443:
472:
between the scape and club, are thickened. Both are features seen in ant species that typically soil or leaf litter dwelling. Due to positioning of the worker in the amber, two of the important features distinguishing the subfamily
Aneuretinae genera are not directly visible. An elongated one segmented petiole is indicated by the lower impression of the petiole in the amber, though the upper portion of the petiole along with the upper portion of the mesosoma after the
324:. The amber was recovered from fossil bearing rocks in the Sakhalin region of Eastern Russia. Sakhalin amber is noted for having undergone high temperatures and pressures after the resin was buried. As a result insects and other inclusions in the amber are not as well preserved as those of other amber locations, even those of older ambers. The inclusions are most of the time carbonized reducing fine detail preservation, and the amber has been subjected to
110:
471:
in shape, with a notably convex rear margin. The antennae have a basal segment, that does not reach past the occipital margin, eleven additional segments, the last three of which are enlarged and form a terminal club. The eyes are shifted forward on the head, while the funicular segments, those
348:
suggested a tentative
Paleocene age, which was followed by subsequent authors through 2013. However research published in 1999 on the Naibuchi Formation, in which Sakhalin amber is directly preserved, gives a Middle Eocene age based on geological and paleobotanical context. The Sakhalin amber
328:
changing the shapes and features of the inclusions. The distortion is visible in the outlines of trapped air bubbles, which are elongated along the plane of distortion, rather than spherical as seen with bubbles in Eocene
680:
Dlussky, G.M.; Wedmann, S. (2012). "The poneromorph ants (Hymenoptera, Formicidae: Amblyoponinae, Ectatomminae, Ponerinae) of Grube Messel, Germany: High biodiversity in the Eocene".
510:
317:
349:
forest had a variety of plants living in a mixed coastal swamp, river, and lake environment. The river and lake system had numerous swampy areas that resulted in active
802:
661:
Dlussky, G.M.; Rasnitsyn, A.P. (2002). "Ants (Hymenoptera: Formicidae) of formation Green river and some other Middle Eocene deposits of North
America".
789:
815:
874:
539:
sp. nov. (Hymenoptera, Formicidae) from the
Sakhalin amberβthe earliest described species of an extant genus of Myrmicinae".
820:
316:
in a transparent chunk of
Sakhalin amber. When the fossil was described it was part of the amber collections housed in the
388:
The amber fossil was first studied by paleoentomologist
Gennady Dlussky of the Russian Academy of Sciences, with his 1988
109:
321:
889:
879:
442:
894:
884:
477:
731:
841:
232:
630:
Emery, 1893 (Formicidae, Aneuretinae) and other ants in
Indikada Mukalana Forest Reserve in Sri Lanka"
869:
689:
548:
345:
715:
429:
337:. In some cases the inclusions are at least partially filled with amber rather than being hollow.
325:
313:
334:
705:
603:
564:
463:
has a body length of 3 mm (0.12 in), a head length of 0.61 mm (0.024 in) and
417:
341:
104:
846:
480:
is missing, the lower impression of the petiole shows it was probably widening at the junction.
768:
828:
807:
776:
464:
282:, and is one of eight genera of the subfamily. The genus contains a single described species
833:
415:
into the ant subfamily
Aneuretinae. The subfamily includes three additional extinct genera,
697:
641:
595:
556:
447:
435:
408:
297:
693:
552:
389:
863:
289:
63:
32:
709:
607:
584:"Orthoclads from Eocene Amber from Sakhalin (Diptera: Chironomidae, Orthocladiinae)"
568:
400:
was coined from the Latin word meaning "deformed". The etymology of the genus name
423:
360:
330:
781:
754:
701:
279:
181:
161:
38:
599:
560:
191:
83:
48:
646:
625:
476:
are missing from the specimen. Though the junction of the petiole with the
468:
364:
141:
121:
88:
748:
473:
369:
350:
293:
275:
78:
73:
58:
53:
43:
794:
355:
344:
in 1978 suggesting dates between 59 and 47 million years old. In 1988,
264:
93:
68:
583:
763:
151:
131:
725:
340:
Sakhalin amber has been attributed a range of geological ages, with
441:
381:
375:
267:
729:
271:
171:
626:"Nest density and other observations on a population of
535:
Radchenko, A. G.; Perkovsky, E. E. (2016). "The ant
738:
582:Baranov, V.; Andersen, T.; Perkovsky, E.E. (2014).
504:
502:
500:
498:
496:
494:
492:
467:with twelve segments. The head is generally
8:
726:
619:
617:
20:
645:
624:Dias, R. K.; Udayakantha, W. S. (2016).
433:, and a single living genus and species
411:and shape of the gaster, Dlussky placed
385:, and other trees populated the forest.
488:
353:formation. The bogs were surrounded by
312:was known from a single fossil insect
511:"Ants of Sakhalin amber (Paleocene?)"
7:
682:Journal of Systematic Palaeontology
588:Insect Systematics & Evolution
14:
108:
634:Journal of Insect Biodiversity
407:Based on the structure of the
404:was not explained by Dlussky.
392:for the species published in
1:
875:Fossil taxa described in 1988
663:Russian Entomological Journal
702:10.1080/14772019.2011.628341
518:Paleontologicheskii Zhurnal
394:Paleontologicheskii Zhurnal
322:Russian Academy of Sciences
911:
304:History and classification
292:fossil which was found in
600:10.1163/1876312X-45032122
561:10.1134/S0031030116090136
318:Paleontological Institute
238:
231:
105:Scientific classification
103:
23:
537:Aphaenogaster dlusskyana
719:(subscription required)
541:Paleontological Journal
509:Dlussky, G. M. (1988).
714: β via
647:10.12976/jib/2016.4.12
451:
439:, found in Sri Lanka.
308:When first described,
288:and is known from one
842:Paleobiology Database
445:
285:Aneuretellus deformis
245:Aneuretellus deformis
716:Taylor & Francis
396:. The species name
694:2012JSPal..10..725D
553:2016PalJ...50..936R
459:The lone worker of
326:plastic deformation
452:
342:Vladimir Zherikhin
890:Fossil ant genera
857:
856:
829:Open Tree of Life
732:Taxon identifiers
256:
255:
212:
902:
850:
849:
837:
836:
824:
823:
811:
810:
798:
797:
785:
784:
772:
771:
759:
758:
757:
727:
721:
720:
713:
677:
671:
670:
658:
652:
651:
649:
628:Aneuretus simoni
621:
612:
611:
579:
573:
572:
532:
526:
525:
515:
506:
448:Aneuretus simoni
436:Aneuretus simoni
390:type description
298:Russian Far East
247:
243:
224:A. deformis
210:
203:
113:
112:
98:
35:
29:Temporal range:
21:
910:
909:
905:
904:
903:
901:
900:
899:
880:Insects of Asia
860:
859:
858:
853:
845:
840:
832:
827:
819:
814:
806:
801:
793:
788:
780:
775:
767:
762:
753:
752:
747:
734:
724:
718:
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674:
660:
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623:
622:
615:
581:
580:
576:
534:
533:
529:
513:
508:
507:
490:
486:
457:
346:Gennady Dlussky
333:and Cretaceous
306:
252:
249:
241:
240:
227:
209:
201:
107:
99:
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96:
91:
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81:
76:
71:
66:
61:
56:
51:
46:
41:
31:
30:
27:
17:
12:
11:
5:
908:
906:
898:
897:
895:Sakhalin amber
892:
887:
885:Eocene insects
882:
877:
872:
862:
861:
855:
854:
852:
851:
838:
825:
812:
799:
786:
773:
760:
744:
742:
736:
735:
730:
723:
722:
688:(4): 725β753.
672:
653:
613:
594:(4): 359β378.
574:
547:(9): 936β946.
527:
520:(in Russian).
487:
485:
482:
456:
453:
367:plants, while
305:
302:
254:
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250:
236:
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139:
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92:
87:
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77:
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67:
62:
57:
52:
47:
42:
37:
36:
28:
15:
13:
10:
9:
6:
4:
3:
2:
907:
896:
893:
891:
888:
886:
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868:
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865:
848:
843:
839:
835:
830:
826:
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756:
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746:
745:
743:
741:
737:
733:
728:
717:
711:
707:
703:
699:
695:
691:
687:
683:
676:
673:
669:(4): 411β436.
668:
664:
657:
654:
648:
643:
639:
635:
631:
629:
620:
618:
614:
609:
605:
601:
597:
593:
589:
585:
578:
575:
570:
566:
562:
558:
554:
550:
546:
542:
538:
531:
528:
523:
519:
512:
505:
503:
501:
499:
497:
495:
493:
489:
483:
481:
479:
475:
470:
466:
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454:
450:
449:
444:
440:
438:
437:
432:
431:
430:Protaneuretus
426:
425:
420:
419:
414:
410:
405:
403:
399:
395:
391:
386:
384:
383:
378:
377:
372:
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366:
362:
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357:
352:
347:
343:
338:
336:
332:
327:
323:
319:
315:
311:
303:
301:
299:
295:
291:
290:Middle Eocene
287:
286:
281:
277:
273:
269:
266:
262:
261:
251:Dlussky, 1988
248:
246:
237:
234:
233:Binomial name
230:
226:
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219:
216:
215:
211:Dlussky, 1988
208:
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106:
102:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
34:
33:Middle Eocene
26:
22:
19:
16:Genus of ants
769:Aneuretellus
740:Aneuretellus
739:
685:
681:
675:
666:
662:
656:
637:
633:
627:
591:
587:
577:
544:
540:
536:
530:
521:
517:
461:Aneuretellus
460:
458:
446:
434:
428:
424:Paraneuretus
422:
416:
412:
406:
402:Aneuretellus
401:
397:
393:
387:
380:
374:
368:
361:Nymphaeaceae
354:
339:
335:Taymyr amber
331:Baltic amber
309:
307:
284:
283:
260:Aneuretellus
259:
258:
257:
244:
239:
223:
222:
206:Aneuretellus
205:
204:
25:Aneuretellus
24:
18:
870:Aneuretinae
640:(12): 1β9.
524:(1): 50β61.
455:Description
418:Mianeuretus
413:A. deformis
310:A. deformis
280:Aneuretinae
182:Aneuretinae
178:Subfamily:
162:Hymenoptera
864:Categories
484:References
278:subfamily
192:Aneuretini
172:Formicidae
142:Arthropoda
755:Q18102276
469:trapezoid
365:Ericaceae
217:Species:
128:Kingdom:
122:Eukaryota
749:Wikidata
710:83928415
608:81975987
569:90471180
474:pronotum
465:antennae
398:deformis
370:Taxodium
351:peat bog
314:included
294:Sakhalin
276:formicid
168:Family:
138:Phylum:
132:Animalia
118:Domain:
834:4539098
808:1007752
795:1320412
690:Bibcode
549:Bibcode
409:petiole
356:Osmunda
320:of the
296:in the
274:in the
265:extinct
242:†
221:†
198:Genus:
188:Tribe:
158:Order:
152:Insecta
148:Class:
847:255876
821:573890
764:AntWeb
708:
606:
567:
478:gaster
427:, and
263:is an
803:IRMNG
706:S2CID
604:S2CID
565:S2CID
514:(PDF)
382:Salix
376:Alnus
268:genus
816:ITIS
790:GBIF
522:1988
363:and
39:Preκ
782:W2K
777:CoL
698:doi
642:doi
596:doi
557:doi
379:,
359:,
272:ant
270:of
866::
844::
831::
818::
805::
792::
779::
766::
751::
704:.
696:.
686:10
684:.
667:11
665:.
636:.
632:.
616:^
602:.
592:46
590:.
586:.
563:.
555:.
545:50
543:.
516:.
491:^
421:,
373:,
300:.
89:Pg
712:.
700::
692::
650:.
644::
638:4
610:.
598::
571:.
559::
551::
202:β
94:N
84:K
79:J
74:T
69:P
64:C
59:D
54:S
49:O
44:κ
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