714:
dorsum; the presence of hollow pup-tent-like osteoderms over neck and shoulders; triangular, dorsally compressed caudal osteoderms that are highly asymmetrical from top to bottom; and hollow and thin osteoderms. However, Arbour & Currie (2015) later diagnosed
Aletopelta based on the presence of hexagonal pelvic osteoderms that form a semi-continuous sheet over the pelvis; the pelvic osteoderms lack a dendritic surface texture; the first cervical half ring is composed of osteoderms fused to an underlying bony band rather than being closely appressed adjacent osteoderms.
3031:
3026:
154:
3054:
683:
723:
1574:
481:
129:
713:
based on the teeth being wider than tall; the femur being much longer than both the tibia and fibula; the presence of three metatarsals; a pelvic shield composed of polygonal, low-peaked osteoderms; a large short-pointed spike in the shoulder region; the presence of hollow cap-like osteoderms across
768:
based on the presence of a pelvic shield composed of co-ossified hexagonal osteoderms. Ford & Kirkland (2001) also reinterpreted it as an ankylosaurid and considered it to be taxonomically distinct enough to be warranted as a new genus and species. Vickaryous et al. (2004) considered
1893:
Park, J. Y.; Lee, Y. N.; Currie, P. J.; Kobayashi, Y.; Koppelhus, E.; Barsbold, R.; Mateus, O.; Lee, S.; Kim, S. H. (2019). "Additional skulls of
Talarurus plicatospineus (Dinosauria: Ankylosauridae) and implications for paleobiogeography and paleoecology of armored dinosaurs".
779:
due to the lack of diagnostic characteristics. Arbour & Currie (2015), however, considered it as a valid genus based on new characteristics and found
Stegopeltinae to not be a valid clade. Arbour & Currie (2015) additionally recovered
492:
at the
Californian coast. While paleontologically surveying the work, Bradford Riney noted that a skeleton had been uncovered by a ditch dug for a sewage pipe. Within days, the specimen was secured by the
497:. It was dubbed the "Carlsbad Ankylosaur". The skeleton was later described, but not named, in 1996 by Thomas Deméré and Walter Preston Coombs before being named in 2001 by Tracy Lee Ford and
674:, carrying the holotype specimen with it. The specific name, coombsi, honours the vertebrate palaeontologist Walter Preston Coombs, Jr., for their work and years of research.
3191:
3132:
1929:
G. Frauenfelder, Timothy; R. Bell, Phil; Brougham, Tom; J. Bevitt, Joseph; D. C. Bicknell, Russell; P. Kear, Benjamin; Wroe, Stephen; E. Campione, Nicolás (2022).
537:
was once a bloated carcass that had been washed out to sea, likely by a stream, which sank to the bottom with its underside facing upwards and became a miniature
1733:
T.L. Ford. (2000). "A review of ankylosaur osteoderms from New Mexico and a preliminary review of ankylosaur armor", In: S. G. Lucas and A. B. Heckert (eds.),
513:, foraminifers, scaphopods, pelecypods, gastropods, cephalopods, ostracods, decapods, echinoids, elasmobranchs, and actinopterygians, with the addition of a
1615:
W.P. Coombs, Jr. and T.A. Deméré, 1996, "A Late
Cretaceous nodosaurid ankylosaur (Dinosauria: Ornithischia) from marine sediments of coastal California",
754:
but refrained from naming it as they considered it to be undiagnostic. Ford (2000) considered that SDNHM 33909 formed the clade
Stegopeltinae, along with
3211:
3201:
3119:
3030:
3196:
1985:
1781:"A new ankylosaurine dinosaur from the Judith River Formation of Montana, USA, based on an exceptional skeleton with soft tissue preservation"
806:
and more derived ankylosaurids. A similar position was recovered by Arbour & Evans (2017) and Rivera-Sylva et al. (2018) but also placed
448:
preserving osteoderms. It was originally described in 1996 by W. P. Coombs, Jr. and T.A. Deméré before being named in 2001 by Tracy Ford and
3216:
3206:
559:
and gastropods found in association with the skeleton. The carcass was also scavenged upon by marine invertebrates and sharks such as
1763:
852:. Although most often being recovered as a basal ankylosaurid, Frauenfelder et al. (2022) found it to be the basalmost nodosaurid.
494:
137:
3047:
1587:
283:
3221:
3025:
1165:
1931:"New Ankylosaurian Cranial Remains From the Lower Cretaceous (Upper Albian) Toolebuc Formation of Queensland, Australia"
1836:
Rivera-Sylva, H.E.; Frey, E.; Stinnesbeck, W.; Carbot-Chanona, G.; Sanchez-Uribe, I.E.; Guzmán-Gutiérrez, J.R. (2018).
1691:
Arbour, V. M.; Currie, P. J. (2015). "Systematics, phylogeny and palaeobiogeography of the ankylosaurid dinosaurs".
153:
1978:
571:, which resulted with most of the long bones being hollow and many shallow pits on the osteoderms and ribs. The
2076:
510:
1754:& Weishampel, D.B. (2004). "Ankylosauria". In Weishampel, D. B.; Dodson, P. & Osmólska, H. (eds.).
639:
shield and cervical half ring. The type specimen may represent an immature individual based on the unfused
3186:
3074:
3158:
2103:
1971:
1148:
310:
3106:
3057:
1849:
1792:
1700:
502:
489:
435:
287:
855:
The results of a phylogenetic analysis conducted by Arbour & Currie (2015) is reproduced below.
2463:
2360:
1751:
883:
2199:
1911:
1875:
1716:
148:
1838:"Paleodiversity of Late Cretaceous Ankylosauria from Mexico and their phylogenetic significance"
3163:
3145:
3137:
2956:
2220:
1867:
1818:
1759:
1643:
1336:
498:
449:
3150:
706:
gave a lower estimate of 5 metres (16 feet) long and weight of 2 tonnes (4,409 lbs) in 2010.
2421:
2368:
2206:
1942:
1903:
1857:
1808:
1800:
1708:
1579:
760:
726:
344:
2886:
2850:
2790:
2756:
2749:
2642:
2583:
2455:
2352:
2268:
2228:
2061:
1123:
950:
790:
703:
671:
567:
427:
41:
1853:
1796:
1704:
1644:"Carlsbad ankylosaur (Ornithischia, Ankylosauria): an ankylosaurid and not a nodosaurid"
2949:
2935:
2921:
2829:
2802:
2698:
2679:
2594:
2540:
2405:
2375:
2345:
2308:
2284:
1813:
1780:
1419:
1393:
862:
848:
765:
663:
416:
262:
3180:
2912:
2822:
2815:
2767:
2711:
2658:
2624:
2525:
2471:
2446:
2428:
2331:
2277:
2250:
2242:
1915:
1879:
1720:
1081:
900:
814:
744:
651:
596:
530:
465:
79:
456:
has an estimated size of 5 metres (16 feet) and weight of 2 tonnes (4,409 lbs). The
2942:
2928:
2857:
2741:
2665:
2610:
2561:
2533:
2518:
2505:
2478:
2397:
2382:
2338:
2318:
2185:
2167:
2145:
2115:
2088:
2036:
1994:
1376:
1288:
808:
775:
733:
409:
246:
214:
1907:
1735:
Dinosaurs of New Mexico. New Mexico Museum of
Natural History and Science Bulletin
1712:
732:
Deméré & Coombs (1996) originally interpreted SDNHM 33909 as an indeterminate
17:
2977:
2894:
2843:
2783:
2734:
2576:
2554:
2486:
2435:
2235:
2213:
2045:
1443:
1105:
1048:
991:
836:
826:
802:
561:
230:
54:
682:
2970:
2963:
2901:
2776:
2727:
2672:
2617:
2603:
2569:
2547:
2413:
2257:
2192:
1947:
1930:
1862:
1837:
1569:
1246:
1031:
974:
842:
820:
796:
750:
738:
666:
containing the
Peninsular Ranges Terrane was somewhere opposite the middle of
640:
632:
423:
141:
99:
64:
3097:
1871:
2984:
2864:
2651:
2631:
2493:
2390:
2292:
1319:
1205:
620:
556:
550:
526:
506:
431:
165:
104:
48:
1822:
722:
480:
3091:
2991:
2027:
2018:
1460:
785:
572:
457:
445:
419:
198:
185:
94:
89:
74:
69:
59:
1804:
3124:
2879:
2871:
1229:
604:
588:
584:
518:
128:
109:
84:
2012:
1758:(Second ed.). University of California Press. pp. 363–392.
667:
636:
624:
616:
543:
469:
175:
3068:
662:" (small shield), in reference to its armour and the fact that the
2836:
721:
681:
612:
608:
580:
522:
514:
509:
stage, 75.5 Ma. The formation has yielded specimens pertaining to
488:
In 1987, construction work was done on the
College Boulevard near
479:
412:
3111:
2998:
600:
592:
538:
461:
3072:
2005:
1967:
1963:
628:
356:
400:
391:
380:
371:
362:
347:
702:
an estimated length of 6 metres (19.7 feet). However,
501:. The skeleton originated from a layer of the marine
397:
388:
350:
830:
within the polytomy. Park et al. (2019) also placed
377:
374:
368:
359:
353:
3081:
2911:
2801:
2766:
2697:
2641:
2593:
2504:
2445:
2317:
2306:
2267:
2165:
2114:
2087:
2060:
1661:
Dinosaurs and Other
Mesozoic Reptiles of California
394:
385:
365:
1650:. Indiana University Press, Bloomington pp 239-260
643:, partly fused scutes and unfused neural spines.
670:in the Cretaceous, only moving northward due to
555:being attached to the bones and the presence of
709:Ford and Kirkland (2001) originally diagnosed
1979:
1779:Arbour, Victoria M.; Evans, David C. (2017).
8:
1686:
1684:
1682:
1611:
1609:
1607:
1605:
1603:
834:within a polytomy but with the inclusion of
698:Ford & Kirkland (2001) originally gave
623:, dorsal neural arch, neural arches of the
484:Point Loma Formation in Southern California
3192:Late Cretaceous dinosaurs of North America
3069:
2807:
2703:
2510:
2323:
2314:
2177:
2173:
2162:
2093:
2066:
2057:
2002:
1986:
1972:
1964:
1638:
1636:
1634:
1632:
1630:
1628:
127:
31:
1946:
1861:
1812:
1599:
1674:The Princeton Field Guide to Dinosaurs
533:. The type and only known specimen of
7:
541:, as evidence by Pelecypoda such as
136:Holotype specimen on display at the
1693:Journal of Systematic Palaeontology
1676:, Princeton University Press p. 233
1642:T.L. Ford and J.I. Kirkland, 2001,
1435:
1368:
1361:
1311:
1304:
1280:
1221:
1197:
1190:
1140:
1097:
1073:
1023:
1016:
966:
925:
875:
868:
858:
434:stage, 75.5 Ma) in what is now the
603:, partial left and possibly right
408:; meaning 'wanderer shield') is a
25:
3212:Campanian genus first appearances
3053:
3052:
3029:
3024:
1572:
495:San Diego Natural History Museum
343:
152:
138:San Diego Natural History Museum
3202:Taxa named by James I. Kirkland
1588:Timeline of ankylosaur research
1842:Swiss Journal of Palaeontology
736:that showed similarities with
694:Size and distinguishing traits
1:
3197:Fossil taxa described in 2001
1908:10.1016/j.cretres.2019.104340
1713:10.1080/14772019.2015.1059985
1166:Pinacosaurus mephistocephalus
595:, possible fragment of right
438:. The type and only species,
3217:Campanian genus extinctions
3238:
3207:Paleontology in California
1935:Frontiers in Earth Science
1785:Royal Society Open Science
464:and was scavenged upon by
444:, is known from a partial
27:Extinct genus of dinosaurs
3042:
3022:
2810:
2706:
2513:
2326:
2180:
2176:
2161:
2142:
2096:
2069:
2056:
2001:
1948:10.3389/feart.2022.803505
1863:10.1007/s13358-018-0153-1
1457:
1440:
1433:
1416:
1390:
1373:
1366:
1359:
1333:
1316:
1309:
1302:
1285:
1278:
1243:
1226:
1219:
1202:
1195:
1188:
1162:
1145:
1138:
1119:
1102:
1095:
1078:
1071:
1045:
1028:
1021:
1014:
988:
971:
964:
947:
930:
923:
897:
880:
873:
866:
505:which dates to the upper
329:Ford & Kirkland, 2001
316:
309:
149:Scientific classification
147:
135:
126:
34:
511:calcareous nannoplankton
1617:Journal of Paleontology
619:, four or five partial
729:
690:
485:
426:that lived during the
3159:Paleobiology Database
1648:The Armored Dinosaurs
1149:Pinacosaurus grangeri
725:
685:
650:, is composed of the
483:
3222:Ornithischian genera
1663:, 2003), pp. 233-36.
503:Point Loma Formation
476:Discovery and naming
436:Point Loma Formation
1896:Cretaceous Research
1854:2018SwJP..137...83R
1805:10.1098/rsos.161086
1797:2017RSOS....461086A
1705:2016JSPal..14..385A
686:Size comparison of
460:formed a miniature
1750:Vickaryous, M.K.;
1672:Paul, G.S., 2010,
730:
691:
658:" (wanderer) and "
646:The generic name,
486:
441:Aletopelta coombsi
323:Aletopelta coombsi
18:Aletopelta coombsi
3174:
3173:
3146:Open Tree of Life
3075:Taxon identifiers
3066:
3065:
3038:
3037:
3020:
3019:
3016:
3015:
3012:
3011:
3008:
3007:
2957:Nodocephalosaurus
2693:
2692:
2689:
2688:
2302:
2301:
2221:Serendipaceratops
2157:
2156:
2153:
2152:
2138:
2137:
2134:
2133:
1562:
1561:
1553:
1552:
1544:
1543:
1535:
1534:
1526:
1525:
1517:
1516:
1508:
1507:
1499:
1498:
1490:
1489:
1481:
1480:
1472:
1471:
1405:
1404:
1348:
1347:
1337:Nodocephalosaurus
1267:
1266:
1258:
1257:
1177:
1176:
1060:
1059:
1003:
1002:
912:
911:
334:
333:
290:
16:(Redirected from
3229:
3167:
3166:
3154:
3153:
3141:
3140:
3128:
3127:
3115:
3114:
3102:
3101:
3100:
3070:
3056:
3055:
3033:
3028:
2808:
2704:
2511:
2422:Propanoplosaurus
2369:Glyptodontopelta
2324:
2315:
2207:Priodontognathus
2178:
2174:
2163:
2094:
2067:
2058:
2051:
2050:
2003:
1988:
1981:
1974:
1965:
1953:
1952:
1950:
1926:
1920:
1919:
1890:
1884:
1883:
1865:
1833:
1827:
1826:
1816:
1776:
1770:
1769:
1747:
1741:
1731:
1725:
1724:
1688:
1677:
1670:
1664:
1657:
1651:
1640:
1623:
1613:
1582:
1580:Dinosaurs portal
1577:
1576:
1575:
1436:
1369:
1362:
1312:
1305:
1281:
1222:
1198:
1191:
1141:
1098:
1074:
1024:
1017:
967:
926:
876:
869:
859:
761:Glyptodontopelta
727:Life restoration
407:
406:
403:
402:
399:
396:
393:
390:
387:
383:
382:
379:
376:
373:
370:
367:
364:
361:
358:
355:
352:
349:
325:
321:
281:
274:
261:
245:
229:
213:
157:
156:
131:
121:
51:
40:Temporal range:
32:
21:
3237:
3236:
3232:
3231:
3230:
3228:
3227:
3226:
3177:
3176:
3175:
3170:
3162:
3157:
3149:
3144:
3136:
3131:
3123:
3118:
3110:
3105:
3096:
3095:
3090:
3077:
3067:
3062:
3034:
3004:
2907:
2887:Tianzhenosaurus
2851:Minotaurasaurus
2797:
2791:Zhongyuansaurus
2762:
2757:Zhejiangosaurus
2750:Liaoningosaurus
2685:
2643:Struthiosaurini
2637:
2589:
2584:Tatankacephalus
2500:
2456:Gargoyleosaurus
2441:
2353:Gargoyleosaurus
2311:
2298:
2269:Parankylosauria
2263:
2229:Sinankylosaurus
2170:
2149:
2130:
2110:
2083:
2077:Avemetatarsalia
2062:Avemetatarsalia
2052:
2008:
2007:
1997:
1992:
1962:
1957:
1956:
1928:
1927:
1923:
1892:
1891:
1887:
1835:
1834:
1830:
1778:
1777:
1773:
1766:
1749:
1748:
1744:
1732:
1728:
1690:
1689:
1680:
1671:
1667:
1658:
1654:
1641:
1626:
1614:
1601:
1596:
1578:
1573:
1571:
1568:
1563:
1554:
1545:
1536:
1527:
1518:
1509:
1500:
1491:
1482:
1473:
1406:
1349:
1268:
1259:
1178:
1124:Zhejiangosaurus
1061:
1004:
951:Liaoningosaurus
913:
791:Liaoningosaurus
720:
704:Gregory S. Paul
696:
680:
672:plate tectonics
568:Scapanorhynchus
478:
428:Late Cretaceous
384:
346:
342:
330:
327:
319:
318:
305:
302:A. coombsi
280:
272:
259:
243:
227:
211:
151:
122:
120:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
46:
45:
42:Late Cretaceous
38:
28:
23:
22:
15:
12:
11:
5:
3235:
3233:
3225:
3224:
3219:
3214:
3209:
3204:
3199:
3194:
3189:
3179:
3178:
3172:
3171:
3169:
3168:
3155:
3142:
3129:
3116:
3103:
3087:
3085:
3079:
3078:
3073:
3064:
3063:
3061:
3060:
3050:
3043:
3040:
3039:
3036:
3035:
3023:
3021:
3018:
3017:
3014:
3013:
3010:
3009:
3006:
3005:
3003:
3002:
2995:
2988:
2981:
2974:
2967:
2960:
2953:
2950:Euoplocephalus
2946:
2939:
2936:Anodontosaurus
2932:
2925:
2922:Akainacephalus
2917:
2915:
2909:
2908:
2906:
2905:
2898:
2891:
2883:
2876:
2868:
2861:
2854:
2847:
2840:
2833:
2830:Crichtonsaurus
2826:
2819:
2811:
2805:
2803:Ankylosaurinae
2799:
2798:
2796:
2795:
2787:
2780:
2772:
2770:
2764:
2763:
2761:
2760:
2753:
2746:
2738:
2731:
2724:
2716:
2707:
2701:
2699:Ankylosauridae
2695:
2694:
2691:
2690:
2687:
2686:
2684:
2683:
2680:Struthiosaurus
2676:
2669:
2662:
2655:
2647:
2645:
2639:
2638:
2636:
2635:
2628:
2621:
2614:
2607:
2599:
2597:
2595:Panoplosaurini
2591:
2590:
2588:
2587:
2580:
2573:
2566:
2558:
2551:
2544:
2541:Niobrarasaurus
2537:
2530:
2522:
2514:
2508:
2502:
2501:
2499:
2498:
2490:
2483:
2475:
2468:
2460:
2451:
2449:
2443:
2442:
2440:
2439:
2432:
2425:
2418:
2410:
2406:Polacanthoides
2402:
2394:
2387:
2379:
2376:Horshamosaurus
2372:
2365:
2357:
2349:
2346:Dongyangopelta
2342:
2335:
2327:
2321:
2312:
2309:Euankylosauria
2307:
2304:
2303:
2300:
2299:
2297:
2296:
2289:
2285:Kunbarrasaurus
2281:
2273:
2271:
2265:
2264:
2262:
2261:
2254:
2247:
2239:
2232:
2225:
2217:
2210:
2203:
2196:
2189:
2181:
2171:
2166:
2159:
2158:
2155:
2154:
2151:
2150:
2143:
2140:
2139:
2136:
2135:
2132:
2131:
2129:
2128:
2127:
2126:
2120:
2118:
2112:
2111:
2109:
2108:
2107:
2106:
2097:
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766:Ankylosauridae
719:
718:Classification
716:
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679:
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664:tectonic plate
627:, fragmentary
583:, fragmentary
579:, consists of
499:James Kirkland
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2823:Crichtonpelta
2820:
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2816:Bissektipelta
2813:
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2809:
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2800:
2793:
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2768:Shamosaurinae
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2709:
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2700:
2696:
2682:
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2677:
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2670:
2668:
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2659:Hungarosaurus
2656:
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2523:
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2496:
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2472:Hoplitosaurus
2469:
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2452:
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2447:Polacanthinae
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2255:
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1082:Crichtonpelta
1076:
1075:
1069:
1068:
1065:
1064:
1056:
1055:
1052:
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1050:
1043:
1042:
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1019:
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936:
935:
928:
927:
921:
920:
917:
916:
908:
907:
904:
903:
902:
901:Ahshislepelta
895:
894:
891:
890:
887:
886:
885:
878:
877:
871:
870:
864:
861:
860:
856:
853:
851:
850:
845:
844:
839:
838:
833:
829:
828:
823:
822:
817:
816:
815:Crichtonpelta
811:
810:
805:
804:
799:
798:
793:
792:
787:
783:
778:
777:
772:
767:
763:
762:
757:
753:
752:
747:
746:
745:Panoplosaurus
741:
740:
735:
728:
724:
717:
715:
712:
707:
705:
701:
693:
689:
684:
677:
675:
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669:
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618:
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598:
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590:
586:
582:
578:
574:
570:
569:
564:
563:
558:
554:
552:
547:
545:
540:
536:
532:
528:
525:and cervical
524:
520:
516:
512:
508:
504:
500:
496:
491:
482:
475:
473:
471:
467:
466:invertebrates
463:
459:
455:
451:
447:
443:
442:
437:
433:
429:
425:
421:
418:
414:
411:
405:
340:
339:
326:
324:
315:
312:
311:Binomial name
308:
304:
303:
297:
294:
293:
289:
285:
279:
278:
271:
268:
267:
264:
258:
255:
252:
251:
248:
242:
239:
236:
235:
232:
226:
223:
220:
219:
216:
210:
207:
204:
203:
200:
197:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
155:
150:
146:
143:
139:
134:
130:
125:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
50:
43:
37:
33:
30:
19:
3082:
2997:
2990:
2983:
2976:
2969:
2962:
2955:
2948:
2943:Dyoplosaurus
2941:
2934:
2929:Ankylosaurus
2927:
2920:
2900:
2893:
2885:
2878:
2870:
2863:
2858:Pinacosaurus
2856:
2849:
2842:
2835:
2828:
2821:
2814:
2789:
2782:
2775:
2755:
2748:
2742:Hylaeosaurus
2740:
2733:
2726:
2719:
2718:
2710:
2678:
2671:
2666:Pawpawsaurus
2664:
2657:
2650:
2630:
2623:
2616:
2611:Denversaurus
2609:
2602:
2582:
2575:
2568:
2562:Peloroplites
2560:
2553:
2546:
2539:
2534:Borealopelta
2532:
2524:
2519:Acantholipan
2517:
2506:Nodosaurinae
2492:
2485:
2479:Peloroplites
2477:
2470:
2462:
2454:
2434:
2427:
2420:
2412:
2404:
2398:Mymoorapelta
2396:
2389:
2383:Hylaeosaurus
2381:
2374:
2367:
2359:
2351:
2344:
2339:Anoplosaurus
2337:
2330:
2319:Nodosauridae
2291:
2283:
2276:
2256:
2249:
2241:
2234:
2227:
2219:
2212:
2205:
2198:
2191:
2186:Cryptosaurus
2184:
2168:Ankylosauria
2146:Ankylosauria
2144:
2116:Ankylosauria
2104:Ornithischia
2089:Ornithischia
2041:
2037:Ornithischia
2032:
2023:
1995:Ankylosauria
1938:
1934:
1924:
1899:
1895:
1888:
1848:(1): 83–93.
1845:
1841:
1831:
1788:
1784:
1774:
1755:
1745:
1737:
1734:
1729:
1696:
1692:
1673:
1668:
1660:
1655:
1647:
1622:(2): 311-326
1619:
1616:
1459:
1458:
1442:
1441:
1418:
1417:
1392:
1391:
1377:Ankylosaurus
1375:
1374:
1335:
1334:
1318:
1317:
1289:Dyoplosaurus
1287:
1286:
1245:
1244:
1228:
1227:
1204:
1203:
1164:
1163:
1147:
1146:
1121:
1120:
1104:
1103:
1080:
1079:
1047:
1046:
1030:
1029:
990:
989:
973:
972:
949:
948:
933:
932:
931:
899:
898:
882:
881:
854:
847:
841:
835:
831:
825:
819:
813:
809:Hylaeosaurus
807:
801:
795:
789:
781:
776:nomen dubium
774:
770:
759:
755:
749:
743:
737:
731:
710:
708:
699:
697:
687:
659:
655:
647:
645:
576:
566:
560:
549:
542:
534:
487:
453:
440:
439:
417:ankylosaurid
410:monospecific
337:
336:
335:
322:
317:
301:
300:
276:
275:
253:
247:Ankylosauria
237:
221:
215:Ornithischia
205:
192:
35:
29:
2978:Scolosaurus
2895:Tsagantegia
2844:Jinyunpelta
2784:Shamosaurus
2735:Chuanqilong
2577:Silvisaurus
2555:Patagopelta
2487:Polacanthus
2436:Sauroplites
2236:Spicomellus
2214:Sarcolestes
2046:Thyreophora
1699:(5): 1–60.
1444:Scolosaurus
1106:Tsagantegia
1049:Shamosaurus
992:Chuanqilong
837:Jinyunpelta
827:Shamosaurus
803:Chuanqilong
788:containing
678:Description
577:SDNHM 33909
562:Squalicorax
521:containing
282:Ford &
231:Thyreophora
3181:Categories
3083:Aletopelta
3046:See also:
2971:Platypelta
2964:Oohkotokia
2902:Zaraapelta
2777:Gobisaurus
2728:Cedarpelta
2720:Aletopelta
2673:Stegopelta
2618:Edmontonia
2604:Animantarx
2570:Sauropelta
2548:Nodosaurus
2414:Priconodon
2258:Vectipelta
2193:Dracopelta
2125:see below↓
2028:Dinosauria
1902:: 104340.
1594:References
1247:Zaraapelta
1032:Gobisaurus
975:Cedarpelta
934:Aletopelta
843:Nodosaurus
832:Aletopelta
821:Gobisaurus
797:Cedarpelta
782:Aletopelta
771:Aletopelta
756:Stegopelta
751:Stegopelta
739:Edmontonia
734:nodosaurid
711:Aletopelta
700:Aletopelta
688:Aletopelta
648:Aletopelta
641:astragalus
635:including
633:osteoderms
591:, partial
587:, partial
575:specimen,
535:Aletopelta
454:Aletopelta
424:California
338:Aletopelta
277:Aletopelta
199:Dinosauria
142:California
47:75.5
36:Aletopelta
2985:Talarurus
2865:Saichania
2652:Europelta
2632:Texasetes
2494:Taohelong
2391:Invictarx
2293:Stegouros
2011:Kingdom:
1916:212423361
1880:134924657
1872:1664-2376
1740:: 157-176
1721:214625754
1320:Talarurus
1206:Saichania
784:within a
764:, within
621:vertebrae
557:ammonites
551:Spondylus
531:hadrosaur
527:vertebrae
507:Campanian
432:Campanian
415:of basal
295:Species:
172:Kingdom:
166:Eukaryota
3112:52571830
3092:Wikidata
3058:Category
3048:Timeline
2992:Ziapelta
2464:Gastonia
2361:Gastonia
2019:Chordata
2017:Phylum:
2013:Animalia
1941:: 1–17.
1823:28573004
1659:Hilton (
1566:See also
1461:Ziapelta
884:Gastonia
786:polytomy
585:scapulae
573:holotype
517:, right
490:Carlsbad
458:holotype
446:skeleton
420:dinosaur
284:Kirkland
186:Chordata
182:Phylum:
176:Animalia
162:Domain:
3151:4128538
3138:1392218
3125:4823220
3098:Q134299
2880:Tarchia
2872:Shanxia
1850:Bibcode
1814:5451805
1793:Bibcode
1701:Bibcode
1230:Tarchia
654:words "
617:fibulae
605:ischium
589:humerus
519:dentary
430:(upper
320:†
299:†
269:Genus:
116:↓
1914:
1878:
1870:
1821:
1811:
1762:
1719:
1646:, In:
668:Mexico
656:aletes
637:pelvic
625:sacrum
613:tibiae
609:femora
597:radius
544:Ostrea
470:sharks
3164:81277
3133:IRMNG
2837:Datai
2200:Minmi
2042:Clade
2033:Clade
2024:Clade
1912:S2CID
1876:S2CID
1717:S2CID
773:as a
660:pelte
652:Greek
581:teeth
529:of a
523:teeth
515:femur
413:genus
254:Clade
238:Clade
222:Clade
206:Clade
193:Clade
3120:GBIF
2999:Zuul
2102:see
2075:see
1868:ISSN
1819:PMID
1760:ISBN
846:and
824:and
758:and
748:and
629:ribs
601:ulna
593:ulna
565:and
548:and
539:reef
468:and
462:reef
288:2001
55:PreꞒ
3107:EoL
1943:doi
1904:doi
1900:108
1858:doi
1846:137
1809:PMC
1801:doi
1709:doi
553:sp.
546:sp.
3183::
3161::
3148::
3135::
3122::
3109::
3094::
2044::
2035::
2026::
1939:10
1937:.
1933:.
1910:.
1898:.
1874:.
1866:.
1856:.
1844:.
1840:.
1817:.
1807:.
1799:.
1787:.
1783:.
1738:17
1715:.
1707:.
1697:14
1695:.
1681:^
1627:^
1620:70
1602:^
840:,
818:,
812:,
800:,
794:,
742:,
631:,
615:,
611:,
607:,
599:,
472:.
452:.
392:oʊ
363:oʊ
357:iː
286:,
256::
240::
224::
208::
195::
140:,
105:Pg
49:Ma
44:,
2890:?
2875:?
2794:?
2745:?
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2565:?
2529:?
2497:?
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2356:?
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2246:?
2224:?
1987:e
1980:t
1973:v
1951:.
1945::
1918:.
1906::
1882:.
1860::
1852::
1825:.
1803::
1795::
1789:4
1768:.
1723:.
1711::
1703::
1126:"
1122:"
404:/
401:i
398:z
395:m
389:k
386:ˈ
381:ə
378:t
375:l
372:ɛ
369:p
366:ˈ
360:t
354:l
351:ˌ
348:ə
345:/
341:(
273:†
260:†
244:†
228:†
212:†
110:N
100:K
95:J
90:T
85:P
80:C
75:D
70:S
65:O
60:Ꞓ
20:)
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