761:
6641:
394:
5408:
6776:
4210:
672:
772:
111:
132:
6800:
4216:
944:
690:, VF 1, consists of a very elongated cervical vertebra, probably the fifth. Today the middle section is missing; the original find was about 62 cm (24 in) long, but had been sawed into three parts. Most of the fossil consists of an internal infilling or mold; the thin bone walls are missing on most of the surface. The find had not presented the whole vertebra; a piece was absent from its posterior end as well.
3593:
488:
5424:
6648:
3598:
3052:
6788:
6654:
3060:
1309:
3067:
5416:
560:
they discovered some other pterosaur bones: a smaller vertebra and the proximal and distal extremities of a wing phalanx — but not the original find. However, after their departure to Europe engineer
Rashdie Sadaqah of the mine investigated further and in 1996 established it had been bought from the
729:
was more likely at 7 m (23 ft), but this estimate was not given a rationale. In 2010, Mark Witton and
Michael Habib argued that a 7 m (23 ft) wingspan is an underestimate, while a 11–13 m (36–43 ft) wingspan is an overestimate. In 2022, Gregory S. Paul proposed that
1938:
Rosenbach, K. L.; Goodvin, D. M.; Albshysh, M. G.; Azzam, H. A.; Smadi, A. A.; Mustafa, H. A.; Zalmout, I. S. A.; Wilson
Mantilla, J. A. (2024). "New pterosaur remains from the Late Cretaceous of Afro-Arabia provide insight into flight capacity of large pterosaurs". Journal of Vertebrate
629:
In 2018, topotype specimens were located in
Bavarian State Collection for Palaeontology and Geology in Munich, Germany that were placed there in 1966 from Jordan and probably represent additional elements of the holotype individual. These are identified as cf.
718:
However, the estimate proposed by Frey and
Martill was taken into question and the later estimates of wingspan have been more moderate, for the remains are too fragmentary to estimate a gigantic size. The researchers who described
699:. From this again the total neck length was extrapolated at about 3 m (9.8 ft). From the relatively slender vertebra the length dimension was then selected to be compared to that of
693:
Frey and
Martill estimated the total length to have been 78 cm (31 in), using for comparison the relative position of the smallest shaft diameter of the fifth cervical vertebra of
703:, estimated at 66 cm (26 in) long, resulting in a ratio of 1.18. Applying that ratio to the overall size, Frey and Martill in the late 1990s concluded that the wingspan of
6835:
471:: Philadelphia. Arambourg let a plaster cast be made and then sent the fossil back to the phosphate mine; this last aspect was later forgotten and the bone was assumed lost.
634:
and include the "fragments of two cervical vertebrae, a neural arch, a left femur, a ?radius, and a metacarpal IV" and other indeterminate fragments. The incomplete left
659:
from the
Ruseifa Phosphate Mines, near the Jordanian capital of Amman, where the holotype was recovered. The specimen is comparable in size and shape to the humerus of
436:
mine, Amin Kawar, who brought it to the attention of a
British archeologist, Fielding, after the war. This generated some publicity — the bone was even shown to
6830:
539:, honoring Arambourg. However, the name "Titanopteryx" was informally kept in use in the West, partially because the new name was assumed by many to be a
452:
6733:
2005:
Nessov, L.A., and Yarkov, A.A. (1989). "New
Cretaceous-Paleogene birds of the USSR and some remarks on the origin and evolution of the class Aves".
1542:
Rosenbach, K. L.; Goodvin, D. M.; Albshysh, M. G.; Azzam, H. A.; Smadi, A. A.; Mustafa, H. A.; Zalmout, I. S. A.; Wilson
Mantilla, J. A. (2024).
6825:
2053:
1806:"On the Size and Flight Diversity of Giant Pterosaurs, the Use of Birds as Pterosaur Analogues and Comments on Pterosaur Flightlessness"
6640:
1880:
1466:(Pterosauria, Azhdarchidae) from the Late Campanian micaceous facies of the Coon Creek Formation in McNairy County, Tennessee, USA"
1435:
557:
6855:
6850:
6845:
5407:
4215:
1634:"Topotype specimens probably attributable to the giant azhdarchid pterosaur Arambourgiania philadelphiae (Arambourg 1959)"
760:
6766:
4209:
1757:
Suberbiola, Xabier Pereda; Bardet, Nathalie; Jouve, Stéphane; Iarochène, Mohamed; Bouya, Baâdi; Amaghzaz, Mbarek (2003).
1488:"Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary"
131:
1544:"New pterosaur remains from the Late Cretaceous of Afro-Arabia provide insight into flight capacity of large pterosaurs"
1097:
1262:
6860:
6840:
3592:
2046:
1982:
Martill, D.M., E. Frey, R.M. Sadaqah & H.N. Khoury (1998). "Discovery of the holotype of the giant pterosaur
611:
2150:
1718:
Martill, D.M.; Frey, E.; Sadaqah, R.M.; Khoury, H.N. (1998). "Discovery of the holotype of the giant pterosaur
1430:
1303:
938:
884:
3051:
3628:
1189:
1073:
993:
464:
3597:
3059:
565:; it was still present in the collection of this institute and now could be restudied by Martill and Frey.
6673:
6548:
4706:
1325:
1121:
6647:
6746:
6653:
1218:
843:
393:
260:
5423:
6820:
6804:
6720:
4801:
3240:
3150:
3066:
2039:
1817:
1770:
1645:
1588:
607:
562:
528:
437:
5415:
6780:
6711:
6305:
5994:
5681:
4980:
4907:
2675:
2395:
960:
867:
553:
707:
had been 11–13 m (36–43 ft), compared with the 10–11 m (33–36 ft) wingspan of
5834:
4308:
4151:
3294:
3097:
3084:
2998:
2840:
2471:
2446:
1921:
1886:
1786:
1669:
1614:
1138:
779:
289:
126:
6751:
2016:
Steel, L., D.M. Martill., J. Kirk, A. Anders, R.F. Loveridge, E. Frey, and J.G. Martin (1997). "
771:
671:
514:
110:
6260:
5075:
5004:
4246:
4233:
4176:
3939:
3837:
3739:
3657:
3435:
3412:
3378:
3308:
2849:
2765:
2717:
2691:
2404:
2309:
1876:
1845:
1661:
1606:
1519:
1058:
1010:
475:
448:
280:
1462:
Harrell, T. Lynn Jr.; Gibson, Michael A.; Langston, Wann Jr. (2016). "A cervical vertebra of
6489:
6372:
6364:
6156:
6085:
6010:
5552:
5455:
5442:
5374:
5254:
5243:
4493:
4350:
4287:
4062:
4044:
3882:
3706:
3522:
3508:
3466:
2972:
2958:
2890:
2569:
2119:
1913:
1868:
1835:
1825:
1778:
1739:
1700:
1653:
1596:
1551:
1509:
1499:
916:
828:
801:. The cladogram is based on a topology recovered by Brian Andres and Timothy Myers in 2013.
596:
215:
189:
1633:
6578:
6421:
6270:
6055:
5959:
5934:
5927:
5914:
5847:
5733:
5631:
5597:
5493:
5473:
5327:
5185:
5126:
5113:
5084:
4960:
4871:
4730:
4603:
4558:
4430:
4359:
4338:
4164:
3825:
3805:
3777:
3753:
3689:
3681:
3674:
3666:
3421:
3401:
3231:
3207:
3157:
2965:
2948:
2938:
2915:
2871:
2863:
2831:
2820:
2799:
2753:
2745:
2684:
2600:
2540:
2459:
2428:
2318:
2297:
2250:
2135:
2110:
588:
468:
358:
599:. Both concepts have fallen into disuse now that such forms are commonly assigned to the
432:
found a two foot long fossil bone. In 1943 this was acquired by the director of a nearby
1821:
1774:
1649:
1592:
1408:
A 2024 description of its humerus compared it to that of soaring birds, suggesting that
556:
traveled to Jordan in an attempt to clarify matters. In a cupboard of the office of the
6555:
6528:
6506:
6330:
6244:
6229:
6181:
6120:
6105:
6071:
5896:
5880:
5868:
5854:
5807:
5764:
5724:
5638:
5584:
5312:
5305:
5298:
5177:
5098:
5025:
4936:
4692:
4584:
4519:
4460:
4400:
4366:
4086:
4013:
3985:
3904:
3854:
3793:
3762:
3362:
3338:
3316:
3262:
3220:
3164:
3132:
2813:
2730:
2707:
2586:
2512:
2480:
2333:
2282:
2230:
1840:
1805:
1514:
1487:
1440:
721:
695:
511:
480:
409:
6814:
6792:
6592:
6563:
6539:
6520:
6478:
6463:
6412:
6401:
6294:
6286:
6236:
6173:
6144:
6128:
6112:
6098:
6039:
6017:
5968:
5861:
5622:
5565:
5484:
5319:
5291:
5263:
5134:
5017:
4808:
4738:
4610:
4593:
4449:
4386:
4330:
4095:
4051:
3932:
3925:
3918:
3911:
3896:
3846:
3770:
3727:
3720:
3646:
3493:
3486:
3479:
3328:
3270:
3255:
3248:
3196:
2979:
2648:
2608:
2558:
2527:
2488:
2383:
2375:
2353:
2257:
2241:
1890:
1790:
1758:
1704:
1673:
1618:
809:
798:
740:
549:
503:
497:
403:
370:
362:
68:
1925:
1743:
487:
6585:
6513:
6384:
6279:
6213:
6165:
5975:
5903:
5781:
5749:
5517:
5284:
5212:
5153:
5048:
4685:
4670:
4654:
4645:
4634:
4474:
4437:
4323:
4273:
4116:
4029:
4022:
4002:
3974:
3967:
3953:
3946:
3861:
3818:
3785:
3501:
3443:
3386:
3371:
3351:
3176:
2907:
2899:
2879:
2806:
2785:
2778:
2633:
2421:
2367:
2326:
1203:
943:
600:
592:
584:
541:
523:
518:
382:
312:
202:
1782:
1601:
1572:
1556:
1543:
1830:
1504:
606:
In 2016, an azhdarchid cervical vertebra (MPPM 2000.23.1) was described from the
6705:
6471:
6431:
6350:
6314:
6048:
5823:
5756:
5699:
5645:
5502:
5170:
5041:
5033:
4993:
4952:
4944:
4819:
4786:
4754:
4746:
4717:
4678:
4662:
4621:
4550:
4542:
4535:
4481:
4422:
4414:
4075:
4037:
3960:
3615:
3548:
3515:
3451:
2792:
2661:
2616:
2593:
2268:
765:
43:
20:
6696:
1308:
6439:
6394:
6322:
6197:
6189:
6024:
6003:
5948:
5816:
5791:
5741:
5706:
5652:
5540:
5532:
5525:
5277:
5200:
5192:
5091:
5055:
4893:
4794:
4773:
4762:
4699:
4316:
4109:
4102:
3537:
3428:
2435:
2092:
1917:
1906:
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
1486:
Longrich, Nicholas R.; Martill, David M.; Andres, Brian; Penny, David (2018).
561:
company in 1969 by geologist Hani N. Khoury who had donated it in 1973 to the
456:
351:
88:
53:
1971:(Pterosauria, Pterodactyloidea): One of the world's largest flying animals."
1691:(Pterosauria, pterodactyloidea): one of the world's largest flying animals".
1665:
1610:
6499:
5941:
5889:
5611:
5604:
5572:
5510:
5236:
5229:
5221:
5141:
4919:
4885:
4835:
4828:
4565:
4503:
4407:
3995:
3394:
3111:
2738:
2578:
2413:
2290:
2212:
2199:
2162:
2101:
2062:
1414:
790:
775:
651:
532:
440:— but more importantly, it made the scientific community aware of the find.
433:
354:
176:
143:
93:
1849:
1759:"A new azhdarchid pterosaur from the Late Cretaceous phosphates of Morocco"
1523:
1872:
744:. In a 2024 study, based on a large humerus comparable in size to that of
6690:
6571:
6455:
6447:
6221:
6205:
6136:
5713:
5063:
4968:
4900:
4572:
4527:
4300:
3529:
3189:
2086:
687:
680:
429:
425:
163:
118:
83:
78:
63:
58:
48:
6738:
639:
374:
98:
73:
5162:
4929:
4511:
3119:
2986:
2080:
366:
153:
6667:
1657:
734:
had a wingspan of 8–9 m (26–30 ft), smaller than that of
484:, concluded the bone was not a metacarpal but a cervical vertebra.
4260:
2856:
1418:, whose condition is more similar to those of continous flappers.
770:
759:
670:
486:
444:
421:
392:
6725:
459:
of a giant pterosaur. In 1959, he named a new genus and species:
635:
6671:
5440:
4231:
3613:
3082:
2197:
2073:
2035:
385:, and it is also one of the largest flying animals ever known.
2031:
622:
to North America. This specimen was later referred to as cf.
535:
family in 1935. Therefore, in 1989 he renamed the genus into
1732:
Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen
1693:
Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen
16:
Genus of large azhdarchid pterosaur from the Late Cretaceous
420:
In the early 1940s, a railway worker during repairs on the
1939:
Paleontology. e2385068. doi:10.1080/02724634.2024.2385068.
764:
Holotype cervical vertebra (F) compared to those of other
1904:
Andres, B.; Myers, T. S. (2013). "Lone Star Pterosaurs".
580:
and affirmed its validity in relation to "Titanopteryx".
1967:
Frey, E. & Martill, D.M. (1996). "A reappraisal of
618:, which may potentially extend the geographic range of
1412:
was a soarer. This is in contrast to the contemporary
6764:
377:. With a wingspan of around 10 metres (33 ft),
6680:
6538:
6488:
6420:
6411:
6383:
6348:
6304:
6269:
6259:
6155:
6084:
6069:
6038:
5993:
5958:
5913:
5879:
5833:
5806:
5779:
5723:
5679:
5621:
5583:
5551:
5492:
5483:
5453:
5373:
5253:
5211:
5152:
5112:
5074:
5003:
4979:
4918:
4869:
4818:
4772:
4716:
4644:
4620:
4592:
4583:
4492:
4458:
4447:
4384:
4349:
4286:
4271:
4244:
4175:
4150:
4085:
4061:
4012:
3984:
3880:
3836:
3804:
3752:
3737:
3704:
3656:
3626:
3547:
3465:
3411:
3361:
3337:
3326:
3292:
3230:
3206:
3175:
3130:
3095:
2997:
2947:
2937:
2889:
2839:
2830:
2764:
2716:
2705:
2674:
2659:
2631:
2568:
2538:
2510:
2470:
2445:
2403:
2394:
2351:
2308:
2267:
2240:
2210:
2161:
2134:
1804:Witton, M.; Habib, M.B. (2010). Laudet, V. (ed.).
1687:Frey, E.; Martill, D.M. (1996). "A reappraisal of
463:. The genus name meant "titan wing" in Greek; the
381:was among the largest members of its family, the
1763:Geological Society, London, Special Publications
1638:Geological Society, London, Special Publications
121:fossil cast at Museum Histoire Naturelle, Paris
1577:Lawson 1975 (Pterodactyloidea: Azhdarchoidea)"
1481:
1479:
568:Frey and Martill rejected the suggestion that
455:. In 1954, he concluded the bone was the wing
2047:
1996:Neues Jahrbuch fur Geologie und Paläontologie
752:was estimated around 10 metres (33 ft).
8:
655:discovered a large partial right humerus of
583:Nesov in 1984 had placed the species within
6836:Late Cretaceous pterosaurs of North America
1867:. Princeton University Press. p. 161.
1537:
1535:
1533:
1458:
1456:
715:was thus the largest pterosaur then known.
6668:
6417:
6356:
6266:
6090:
6081:
6077:
6035:
6031:
5919:
5839:
5803:
5799:
5691:
5687:
5589:
5557:
5489:
5465:
5461:
5450:
5437:
5370:
5208:
5149:
5118:
5009:
4985:
4877:
4778:
4722:
4641:
4626:
4589:
4466:
4455:
4392:
4292:
4283:
4279:
4252:
4241:
4228:
4156:
4147:
4067:
3888:
3810:
3749:
3745:
3712:
3638:
3634:
3623:
3610:
3471:
3343:
3334:
3300:
3212:
3181:
3142:
3138:
3103:
3092:
3079:
2944:
2836:
2770:
2722:
2713:
2671:
2667:
2640:
2550:
2546:
2519:
2451:
2400:
2359:
2274:
2222:
2218:
2207:
2194:
2140:
2131:
2070:
2054:
2040:
2032:
109:
27:
1956:nov. gen., nov. sp. Ptérosaurien géant."
1839:
1829:
1632:Martill, David M.; Moser, Markus (2018).
1600:
1555:
1513:
1503:
6771:
2007:Trudy Zoologicheskogo Instituta AN SSSR
1865:The Princeton Field Guide to Pterosaurs
1452:
638:of the "Sidi Chennane azhdarchid" from
328:
304:
793:showing the phylogenetic placement of
1571:Andres, B.; Langston, W. Jr. (2021).
7:
453:Muséum national d'histoire naturelle
1722:Arambourg, 1959, and the status of
1278:
1254:
1234:
1210:
1181:
1113:
1089:
1065:
1053:
985:
930:
923:
859:
835:
823:
816:
804:
6831:Late Cretaceous pterosaurs of Asia
1986:Arambourg 1959, and the status of
1581:Journal of Vertebrate Paleontology
1548:Journal of Vertebrate Paleontology
14:
495:(far right) compared to a human,
6798:
6786:
6774:
6652:
6646:
6639:
5422:
5414:
5406:
4214:
4208:
3596:
3591:
3065:
3058:
3050:
1307:
942:
642:has also been identified as cf.
130:
2020:: giant wings in small halls."
467:refers to the name of Amman in
443:In 1953 the fossil was sent to
1436:Timeline of pterosaur research
782:of Morocco that may belong to
558:Jordan Phosphate Mines Company
1:
6826:Fossil taxa described in 1989
1783:10.1144/GSL.SP.2003.217.01.08
1602:10.1080/02724634.2021.1907587
1557:10.1080/02724634.2024.2385068
1468:Bull. Alabama Mus. Nat. Hist.
416:(D), with known parts in gray
2018:Arambourgiania philadelphiae
1831:10.1371/journal.pone.0013982
1573:"Morphology and taxonomy of
1505:10.1371/journal.pbio.2001663
1464:Arambourgiania philadelphiae
1287:Arambourgiania philadelphiae
1098:Shenzhoupterus chaoyangensis
725:stated that the wingspan of
616:Arambourgiania philadelphiae
548:Early 1995, paleontologists
397:Skeletal reconstructions of
273:Arambourgiania philadelphiae
1263:Zhejiangopterus linhaiensis
649:In 2024, the describers of
447:, where it was examined by
6877:
1984:Titanopteryx philadelphiae
1954:Titanopteryx philadelphiae
1720:Titanopteryx philadelphiae
1705:10.1127/njgpa/199/1996/221
527:had already been given by
461:Titanopteryx philadelphiae
331:Titanopteryx philadelphiae
18:
6636:
6359:
6093:
6080:
6034:
5922:
5842:
5802:
5694:
5690:
5592:
5560:
5468:
5464:
5449:
5436:
5403:
5369:
5121:
5012:
4988:
4880:
4781:
4725:
4629:
4469:
4395:
4295:
4282:
4255:
4240:
4227:
4205:
4159:
4146:
4070:
3891:
3813:
3748:
3715:
3641:
3637:
3622:
3609:
3589:
3474:
3346:
3303:
3215:
3184:
3145:
3141:
3106:
3091:
3078:
3047:
2773:
2725:
2670:
2643:
2553:
2549:
2522:
2454:
2362:
2277:
2225:
2221:
2206:
2193:
2143:
2130:
2069:
1918:10.1017/S1755691013000303
1863:Paul, Gregory S. (2022).
1744:10.1127/njgpa/207/1998/57
1322:
1300:
1283:
1276:
1259:
1252:
1239:
1232:
1215:
1208:
1186:
1179:
1135:
1118:
1111:
1094:
1087:
1070:
1063:
1051:
1007:
990:
983:
957:
935:
928:
921:
881:
864:
857:
840:
833:
821:
814:
612:McNairy County, Tennessee
510:In the eighties, Russian
295:
288:
266:
259:
239:Nessov & Yarkov, 1989
127:Scientific classification
125:
117:
108:
30:
1431:List of pterosaur genera
1304:Quetzalcoatlus northropi
939:Domeykodactylus ceciliae
885:Tupuxuara longicristatus
746:Quetzalcoatlus northropi
661:Quetzalcoatlus northropi
587:, back then part of the
19:Not to be confused with
3629:Archaeopterodactyloidea
1958:Notes Mém. Moyen-Orient
1952:Arambourg, C. (1959). "
1190:Radiodactylus langstoni
1074:Eoazhdarcho liaoxiensis
994:Noripterus complicidens
478:, studying the related
350:is an extinct genus of
2022:The Geological Curator
1973:N.Jb.Geol.Paläont.Abh.
1326:Quetzalcoatlus lawsoni
1122:Chaoyangopterus zhangi
786:
768:
683:
679:fighting over a small
507:
417:
6747:Paleobiology Database
1873:10.1515/9780691232218
1219:Azhdarcho lancicollis
844:Thalassodromeus sethi
774:
763:
674:
490:
396:
252:A. philadelphiae
6856:Cretaceous Tennessee
6851:Fossils of Tennessee
6846:Maastrichtian genera
3241:Archaeoistiodactylus
3151:Archaeoistiodactylus
608:Coon Creek Formation
563:University of Jordan
438:Abdullah I of Jordan
389:History of discovery
6306:Targaryendraconidae
5995:Ornithocheiriformes
5682:Ornithocheiromorpha
4981:Neopterodactyloidea
4908:Dsungaripteromorpha
2676:Campylognathoididae
2396:Eudimorphodontoidea
1822:2010PLoSO...513982W
1775:2003GSLSP.217...79S
1650:2018GSLSP.455..159M
1593:2021JVPal..41S..46A
961:Dsungaripterus weii
868:Tupuxuara leonardii
675:Restoration of two
517:was informed by an
369:, and possibly the
5835:Istiodactyliformes
4309:Lonchognathosaurus
4152:Eupterodactyloidea
3295:Pterodactyliformes
3098:Pterodactylomorpha
3085:Pterodactylomorpha
2999:Pterodactylomorpha
2841:Angustinaripterini
2472:Eudimorphodontinae
2447:Eudimorphodontidae
1139:Jidapterus edentus
787:
780:Ouled Abdoun Basin
769:
748:, the wingspan of
684:
531:to a fly from the
508:
418:
6861:Cretaceous Jordan
6841:Fossils of Jordan
6762:
6761:
6674:Taxon identifiers
6665:
6664:
6661:
6660:
6634:
6633:
6630:
6629:
6626:
6625:
6622:
6621:
6618:
6617:
6614:
6613:
6610:
6609:
6606:
6605:
6602:
6601:
6344:
6343:
6340:
6339:
6261:Targaryendraconia
6255:
6254:
6065:
6064:
5989:
5988:
5985:
5984:
5775:
5774:
5675:
5674:
5671:
5670:
5667:
5666:
5663:
5662:
5432:
5431:
5401:
5400:
5397:
5396:
5393:
5392:
5365:
5364:
5361:
5360:
5357:
5356:
5353:
5352:
5349:
5348:
5345:
5344:
5341:
5340:
5337:
5336:
5108:
5107:
5076:Chaoyangopterinae
5005:Chaoyangopteridae
4865:
4864:
4861:
4860:
4857:
4856:
4853:
4852:
4849:
4848:
4845:
4844:
4380:
4379:
4376:
4375:
4247:Ornithocheiroidea
4234:Ornithocheiroidea
4223:
4222:
4203:
4202:
4199:
4198:
4195:
4194:
4177:Ornithocheiroidea
4142:
4141:
4138:
4137:
4134:
4133:
4130:
4129:
4126:
4125:
3940:Gladocephaloideus
3876:
3875:
3872:
3871:
3838:Aurorazhdarchidae
3740:Ctenochasmatoidea
3700:
3699:
3658:Germanodactylidae
3605:
3604:
3587:
3586:
3583:
3582:
3579:
3578:
3575:
3574:
3571:
3570:
3567:
3566:
3461:
3460:
3436:Dendrorhynchoides
3413:Batrachognathinae
3379:Dendrorhynchoides
3309:Changchengopterus
3288:
3287:
3284:
3283:
3280:
3279:
3074:
3073:
3045:
3044:
3041:
3040:
3037:
3036:
3033:
3032:
3029:
3028:
3025:
3024:
3021:
3020:
3017:
3016:
2933:
2932:
2929:
2928:
2925:
2924:
2850:Angustinaripterus
2766:Rhamphorhynchinae
2718:Rhamphorhynchidae
2701:
2700:
2692:Campylognathoides
2627:
2626:
2506:
2505:
2502:
2501:
2498:
2497:
2405:Raeticodactylidae
2347:
2346:
2343:
2342:
2310:Austriadraconidae
2189:
2188:
2185:
2184:
2181:
2180:
1400:
1399:
1391:
1390:
1382:
1381:
1373:
1372:
1364:
1363:
1355:
1354:
1346:
1345:
1337:
1336:
1168:
1167:
1159:
1158:
1150:
1149:
1059:Chaoyangopteridae
1040:
1039:
1031:
1030:
1022:
1021:
1011:Noripterus parvus
972:
971:
905:
904:
896:
895:
797:within the clade
595:placed it within
529:Günther Enderlein
476:Douglas A. Lawson
449:Camille Arambourg
343:
342:
336:
325:
316:
311:Arambourg, 1959 (
301:
240:
6868:
6803:
6802:
6801:
6791:
6790:
6789:
6779:
6778:
6777:
6770:
6755:
6754:
6742:
6741:
6729:
6728:
6716:
6715:
6714:
6701:
6700:
6699:
6669:
6656:
6650:
6644:
6643:
6490:Coloborhynchinae
6418:
6373:Ornithocheiridae
6365:Brasileodactylus
6357:
6267:
6157:Ornithocheirinae
6091:
6086:Ornithocheiridae
6082:
6078:
6036:
6032:
6011:Brasileodactylus
5920:
5840:
5808:Lonchodraconidae
5804:
5800:
5692:
5688:
5590:
5558:
5553:Nyctosauromorpha
5490:
5466:
5462:
5456:Pteranodontoidea
5451:
5443:Pteranodontoidea
5438:
5427:
5426:
5419:
5418:
5411:
5410:
5375:Pteranodontoidea
5371:
5255:Quetzalcoatlinae
5244:Mistralazhdarcho
5209:
5150:
5119:
5010:
4986:
4878:
4779:
4723:
4642:
4627:
4590:
4494:Thalassodromidae
4467:
4456:
4393:
4351:Dsungaripterinae
4293:
4288:Dsungaripteridae
4284:
4280:
4253:
4242:
4229:
4218:
4213:
4212:
4157:
4148:
4068:
4063:Ctenochasmatinae
4045:Plataleorhynchus
3889:
3883:Ctenochasmatidae
3811:
3750:
3746:
3713:
3707:Euctenochasmatia
3639:
3635:
3624:
3611:
3600:
3595:
3523:Propterodactylus
3509:Ningchengopterus
3472:
3467:Pterodactyloidea
3344:
3335:
3301:
3213:
3182:
3143:
3139:
3104:
3093:
3080:
3069:
3063:
3062:
3055:
3054:
2973:Jianchangopterus
2959:Fenghuangopterus
2945:
2891:Rhamphorhynchini
2837:
2771:
2723:
2714:
2672:
2668:
2641:
2570:Dimorphodontidae
2551:
2547:
2520:
2452:
2401:
2360:
2275:
2223:
2219:
2208:
2195:
2141:
2132:
2125:
2124:
2120:Pterosauromorpha
2071:
2056:
2049:
2042:
2033:
1940:
1936:
1930:
1929:
1901:
1895:
1894:
1860:
1854:
1853:
1843:
1833:
1801:
1795:
1794:
1754:
1748:
1747:
1715:
1709:
1708:
1684:
1678:
1677:
1629:
1623:
1622:
1604:
1568:
1562:
1561:
1559:
1539:
1528:
1527:
1517:
1507:
1483:
1474:
1460:
1311:
1279:
1255:
1235:
1211:
1182:
1114:
1090:
1066:
1054:
986:
946:
931:
924:
917:Dsungaripteridae
860:
836:
829:Thalassodrominae
824:
817:
805:
657:A. philadelphiae
614:and referred to
597:Titanopterygidae
591:; the same year
576:or identical to
334:
324:Species synonymy
323:
310:
299:
275:
271:
234:
227:
216:Quetzalcoatlinae
214:
201:
190:Pterodactyloidea
188:
175:
135:
134:
113:
103:
40:
36:Temporal range:
28:
6876:
6875:
6871:
6870:
6869:
6867:
6866:
6865:
6811:
6810:
6809:
6799:
6797:
6787:
6785:
6775:
6773:
6765:
6763:
6758:
6750:
6745:
6737:
6732:
6724:
6719:
6710:
6709:
6704:
6695:
6694:
6689:
6676:
6666:
6657:
6638:
6598:
6579:Liaoningopterus
6534:
6484:
6422:Tropeognathinae
6407:
6379:
6353:
6336:
6300:
6271:Cimoliopteridae
6251:
6151:
6074:
6061:
6056:Zhenyuanopterus
6030:
5981:
5960:Istiodactylinae
5954:
5935:Longchengpterus
5928:Lingyuanopterus
5915:Istiodactylidae
5909:
5875:
5848:Hongshanopterus
5829:
5798:
5784:
5771:
5734:Hongshanopterus
5719:
5684:
5659:
5632:Barbaridactylus
5617:
5579:
5547:
5494:Pteranodontidae
5479:
5474:Santanadactylus
5458:
5445:
5428:
5421:
5413:
5405:
5389:
5333:
5328:Zhejiangopterus
5249:
5207:
5186:Palaeocursornis
5148:
5127:Montanazhdarcho
5114:Azhdarchiformes
5104:
5085:Chaoyangopterus
5070:
4999:
4975:
4961:Montanazhdarcho
4914:
4874:
4872:Azhdarchomorpha
4841:
4814:
4768:
4731:Aymberedactylus
4712:
4640:
4616:
4604:Aymberedactylus
4579:
4559:Thalassodromeus
4488:
4463:
4452:
4443:
4431:Montanazhdarcho
4389:
4372:
4360:Domeykodactylus
4345:
4339:Tendaguripterus
4276:
4267:
4249:
4236:
4219:
4207:
4191:
4171:
4165:Altmuehlopterus
4122:
4081:
4057:
4008:
3980:
3885:
3868:
3832:
3826:Gallodactylidae
3806:Aurorazhdarchia
3800:
3778:Normannognathus
3754:Gallodactylidae
3742:
3733:
3709:
3696:
3690:Tendaguripterus
3682:Normannognathus
3675:Germanodactylus
3667:Altmuehlopterus
3652:
3631:
3618:
3601:
3563:
3543:
3457:
3422:Batrachognathus
3407:
3402:Vesperopterylus
3357:
3331:
3322:
3297:
3276:
3232:Wukongopterinae
3226:
3208:Wukongopteridae
3202:
3171:
3158:Normannognathus
3135:
3126:
3100:
3087:
3070:
3057:
3049:
3013:
2993:
2966:Jianchangnathus
2949:Scaphognathidae
2939:Digibrevisauria
2921:
2916:Rhamphorhynchus
2885:
2872:Qinglongopterus
2864:Harpactognathus
2832:Rhamphorhynchae
2826:
2821:Qinglongopterus
2800:Harpactognathus
2760:
2754:Scaphognathinae
2746:Parapsicephalus
2710:
2697:
2685:Bergamodactylus
2664:
2655:
2637:
2623:
2601:Parapsicephalus
2564:
2543:
2541:Macronychoptera
2534:
2516:
2494:
2466:
2460:Arcticodactylus
2441:
2429:Raeticodactylus
2390:
2356:
2339:
2319:Arcticodactylus
2304:
2298:Raeticodactylus
2263:
2251:Austriadactylus
2236:
2215:
2202:
2177:
2157:
2151:Avemetatarsalia
2136:Avemetatarsalia
2126:
2111:Avemetatarsalia
2076:
2075:
2065:
2060:
1949:
1944:
1943:
1937:
1933:
1903:
1902:
1898:
1883:
1862:
1861:
1857:
1803:
1802:
1798:
1756:
1755:
1751:
1717:
1716:
1712:
1686:
1685:
1681:
1658:10.1144/SP455.6
1631:
1630:
1626:
1570:
1569:
1565:
1541:
1540:
1531:
1498:(3): e2001663.
1485:
1484:
1477:
1461:
1454:
1449:
1427:
1421:
1406:
1401:
1392:
1383:
1374:
1365:
1356:
1347:
1338:
1169:
1160:
1151:
1041:
1032:
1023:
973:
906:
897:
758:
669:
589:Pteranodontidae
391:
359:Late Cretaceous
339:
335:Arambourg, 1959
333:
327:
326:
319:
309:
303:
302:
284:
277:
269:
268:
255:
233:
225:
212:
199:
186:
173:
129:
104:
102:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
38:
37:
34:
24:
17:
12:
11:
5:
6874:
6872:
6864:
6863:
6858:
6853:
6848:
6843:
6838:
6833:
6828:
6823:
6813:
6812:
6808:
6807:
6795:
6783:
6760:
6759:
6757:
6756:
6743:
6730:
6717:
6712:Arambourgiania
6702:
6686:
6684:
6682:Arambourgiania
6678:
6677:
6672:
6663:
6662:
6659:
6658:
6637:
6635:
6632:
6631:
6628:
6627:
6624:
6623:
6620:
6619:
6616:
6615:
6612:
6611:
6608:
6607:
6604:
6603:
6600:
6599:
6597:
6596:
6589:
6582:
6575:
6568:
6560:
6556:Caulkicephalus
6552:
6544:
6542:
6536:
6535:
6533:
6532:
6529:Uktenadactylus
6525:
6517:
6510:
6507:Coloborhynchus
6503:
6495:
6493:
6486:
6485:
6483:
6482:
6475:
6468:
6460:
6452:
6444:
6436:
6427:
6425:
6415:
6409:
6408:
6406:
6405:
6398:
6390:
6388:
6381:
6380:
6378:
6377:
6369:
6360:
6354:
6349:
6346:
6345:
6342:
6341:
6338:
6337:
6335:
6334:
6331:Targaryendraco
6327:
6319:
6310:
6308:
6302:
6301:
6299:
6298:
6291:
6283:
6275:
6273:
6264:
6257:
6256:
6253:
6252:
6250:
6249:
6245:Uktenadactylus
6241:
6233:
6230:Ornithocheirus
6226:
6218:
6210:
6202:
6194:
6186:
6182:Coloborhynchus
6178:
6170:
6161:
6159:
6153:
6152:
6150:
6149:
6141:
6133:
6125:
6121:Caulkicephalus
6117:
6109:
6106:Arthurdactylus
6102:
6094:
6088:
6075:
6072:Ornithocheirae
6070:
6067:
6066:
6063:
6062:
6060:
6059:
6052:
6044:
6042:
6029:
6028:
6021:
6014:
6007:
5999:
5997:
5991:
5990:
5987:
5986:
5983:
5982:
5980:
5979:
5972:
5964:
5962:
5956:
5955:
5953:
5952:
5945:
5938:
5931:
5923:
5917:
5911:
5910:
5908:
5907:
5900:
5897:Linlongopterus
5893:
5885:
5883:
5881:Mimodactylidae
5877:
5876:
5874:
5873:
5869:Lonchodectidae
5865:
5858:
5855:Linlongopterus
5851:
5843:
5837:
5831:
5830:
5828:
5827:
5820:
5812:
5810:
5797:
5796:
5787:
5785:
5780:
5777:
5776:
5773:
5772:
5770:
5769:
5765:Targaryendraco
5761:
5753:
5746:
5738:
5729:
5727:
5725:Lonchodectidae
5721:
5720:
5718:
5717:
5710:
5703:
5695:
5685:
5680:
5677:
5676:
5673:
5672:
5669:
5668:
5665:
5664:
5661:
5660:
5658:
5657:
5649:
5642:
5639:Muzquizopteryx
5635:
5627:
5625:
5619:
5618:
5616:
5615:
5608:
5601:
5593:
5587:
5585:Aponyctosauria
5581:
5580:
5578:
5577:
5569:
5561:
5555:
5549:
5548:
5546:
5545:
5537:
5529:
5522:
5514:
5507:
5498:
5496:
5487:
5481:
5480:
5478:
5477:
5469:
5459:
5454:
5447:
5446:
5441:
5434:
5433:
5430:
5429:
5404:
5402:
5399:
5398:
5395:
5394:
5391:
5390:
5388:
5387:
5386:
5385:
5379:
5377:
5367:
5366:
5363:
5362:
5359:
5358:
5355:
5354:
5351:
5350:
5347:
5346:
5343:
5342:
5339:
5338:
5335:
5334:
5332:
5331:
5324:
5316:
5313:Thanatosdrakon
5309:
5306:Quetzalcoatlus
5302:
5299:Phosphatodraco
5295:
5288:
5281:
5274:
5271:Arambourgiania
5267:
5259:
5257:
5251:
5250:
5248:
5247:
5240:
5233:
5226:
5217:
5215:
5206:
5205:
5197:
5189:
5182:
5178:Navajodactylus
5174:
5167:
5158:
5156:
5147:
5146:
5138:
5131:
5122:
5116:
5110:
5109:
5106:
5105:
5103:
5102:
5099:Shenzhoupterus
5095:
5088:
5080:
5078:
5072:
5071:
5069:
5068:
5060:
5052:
5045:
5038:
5030:
5026:Argentinadraco
5022:
5013:
5007:
5001:
5000:
4998:
4997:
4989:
4983:
4977:
4976:
4974:
4973:
4965:
4957:
4949:
4941:
4937:Argentinadraco
4933:
4925:
4923:
4916:
4915:
4913:
4912:
4904:
4897:
4890:
4881:
4875:
4870:
4867:
4866:
4863:
4862:
4859:
4858:
4855:
4854:
4851:
4850:
4847:
4846:
4843:
4842:
4840:
4839:
4832:
4824:
4822:
4816:
4815:
4813:
4812:
4805:
4798:
4791:
4782:
4776:
4770:
4769:
4767:
4766:
4759:
4751:
4743:
4735:
4726:
4720:
4714:
4713:
4711:
4710:
4703:
4696:
4693:Nemicolopterus
4689:
4682:
4675:
4667:
4659:
4650:
4648:
4639:
4638:
4630:
4624:
4618:
4617:
4615:
4614:
4607:
4599:
4597:
4587:
4585:Tapejariformes
4581:
4580:
4578:
4577:
4569:
4562:
4555:
4547:
4539:
4532:
4524:
4520:Argentinadraco
4516:
4508:
4499:
4497:
4490:
4489:
4487:
4486:
4478:
4470:
4464:
4461:Tapejaromorpha
4459:
4453:
4448:
4445:
4444:
4442:
4441:
4434:
4427:
4419:
4411:
4404:
4401:Argentinadraco
4396:
4390:
4385:
4382:
4381:
4378:
4377:
4374:
4373:
4371:
4370:
4367:Dsungaripterus
4363:
4355:
4353:
4347:
4346:
4344:
4343:
4335:
4327:
4320:
4313:
4305:
4296:
4290:
4277:
4272:
4269:
4268:
4266:
4265:
4256:
4250:
4245:
4238:
4237:
4232:
4225:
4224:
4221:
4220:
4206:
4204:
4201:
4200:
4197:
4196:
4193:
4192:
4190:
4189:
4188:
4187:
4181:
4179:
4173:
4172:
4170:
4169:
4160:
4154:
4144:
4143:
4140:
4139:
4136:
4135:
4132:
4131:
4128:
4127:
4124:
4123:
4121:
4120:
4113:
4106:
4099:
4091:
4089:
4087:Pterodaustrini
4083:
4082:
4080:
4079:
4071:
4065:
4059:
4058:
4056:
4055:
4048:
4041:
4034:
4026:
4018:
4016:
4014:Gnathosaurinae
4010:
4009:
4007:
4006:
3999:
3991:
3989:
3986:Moganopterinae
3982:
3981:
3979:
3978:
3971:
3964:
3957:
3950:
3943:
3936:
3929:
3922:
3915:
3908:
3905:Balaenognathus
3901:
3892:
3886:
3881:
3878:
3877:
3874:
3873:
3870:
3869:
3867:
3866:
3858:
3855:Aurorazhdarcho
3851:
3842:
3840:
3834:
3833:
3831:
3830:
3822:
3814:
3808:
3802:
3801:
3799:
3798:
3794:Moganopterinae
3790:
3782:
3774:
3767:
3763:Aurorazhdarcho
3758:
3756:
3743:
3738:
3735:
3734:
3732:
3731:
3724:
3716:
3710:
3705:
3702:
3701:
3698:
3697:
3695:
3694:
3686:
3678:
3671:
3662:
3660:
3654:
3653:
3651:
3650:
3642:
3632:
3627:
3620:
3619:
3614:
3607:
3606:
3603:
3602:
3590:
3588:
3585:
3584:
3581:
3580:
3577:
3576:
3573:
3572:
3569:
3568:
3565:
3564:
3562:
3561:
3560:
3559:
3553:
3551:
3545:
3544:
3542:
3541:
3534:
3526:
3519:
3512:
3505:
3498:
3490:
3483:
3475:
3469:
3463:
3462:
3459:
3458:
3456:
3455:
3448:
3440:
3432:
3425:
3417:
3415:
3409:
3408:
3406:
3405:
3398:
3391:
3383:
3375:
3367:
3365:
3363:Anurognathinae
3359:
3358:
3356:
3355:
3347:
3341:
3339:Anurognathidae
3332:
3327:
3324:
3323:
3321:
3320:
3317:Douzhanopterus
3313:
3304:
3298:
3293:
3290:
3289:
3286:
3285:
3282:
3281:
3278:
3277:
3275:
3274:
3267:
3263:Douzhanopterus
3259:
3252:
3245:
3236:
3234:
3228:
3227:
3225:
3224:
3221:Kunpengopterus
3216:
3210:
3204:
3203:
3201:
3200:
3193:
3185:
3179:
3173:
3172:
3170:
3169:
3165:Anurognathidae
3161:
3154:
3146:
3136:
3133:Monofenestrata
3131:
3128:
3127:
3125:
3124:
3116:
3107:
3101:
3096:
3089:
3088:
3083:
3076:
3075:
3072:
3071:
3048:
3046:
3043:
3042:
3039:
3038:
3035:
3034:
3031:
3030:
3027:
3026:
3023:
3022:
3019:
3018:
3015:
3014:
3012:
3011:
3010:
3009:
3003:
3001:
2995:
2994:
2992:
2991:
2983:
2976:
2969:
2962:
2954:
2952:
2942:
2935:
2934:
2931:
2930:
2927:
2926:
2923:
2922:
2920:
2919:
2912:
2904:
2895:
2893:
2887:
2886:
2884:
2883:
2876:
2868:
2860:
2853:
2845:
2843:
2834:
2828:
2827:
2825:
2824:
2817:
2814:Orientognathus
2810:
2803:
2796:
2789:
2782:
2774:
2768:
2762:
2761:
2759:
2758:
2750:
2742:
2735:
2731:Dolicorhamphus
2726:
2720:
2711:
2708:Breviquartossa
2706:
2703:
2702:
2699:
2698:
2696:
2695:
2688:
2680:
2678:
2665:
2660:
2657:
2656:
2654:
2653:
2644:
2638:
2632:
2629:
2628:
2625:
2624:
2622:
2621:
2613:
2605:
2597:
2590:
2587:Caelestiventus
2583:
2574:
2572:
2566:
2565:
2563:
2562:
2554:
2544:
2539:
2536:
2535:
2533:
2532:
2523:
2517:
2513:Zambellisauria
2511:
2508:
2507:
2504:
2503:
2500:
2499:
2496:
2495:
2493:
2492:
2485:
2481:Carniadactylus
2476:
2474:
2468:
2467:
2465:
2464:
2455:
2449:
2443:
2442:
2440:
2439:
2432:
2425:
2418:
2409:
2407:
2398:
2392:
2391:
2389:
2388:
2380:
2372:
2363:
2357:
2352:
2349:
2348:
2345:
2344:
2341:
2340:
2338:
2337:
2334:Seazzadactylus
2330:
2323:
2314:
2312:
2306:
2305:
2303:
2302:
2294:
2287:
2283:Carniadactylus
2278:
2272:
2265:
2264:
2262:
2261:
2254:
2246:
2244:
2238:
2237:
2235:
2234:
2231:Daohugoupterus
2226:
2216:
2211:
2204:
2203:
2198:
2191:
2190:
2187:
2186:
2183:
2182:
2179:
2178:
2176:
2175:
2174:
2173:
2167:
2165:
2159:
2158:
2156:
2155:
2154:
2153:
2144:
2138:
2128:
2127:
2123:
2122:
2113:
2104:
2095:
2089:
2083:
2074:
2067:
2066:
2061:
2059:
2058:
2051:
2044:
2036:
2030:
2029:
2014:
2003:
1992:Quetzalcoatlus
1988:Arambourgiania
1980:
1969:Arambourgiania
1965:
1948:
1945:
1942:
1941:
1931:
1896:
1881:
1855:
1816:(11): e13982.
1796:
1749:
1728:Quetzalcoatlus
1724:Arambourgiania
1710:
1699:(2): 221–247.
1689:Arambourgiania
1679:
1644:(1): 159–169.
1624:
1575:Quetzalcoatlus
1563:
1529:
1475:
1451:
1450:
1448:
1445:
1444:
1443:
1441:Pterosaur size
1438:
1433:
1426:
1423:
1410:Arambourgiania
1405:
1402:
1398:
1397:
1394:
1393:
1389:
1388:
1385:
1384:
1380:
1379:
1376:
1375:
1371:
1370:
1367:
1366:
1362:
1361:
1358:
1357:
1353:
1352:
1349:
1348:
1344:
1343:
1340:
1339:
1335:
1334:
1331:
1330:
1321:
1318:
1317:
1314:
1313:
1299:
1296:
1295:
1292:
1291:
1282:
1277:
1275:
1272:
1271:
1268:
1267:
1258:
1253:
1251:
1248:
1247:
1244:
1243:
1238:
1233:
1231:
1228:
1227:
1224:
1223:
1214:
1209:
1207:
1199:
1198:
1195:
1194:
1185:
1180:
1178:
1175:
1174:
1171:
1170:
1166:
1165:
1162:
1161:
1157:
1156:
1153:
1152:
1148:
1147:
1144:
1143:
1134:
1131:
1130:
1127:
1126:
1117:
1112:
1110:
1107:
1106:
1103:
1102:
1093:
1088:
1086:
1083:
1082:
1079:
1078:
1069:
1064:
1062:
1052:
1050:
1047:
1046:
1043:
1042:
1038:
1037:
1034:
1033:
1029:
1028:
1025:
1024:
1020:
1019:
1016:
1015:
1006:
1003:
1002:
999:
998:
989:
984:
982:
979:
978:
975:
974:
970:
969:
966:
965:
956:
953:
952:
949:
948:
934:
929:
927:
922:
920:
912:
911:
908:
907:
903:
902:
899:
898:
894:
893:
890:
889:
880:
877:
876:
873:
872:
863:
858:
856:
853:
852:
849:
848:
839:
834:
832:
822:
820:
815:
813:
803:
795:Arambourgiania
784:Arambourgiania
757:
756:Classification
754:
750:Arambourgiania
736:Quetzalcoatlus
732:Arambourgiania
727:Arambourgiania
722:Phosphatodraco
713:Arambourgiania
709:Quetzalcoatlus
705:Arambourgiania
701:Quetzalcoatlus
696:Quetzalcoatlus
677:Arambourgiania
668:
665:
644:Arambourgiania
632:Arambourgiania
624:Arambourgiania
620:Arambourgiania
578:Quetzalcoatlus
570:Arambourgiania
537:Arambourgiania
521:that the name
512:paleontologist
493:Arambourgiania
481:Quetzalcoatlus
428:railroad near
410:Quetzalcoatlus
399:Arambourgiania
390:
387:
379:Arambourgiania
347:Arambourgiania
341:
340:
338:
337:
322:
321:
320:
318:
317:
300:Genus synonymy
298:
297:
296:
293:
292:
286:
285:
278:
264:
263:
257:
256:
248:
246:
242:
241:
230:Arambourgiania
223:
219:
218:
210:
206:
205:
197:
193:
192:
184:
180:
179:
171:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
123:
122:
115:
114:
106:
105:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
35:
32:Arambourgiania
15:
13:
10:
9:
6:
4:
3:
2:
6873:
6862:
6859:
6857:
6854:
6852:
6849:
6847:
6844:
6842:
6839:
6837:
6834:
6832:
6829:
6827:
6824:
6822:
6819:
6818:
6816:
6806:
6805:United States
6796:
6794:
6784:
6782:
6772:
6768:
6753:
6748:
6744:
6740:
6735:
6731:
6727:
6722:
6718:
6713:
6707:
6703:
6698:
6692:
6688:
6687:
6685:
6683:
6679:
6675:
6670:
6655:
6651:
6649:
6642:
6595:
6594:
6593:Maaradactylus
6590:
6588:
6587:
6583:
6581:
6580:
6576:
6574:
6573:
6569:
6566:
6565:
6564:Cearadactylus
6561:
6558:
6557:
6553:
6551:
6550:
6546:
6545:
6543:
6541:
6540:Anhanguerinae
6537:
6531:
6530:
6526:
6523:
6522:
6521:Siroccopteryx
6518:
6516:
6515:
6511:
6509:
6508:
6504:
6502:
6501:
6497:
6496:
6494:
6491:
6487:
6481:
6480:
6479:Tropeognathus
6476:
6474:
6473:
6469:
6466:
6465:
6464:Siroccopteryx
6461:
6458:
6457:
6453:
6450:
6449:
6445:
6442:
6441:
6437:
6434:
6433:
6429:
6428:
6426:
6423:
6419:
6416:
6414:
6413:Anhangueridae
6410:
6404:
6403:
6402:Iberodactylus
6399:
6397:
6396:
6392:
6391:
6389:
6386:
6382:
6375:
6374:
6370:
6367:
6366:
6362:
6361:
6358:
6355:
6352:
6347:
6333:
6332:
6328:
6325:
6324:
6320:
6317:
6316:
6312:
6311:
6309:
6307:
6303:
6297:
6296:
6295:Cimoliopterus
6292:
6289:
6288:
6287:Camposipterus
6284:
6282:
6281:
6277:
6276:
6274:
6272:
6268:
6265:
6262:
6258:
6247:
6246:
6242:
6239:
6238:
6237:Siroccopteryx
6234:
6232:
6231:
6227:
6224:
6223:
6219:
6216:
6215:
6211:
6208:
6207:
6203:
6200:
6199:
6195:
6192:
6191:
6187:
6184:
6183:
6179:
6176:
6175:
6174:Camposipterus
6171:
6168:
6167:
6163:
6162:
6160:
6158:
6154:
6147:
6146:
6145:Tropeognathus
6142:
6139:
6138:
6134:
6131:
6130:
6129:Cimoliopterus
6126:
6123:
6122:
6118:
6115:
6114:
6113:Camposipterus
6110:
6108:
6107:
6103:
6101:
6100:
6099:Araripesaurus
6096:
6095:
6092:
6089:
6087:
6083:
6079:
6076:
6073:
6068:
6058:
6057:
6053:
6051:
6050:
6046:
6045:
6043:
6041:
6040:Boreopteridae
6037:
6033:
6027:
6026:
6022:
6020:
6019:
6018:Cearadactylus
6015:
6013:
6012:
6008:
6006:
6005:
6001:
6000:
5998:
5996:
5992:
5978:
5977:
5973:
5971:
5970:
5969:Istiodactylus
5966:
5965:
5963:
5961:
5957:
5951:
5950:
5946:
5944:
5943:
5939:
5937:
5936:
5932:
5930:
5929:
5925:
5924:
5921:
5918:
5916:
5912:
5906:
5905:
5901:
5899:
5898:
5894:
5892:
5891:
5887:
5886:
5884:
5882:
5878:
5871:
5870:
5866:
5864:
5863:
5862:Yixianopterus
5859:
5857:
5856:
5852:
5850:
5849:
5845:
5844:
5841:
5838:
5836:
5832:
5826:
5825:
5821:
5819:
5818:
5814:
5813:
5811:
5809:
5805:
5801:
5794:
5793:
5789:
5788:
5786:
5783:
5778:
5767:
5766:
5762:
5759:
5758:
5754:
5752:
5751:
5747:
5744:
5743:
5739:
5736:
5735:
5731:
5730:
5728:
5726:
5722:
5716:
5715:
5711:
5709:
5708:
5704:
5702:
5701:
5697:
5696:
5693:
5689:
5686:
5683:
5678:
5655:
5654:
5650:
5648:
5647:
5643:
5641:
5640:
5636:
5634:
5633:
5629:
5628:
5626:
5624:
5623:Nyctosauridae
5620:
5614:
5613:
5609:
5607:
5606:
5602:
5600:
5599:
5595:
5594:
5591:
5588:
5586:
5582:
5575:
5574:
5570:
5568:
5567:
5566:Alamodactylus
5563:
5562:
5559:
5556:
5554:
5550:
5543:
5542:
5538:
5535:
5534:
5530:
5528:
5527:
5523:
5520:
5519:
5515:
5513:
5512:
5508:
5505:
5504:
5500:
5499:
5497:
5495:
5491:
5488:
5486:
5485:Pteranodontia
5482:
5476:
5475:
5471:
5470:
5467:
5463:
5460:
5457:
5452:
5448:
5444:
5439:
5435:
5425:
5420:
5417:
5409:
5383:
5382:
5381:
5380:
5378:
5376:
5372:
5368:
5330:
5329:
5325:
5322:
5321:
5320:Wellnhopterus
5317:
5315:
5314:
5310:
5308:
5307:
5303:
5301:
5300:
5296:
5294:
5293:
5292:Hatzegopteryx
5289:
5287:
5286:
5282:
5280:
5279:
5275:
5273:
5272:
5268:
5266:
5265:
5264:Aralazhdarcho
5261:
5260:
5258:
5256:
5252:
5246:
5245:
5241:
5239:
5238:
5234:
5232:
5231:
5227:
5224:
5223:
5219:
5218:
5216:
5214:
5210:
5203:
5202:
5198:
5195:
5194:
5190:
5188:
5187:
5183:
5180:
5179:
5175:
5173:
5172:
5168:
5165:
5164:
5160:
5159:
5157:
5155:
5151:
5144:
5143:
5139:
5137:
5136:
5135:Radiodactylus
5132:
5129:
5128:
5124:
5123:
5120:
5117:
5115:
5111:
5101:
5100:
5096:
5094:
5093:
5089:
5087:
5086:
5082:
5081:
5079:
5077:
5073:
5066:
5065:
5061:
5058:
5057:
5053:
5051:
5050:
5046:
5044:
5043:
5039:
5036:
5035:
5031:
5028:
5027:
5023:
5020:
5019:
5018:Apatorhamphus
5015:
5014:
5011:
5008:
5006:
5002:
4996:
4995:
4991:
4990:
4987:
4984:
4982:
4978:
4971:
4970:
4966:
4963:
4962:
4958:
4955:
4954:
4950:
4947:
4946:
4942:
4939:
4938:
4934:
4932:
4931:
4927:
4926:
4924:
4921:
4917:
4910:
4909:
4905:
4903:
4902:
4898:
4896:
4895:
4891:
4888:
4887:
4883:
4882:
4879:
4876:
4873:
4868:
4838:
4837:
4833:
4831:
4830:
4826:
4825:
4823:
4821:
4817:
4811:
4810:
4809:Tupandactylus
4806:
4804:
4803:
4799:
4797:
4796:
4792:
4789:
4788:
4784:
4783:
4780:
4777:
4775:
4771:
4765:
4764:
4760:
4757:
4756:
4752:
4749:
4748:
4744:
4741:
4740:
4739:Caupedactylus
4736:
4733:
4732:
4728:
4727:
4724:
4721:
4719:
4715:
4709:
4708:
4704:
4702:
4701:
4697:
4695:
4694:
4690:
4688:
4687:
4683:
4681:
4680:
4676:
4673:
4672:
4668:
4665:
4664:
4660:
4657:
4656:
4652:
4651:
4649:
4647:
4643:
4637:
4636:
4632:
4631:
4628:
4625:
4623:
4619:
4613:
4612:
4611:Caupedactylus
4608:
4606:
4605:
4601:
4600:
4598:
4595:
4594:Caupedactylia
4591:
4588:
4586:
4582:
4575:
4574:
4570:
4568:
4567:
4563:
4561:
4560:
4556:
4553:
4552:
4548:
4545:
4544:
4540:
4538:
4537:
4533:
4530:
4529:
4525:
4522:
4521:
4517:
4514:
4513:
4509:
4506:
4505:
4501:
4500:
4498:
4495:
4491:
4484:
4483:
4479:
4477:
4476:
4472:
4471:
4468:
4465:
4462:
4457:
4454:
4451:
4450:Neoazhdarchia
4446:
4440:
4439:
4435:
4433:
4432:
4428:
4425:
4424:
4420:
4417:
4416:
4412:
4410:
4409:
4405:
4403:
4402:
4398:
4397:
4394:
4391:
4388:
4387:Azhdarchoidea
4383:
4369:
4368:
4364:
4362:
4361:
4357:
4356:
4354:
4352:
4348:
4341:
4340:
4336:
4333:
4332:
4331:Puntanipterus
4328:
4326:
4325:
4321:
4319:
4318:
4314:
4311:
4310:
4306:
4303:
4302:
4298:
4297:
4294:
4291:
4289:
4285:
4281:
4278:
4275:
4270:
4263:
4262:
4258:
4257:
4254:
4251:
4248:
4243:
4239:
4235:
4230:
4226:
4217:
4211:
4185:
4184:
4183:
4182:
4180:
4178:
4174:
4167:
4166:
4162:
4161:
4158:
4155:
4153:
4149:
4145:
4119:
4118:
4114:
4112:
4111:
4107:
4105:
4104:
4100:
4098:
4097:
4096:Beipiaopterus
4093:
4092:
4090:
4088:
4084:
4078:
4077:
4073:
4072:
4069:
4066:
4064:
4060:
4054:
4053:
4052:Tacuadactylus
4049:
4047:
4046:
4042:
4040:
4039:
4035:
4032:
4031:
4027:
4025:
4024:
4020:
4019:
4017:
4015:
4011:
4005:
4004:
4000:
3998:
3997:
3993:
3992:
3990:
3987:
3983:
3977:
3976:
3972:
3970:
3969:
3965:
3963:
3962:
3958:
3956:
3955:
3951:
3949:
3948:
3944:
3942:
3941:
3937:
3935:
3934:
3933:Forfexopterus
3930:
3928:
3927:
3926:Elanodactylus
3923:
3921:
3920:
3919:Cratonopterus
3916:
3914:
3913:
3912:Cathayopterus
3909:
3907:
3906:
3902:
3899:
3898:
3897:Ardeadactylus
3894:
3893:
3890:
3887:
3884:
3879:
3864:
3863:
3859:
3857:
3856:
3852:
3849:
3848:
3847:Ardeadactylus
3844:
3843:
3841:
3839:
3835:
3828:
3827:
3823:
3821:
3820:
3816:
3815:
3812:
3809:
3807:
3803:
3796:
3795:
3791:
3788:
3787:
3783:
3780:
3779:
3775:
3773:
3772:
3771:Cycnorhamphus
3768:
3765:
3764:
3760:
3759:
3757:
3755:
3751:
3747:
3744:
3741:
3736:
3730:
3729:
3728:Pterodactylus
3725:
3723:
3722:
3721:Diopecephalus
3718:
3717:
3714:
3711:
3708:
3703:
3692:
3691:
3687:
3684:
3683:
3679:
3677:
3676:
3672:
3669:
3668:
3664:
3663:
3661:
3659:
3655:
3649:
3648:
3647:Prejanopterus
3644:
3643:
3640:
3636:
3633:
3630:
3625:
3621:
3617:
3612:
3608:
3599:
3594:
3557:
3556:
3555:
3554:
3552:
3550:
3546:
3540:
3539:
3535:
3532:
3531:
3527:
3525:
3524:
3520:
3518:
3517:
3513:
3511:
3510:
3506:
3504:
3503:
3499:
3496:
3495:
3494:Herbstosaurus
3491:
3489:
3488:
3487:Eurolimnornis
3484:
3482:
3481:
3480:Dermodactylus
3477:
3476:
3473:
3470:
3468:
3464:
3454:
3453:
3449:
3446:
3445:
3441:
3438:
3437:
3433:
3431:
3430:
3426:
3424:
3423:
3419:
3418:
3416:
3414:
3410:
3404:
3403:
3399:
3397:
3396:
3392:
3389:
3388:
3384:
3381:
3380:
3376:
3374:
3373:
3369:
3368:
3366:
3364:
3360:
3354:
3353:
3349:
3348:
3345:
3342:
3340:
3336:
3333:
3330:
3329:Caelidracones
3325:
3319:
3318:
3314:
3311:
3310:
3306:
3305:
3302:
3299:
3296:
3291:
3273:
3272:
3271:Wukongopterus
3268:
3265:
3264:
3260:
3258:
3257:
3256:Darwinopterus
3253:
3251:
3250:
3249:Cuspicephalus
3246:
3243:
3242:
3238:
3237:
3235:
3233:
3229:
3223:
3222:
3218:
3217:
3214:
3211:
3209:
3205:
3199:
3198:
3197:Pterorhynchus
3194:
3192:
3191:
3187:
3186:
3183:
3180:
3178:
3174:
3167:
3166:
3162:
3160:
3159:
3155:
3153:
3152:
3148:
3147:
3144:
3140:
3137:
3134:
3129:
3122:
3121:
3117:
3114:
3113:
3109:
3108:
3105:
3102:
3099:
3094:
3090:
3086:
3081:
3077:
3068:
3064:
3061:
3053:
3007:
3006:
3005:
3004:
3002:
3000:
2996:
2989:
2988:
2984:
2982:
2981:
2980:Scaphognathus
2977:
2975:
2974:
2970:
2968:
2967:
2963:
2961:
2960:
2956:
2955:
2953:
2950:
2946:
2943:
2940:
2936:
2918:
2917:
2913:
2910:
2909:
2905:
2902:
2901:
2897:
2896:
2894:
2892:
2888:
2882:
2881:
2877:
2874:
2873:
2869:
2866:
2865:
2861:
2859:
2858:
2854:
2852:
2851:
2847:
2846:
2844:
2842:
2838:
2835:
2833:
2829:
2823:
2822:
2818:
2816:
2815:
2811:
2809:
2808:
2804:
2802:
2801:
2797:
2795:
2794:
2790:
2788:
2787:
2783:
2781:
2780:
2776:
2775:
2772:
2769:
2767:
2763:
2756:
2755:
2751:
2748:
2747:
2743:
2741:
2740:
2736:
2733:
2732:
2728:
2727:
2724:
2721:
2719:
2715:
2712:
2709:
2704:
2694:
2693:
2689:
2687:
2686:
2682:
2681:
2679:
2677:
2673:
2669:
2666:
2663:
2658:
2651:
2650:
2649:Eudimorphodon
2646:
2645:
2642:
2639:
2635:
2630:
2619:
2618:
2614:
2611:
2610:
2609:Peteinosaurus
2606:
2603:
2602:
2598:
2596:
2595:
2591:
2589:
2588:
2584:
2581:
2580:
2576:
2575:
2573:
2571:
2567:
2561:
2560:
2559:Herbstosaurus
2556:
2555:
2552:
2548:
2545:
2542:
2537:
2530:
2529:
2528:Peteinosaurus
2525:
2524:
2521:
2518:
2514:
2509:
2491:
2490:
2489:Eudimorphodon
2486:
2483:
2482:
2478:
2477:
2475:
2473:
2469:
2462:
2461:
2457:
2456:
2453:
2450:
2448:
2444:
2438:
2437:
2433:
2431:
2430:
2426:
2424:
2423:
2419:
2416:
2415:
2411:
2410:
2408:
2406:
2402:
2399:
2397:
2393:
2386:
2385:
2384:Preondactylia
2381:
2378:
2377:
2376:Peteinosaurus
2373:
2370:
2369:
2365:
2364:
2361:
2358:
2355:
2354:Eopterosauria
2350:
2336:
2335:
2331:
2329:
2328:
2324:
2321:
2320:
2316:
2315:
2313:
2311:
2307:
2300:
2299:
2295:
2293:
2292:
2288:
2285:
2284:
2280:
2279:
2276:
2273:
2270:
2266:
2260:
2259:
2258:Preondactylus
2255:
2253:
2252:
2248:
2247:
2245:
2243:
2242:Preondactylia
2239:
2233:
2232:
2228:
2227:
2224:
2220:
2217:
2214:
2209:
2205:
2201:
2196:
2192:
2171:
2170:
2169:
2168:
2166:
2164:
2160:
2152:
2148:
2147:
2146:
2145:
2142:
2139:
2137:
2133:
2129:
2121:
2117:
2114:
2112:
2108:
2105:
2103:
2099:
2096:
2094:
2090:
2088:
2084:
2082:
2078:
2077:
2072:
2068:
2064:
2057:
2052:
2050:
2045:
2043:
2038:
2037:
2034:
2028:(8): 305–313.
2027:
2023:
2019:
2015:
2012:
2008:
2004:
2001:
1997:
1993:
1989:
1985:
1981:
1979:(2): 221–247.
1978:
1974:
1970:
1966:
1963:
1959:
1955:
1951:
1950:
1946:
1935:
1932:
1927:
1923:
1919:
1915:
1911:
1907:
1900:
1897:
1892:
1888:
1884:
1882:9780691232218
1878:
1874:
1870:
1866:
1859:
1856:
1851:
1847:
1842:
1837:
1832:
1827:
1823:
1819:
1815:
1811:
1807:
1800:
1797:
1792:
1788:
1784:
1780:
1776:
1772:
1768:
1764:
1760:
1753:
1750:
1745:
1741:
1737:
1733:
1729:
1725:
1721:
1714:
1711:
1706:
1702:
1698:
1694:
1690:
1683:
1680:
1675:
1671:
1667:
1663:
1659:
1655:
1651:
1647:
1643:
1639:
1635:
1628:
1625:
1620:
1616:
1612:
1608:
1603:
1598:
1594:
1590:
1587:(sup1): 142.
1586:
1582:
1578:
1576:
1567:
1564:
1558:
1553:
1549:
1545:
1538:
1536:
1534:
1530:
1525:
1521:
1516:
1511:
1506:
1501:
1497:
1493:
1489:
1482:
1480:
1476:
1472:
1469:
1465:
1459:
1457:
1453:
1446:
1442:
1439:
1437:
1434:
1432:
1429:
1428:
1424:
1422:
1419:
1417:
1416:
1411:
1403:
1396:
1395:
1387:
1386:
1378:
1377:
1369:
1368:
1360:
1359:
1351:
1350:
1342:
1341:
1333:
1332:
1329:
1328:
1327:
1320:
1319:
1316:
1315:
1312:
1310:
1306:
1305:
1298:
1297:
1294:
1293:
1290:
1289:
1288:
1281:
1280:
1274:
1273:
1270:
1269:
1266:
1265:
1264:
1257:
1256:
1250:
1249:
1246:
1245:
1242:
1237:
1236:
1230:
1229:
1226:
1225:
1222:
1221:
1220:
1213:
1212:
1205:
1201:
1200:
1197:
1196:
1193:
1192:
1191:
1184:
1183:
1177:
1176:
1173:
1172:
1164:
1163:
1155:
1154:
1146:
1145:
1142:
1141:
1140:
1133:
1132:
1129:
1128:
1125:
1124:
1123:
1116:
1115:
1109:
1108:
1105:
1104:
1101:
1100:
1099:
1092:
1091:
1085:
1084:
1081:
1080:
1077:
1076:
1075:
1068:
1067:
1060:
1056:
1055:
1049:
1048:
1045:
1044:
1036:
1035:
1027:
1026:
1018:
1017:
1014:
1013:
1012:
1005:
1004:
1001:
1000:
997:
996:
995:
988:
987:
981:
980:
977:
976:
968:
967:
964:
963:
962:
955:
954:
951:
950:
947:
945:
941:
940:
933:
932:
926:
925:
918:
914:
913:
910:
909:
901:
900:
892:
891:
888:
887:
886:
879:
878:
875:
874:
871:
870:
869:
862:
861:
855:
854:
851:
850:
847:
846:
845:
838:
837:
830:
826:
825:
819:
818:
811:
810:Neoazhdarchia
807:
806:
802:
800:
799:Neoazhdarchia
796:
792:
785:
781:
777:
773:
767:
762:
755:
753:
751:
747:
743:
742:
741:Hatzegopteryx
737:
733:
728:
724:
723:
716:
714:
710:
706:
702:
698:
697:
691:
689:
682:
678:
673:
666:
664:
662:
658:
654:
653:
647:
645:
641:
637:
633:
627:
625:
621:
617:
613:
609:
604:
602:
598:
594:
590:
586:
581:
579:
575:
571:
566:
564:
559:
555:
554:Eberhard Frey
551:
550:David Martill
546:
544:
543:
538:
534:
530:
526:
525:
520:
516:
513:
506:
505:
500:
499:
498:Tyrannosaurus
494:
489:
485:
483:
482:
477:
472:
470:
466:
465:specific name
462:
458:
454:
450:
446:
441:
439:
435:
431:
427:
423:
415:
412:
411:
406:
405:
404:Hatzegopteryx
400:
395:
388:
386:
384:
380:
376:
372:
371:United States
368:
364:
363:Maastrichtian
360:
356:
353:
349:
348:
332:
329:
314:
308:
305:
294:
291:
287:
282:
276:
274:
265:
262:
261:Binomial name
258:
254:
253:
247:
244:
243:
238:
232:
231:
224:
221:
220:
217:
211:
208:
207:
204:
198:
195:
194:
191:
185:
182:
181:
178:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
133:
128:
124:
120:
116:
112:
107:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:Maastrichtian
33:
29:
26:
22:
6781:Paleontology
6681:
6645:
6591:
6586:Ludodactylus
6584:
6577:
6570:
6562:
6554:
6547:
6527:
6519:
6514:Nicorhynchus
6512:
6505:
6498:
6477:
6470:
6462:
6454:
6446:
6438:
6430:
6400:
6393:
6385:Hamipteridae
6371:
6363:
6329:
6321:
6313:
6293:
6285:
6280:Aetodactylus
6278:
6243:
6235:
6228:
6220:
6214:Ludodactylus
6212:
6204:
6196:
6188:
6180:
6172:
6166:Aetodactylus
6164:
6143:
6135:
6127:
6119:
6111:
6104:
6097:
6054:
6047:
6023:
6016:
6009:
6002:
5976:Liaoxipterus
5974:
5967:
5947:
5940:
5933:
5926:
5904:Mimodactylus
5902:
5895:
5888:
5867:
5860:
5853:
5846:
5822:
5815:
5790:
5782:Lanceodontia
5763:
5755:
5750:Lonchodectes
5748:
5740:
5732:
5712:
5705:
5698:
5651:
5644:
5637:
5630:
5610:
5603:
5596:
5571:
5564:
5539:
5531:
5524:
5518:Ornithostoma
5516:
5509:
5501:
5472:
5412:
5326:
5318:
5311:
5304:
5297:
5290:
5285:Eurazhdarcho
5283:
5276:
5270:
5269:
5262:
5242:
5235:
5228:
5220:
5213:Azhdarchinae
5199:
5191:
5184:
5176:
5169:
5161:
5154:Azhdarchidae
5140:
5133:
5125:
5097:
5090:
5083:
5062:
5054:
5049:Meilifeilong
5047:
5040:
5032:
5024:
5016:
4992:
4967:
4959:
4951:
4943:
4935:
4928:
4906:
4899:
4892:
4884:
4834:
4827:
4807:
4800:
4793:
4785:
4761:
4753:
4745:
4737:
4729:
4705:
4698:
4691:
4686:Huaxiapterus
4684:
4677:
4671:Eopteranodon
4669:
4661:
4655:Afrotapejara
4653:
4646:Sinopterinae
4635:Afrotapejara
4633:
4609:
4602:
4571:
4564:
4557:
4549:
4541:
4534:
4526:
4518:
4510:
4502:
4480:
4475:Bennettazhia
4473:
4438:Ornithostoma
4436:
4429:
4421:
4413:
4406:
4399:
4365:
4358:
4337:
4329:
4324:Ordosipterus
4322:
4315:
4307:
4299:
4274:Tapejaroidea
4259:
4163:
4117:Pterodaustro
4115:
4108:
4101:
4094:
4074:
4050:
4043:
4036:
4030:Huanhepterus
4028:
4023:Gnathosaurus
4021:
4003:Moganopterus
4001:
3994:
3975:Pterofiltrus
3973:
3968:Petrodactyle
3966:
3959:
3954:Liaodactylus
3952:
3947:Kepodactylus
3945:
3938:
3931:
3924:
3917:
3910:
3903:
3895:
3862:Huanhepterus
3860:
3853:
3845:
3824:
3819:Aerodactylus
3817:
3792:
3786:Petrodactyle
3784:
3776:
3769:
3761:
3726:
3719:
3688:
3680:
3673:
3665:
3645:
3536:
3528:
3521:
3514:
3507:
3502:Kryptodrakon
3500:
3492:
3485:
3478:
3450:
3444:Jeholopterus
3442:
3434:
3427:
3420:
3400:
3393:
3387:Jeholopterus
3385:
3377:
3372:Anurognathus
3370:
3352:Mesadactylus
3350:
3315:
3307:
3269:
3261:
3254:
3247:
3239:
3219:
3195:
3188:
3177:Darwinoptera
3163:
3156:
3149:
3118:
3110:
3056:
2985:
2978:
2971:
2964:
2957:
2914:
2908:Nesodactylus
2906:
2900:Cacibupteryx
2898:
2880:Sericipterus
2878:
2870:
2862:
2855:
2848:
2819:
2812:
2807:Nesodactylus
2805:
2798:
2791:
2786:Cacibupteryx
2784:
2779:Bellubrunnus
2777:
2752:
2744:
2737:
2729:
2690:
2683:
2647:
2634:Lonchognatha
2615:
2607:
2599:
2592:
2585:
2577:
2557:
2526:
2487:
2479:
2458:
2434:
2427:
2422:Pachagnathus
2420:
2412:
2382:
2374:
2368:Austriadraco
2366:
2332:
2327:Austriadraco
2325:
2317:
2296:
2289:
2281:
2256:
2249:
2229:
2115:
2106:
2097:
2025:
2021:
2017:
2010:
2006:
1999:
1995:
1991:
1987:
1983:
1976:
1972:
1968:
1961:
1957:
1953:
1934:
1909:
1905:
1899:
1864:
1858:
1813:
1809:
1799:
1769:(1): 79–90.
1766:
1762:
1752:
1735:
1731:
1727:
1723:
1719:
1713:
1696:
1692:
1688:
1682:
1641:
1637:
1627:
1584:
1580:
1574:
1566:
1550:. e2385068.
1547:
1495:
1492:PLOS Biology
1491:
1470:
1467:
1463:
1420:
1413:
1409:
1407:
1324:
1323:
1302:
1301:
1286:
1285:
1284:
1261:
1260:
1240:
1217:
1216:
1204:Azhdarchidae
1188:
1187:
1137:
1136:
1120:
1119:
1096:
1095:
1072:
1071:
1009:
1008:
992:
991:
959:
958:
937:
936:
883:
882:
866:
865:
842:
841:
794:
788:
783:
749:
745:
739:
735:
731:
726:
720:
717:
712:
708:
704:
700:
694:
692:
685:
676:
660:
656:
650:
648:
643:
631:
628:
623:
619:
615:
605:
601:Azhdarchidae
593:Kevin Padian
585:Azhdarchinae
582:
577:
574:nomen dubium
573:
569:
567:
547:
542:nomen dubium
540:
536:
524:Titanopteryx
522:
519:entomologist
509:
502:
496:
492:
479:
473:
460:
442:
419:
413:
408:
402:
398:
383:Azhdarchidae
378:
346:
345:
344:
330:
307:Titanopteryx
306:
272:
267:
251:
250:
236:
229:
228:
203:Azhdarchidae
31:
25:
6821:Azhdarchids
6706:Wikispecies
6472:Thapunngaka
6432:Amblydectes
6351:Anhangueria
6315:Aussiedraco
6049:Boreopterus
5824:Lonchodraco
5757:Lonchodraco
5700:Aussiedraco
5646:Nyctosaurus
5503:Bogolubovia
5171:Bogolubovia
5042:Lacusovagus
5034:Eoazhdarcho
4994:Eoazhdarcho
4953:Leptostomia
4945:Keresdrakon
4820:Caiuajarina
4787:Bakonydraco
4755:Lacusovagus
4747:Keresdrakon
4718:Tapejarinae
4679:Huaxiadraco
4663:Bakonydraco
4622:Tapejaridae
4551:Leptostomia
4543:Lacusovagus
4536:Kariridraco
4482:Keresdrakon
4423:Leptostomia
4415:Keresdrakon
4076:Ctenochasma
4038:Lusognathus
3961:Otogopterus
3616:Lophocratia
3549:Lophocratia
3516:Pangupterus
3452:Sinomacrops
2793:Dorygnathus
2662:Novialoidea
2617:Rhamphinion
2594:Dimorphodon
2269:Caviramidae
2213:Pterosauria
2200:Pterosauria
2163:Pterosauria
2102:Archosauria
2063:Pterosauria
2002:(1): 57–76.
789:Below is a
766:azhdarchids
711:, and that
667:Description
313:preoccupied
209:Subfamily:
177:Pterosauria
21:Arambourgia
6815:Categories
6549:Anhanguera
6440:Ferrodraco
6395:Hamipterus
6323:Barbosania
6198:Ferrodraco
6190:Draigwenia
6025:Hamipterus
6004:Barbosania
5949:Nurhachius
5817:Ikrandraco
5792:Draigwenia
5742:Ikrandraco
5707:Serradraco
5653:Volgadraco
5541:Volgadraco
5533:Tethydraco
5526:Pteranodon
5384:see below↓
5278:Cryodrakon
5201:Volgadraco
5193:Tethydraco
5092:Jidapterus
5056:Microtuban
4894:Microtuban
4795:Europejara
4774:Tapejarini
4763:Vectidraco
4700:Sinopterus
4317:Noripterus
4186:see below↓
4110:Gegepterus
4103:Eosipterus
3558:see below↓
3538:Wenupteryx
3429:Cascocauda
3008:see below↓
2436:Yelaphomte
2172:see below↓
2093:Sauropsida
1964:: 229–234.
1947:References
1912:(3–4): 1.
1241:TMM 42489
457:metacarpal
365:stage) of
352:azhdarchid
183:Suborder:
6500:Aerodraco
5942:Luchibang
5890:Haopterus
5612:Simurghia
5605:Epapatelo
5573:Cretornis
5511:Dawndraco
5237:Azhdarcho
5230:Albadraco
5222:Aerotitan
5142:Alanqidae
4920:Alanqidae
4886:Cretornis
4836:Torukjara
4829:Caiuajara
4566:Tupuxuara
4504:Aerotitan
4408:Inabtanin
3996:Feilongus
3395:Luopterus
3112:Allkaruen
2739:Klobiodon
2579:Allkaruen
2414:Caviramus
2291:Caviramus
2079:Kingdom:
2013:: 78–97.
1891:249332375
1791:135043714
1738:: 57–76.
1674:132649124
1666:0305-8719
1619:245125409
1611:0272-4634
1415:Inabtanin
791:cladogram
778:from the
776:Ulna bone
652:Inabtanin
533:Simulidae
515:Lev Nesov
469:Antiquity
434:phosphate
407:(A), and
357:from the
355:pterosaur
281:Arambourg
245:Species:
150:Kingdom:
144:Eukaryota
6726:10583860
6697:Q2317988
6691:Wikidata
6572:Guidraco
6456:Mythunga
6448:Haliskia
6222:Mythunga
6206:Guidraco
6137:Haliskia
5714:Unwindia
5064:Xericeps
4969:Xericeps
4901:Xericeps
4802:Tapejara
4573:Xericeps
4528:Banguela
4301:Banguela
3530:Samrukia
3190:Ceoptera
2087:Chordata
2085:Phylum:
2081:Animalia
1926:84617119
1850:21085624
1810:PLOS ONE
1524:29534059
1425:See also
688:holotype
681:theropod
491:Size of
474:In 1975
430:Russeifa
426:Damascus
361:period (
290:Synonyms
196:Family:
164:Chordata
160:Phylum:
154:Animalia
140:Domain:
119:Holotype
6767:Portals
6739:8608430
5598:Alcione
4707:Wightia
2091:Class:
1998:, Abh.
1841:2981443
1818:Bibcode
1771:Bibcode
1646:Bibcode
1589:Bibcode
1515:5849296
1473::94–103
1404:Biology
1206:
1061:
919:
831:
812:
640:Morocco
451:of the
414:lawsoni
375:Morocco
283:, 1959)
270:†
249:†
235:Nessov
222:Genus:
170:Order:
6793:Jordan
6752:156655
5163:Alanqa
4930:Alanqa
4512:Alanqa
3120:Sordes
2987:Sordes
1924:
1889:
1879:
1848:
1838:
1789:
1672:
1664:
1617:
1609:
1522:
1512:
1202:
1057:
915:
827:
808:
572:was a
504:Balaur
501:, and
367:Jordan
4261:Piksi
2857:Dearc
2116:Clade
2107:Clade
2098:Clade
1922:S2CID
1887:S2CID
1787:S2CID
1670:S2CID
1615:S2CID
1447:Notes
445:Paris
422:Amman
401:(C),
6734:GBIF
2149:see
1990:and
1877:ISBN
1846:PMID
1726:and
1662:ISSN
1607:ISSN
1520:PMID
686:The
636:ulna
552:and
373:and
237:vide
44:PreꞒ
6721:EoL
2011:197
2000:207
1994:."
1977:199
1914:doi
1910:103
1869:doi
1836:PMC
1826:doi
1779:doi
1767:217
1740:doi
1736:207
1730:".
1701:doi
1697:199
1654:doi
1642:455
1597:doi
1552:doi
1510:PMC
1500:doi
738:or
610:of
6817::
6749::
6736::
6723::
6708::
6693::
2118::
2109::
2100::
2024:,
2009:,
1975:,
1960:,
1920:.
1908:.
1885:.
1875:.
1844:.
1834:.
1824:.
1812:.
1808:.
1785:.
1777:.
1765:.
1761:.
1734:.
1695:.
1668:.
1660:.
1652:.
1640:.
1636:.
1613:.
1605:.
1595:.
1585:41
1583:.
1579:.
1546:.
1532:^
1518:.
1508:.
1496:16
1494:.
1490:.
1478:^
1471:33
1455:^
663:.
646:.
626:.
603:.
545:.
94:Pg
6769::
6567:?
6559:?
6524:?
6492:?
6467:?
6459:?
6451:?
6443:?
6435:?
6424:?
6387:?
6376:?
6368:?
6326:?
6318:?
6290:?
6263:?
6248:?
6240:?
6225:?
6217:?
6209:?
6201:?
6193:?
6185:?
6177:?
6169:?
6148:?
6140:?
6132:?
6124:?
6116:?
5872:?
5795:?
5768:?
5760:?
5745:?
5737:?
5656:?
5576:?
5544:?
5536:?
5521:?
5506:?
5323:?
5225:?
5204:?
5196:?
5181:?
5166:?
5145:?
5130:?
5067:?
5059:?
5037:?
5029:?
5021:?
4972:?
4964:?
4956:?
4948:?
4940:?
4922:?
4911:?
4889:?
4790:?
4758:?
4750:?
4742:?
4734:?
4674:?
4666:?
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4523:?
4515:?
4507:?
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4426:?
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4342:?
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2055:e
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2041:v
2026:6
1962:7
1928:.
1916::
1893:.
1871::
1852:.
1828::
1820::
1814:5
1793:.
1781::
1773::
1746:.
1742::
1707:.
1703::
1676:.
1656::
1648::
1621:.
1599::
1591::
1560:.
1554::
1526:.
1502::
424:-
315:)
279:(
226:†
213:†
200:†
187:†
174:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
23:.
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