537:
536:
535:
534:
518:
515:
511:puertoricensis
429:Late Oligocene
317:
316:
315:
314:
294:
271:
270:
266:
265:
244:
240:
239:
231:
227:
226:
221:
217:
216:
211:
207:
206:
201:
194:
193:
188:
181:
180:
175:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3025:
3014:
3011:
3009:
3006:
3004:
3001:
2999:
2996:
2995:
2993:
2976:
2971:
2967:
2963:
2958:
2954:
2950:
2945:
2941:
2936:
2930:
2926:
2925:
2923:
2921:
2917:
2909:
2904:
2900:
2896:
2891:
2887:
2883:
2878:
2874:
2870:
2865:
2861:
2856:
2850:
2846:
2845:
2843:
2841:
2837:
2829:
2824:
2820:
2816:
2811:
2807:
2803:
2798:
2794:
2790:
2785:
2781:
2777:
2772:
2768:
2763:
2757:
2753:
2752:
2750:
2748:
2747:Aktiogavialis
2744:
2740:
2735:
2723:
2722:Gryposuchinae
2720:
2718:
2714:
2713:
2710:
2701:
2696:
2669:
2668:
2664:
2661:
2660:
2656:
2653:
2652:
2648:
2645:
2644:
2640:
2637:
2636:
2632:
2629:
2628:
2624:
2621:
2620:
2616:
2614:
2613:
2609:
2608:
2606:
2604:
2603:
2598:
2592:
2591:
2587:
2585:
2584:
2580:
2578:
2577:
2573:
2570:
2569:
2565:
2563:
2562:
2558:
2556:
2555:
2554:Rhamphosuchus
2551:
2549:
2548:
2547:Piscogavialis
2544:
2542:
2541:
2537:
2534:
2533:
2532:Paratomistoma
2529:
2526:
2525:
2521:
2519:
2518:
2517:Ikanogavialis
2514:
2512:
2511:
2507:
2504:
2503:
2499:
2497:
2496:
2492:
2490:
2489:
2485:
2482:
2481:
2480:Gavialosuchus
2477:
2475:
2474:
2470:
2468:
2467:
2466:Aktiogavialis
2463:
2462:
2459:
2456:
2454:
2450:
2446:
2435:
2434:
2433:T. taiwanicum
2430:
2427:
2426:
2422:
2419:
2418:
2414:
2411:
2410:
2406:
2405:
2403:
2401:
2400:
2395:
2388:
2387:
2383:
2380:
2379:
2378:Paratomistoma
2375:
2373:
2372:
2368:
2365:
2364:
2363:Gavialosuchus
2360:
2357:
2356:
2355:Brasilosuchus
2352:
2351:
2348:
2345:
2343:
2342:sensu stricto
2339:
2335:
2329:
2328:
2324:
2323:
2320:
2317:
2314:
2309:
2303:
2302:
2298:
2296:
2295:
2291:
2288:
2287:
2283:
2281:
2280:
2276:
2274:
2273:
2272:Paratomistoma
2269:
2267:
2266:
2262:
2260:
2259:
2255:
2253:
2252:
2251:Maroccosuchus
2248:
2246:
2245:
2241:
2238:
2237:
2233:
2231:
2230:
2226:
2224:
2223:
2222:Gunggamarandu
2219:
2217:
2216:
2215:Gavialosuchus
2212:
2210:
2209:
2208:Ferganosuchus
2205:
2203:
2202:
2198:
2196:
2195:
2191:
2190:
2187:
2184:
2181:
2176:
2158:
2157:
2153:
2151:
2150:
2146:
2144:
2143:
2139:
2137:
2136:
2132:
2130:
2129:
2125:
2124:
2122:
2120:
2119:
2114:
2108:
2107:
2103:
2100:
2099:
2098:Tzaganosuchus
2095:
2092:
2091:
2087:
2084:
2083:
2079:
2076:
2075:
2071:
2070:
2067:
2064:
2062:
2058:
2051:
2050:
2046:
2044:
2043:
2039:
2037:
2036:
2032:
2030:
2029:
2025:
2023:
2022:
2018:
2017:
2015:
2013:
2012:Osteolaeminae
2009:
2002:
2001:
2000:Oxysdonsaurus
1997:
1994:
1993:
1989:
1987:
1986:
1982:
1981:
1978:
1975:
1972:
1967:
1960:
1959:
1955:
1952:
1951:
1950:Jiangxisuchus
1947:
1945:
1944:
1940:
1937:
1936:
1932:
1929:
1928:
1924:
1922:
1921:
1920:Astorgosuchus
1917:
1915:
1914:
1910:
1907:
1906:
1905:Albertosuchus
1902:
1901:
1898:
1895:
1892:
1891:Crocodyloidea
1887:
1883:
1879:
1874:
1870:
1861:
1856:
1826:
1825:
1821:
1819:
1818:
1814:
1812:
1811:
1810:C. gasparinae
1807:
1805:
1804:
1800:
1798:
1797:
1793:
1792:
1790:
1788:
1787:
1782:
1776:
1775:
1771:
1769:
1768:
1764:
1763:
1761:
1759:
1758:
1753:
1746:
1745:
1741:
1739:
1738:
1734:
1731:
1730:
1726:
1724:
1723:
1719:
1717:
1716:
1712:
1710:
1709:
1705:
1703:
1702:
1698:
1696:
1695:
1691:
1689:
1688:
1684:
1682:
1681:
1677:
1675:
1674:
1673:Kuttanacaiman
1670:
1668:
1667:
1663:
1661:
1660:
1656:
1653:
1652:
1648:
1646:
1645:
1644:Culebrasuchus
1641:
1639:
1638:
1634:
1631:
1630:
1626:
1624:
1623:
1619:
1616:
1615:
1614:Brachychampsa
1611:
1609:
1608:
1604:
1602:
1601:
1597:
1596:
1593:
1590:
1588:
1584:
1574:
1573:
1569:
1567:
1566:
1565:A. prenasalis
1562:
1560:
1559:
1555:
1553:
1552:
1548:
1546:
1545:
1541:
1539:
1538:
1534:
1532:
1531:
1527:
1526:
1524:
1522:
1521:
1516:
1509:
1508:
1504:
1501:
1500:
1496:
1493:
1492:
1488:
1485:
1484:
1480:
1478:
1477:
1476:Chrysochampsa
1473:
1470:
1469:
1465:
1462:
1461:
1457:
1455:
1454:
1450:
1447:
1446:
1445:Akanthosuchus
1442:
1441:
1438:
1435:
1433:
1432:Alligatorinae
1429:
1422:
1421:
1420:Sajkanosuchus
1417:
1414:
1413:
1409:
1406:
1405:
1404:Lianghusuchus
1401:
1398:
1397:
1393:
1392:
1390:
1387:
1386:Alligatoridae
1382:
1372:
1371:
1367:
1365:
1364:
1360:
1358:
1357:
1353:
1351:
1350:
1349:Jiangxisuchus
1346:
1344:
1343:
1339:
1337:
1336:
1332:
1330:
1329:
1325:
1324:
1322:
1320:
1316:
1310:
1309:
1305:
1302:
1301:
1297:
1295:
1294:
1290:
1287:
1286:
1282:
1280:
1279:
1275:
1273:
1272:
1268:
1265:
1264:
1260:
1258:
1257:
1253:
1251:
1250:
1249:Brachychampsa
1246:
1244:
1243:
1239:
1237:
1236:
1232:
1231:
1228:
1224:
1221:
1218:
1213:
1207:
1206:
1202:
1200:
1199:
1195:
1193:
1192:
1188:
1186:
1185:
1181:
1180:
1177:
1174:
1171:
1166:
1162:
1158:
1153:
1149:
1134:
1133:
1129:
1127:
1126:
1122:
1119:
1118:
1114:
1111:
1110:
1106:
1104:
1103:
1099:
1097:
1096:
1092:
1090:
1089:
1085:
1083:
1082:
1078:
1075:
1074:
1073:Borealosuchus
1070:
1068:
1067:
1063:
1060:
1059:
1055:
1053:
1052:
1051:Albertosuchus
1048:
1047:
1045:
1041:
1035:
1034:
1030:
1028:
1027:
1023:
1021:
1020:
1016:
1013:
1012:
1008:
1006:
1005:
1001:
999:
998:
994:
992:
991:
987:
985:
984:
980:
977:
976:
972:
970:
969:
965:
962:
961:
957:
956:
954:
952:
948:
944:
939:
934:
930:
905:
904:
903:
902:
900:
898:
894:
886:
882:
881:
880:
879:
876:
873:
871:
867:
859:
855:
854:
853:
852:
849:
846:
844:
840:
836:
828:
824:
821:
819:
815:
812:
810:
806:
804:
800:
798:
794:
793:
788:
784:
780:
772:
767:
765:
760:
758:
753:
752:
749:
741:
737:
732:
727:
723:
719:
715:
711:
707:
702:
701:
697:
681:
677:
673:
667:
664:
653:on 2008-12-20
652:
648:
644:
638:
635:
630:
626:
622:
618:
614:
610:
602:
599:
594:
590:
585:
580:
576:
572:
568:
564:
560:
553:
551:
549:
547:
545:
543:
539:
532:
521:
516:
514:
512:
508:
504:
500:
496:
492:
491:Aktiogavialis
487:
485:
481:
477:
476:Aktiogavialis
473:
469:
465:
461:
456:
454:
450:
446:
445:Aktiogavialis
442:
438:
437:Río Guatemala
434:
433:San Sebastián
430:
426:
422:
418:
413:
411:
410:
405:
404:
399:
395:
391:
390:Aktiogavialis
387:
383:
379:
378:Aktiogavialis
375:
372:As a typical
370:
368:
364:
360:
356:
352:
348:
344:
340:
336:
332:
329:
325:
324:
323:Aktiogavialis
312:
308:
303:
302:
295:
291:
287:
283:
282:
275:
274:
272:
267:
263:
259:
256:Vélez-Juarbe
253:
252:
251:Aktiogavialis
245:
242:
241:
238:
237:Gryposuchinae
232:
229:
228:
225:
222:
219:
218:
215:
212:
209:
208:
205:
202:
199:
196:
195:
192:
189:
186:
183:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
125:
120:
116:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
43:
36:
32:
26:
25:Aktiogavialis
22:
19:
2919:
2839:
2746:
2717:Brevirostres
2667:G. papuensis
2665:
2657:
2649:
2641:
2633:
2625:
2619:G. breviceps
2617:
2610:
2600:
2588:
2581:
2574:
2566:
2559:
2552:
2545:
2540:Penghusuchus
2538:
2530:
2522:
2515:
2508:
2500:
2493:
2486:
2478:
2473:Dadagavialis
2471:
2465:
2464:
2452:
2431:
2423:
2415:
2407:
2397:
2386:Thecachampsa
2384:
2376:
2371:Melitosaurus
2369:
2361:
2353:
2341:
2338:Tomistominae
2327:Sacacosuchus
2325:
2299:
2292:
2284:
2279:Thecachampsa
2277:
2270:
2263:
2256:
2249:
2242:
2234:
2227:
2220:
2213:
2206:
2199:
2192:
2154:
2147:
2140:
2135:C. checchiai
2133:
2126:
2116:
2104:
2096:
2088:
2080:
2072:
2061:Crocodylinae
2047:
2040:
2033:
2026:
2019:
1998:
1990:
1983:
1971:Crocodylidae
1958:Mekosuchinae
1956:
1948:
1941:
1933:
1925:
1918:
1911:
1903:
1878:Longirostres
1822:
1815:
1808:
1801:
1796:C. australis
1794:
1784:
1772:
1765:
1757:Melanosuchus
1755:
1742:
1735:
1727:
1720:
1713:
1708:Paranasuchus
1706:
1701:Paranacaiman
1699:
1692:
1685:
1678:
1671:
1664:
1657:
1649:
1642:
1635:
1629:Ceratosuchus
1627:
1620:
1612:
1605:
1598:
1570:
1563:
1556:
1549:
1542:
1535:
1530:A. hailensis
1528:
1518:
1505:
1497:
1491:Navajosuchus
1489:
1481:
1474:
1468:Ceratosuchus
1466:
1458:
1451:
1443:
1418:
1410:
1402:
1394:
1368:
1361:
1354:
1347:
1340:
1333:
1326:
1306:
1298:
1293:Navajosuchus
1291:
1285:Leidyosuchus
1283:
1276:
1269:
1261:
1256:Ceratosuchus
1254:
1247:
1240:
1233:
1203:
1198:Leidyosuchus
1196:
1191:Diplocynodon
1189:
1182:
1130:
1123:
1115:
1107:
1100:
1093:
1086:
1079:
1071:
1066:Asiatosuchus
1064:
1056:
1049:
1031:
1024:
1017:
1009:
1002:
995:
988:
981:
973:
966:
958:
951:Mekosuchinae
940:crocodilians
858:Pseudosuchia
843:Pseudosuchia
822:
818:Pseudosuchia
813:
779:crocodilians
713:
709:
698:Bibliography
684:. Retrieved
680:the original
675:
666:
655:. Retrieved
651:the original
646:
637:
612:
608:
601:
566:
562:
510:
502:
501:(river) and
498:
490:
488:
475:
463:
460:nannofossils
457:
444:
425:Puerto Rican
416:
414:
407:
401:
398:Phylogenetic
389:
377:
371:
358:
350:
322:
321:
320:
306:
300:
299:
280:
279:
257:
250:
249:
197:
184:
123:
40:28–6.8
24:
18:
2643:G. leptodus
2495:Hanyusuchus
2488:Gryposuchus
2417:T. gaudense
2286:Tienosuchus
2265:Ocepesuchus
2229:Kentisuchus
2201:Dollosuchus
2180:Gavialoidea
1722:Purussaurus
1715:Protocaiman
1687:Necrosuchus
1680:Mourasuchus
1666:Gnatusuchus
1607:Bottosaurus
1572:A. thomsoni
1551:A. munensis
1544:A. mefferdi
1460:Arambourgia
1396:Balanerodus
1356:Krabisuchus
1335:Eoalligator
1263:Deinosuchus
1242:Arambourgia
1184:Deinosuchus
1058:Arenysuchus
1026:Ultrastenos
983:Kalthifrons
441:crocodylian
403:Gryposuchus
394:eye sockets
386:quadrupedal
382:crocodilian
363:Huayquerian
355:Puerto Rico
335:crocodylian
301:A. caribesi
230:Subfamily:
2998:Gavialidae
2992:Categories
2715:See also:
2453:sensu lato
2449:Gavialinae
2313:Gavialidae
2118:Crocodylus
2042:Rimasuchus
2035:Euthecodon
2021:Brochuchus
1767:M. fisheri
1600:Acresuchus
1587:Caimaninae
1537:A. mcgrewi
1217:Globidonta
997:Mekosuchus
906:see below↓
897:Crocodilia
809:Sauropsida
686:2008-12-26
657:2008-12-26
517:References
468:South Asia
341:until the
224:Gavialidae
214:Crocodilia
2935:Q64831015
2762:Q20718019
2651:G. lewisi
2627:G. browni
2399:Tomistoma
1737:Tsoabichi
1558:A. olseni
1520:Alligator
1004:Paludirex
885:Neosuchia
870:Neosuchia
795:Kingdom:
676:WWF-India
647:WWF-India
643:"Gharial"
507:estuarine
486:species.
453:braincase
443:fossils.
374:gavialoid
367:Venezuela
361:from the
347:Caribbean
339:Oligocene
154:Kingdom:
148:Eukaryota
31:Oligocene
2962:10088914
2949:60776727
2929:Wikidata
2869:47447306
2855:Q4701881
2849:Wikidata
2776:61147059
2756:Wikidata
2602:Gavialis
2090:Quinkana
1651:Eocaiman
1271:Eocaiman
1011:Quinkana
827:Eusuchia
803:Chordata
801:Phylum:
797:Animalia
777:Extinct
740:17341454
629:91495532
593:17341454
503:gavialis
480:islandic
421:holotype
269:Species
220:Family:
178:Reptilia
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
2895:6148475
2882:8528607
2815:6148474
2802:1470112
2789:8541207
2082:Kinyang
2049:Kinyang
990:Kambara
807:Class:
731:2176176
584:2176176
472:deltaic
343:Miocene
328:extinct
243:Genus:
210:Order:
174:Class:
35:Miocene
2975:377478
2908:265607
2828:265606
1786:Caiman
1043:Others
738:
728:
627:
591:
581:
499:aktios
497:words
484:extant
326:is an
307:et al.
258:et al.
2797:IRMNG
1033:Volia
938:Basal
823:Clade
814:Clade
625:S2CID
495:Greek
449:skull
331:genus
198:Clade
185:Clade
2957:GBIF
2877:GBIF
2784:GBIF
2106:Voay
968:Baru
883:see
856:see
736:PMID
589:PMID
406:and
357:and
311:2018
290:type
262:2007
48:PreꞒ
2944:EoL
2864:EoL
2771:EoL
726:PMC
718:doi
714:274
617:doi
579:PMC
571:doi
567:274
333:of
2994::
2972::
2959::
2946::
2931::
2905::
2892::
2879::
2866::
2851::
2825::
2812::
2799::
2786::
2773::
2758::
825::
816::
734:.
724:.
712:.
708:.
674:.
645:.
623:.
613:17
611:.
587:.
577:.
565:.
561:.
541:^
412:.
376:,
369:.
309:,
260:,
200::
187::
98:Pg
42:Ma
37:,
2670:?
2662:?
2654:?
2646:?
2638:?
2630:?
2622:?
2571:?
2535:?
2527:?
2505:?
2483:?
2436:?
2428:?
2420:?
2412:?
2389:?
2381:?
2366:?
2358:?
2289:?
2239:?
2101:?
2093:?
2085:?
2077:?
2052:?
2003:?
1995:?
1961:?
1953:?
1938:?
1930:?
1908:?
1747:?
1732:?
1654:?
1632:?
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631:.
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33:-
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