1874:
1896:
4512:
1801:
4507:
557:
3667:
148:
1542:
127:
613:
568:
3672:
937:
2345:
Christopher A. Brochu; Ane de Celis; Amanda J. Adams; Stephanie K. Drumheller; Jennifer H. Nestler; Brenda R. Benefit; Aryeh
Grossman; Francis Kirera; Thomas Lehmann; Cynthia Liutkus-Pierce; Fredrick K. Manthi; Monte L. McCrossin; Kieran P. McNulty; Rose Nyaboke Juma (June 2022).
987:
because they superficially resemble modern crocodiles. However, the majority of early crocodilians, even some early alligatoroids, resembled modern crocodiles because a triangular, crocodile-shaped head is a primitive condition for crocodilians.
654:
in the skull, a feature that is shared by all fossils attributed to the species. The fossils are also similar in having 6 pairs of teeth lining the symphysis at the tip of the lower jaw. As was the case for many other
Paleogene crocodyloids,
400:
between the two halves of the lower jaw. Recent studies of the evolutionary relationships of early crocodilians along with closer examinations of the morphology of fossil specimens suggest that only the first named species of
463:
can be distinguished from other early crocodyloids by its extended mandibular symphysis, the region where the two halves of the lower jaws connect. In many crocodyloids this joint is formed from two pairs of bones, the
927:
in marine deposits suggests that these crocodilians could have tolerated prolonged periods of time in the ocean, an ability that would have aided in the dispersal of early crocodyloids across Europe and Asia.
1110:
species represent a non-monophyletic grouping. Delfino and Smith considered these relationships to have very weak support because only a few characteristics entered into the data matrix differed between
2400:
Salisbury, S. W.; Willis, P. M. A. (1996). "A new crocodylian from the Early Eocene of south-eastern
Queensland and a preliminary investigation of the phylogenetic relationships of crocodyloids".
1029:
is an unlikely candidate for the ancestor of mekosuchines because it lived very far from
Australia and the likelihood that it could have reached Australia from Europe is very low.
455:
species have teeth in the upper jaw that completely overlap the teeth in the lower jaw, giving them overbites. An overbite is a primitive feature among crocodyloids because modern
2390:
Kotsakis T, Delfino M, Piras P. 2004. Italian
Cenozoic crocodilians: taxa, timing and biogeographic implications. Palaeogeography, Palaeoclimatology, Palaeoecology 210: 67–87.
2093:"A reassessment of the morphology and taxonomic status of 'Crocodylus'depressifronsBlainville, 1855 (Crocodylia, Crocodyloidea) based on the Early Eocene remains from Belgium"
4608:
2483:
Hekkala, E.; Gatesy, J.; Narechania, A.; Meredith, R.; Russello, M.; Aardema, M. L.; Jensen, E.; Montanari, S.; Brochu, C.; Norell, M.; Amato, G. (2021-04-27).
2580:
546:(modern crocodiles) but different in having 17 teeth in each half of the lower jaw and a splenial bone that does not form part of the mandibular symphysis.
1003:. Mekosuchines are a group of crocodyloids from Australia and the South Pacific that are unusual in that they were highly specialized for life on land. If
638:
that dates back to the Early Eocene. The skull was illustrated in the 1855 paper but it was not thoroughly described. The fossil has since become heavily
2019:"Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem"
4837:
2124:
Farion-Demaret, M. & Smith, T. (2002). "Fruits and seeds from the Tienen
Formation at Dormaal, Palaeocene–Eocene transition in eastern Belgium".
4768:
4688:
4595:
642:, losing much of its original anatomical detail. After its naming, several other crocodilian fossils in European museum collections were labeled as
1025:-like crocodyloid could have dispersed into Australia as the ancestor of mekosuchines. Despite the results of the phylogenetic analysis,
1007:
is the sister taxon of
Mekosuchinae, it may have been close to the ancestry of the group. The earliest known and most basal mekosuchine,
4827:
2436:
523:
2553:
2545:
1873:
4506:
2573:
4511:
4832:
3839:
3666:
3885:
3754:
2485:"Paleogenomics illuminates the evolutionary history of the extinct Holocene "horned" crocodile of Madagascar, Voay robustus"
1777:
578:
Well-preserved remains of a crocodyloid were first described from
Germany and France in 1966 and placed in a new species of
538:, a vertebrate paleontologist with the American Museum of Natural History and a member of the expedition. Mook thought that
514:, was named by paleontologist Charles Mook in 1940. It was named on the basis of a lower jaw and pieces of a skull from the
2319:"Taxonomic reassessment and phylogenetic test of Asiatosuchus nanlingensis Young, 1964 and Eoalligator chunyii Young, 1964"
2266:"Taxonomic revision of Eoalligator (Crocodylia, Brevirostres) and the paleogeographic origins of the Chinese alligatoroids"
812:
by
Angielczyk and Gingerich (1998) because they are based on fossil specimens that preserve very little anatomical detail.
684:
have been found from Early Eocene deposits in
Belgium. Together these specimens provide details on most of the skeleton.
423:
lack the unique anatomical features necessary to justify their classification as distinct species. Other species such as
1342:
631:
147:
2566:
3746:
2913:
2348:"Giant dwarf crocodiles from the Miocene of Kenya and crocodylid faunal dynamics in the late Cenozoic of East Africa"
1619:
2174:
3967:
2432:"Tip-dating and homoplasy: reconciling the shallow molecular divergences of modern gharials with their long fossil"
3939:
704:
behind the eye socket that is visible when the skull is viewed from the side. Another distinguishing feature of
3585:
2906:
2204:
1680:
3671:
2152:
3960:
1895:
764:. A subsequent paper argued otherwise, but other researchers agree with the initial assessment synonymizing
4446:
4220:
4550:
4470:
4244:
3953:
2323:
851:
in Wyoming has a similarly shaped splenial bone in the lower jaw and frontal bone in the skull. Although
392:
typified by flat, triangular skulls. The feature that traditionally united these species under the genus
4822:
4794:
4714:
4634:
4430:
4423:
4236:
4228:
4112:
2749:
2669:
2352:
1838:
1506:
1243:
964:
916:
515:
4454:
1800:
731:
665:
because the overall shape of its skull is similar to those of living crocodiles. Soon after Mook named
4755:
4675:
4582:
3946:
3635:
3628:
3578:
3376:
2238:
1457:
833:
Several crocodilian fossils from the Paleogene of North America have also been proposed to belong to
594:
and surrounding subtropical forests during the Eocene. Of all the species that have been assigned to
389:
256:
1056:
crocodyloids rather than its own separate lineage. Since a genus name is normally only applied to a
352:
3614:
3341:
2179:(Crocodyloidea) from the Middle Eocene Drazinda Formation of the Sulaiman Range, Punjab (Pakistan)"
959:, which includes living crocodiles and their extinct relatives. Recent phylogenetic analyses place
756:
556:
4478:
4387:
4379:
4105:
3383:
3362:
3355:
3016:
2892:
2696:
1700:
1318:
1049:
908:
843:
761:
739:
142:
4799:
886:
in 1993, although subsequent authors questioned this referral and considered it synonymous with
4719:
4639:
979:, and their extinct relatives. Many of the species that are most closely related to species of
692:
by a combination of several characteristics including a large hole and a depressed area on the
4781:
4701:
4621:
4613:
4394:
3893:
3860:
3621:
3348:
3153:
2524:
2506:
2465:
2299:
2052:
848:
697:
535:
4786:
4706:
4626:
4372:
3607:
3470:
3433:
3369:
3264:
3130:
2514:
2496:
2455:
2445:
2409:
2371:
2361:
2289:
2279:
2246:
2133:
2104:
2042:
2032:
1541:
1195:
701:
215:
202:
459:
have teeth in the upper and lower jaws that interlock with each other with little overlap.
126:
4462:
4438:
4401:
4321:
4069:
3796:
3555:
3540:
3505:
3448:
3318:
3174:
3119:
2936:
2920:
1565:
1359:
612:
397:
2242:
904:
sp. by Kotsakis et al. (2004) pending revision of the Mount Bolca crocodilian material.
567:
451:
has a generalized crocodilian skull that is triangular in shape when viewed from above.
4335:
4313:
4055:
4047:
4005:
3803:
3738:
3724:
3597:
3310:
3294:
3223:
3181:
3139:
3111:
3089:
3046:
2981:
2968:
2943:
2928:
2899:
2830:
2786:
2771:
2519:
2484:
2460:
2431:
2376:
2347:
2294:
2265:
2047:
2018:
1536:
1461:
1217:
968:
719:
590:
quarry, a fossil site that has preserved many forms of life that inhabited a series of
431:
are still considered valid species, but they do not form an evolutionary grouping with
338:
17:
2137:
4816:
4533:
4365:
4358:
4343:
4328:
4291:
4277:
4189:
4174:
4166:
4083:
4062:
4033:
4026:
4019:
3909:
3823:
3811:
3761:
3731:
3716:
3702:
3484:
3455:
3425:
3287:
3256:
3243:
3231:
3215:
3197:
3160:
3060:
2884:
2862:
2318:
2109:
2092:
1762:
1502:
1481:
1465:
1229:
1168:
1053:
956:
591:
519:
420:
235:
84:
4528:
4351:
4284:
4197:
4182:
4149:
4138:
4090:
3872:
3782:
3769:
3689:
3568:
3519:
3512:
3440:
3302:
3279:
3104:
3096:
3067:
3009:
3002:
2762:
2654:
2629:
1796:
1750:
1736:
1510:
1375:
1057:
1041:
1000:
996:
944:
888:
777:
773:
651:
507:
465:
415:
302:
264:
4666:
967:("primitive") member of this clade, close to the split between Crocodyloidea and
4306:
4299:
4097:
4076:
4040:
4012:
3991:
3533:
3526:
3498:
3491:
3477:
3418:
3271:
3207:
3167:
3146:
3074:
3053:
2995:
2869:
2837:
2794:
1823:
1142:
920:
897:
823:
818:
388:
since the genus was named in 1940. These species have a generalized crocodilian
377:
374:
59:
4746:
4573:
2501:
2229:
Waldman, M. (1970). "Comments on a Cretaceous coprolite from Alberta, Canada".
476:
it is only formed by the dentary bones. Based on largely complete skeletons of
4260:
4124:
3929:
3853:
3846:
3832:
3411:
3028:
2808:
2708:
2620:
2590:
2413:
1854:
1521:
1473:
1453:
1129:
879:
693:
661:
587:
225:
104:
69:
2510:
1099:
have enough distinguishing features to be included in phylogenetic analyses.
1060:
grouping by researchers who study prehistoric crocodilians, the type species
4210:
3548:
3331:
2815:
2681:
1890:
1494:
1103:
972:
936:
875:
743:
456:
381:
159:
109:
53:
41:
2528:
2469:
2450:
2303:
2056:
772:. Since then, different analyses have recovered the species as a member of
384:. Many Paleogene crocodilians from Europe and Asia have been attributed to
4740:
4660:
4567:
4413:
3901:
3462:
3082:
2822:
2638:
2614:
2186:
Contributions from the Museum of Paleontology, the University of Michigan
1868:
735:
469:
179:
99:
94:
79:
74:
64:
2037:
4773:
4693:
4600:
2801:
2558:
2284:
1032:
Most phylogenetic analyses do not support the idea that all species of
1009:
527:
189:
114:
89:
3398:
2608:
2366:
1469:
976:
639:
635:
410:
169:
45:
4544:
2250:
413:
of Mongolia, belongs in the genus. Most species are now regarded as
2844:
2270:
2023:
952:
935:
700:(two holes at the top of the skull behind the eye sockets), and a
611:
566:
555:
370:
4760:
4680:
4587:
947:
analyses of the evolutionary relationships of crocodilians place
630:
The naming of this new species was based on a skull found in the
3917:
2779:
1486:
4548:
3687:
2966:
2747:
2601:
2562:
746:(fossilized feces) were found alongside the type specimen of
919:, a marine deposit which has also preserved the remains of
435:
that would warrant them being placed together in the genus
826:
crocodilians, by Efimov (1988), but Brochu (2007) treated
2017:
Rio, Jonathan P.; Mannion, Philip D. (6 September 2021).
1497:
shows the results of the latest studies, which placed
855:
is known from a complete skull, the skull material of
1472:
age) data established the inter-relationships within
696:
of the skull, a frontal bone that does not touch the
2402:
Alcheringa: An Australasian Journal of Palaeontology
1842:
1781:
1704:
1684:
1650:
1623:
1596:
1569:
1192:
1164:
1076:
are written in quotes because they do not belong to
776:, or as basal to a clade including mekosuchines and
345:
331:
320:
309:
294:
271:
4730:
4650:
4557:
4411:
4259:
4208:
4148:
4122:
3989:
3927:
3871:
3822:
3780:
3700:
3595:
3566:
3397:
3329:
3242:
3195:
3129:
3026:
2979:
2854:
2761:
2707:
2680:
2653:
1513:(the combined group of crocodiles and gavialids).
1080:and have not yet been given different genus names.
1036:belong to their own clade. Instead they find that
518:of Inner Mongolia, China, which dates back to the
2430:Michael S. Y. Lee; Adam M. Yates (27 June 2018).
488:may have grown up to 4 metres (13 ft) long.
1064:is now considered the only valid species within
659:was originally placed in the still-living genus
2425:
2423:
2264:Yan-Yin Wang; Corwin Sullivan; Jun Liu (2016).
2173:Angielczyk, K.D. & Gingerich, P.D. (1998).
923:whales. The presence of a possible specimen of
688:can be distinguished from all other species of
1456:study by Lee & Yates simultaneously using
2574:
2340:
2338:
1476:, which was expanded upon in 2021 by Hekkala
907:A partial skeleton of a crocodyloid from the
754:was discovered concurrently with the similar
8:
2317:Xiao-Chun Wu; Chun Li; Yan-Yin Wang (2017).
915:. The fossil was found in the Middle Eocene
1285:"Asiatosuchus" / "Crocodylus" depressifrons
1106:from Delfino and Smith (2009) showing that
680:Many new and much more complete fossils of
526:'s Central Asiatic Expedition of 1930 near
4545:
4269:
4158:
4130:
3997:
3877:
3788:
3708:
3697:
3684:
3403:
3248:
3239:
3038:
3034:
2987:
2976:
2963:
2758:
2744:
2686:
2659:
2650:
2598:
2581:
2567:
2559:
911:of Pakistan was tentatively attributed to
760:, and a 2016 study proposed that they are
602:is known from the most complete material.
125:
31:
2518:
2500:
2459:
2449:
2375:
2365:
2293:
2283:
2108:
2097:Zoological Journal of the Linnean Society
2046:
2036:
2012:
2010:
730:, based on fragmentary material from the
2153:"A new fossil crocodilian from Mongolia"
983:were originally classified in the genus
447:Like most other Paleogene crocodyloids,
2086:
2006:
991:Some phylogenetic analyses have placed
522:. These fossils were discovered by the
419:or "dubious names", meaning that their
2168:
2166:
2084:
2082:
2080:
2078:
2076:
2074:
2072:
2070:
2068:
2066:
999:or closest relative of a group called
2126:Review of Palaeobotany and Palynology
650:refers to the flattened shape of the
7:
1083:The relationships of other putative
1860:
1830:
1769:
1757:
1728:
1672:
1665:
1638:
1611:
1584:
1557:
1528:
1516:
1484:by extracting DNA from the extinct
1334:
1276:
1235:
1210:
1185:
1157:
1135:
1125:
1115:species, and none differed between
1087:species are uncertain because only
896:from the Middle Eocene locality of
2437:Proceedings of the Royal Society B
2231:Canadian Journal of Earth Sciences
2214:(in Chinese (China) and English).
2205:"New fossil crocodiles from China"
882:, Italy in 1914 and reassigned to
542:was closely related to species of
524:American Museum of Natural History
25:
971:, the group that includes living
586:The German remains came from the
380:that lived in Eurasia during the
4838:Prehistoric pseudosuchian genera
4510:
4505:
3670:
3665:
2110:10.1111/j.1096-3642.2008.00478.x
1894:
1872:
1799:
1540:
1013:, lived during the same time as
955:or evolutionary grouping called
146:
2091:Delfino, M.; Smith, T. (2009).
718:In 1964 Chinese paleontologist
859:is too fragmentary to support
1:
2138:10.1016/S0034-6667(02)00103-3
624:"Asiatosuchus" depressifrons
2554:Paleopedia - List of species
2549:at the Paleobiology Database
1343:Brachyuranochampsa eversolei
708:is its lack of an overbite.
632:Sables du Castrais Formation
626:was first named in 1855 as
4854:
2502:10.1038/s42003-021-02017-0
1647:"Crocodylus" depressifrons
872:"Crocodylus" monsvialensis
4828:Paleocene crocodylomorphs
4523:
4503:
4272:
4161:
4133:
4000:
3880:
3791:
3711:
3696:
3683:
3663:
3406:
3251:
3041:
3037:
2990:
2975:
2962:
2757:
2743:
2689:
2662:
2649:
2597:
2414:10.1080/03115519608619189
2157:American Museum Novitates
1887:
1865:
1858:
1835:
1828:
1794:
1774:
1767:
1755:
1733:
1726:
1697:
1677:
1670:
1663:
1643:
1636:
1616:
1609:
1589:
1582:
1562:
1555:
1533:
1526:
1373:
1356:
1339:
1332:
1315:
1298:
1281:
1274:
1261:"Asiatosuchus" germanicus
1257:
1240:
1233:
1215:
1208:
1190:
1183:
1162:
1155:
1140:
1133:
993:"Asiatosuchus" germanicus
628:Crocodilus depressifrons.
561:"Asiatosuchus" germanicus
396:is a broad connection or
291:
286:
270:
263:
143:Scientific classification
141:
133:
124:
34:
27:Extinct genus of reptiles
3840:"Crocodylus" gariepensis
870:The crocodyloid species
863:being classified within
816:was later reassigned to
792:were named from Russia,
4732:Asiatosuchus germanicus
4113:"Tomistoma" lusitanicum
3886:"Crocodylus" megarhinus
3755:"Crocodylus" megarhinus
1778:"Crocodylus" megarhinus
1593:Asiatosuchus germanicus
1044:grouping, meaning that
722:named a new species of
698:supratemporal fenestrae
18:Asiatosuchus germanicus
4833:Eocene crocodylomorphs
2489:Communications Biology
2451:10.1098/rspb.2018.1071
2324:Vertebrata PalAsiatica
2212:Vertebrata PalAsiatica
941:
894:"Crocodylus" vicetinus
620:
575:
564:
4795:Paleobiology Database
4715:Paleobiology Database
4652:Asiatosuchus grangeri
4635:Paleobiology Database
2353:The Anatomical Record
2203:Young, C.-C. (1964).
2175:"New specimen of cf.
1302:Asiatosuchus grangeri
1244:Prodiplocynodon langi
1052:of successively more
939:
874:was named from Early
667:Asiatosuchus grangeri
615:
570:
559:
540:Asiatosuchus grangeri
532:Asiatosuchus grangeri
516:Irdin Manha Formation
276:Asiatosuchus grangeri
4388:"Tomistoma" coppensi
4380:"Tomistoma" cairense
4106:"Tomistoma" cairense
3747:"Crocodylus" affinis
2914:"Crocodylus" affinis
1620:"Crocodylus" affinis
1319:"Crocodylus" affinis
889:Diplocynodon ratelii
844:"Crocodylus" affinis
4395:"Tomistoma" dowsoni
2243:1970CaJES...7.1008W
2151:Mook, C.C. (1940).
2038:10.7717/peerj.12094
837:. In comparison to
757:Eoalligator chunyii
646:. The species name
3968:C. thorbjarnarsoni
3017:Listrognathosuchus
2893:Brachyuranochampsa
2285:10.7717/peerj.2356
1701:Brachyuranochampsa
1050:evolutionary grade
1017:, suggesting that
942:
917:Drazinda Formation
909:Sulaiman Mountains
900:, was assigned to
740:Guangdong Province
706:"A." depressifrons
686:"A." depressifrons
682:"A." depressifrons
673:was reassigned to
657:"A." depressifrons
621:
617:"A." depressifrons
607:"A." depressifrons
576:
565:
482:"A." depressifrons
429:"A." depressifrons
327:(Blainville, 1855)
324:"A." depressifrons
4810:
4809:
4782:Open Tree of Life
4702:Open Tree of Life
4622:Open Tree of Life
4551:Taxon identifiers
4542:
4541:
4519:
4518:
4501:
4500:
4497:
4496:
4493:
4492:
4489:
4488:
4255:
4254:
3985:
3984:
3981:
3980:
3977:
3976:
3940:C. anthropophagus
3679:
3678:
3661:
3660:
3657:
3656:
3653:
3652:
3649:
3648:
3645:
3644:
3393:
3392:
3191:
3190:
3154:Eurycephalosuchus
2958:
2957:
2954:
2953:
2907:"Crocodylus" acer
2739:
2738:
2735:
2734:
2731:
2730:
2727:
2726:
2360:(10): 2729–2765.
1997:
1996:
1988:
1987:
1979:
1978:
1970:
1969:
1961:
1960:
1952:
1951:
1943:
1942:
1934:
1933:
1925:
1924:
1916:
1915:
1907:
1906:
1812:
1811:
1715:
1714:
1681:"Crocodylus" acer
1449:
1448:
1440:
1439:
1431:
1430:
1422:
1421:
1413:
1412:
1404:
1403:
1395:
1394:
1386:
1385:
1360:"Crocodylus" acer
1097:A." depressifrons
1074:"A. depressifrons
951:as a member of a
849:Bridger Formation
808:were regarded as
766:"A." nanlingensis
732:Shanghu Formation
728:"A." nanlingensis
713:"A." nanlingensis
574:hind leg and tail
536:Walter W. Granger
362:
361:
356:
349:"A." oenotriensis
342:
335:"A." nanlingensis
328:
317:
306:
259:
16:(Redirected from
4845:
4803:
4802:
4790:
4789:
4777:
4776:
4764:
4763:
4751:
4750:
4749:
4723:
4722:
4710:
4709:
4697:
4696:
4684:
4683:
4671:
4670:
4669:
4643:
4642:
4630:
4629:
4617:
4616:
4604:
4603:
4591:
4590:
4578:
4577:
4576:
4546:
4514:
4509:
4471:G. pachyrhynchus
4373:Siquisiquesuchus
4270:
4159:
4131:
3998:
3878:
3789:
3709:
3698:
3685:
3674:
3669:
3636:C. wannlangstoni
3629:C. venezuelensis
3471:Globidentosuchus
3434:Centenariosuchus
3404:
3265:Allognathosuchus
3249:
3240:
3131:Orientalosuchina
3039:
3035:
2988:
2977:
2964:
2759:
2745:
2687:
2660:
2651:
2644:
2643:
2599:
2583:
2576:
2569:
2560:
2533:
2532:
2522:
2504:
2480:
2474:
2473:
2463:
2453:
2427:
2418:
2417:
2397:
2391:
2388:
2382:
2381:
2379:
2369:
2367:10.1002/ar.25005
2342:
2333:
2332:
2314:
2308:
2307:
2297:
2287:
2261:
2255:
2254:
2237:(3): 1008–1012.
2226:
2220:
2219:
2209:
2200:
2194:
2193:
2183:
2170:
2161:
2160:
2148:
2142:
2141:
2121:
2115:
2114:
2112:
2088:
2061:
2060:
2050:
2040:
2014:
1898:
1876:
1861:
1844:
1831:
1803:
1783:
1770:
1758:
1729:
1706:
1686:
1673:
1666:
1652:
1639:
1625:
1612:
1598:
1585:
1571:
1558:
1544:
1529:
1517:
1335:
1277:
1236:
1211:
1196:Pristichampsidae
1194:
1186:
1166:
1158:
1136:
1126:
1093:A." germanicus,
702:postorbital bone
671:C. depressifrons
644:C. depressifrons
351:
347:
337:
333:
326:
322:
315:
311:
300:
296:
273:
254:
247:
216:Archosauriformes
203:Archosauromorpha
151:
150:
129:
119:
56:
40:Temporal range:
32:
21:
4853:
4852:
4848:
4847:
4846:
4844:
4843:
4842:
4813:
4812:
4811:
4806:
4798:
4793:
4785:
4780:
4772:
4767:
4759:
4754:
4745:
4744:
4739:
4726:
4718:
4713:
4705:
4700:
4692:
4687:
4679:
4674:
4665:
4664:
4659:
4646:
4638:
4633:
4625:
4620:
4612:
4607:
4599:
4594:
4586:
4581:
4572:
4571:
4566:
4553:
4543:
4538:
4515:
4485:
4447:G. curvirostris
4424:G. bengawanicus
4407:
4402:Toyotamaphimeia
4322:Hesperogavialis
4263:
4251:
4204:
4152:
4144:
4127:
4118:
4070:Megadontosuchus
3994:
3973:
3961:C. palaeindicus
3923:
3867:
3818:
3797:Aldabrachampsus
3785:
3776:
3705:
3692:
3675:
3641:
3615:C. brevirostris
3591:
3562:
3556:Wannaganosuchus
3541:Stangerochampsa
3506:Orthogenysuchus
3449:Chinatichampsus
3389:
3325:
3319:Wannaganosuchus
3238:
3200:
3187:
3175:Orientalosuchus
3125:
3120:Stangerochampsa
3031:
3022:
2984:
2971:
2950:
2937:Prodiplocynodon
2921:Portugalosuchus
2850:
2753:
2723:
2703:
2676:
2645:
2604:
2603:
2593:
2587:
2542:
2537:
2536:
2482:
2481:
2477:
2429:
2428:
2421:
2399:
2398:
2394:
2389:
2385:
2344:
2343:
2336:
2316:
2315:
2311:
2263:
2262:
2258:
2251:10.1139/e70-093
2228:
2227:
2223:
2207:
2202:
2201:
2197:
2181:
2172:
2171:
2164:
2150:
2149:
2145:
2123:
2122:
2118:
2090:
2089:
2064:
2016:
2015:
2008:
2003:
1998:
1989:
1980:
1971:
1962:
1953:
1944:
1935:
1926:
1917:
1908:
1813:
1716:
1566:Prodiplocynodon
1450:
1441:
1432:
1423:
1414:
1405:
1396:
1387:
1070:"A." germanicus
1040:species form a
1027:"A." germanicus
1005:"A." germanicus
934:
788:Two species of
786:
752:A. nanlingensis
748:A. nanlingensis
742:, China. Small
716:
610:
600:"A." germanicus
572:"A." germanicus
554:
551:"A." germanicus
501:
494:
478:"A." germanicus
445:
425:"A." germanicus
313:"A." germanicus
282:
279:
253:
245:
145:
135:A. nanlingensis
120:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
52:61.6–41.2
51:
50:
49:
38:
28:
23:
22:
15:
12:
11:
5:
4851:
4849:
4841:
4840:
4835:
4830:
4825:
4815:
4814:
4808:
4807:
4805:
4804:
4791:
4778:
4765:
4752:
4736:
4734:
4728:
4727:
4725:
4724:
4711:
4698:
4685:
4672:
4656:
4654:
4648:
4647:
4645:
4644:
4631:
4618:
4605:
4592:
4579:
4563:
4561:
4555:
4554:
4549:
4540:
4539:
4537:
4536:
4531:
4524:
4521:
4520:
4517:
4516:
4504:
4502:
4499:
4498:
4495:
4494:
4491:
4490:
4487:
4486:
4484:
4483:
4475:
4467:
4459:
4451:
4443:
4435:
4427:
4419:
4417:
4409:
4408:
4406:
4405:
4398:
4391:
4384:
4376:
4369:
4362:
4355:
4348:
4340:
4336:Maomingosuchus
4332:
4325:
4318:
4314:Harpacochampsa
4310:
4303:
4296:
4288:
4281:
4273:
4267:
4257:
4256:
4253:
4252:
4250:
4249:
4241:
4237:T. lusitanicum
4233:
4225:
4221:T. calaritanum
4216:
4214:
4206:
4205:
4203:
4202:
4194:
4186:
4179:
4171:
4162:
4156:
4146:
4145:
4143:
4142:
4134:
4128:
4123:
4120:
4119:
4117:
4116:
4109:
4102:
4094:
4087:
4080:
4073:
4066:
4059:
4056:Maomingosuchus
4052:
4048:Leptorrhamphus
4044:
4037:
4030:
4023:
4016:
4009:
4006:Dollosuchoides
4001:
3995:
3990:
3987:
3986:
3983:
3982:
3979:
3978:
3975:
3974:
3972:
3971:
3964:
3957:
3954:C. falconensis
3950:
3943:
3935:
3933:
3925:
3924:
3922:
3921:
3914:
3906:
3898:
3890:
3881:
3875:
3869:
3868:
3866:
3865:
3857:
3850:
3843:
3836:
3828:
3826:
3820:
3819:
3817:
3816:
3808:
3804:Dzungarisuchus
3800:
3792:
3786:
3781:
3778:
3777:
3775:
3774:
3766:
3758:
3751:
3743:
3739:Australosuchus
3735:
3728:
3725:Antecrocodylus
3721:
3712:
3706:
3701:
3694:
3693:
3688:
3681:
3680:
3677:
3676:
3664:
3662:
3659:
3658:
3655:
3654:
3651:
3650:
3647:
3646:
3643:
3642:
3640:
3639:
3632:
3625:
3618:
3611:
3603:
3601:
3593:
3592:
3590:
3589:
3586:M. latrubessei
3582:
3574:
3572:
3564:
3563:
3561:
3560:
3552:
3545:
3537:
3530:
3523:
3516:
3509:
3502:
3495:
3488:
3481:
3474:
3467:
3459:
3452:
3445:
3437:
3430:
3422:
3415:
3407:
3401:
3395:
3394:
3391:
3390:
3388:
3387:
3380:
3373:
3366:
3359:
3352:
3345:
3337:
3335:
3327:
3326:
3324:
3323:
3315:
3311:Procaimanoidea
3307:
3299:
3295:Hassiacosuchus
3291:
3284:
3276:
3268:
3261:
3252:
3246:
3237:
3236:
3228:
3224:Menatalligator
3220:
3212:
3203:
3201:
3196:
3193:
3192:
3189:
3188:
3186:
3185:
3182:Protoalligator
3178:
3171:
3164:
3157:
3150:
3143:
3140:Dongnanosuchus
3135:
3133:
3127:
3126:
3124:
3123:
3116:
3112:Procaimanoidea
3108:
3101:
3093:
3090:Hassiacosuchus
3086:
3079:
3071:
3064:
3057:
3050:
3047:Albertochampsa
3042:
3032:
3027:
3024:
3023:
3021:
3020:
3013:
3006:
2999:
2991:
2985:
2982:Alligatoroidea
2980:
2973:
2972:
2969:Alligatoroidea
2967:
2960:
2959:
2956:
2955:
2952:
2951:
2949:
2948:
2944:Planocraniidae
2940:
2933:
2929:Pristichampsus
2925:
2917:
2910:
2903:
2900:Charactosuchus
2896:
2889:
2881:
2874:
2866:
2858:
2856:
2852:
2851:
2849:
2848:
2841:
2834:
2831:Trilophosuchus
2827:
2819:
2812:
2805:
2798:
2791:
2787:Harpacochampsa
2783:
2776:
2772:Australosuchus
2767:
2765:
2755:
2754:
2748:
2741:
2740:
2737:
2736:
2733:
2732:
2729:
2728:
2725:
2724:
2722:
2721:
2720:
2719:
2713:
2711:
2705:
2704:
2702:
2701:
2700:
2699:
2690:
2684:
2678:
2677:
2675:
2674:
2673:
2672:
2663:
2657:
2647:
2646:
2642:
2641:
2632:
2623:
2617:
2611:
2602:
2595:
2594:
2588:
2586:
2585:
2578:
2571:
2563:
2557:
2556:
2551:
2541:
2540:External links
2538:
2535:
2534:
2475:
2419:
2392:
2383:
2334:
2309:
2256:
2221:
2195:
2162:
2143:
2132:(1–2): 47–62.
2116:
2062:
2005:
2004:
2002:
1999:
1995:
1994:
1991:
1990:
1986:
1985:
1982:
1981:
1977:
1976:
1973:
1972:
1968:
1967:
1964:
1963:
1959:
1958:
1955:
1954:
1950:
1949:
1946:
1945:
1941:
1940:
1937:
1936:
1932:
1931:
1928:
1927:
1923:
1922:
1919:
1918:
1914:
1913:
1910:
1909:
1905:
1904:
1901:
1900:
1886:
1883:
1882:
1879:
1878:
1864:
1859:
1857:
1851:
1850:
1847:
1846:
1834:
1829:
1827:
1819:
1818:
1815:
1814:
1810:
1809:
1806:
1805:
1793:
1790:
1789:
1786:
1785:
1773:
1768:
1766:
1756:
1754:
1746:
1745:
1742:
1741:
1732:
1727:
1725:
1722:
1721:
1718:
1717:
1713:
1712:
1709:
1708:
1696:
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1692:
1689:
1688:
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1669:
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1659:
1658:
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1654:
1642:
1637:
1635:
1632:
1631:
1628:
1627:
1615:
1610:
1608:
1605:
1604:
1601:
1600:
1588:
1583:
1581:
1578:
1577:
1574:
1573:
1561:
1556:
1554:
1551:
1550:
1547:
1546:
1537:Alligatoroidea
1532:
1527:
1525:
1515:
1462:DNA sequencing
1447:
1446:
1443:
1442:
1438:
1437:
1434:
1433:
1429:
1428:
1425:
1424:
1420:
1419:
1416:
1415:
1411:
1410:
1407:
1406:
1402:
1401:
1398:
1397:
1393:
1392:
1389:
1388:
1384:
1383:
1380:
1379:
1372:
1369:
1368:
1365:
1364:
1355:
1352:
1351:
1348:
1347:
1338:
1333:
1331:
1328:
1327:
1324:
1323:
1314:
1311:
1310:
1307:
1306:
1297:
1294:
1293:
1290:
1289:
1280:
1275:
1273:
1270:
1269:
1266:
1265:
1256:
1253:
1252:
1249:
1248:
1239:
1234:
1232:
1226:
1225:
1222:
1221:
1218:Alligatoroidea
1214:
1209:
1207:
1204:
1203:
1200:
1199:
1189:
1184:
1182:
1179:
1178:
1175:
1174:
1161:
1156:
1154:
1151:
1150:
1147:
1146:
1139:
1134:
1132:
1124:
1068:. The species
1048:represents an
969:Alligatoroidea
933:
930:
785:
782:
720:Yang Zhongjian
715:
710:
609:
604:
584:A. germanicus.
553:
548:
500:
495:
493:
490:
470:splenial bones
444:
441:
421:type specimens
369:is an extinct
360:
359:
358:
357:
353:Narváez et al.
343:
329:
318:
307:
289:
288:
284:
283:
280:
268:
267:
261:
260:
243:
239:
238:
233:
229:
228:
223:
219:
218:
213:
206:
205:
200:
193:
192:
187:
183:
182:
177:
173:
172:
167:
163:
162:
157:
153:
152:
139:
138:
131:
130:
122:
121:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
57:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4850:
4839:
4836:
4834:
4831:
4829:
4826:
4824:
4821:
4820:
4818:
4801:
4796:
4792:
4788:
4783:
4779:
4775:
4770:
4766:
4762:
4757:
4753:
4748:
4742:
4738:
4737:
4735:
4733:
4729:
4721:
4716:
4712:
4708:
4703:
4699:
4695:
4690:
4686:
4682:
4677:
4673:
4668:
4662:
4658:
4657:
4655:
4653:
4649:
4641:
4636:
4632:
4628:
4623:
4619:
4615:
4610:
4606:
4602:
4597:
4593:
4589:
4584:
4580:
4575:
4569:
4565:
4564:
4562:
4560:
4556:
4552:
4547:
4535:
4534:Gryposuchinae
4532:
4530:
4526:
4525:
4522:
4513:
4508:
4481:
4480:
4476:
4473:
4472:
4468:
4465:
4464:
4460:
4457:
4456:
4452:
4449:
4448:
4444:
4441:
4440:
4436:
4433:
4432:
4428:
4426:
4425:
4421:
4420:
4418:
4416:
4415:
4410:
4404:
4403:
4399:
4397:
4396:
4392:
4390:
4389:
4385:
4382:
4381:
4377:
4375:
4374:
4370:
4368:
4367:
4366:Rhamphosuchus
4363:
4361:
4360:
4359:Piscogavialis
4356:
4354:
4353:
4349:
4346:
4345:
4344:Paratomistoma
4341:
4338:
4337:
4333:
4331:
4330:
4329:Ikanogavialis
4326:
4324:
4323:
4319:
4316:
4315:
4311:
4309:
4308:
4304:
4302:
4301:
4297:
4294:
4293:
4292:Gavialosuchus
4289:
4287:
4286:
4282:
4280:
4279:
4278:Aktiogavialis
4275:
4274:
4271:
4268:
4266:
4262:
4258:
4247:
4246:
4245:T. taiwanicum
4242:
4239:
4238:
4234:
4231:
4230:
4226:
4223:
4222:
4218:
4217:
4215:
4213:
4212:
4207:
4200:
4199:
4195:
4192:
4191:
4190:Paratomistoma
4187:
4185:
4184:
4180:
4177:
4176:
4175:Gavialosuchus
4172:
4169:
4168:
4167:Brasilosuchus
4164:
4163:
4160:
4157:
4155:
4154:sensu stricto
4151:
4147:
4141:
4140:
4136:
4135:
4132:
4129:
4126:
4121:
4115:
4114:
4110:
4108:
4107:
4103:
4100:
4099:
4095:
4093:
4092:
4088:
4086:
4085:
4084:Paratomistoma
4081:
4079:
4078:
4074:
4072:
4071:
4067:
4065:
4064:
4063:Maroccosuchus
4060:
4058:
4057:
4053:
4050:
4049:
4045:
4043:
4042:
4038:
4036:
4035:
4034:Gunggamarandu
4031:
4029:
4028:
4027:Gavialosuchus
4024:
4022:
4021:
4020:Ferganosuchus
4017:
4015:
4014:
4010:
4008:
4007:
4003:
4002:
3999:
3996:
3993:
3988:
3970:
3969:
3965:
3963:
3962:
3958:
3956:
3955:
3951:
3949:
3948:
3944:
3942:
3941:
3937:
3936:
3934:
3932:
3931:
3926:
3920:
3919:
3915:
3912:
3911:
3910:Tzaganosuchus
3907:
3904:
3903:
3899:
3896:
3895:
3891:
3888:
3887:
3883:
3882:
3879:
3876:
3874:
3870:
3863:
3862:
3858:
3856:
3855:
3851:
3849:
3848:
3844:
3842:
3841:
3837:
3835:
3834:
3830:
3829:
3827:
3825:
3824:Osteolaeminae
3821:
3814:
3813:
3812:Oxysdonsaurus
3809:
3806:
3805:
3801:
3799:
3798:
3794:
3793:
3790:
3787:
3784:
3779:
3772:
3771:
3767:
3764:
3763:
3762:Jiangxisuchus
3759:
3757:
3756:
3752:
3749:
3748:
3744:
3741:
3740:
3736:
3734:
3733:
3732:Astorgosuchus
3729:
3727:
3726:
3722:
3719:
3718:
3717:Albertosuchus
3714:
3713:
3710:
3707:
3704:
3703:Crocodyloidea
3699:
3695:
3691:
3686:
3682:
3673:
3668:
3638:
3637:
3633:
3631:
3630:
3626:
3624:
3623:
3622:C. gasparinae
3619:
3617:
3616:
3612:
3610:
3609:
3605:
3604:
3602:
3600:
3599:
3594:
3588:
3587:
3583:
3581:
3580:
3576:
3575:
3573:
3571:
3570:
3565:
3558:
3557:
3553:
3551:
3550:
3546:
3543:
3542:
3538:
3536:
3535:
3531:
3529:
3528:
3524:
3522:
3521:
3517:
3515:
3514:
3510:
3508:
3507:
3503:
3501:
3500:
3496:
3494:
3493:
3489:
3487:
3486:
3485:Kuttanacaiman
3482:
3480:
3479:
3475:
3473:
3472:
3468:
3465:
3464:
3460:
3458:
3457:
3456:Culebrasuchus
3453:
3451:
3450:
3446:
3443:
3442:
3438:
3436:
3435:
3431:
3428:
3427:
3426:Brachychampsa
3423:
3421:
3420:
3416:
3414:
3413:
3409:
3408:
3405:
3402:
3400:
3396:
3386:
3385:
3381:
3379:
3378:
3377:A. prenasalis
3374:
3372:
3371:
3367:
3365:
3364:
3360:
3358:
3357:
3353:
3351:
3350:
3346:
3344:
3343:
3339:
3338:
3336:
3334:
3333:
3328:
3321:
3320:
3316:
3313:
3312:
3308:
3305:
3304:
3300:
3297:
3296:
3292:
3290:
3289:
3288:Chrysochampsa
3285:
3282:
3281:
3277:
3274:
3273:
3269:
3267:
3266:
3262:
3259:
3258:
3257:Akanthosuchus
3254:
3253:
3250:
3247:
3245:
3244:Alligatorinae
3241:
3234:
3233:
3232:Sajkanosuchus
3229:
3226:
3225:
3221:
3218:
3217:
3216:Lianghusuchus
3213:
3210:
3209:
3205:
3204:
3202:
3199:
3198:Alligatoridae
3194:
3184:
3183:
3179:
3177:
3176:
3172:
3170:
3169:
3165:
3163:
3162:
3161:Jiangxisuchus
3158:
3156:
3155:
3151:
3149:
3148:
3144:
3142:
3141:
3137:
3136:
3134:
3132:
3128:
3122:
3121:
3117:
3114:
3113:
3109:
3107:
3106:
3102:
3099:
3098:
3094:
3092:
3091:
3087:
3085:
3084:
3080:
3077:
3076:
3072:
3070:
3069:
3065:
3063:
3062:
3061:Brachychampsa
3058:
3056:
3055:
3051:
3049:
3048:
3044:
3043:
3040:
3036:
3033:
3030:
3025:
3019:
3018:
3014:
3012:
3011:
3007:
3005:
3004:
3000:
2998:
2997:
2993:
2992:
2989:
2986:
2983:
2978:
2974:
2970:
2965:
2961:
2946:
2945:
2941:
2939:
2938:
2934:
2931:
2930:
2926:
2923:
2922:
2918:
2916:
2915:
2911:
2909:
2908:
2904:
2902:
2901:
2897:
2895:
2894:
2890:
2887:
2886:
2885:Borealosuchus
2882:
2880:
2879:
2875:
2872:
2871:
2867:
2865:
2864:
2863:Albertosuchus
2860:
2859:
2857:
2853:
2847:
2846:
2842:
2840:
2839:
2835:
2833:
2832:
2828:
2825:
2824:
2820:
2818:
2817:
2813:
2811:
2810:
2806:
2804:
2803:
2799:
2797:
2796:
2792:
2789:
2788:
2784:
2782:
2781:
2777:
2774:
2773:
2769:
2768:
2766:
2764:
2760:
2756:
2751:
2746:
2742:
2717:
2716:
2715:
2714:
2712:
2710:
2706:
2698:
2694:
2693:
2692:
2691:
2688:
2685:
2683:
2679:
2671:
2667:
2666:
2665:
2664:
2661:
2658:
2656:
2652:
2648:
2640:
2636:
2633:
2631:
2627:
2624:
2622:
2618:
2616:
2612:
2610:
2606:
2605:
2600:
2596:
2592:
2584:
2579:
2577:
2572:
2570:
2565:
2564:
2561:
2555:
2552:
2550:
2548:
2544:
2543:
2539:
2530:
2526:
2521:
2516:
2512:
2508:
2503:
2498:
2494:
2490:
2486:
2479:
2476:
2471:
2467:
2462:
2457:
2452:
2447:
2443:
2439:
2438:
2433:
2426:
2424:
2420:
2415:
2411:
2407:
2403:
2396:
2393:
2387:
2384:
2378:
2373:
2368:
2363:
2359:
2355:
2354:
2349:
2341:
2339:
2335:
2331:(2): 137–146.
2330:
2326:
2325:
2320:
2313:
2310:
2305:
2301:
2296:
2291:
2286:
2281:
2277:
2273:
2272:
2267:
2260:
2257:
2252:
2248:
2244:
2240:
2236:
2232:
2225:
2222:
2218:(2): 189–208.
2217:
2213:
2206:
2199:
2196:
2192:(5): 163–189.
2191:
2187:
2180:
2178:
2169:
2167:
2163:
2158:
2154:
2147:
2144:
2139:
2135:
2131:
2127:
2120:
2117:
2111:
2106:
2102:
2098:
2094:
2087:
2085:
2083:
2081:
2079:
2077:
2075:
2073:
2071:
2069:
2067:
2063:
2058:
2054:
2049:
2044:
2039:
2034:
2030:
2026:
2025:
2020:
2013:
2011:
2007:
2000:
1993:
1992:
1984:
1983:
1975:
1974:
1966:
1965:
1957:
1956:
1948:
1947:
1939:
1938:
1930:
1929:
1921:
1920:
1912:
1911:
1903:
1902:
1899:
1897:
1893:
1892:
1885:
1884:
1881:
1880:
1877:
1875:
1871:
1870:
1863:
1862:
1856:
1853:
1852:
1849:
1848:
1845:
1840:
1833:
1832:
1826:
1825:
1821:
1820:
1817:
1816:
1808:
1807:
1804:
1802:
1798:
1792:
1791:
1788:
1787:
1784:
1780:
1779:
1772:
1771:
1765:
1764:
1763:Crocodyloidea
1760:
1759:
1753:
1752:
1748:
1747:
1744:
1743:
1740:
1739:
1738:
1731:
1730:
1724:
1723:
1720:
1719:
1711:
1710:
1707:
1703:
1702:
1695:
1694:
1691:
1690:
1687:
1683:
1682:
1675:
1674:
1668:
1667:
1661:
1660:
1657:
1656:
1653:
1649:
1648:
1641:
1640:
1634:
1633:
1630:
1629:
1626:
1622:
1621:
1614:
1613:
1607:
1606:
1603:
1602:
1599:
1595:
1594:
1587:
1586:
1580:
1579:
1576:
1575:
1572:
1568:
1567:
1560:
1559:
1553:
1552:
1549:
1548:
1545:
1543:
1539:
1538:
1531:
1530:
1524:
1523:
1519:
1518:
1514:
1512:
1508:
1504:
1503:Crocodyloidea
1500:
1496:
1491:
1489:
1488:
1483:
1482:paleogenomics
1479:
1475:
1471:
1467:
1466:stratigraphic
1463:
1460:, molecular (
1459:
1458:morphological
1455:
1445:
1444:
1436:
1435:
1427:
1426:
1418:
1417:
1409:
1408:
1400:
1399:
1391:
1390:
1382:
1381:
1378:
1377:
1371:
1370:
1367:
1366:
1363:
1362:
1361:
1354:
1353:
1350:
1349:
1346:
1345:
1344:
1337:
1336:
1330:
1329:
1326:
1325:
1322:
1321:
1320:
1313:
1312:
1309:
1308:
1305:
1304:
1303:
1296:
1295:
1292:
1291:
1288:
1287:
1286:
1279:
1278:
1272:
1271:
1268:
1267:
1264:
1263:
1262:
1255:
1254:
1251:
1250:
1247:
1246:
1245:
1238:
1237:
1231:
1230:Crocodyloidea
1228:
1227:
1224:
1223:
1220:
1219:
1213:
1212:
1206:
1205:
1202:
1201:
1198:
1197:
1188:
1187:
1181:
1180:
1177:
1176:
1173:
1171:
1170:
1169:Borealosuchus
1160:
1159:
1153:
1152:
1149:
1148:
1145:
1144:
1138:
1137:
1131:
1128:
1127:
1123:
1122:
1121:"C." affinis.
1118:
1114:
1113:Asiatosuchus
1109:
1108:Asiatosuchus
1105:
1100:
1098:
1094:
1090:
1086:
1082:
1079:
1075:
1071:
1067:
1063:
1059:
1055:
1051:
1047:
1046:Asiatosuchus
1043:
1039:
1038:Asiatosuchus
1035:
1030:
1028:
1024:
1020:
1016:
1012:
1011:
1006:
1002:
998:
994:
989:
986:
982:
978:
974:
970:
966:
962:
958:
957:Crocodyloidea
954:
950:
946:
938:
931:
929:
926:
922:
918:
914:
910:
905:
903:
899:
895:
891:
890:
885:
881:
877:
873:
868:
866:
862:
858:
854:
850:
846:
845:
840:
836:
831:
829:
825:
822:, a genus of
821:
820:
815:
814:A. zajsanicus
811:
807:
806:A. zajsanicus
803:
799:
795:
794:A. zajsanicus
791:
784:Other species
783:
781:
779:
775:
771:
767:
763:
759:
758:
753:
749:
745:
741:
737:
733:
729:
725:
721:
714:
711:
709:
707:
703:
699:
695:
691:
687:
683:
678:
676:
672:
668:
664:
663:
658:
653:
649:
648:depressifrons
645:
641:
637:
633:
629:
625:
618:
614:
608:
605:
603:
601:
597:
593:
589:
585:
581:
573:
569:
562:
558:
552:
549:
547:
545:
541:
537:
533:
530:. Mook named
529:
525:
521:
520:Middle Eocene
517:
513:
509:
505:
499:
496:
491:
489:
487:
483:
479:
475:
471:
467:
466:dentary bones
462:
458:
454:
450:
442:
440:
438:
434:
430:
426:
422:
418:
417:
412:
408:
404:
399:
395:
391:
387:
383:
379:
376:
372:
368:
367:
354:
350:
344:
340:
336:
330:
325:
319:
314:
308:
304:
299:
293:
292:
290:
285:
278:
277:
269:
266:
262:
258:
252:
251:
244:
241:
240:
237:
236:Crocodyloidea
234:
232:Superfamily:
231:
230:
227:
224:
221:
220:
217:
214:
211:
208:
207:
204:
201:
198:
195:
194:
191:
188:
185:
184:
181:
178:
175:
174:
171:
168:
165:
164:
161:
158:
155:
154:
149:
144:
140:
136:
132:
128:
123:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
55:
47:
43:
37:
33:
30:
19:
4823:Crocodilians
4731:
4651:
4559:Asiatosuchus
4558:
4529:Brevirostres
4479:G. papuensis
4477:
4469:
4461:
4453:
4445:
4437:
4431:G. breviceps
4429:
4422:
4412:
4400:
4393:
4386:
4378:
4371:
4364:
4357:
4352:Penghusuchus
4350:
4342:
4334:
4327:
4320:
4312:
4305:
4298:
4290:
4285:Dadagavialis
4283:
4276:
4264:
4243:
4235:
4227:
4219:
4209:
4198:Thecachampsa
4196:
4188:
4183:Melitosaurus
4181:
4173:
4165:
4153:
4150:Tomistominae
4139:Sacacosuchus
4137:
4111:
4104:
4096:
4091:Thecachampsa
4089:
4082:
4075:
4068:
4061:
4054:
4046:
4039:
4032:
4025:
4018:
4011:
4004:
3966:
3959:
3952:
3947:C. checchiai
3945:
3938:
3928:
3916:
3908:
3900:
3892:
3884:
3873:Crocodylinae
3859:
3852:
3845:
3838:
3831:
3810:
3802:
3795:
3783:Crocodylidae
3770:Mekosuchinae
3768:
3760:
3753:
3745:
3737:
3730:
3723:
3715:
3690:Longirostres
3634:
3627:
3620:
3613:
3608:C. australis
3606:
3596:
3584:
3577:
3569:Melanosuchus
3567:
3554:
3547:
3539:
3532:
3525:
3520:Paranasuchus
3518:
3513:Paranacaiman
3511:
3504:
3497:
3490:
3483:
3476:
3469:
3461:
3454:
3447:
3441:Ceratosuchus
3439:
3432:
3424:
3417:
3410:
3382:
3375:
3368:
3361:
3354:
3347:
3342:A. hailensis
3340:
3330:
3317:
3309:
3303:Navajosuchus
3301:
3293:
3286:
3280:Ceratosuchus
3278:
3270:
3263:
3255:
3230:
3222:
3214:
3206:
3180:
3173:
3166:
3159:
3152:
3145:
3138:
3118:
3110:
3105:Navajosuchus
3103:
3097:Leidyosuchus
3095:
3088:
3081:
3073:
3068:Ceratosuchus
3066:
3059:
3052:
3045:
3015:
3010:Leidyosuchus
3008:
3003:Diplocynodon
3001:
2994:
2942:
2935:
2927:
2919:
2912:
2905:
2898:
2891:
2883:
2878:Asiatosuchus
2877:
2876:
2868:
2861:
2843:
2836:
2829:
2821:
2814:
2807:
2800:
2793:
2785:
2778:
2770:
2763:Mekosuchinae
2752:crocodilians
2670:Pseudosuchia
2655:Pseudosuchia
2634:
2630:Pseudosuchia
2625:
2591:crocodilians
2547:Asiatosuchus
2546:
2492:
2488:
2478:
2441:
2435:
2405:
2401:
2395:
2386:
2357:
2351:
2328:
2322:
2312:
2275:
2269:
2259:
2234:
2230:
2224:
2215:
2211:
2198:
2189:
2185:
2177:Asiatosuchus
2176:
2159:(1097): 1–3.
2156:
2146:
2129:
2125:
2119:
2100:
2096:
2028:
2022:
1889:
1888:
1867:
1866:
1836:
1822:
1797:Crocodylidae
1795:
1776:
1775:
1761:
1751:Longirostres
1749:
1737:Mekosuchinae
1735:
1734:
1699:
1698:
1679:
1678:
1646:
1645:
1644:
1618:
1617:
1592:
1591:
1590:
1564:
1563:
1535:
1534:
1520:
1511:Longirostres
1499:Asiatosuchus
1498:
1492:
1485:
1477:
1451:
1376:Crocodylidae
1374:
1358:
1357:
1341:
1340:
1317:
1316:
1301:
1300:
1299:
1284:
1283:
1282:
1260:
1259:
1258:
1242:
1241:
1216:
1191:
1167:
1163:
1141:
1120:
1117:A. grangeri
1116:
1112:
1107:
1101:
1096:
1092:
1088:
1085:Asiatosuchus
1084:
1081:
1078:Asiatosuchus
1077:
1073:
1069:
1066:Asiatosuchus
1065:
1061:
1058:monophyletic
1045:
1042:paraphyletic
1037:
1034:Asiatosuchus
1033:
1031:
1026:
1023:Asiatosuchus
1022:
1019:Asiatosuchus
1018:
1015:Asiatosuchus
1014:
1008:
1004:
1001:Mekosuchinae
997:sister taxon
992:
990:
984:
981:Asiatosuchus
980:
961:Asiatosuchus
960:
949:Asiatosuchus
948:
945:Phylogenetic
943:
925:Asiatosuchus
924:
913:Asiatosuchus
912:
906:
902:Asiatosuchus
901:
893:
887:
884:Asiatosuchus
883:
878:deposits in
871:
869:
865:Asiatosuchus
864:
861:"C." affinis
860:
856:
853:"C." affinis
852:
842:
838:
835:Asiatosuchus
834:
832:
830:as dubious.
827:
817:
813:
810:nomina dubia
809:
805:
802:A. volgensis
801:
798:A. volgensis
797:
796:in 1982 and
793:
790:Asiatosuchus
789:
787:
778:Crocodylidae
774:Mekosuchinae
770:"E." chunyii
769:
765:
755:
751:
747:
727:
724:Asiatosuchus
723:
717:
712:
705:
690:Asiatosuchus
689:
685:
681:
679:
675:Asiatosuchus
674:
670:
666:
660:
656:
652:frontal bone
647:
643:
627:
623:
622:
616:
606:
599:
596:Asiatosuchus
595:
592:anoxic lakes
583:
580:Asiatosuchus
579:
577:
571:
560:
550:
543:
539:
531:
512:Asiatosuchus
511:
508:type species
503:
502:
497:
486:Asiatosuchus
485:
481:
477:
474:Asiatosuchus
473:
461:Asiatosuchus
460:
453:Asiatosuchus
452:
449:Asiatosuchus
448:
446:
437:Asiatosuchus
436:
432:
428:
424:
416:nomina dubia
414:
406:
403:Asiatosuchus
402:
394:Asiatosuchus
393:
386:Asiatosuchus
385:
378:crocodilians
366:Asiatosuchus
365:
364:
363:
348:
334:
323:
312:
301:Mook, 1940 (
297:
275:
274:
265:Type species
250:Asiatosuchus
249:
248:
209:
196:
134:
36:Asiatosuchus
35:
29:
4455:G. leptodus
4307:Hanyusuchus
4300:Gryposuchus
4229:T. gaudense
4098:Tienosuchus
4077:Ocepesuchus
4041:Kentisuchus
4013:Dollosuchus
3992:Gavialoidea
3534:Purussaurus
3527:Protocaiman
3499:Necrosuchus
3492:Mourasuchus
3478:Gnatusuchus
3419:Bottosaurus
3384:A. thomsoni
3363:A. munensis
3356:A. mefferdi
3272:Arambourgia
3208:Balanerodus
3168:Krabisuchus
3147:Eoalligator
3075:Deinosuchus
3054:Arambourgia
2996:Deinosuchus
2870:Arenysuchus
2838:Ultrastenos
2795:Kalthifrons
2103:: 140–167.
1824:Gavialoidea
1501:outside of
1143:Gavialoidea
1102:Below is a
1089:A. grangeri
1062:A. grangeri
921:archaeocete
898:Monte Bolca
857:A. grangeri
839:A. grangeri
828:Dollosuchus
824:tomistomine
819:Dollosuchus
504:A. grangeri
498:A. grangeri
443:Description
433:A. grangeri
407:A. grangeri
375:crocodyloid
316:Berg, 1966
298:A. grangeri
4817:Categories
4527:See also:
4265:sensu lato
4261:Gavialinae
4125:Gavialidae
3930:Crocodylus
3854:Rimasuchus
3847:Euthecodon
3833:Brochuchus
3579:M. fisheri
3412:Acresuchus
3399:Caimaninae
3349:A. mcgrewi
3029:Globidonta
2809:Mekosuchus
2718:see below↓
2709:Crocodilia
2621:Sauropsida
2495:(1): 505.
2408:(3): 179.
2031:: e12094.
2001:References
1855:Gavialidae
1841:Gavialoids
1522:Crocodylia
1505:, as more
1493:The below
1474:Crocodilia
1454:tip dating
1130:Crocodylia
985:Crocodilus
973:alligators
880:Monteviale
762:synonymous
744:coprolites
694:jugal bone
662:Crocodylus
588:Messel Pit
544:Crocodylus
457:crocodiles
390:morphology
281:Mook, 1940
226:Crocodilia
137:specimens
4667:Q20717084
4463:G. lewisi
4439:G. browni
4211:Tomistoma
3549:Tsoabichi
3370:A. olseni
3332:Alligator
2816:Paludirex
2697:Neosuchia
2682:Neosuchia
2607:Kingdom:
2511:2399-3642
2278:: e2356.
1891:Tomistoma
1495:cladogram
1104:cladogram
932:Phylogeny
876:Oligocene
847:from the
800:in 1993.
640:pyritized
472:, but in
409:from the
398:symphysis
382:Paleogene
166:Kingdom:
160:Eukaryota
42:Paleocene
4761:47447179
4747:Q4806860
4741:Wikidata
4681:47447625
4661:Wikidata
4588:47447475
4574:Q3282861
4568:Wikidata
4414:Gavialis
3902:Quinkana
3463:Eocaiman
3083:Eocaiman
2823:Quinkana
2639:Eusuchia
2615:Chordata
2613:Phylum:
2609:Animalia
2589:Extinct
2529:33907305
2470:30051855
2444:(1881).
2304:27635329
2057:34567843
1869:Gavialis
1837:extinct
736:Nanxiong
468:and the
287:Species
190:Reptilia
180:Chordata
176:Phylum:
170:Animalia
156:Domain:
4787:6148548
4774:8634951
4707:3606026
4694:4967699
4627:3606027
4614:1242654
4601:4819398
3894:Kinyang
3861:Kinyang
2802:Kambara
2619:Class:
2520:8079395
2461:6030529
2377:9541231
2295:5012266
2239:Bibcode
2048:8428266
1464:), and
1452:A 2018
1054:derived
1010:Kambara
995:as the
977:caimans
528:Erenhot
492:Species
341:, 1964
242:Genus:
222:Order:
186:Class:
4800:194881
3598:Caiman
2855:Others
2527:
2517:
2509:
2468:
2458:
2374:
2302:
2292:
2055:
2045:
1480:using
1478:et al.
1470:fossil
1021:or an
636:France
534:after
506:, the
411:Eocene
355:, 2024
255:Mook,
46:Eocene
4720:97254
4640:38438
4609:IRMNG
2845:Volia
2750:Basal
2635:Clade
2626:Clade
2271:PeerJ
2208:(PDF)
2182:(PDF)
2024:PeerJ
1839:basal
1509:than
1507:basal
1172:spp.
1095:and "
965:basal
963:as a
953:clade
940:Snout
619:skull
563:skull
371:genus
339:Young
210:Clade
197:Clade
4769:GBIF
4689:GBIF
4596:GBIF
3918:Voay
2780:Baru
2695:see
2668:see
2525:PMID
2507:ISSN
2466:PMID
2300:PMID
2053:PMID
1487:Voay
1119:and
1072:and
804:and
768:and
480:and
427:and
303:type
257:1940
60:PreꞒ
4756:EoL
4676:EoL
4583:EoL
2515:PMC
2497:doi
2456:PMC
2446:doi
2442:285
2410:doi
2372:PMC
2362:doi
2358:305
2290:PMC
2280:doi
2247:doi
2134:doi
2130:122
2105:doi
2101:156
2043:PMC
2033:doi
1091:, "
734:in
510:of
373:of
4819::
4797::
4784::
4771::
4758::
4743::
4717::
4704::
4691::
4678::
4663::
4637::
4624::
4611::
4598::
4585::
4570::
2637::
2628::
2523:.
2513:.
2505:.
2491:.
2487:.
2464:.
2454:.
2440:.
2434:.
2422:^
2406:20
2404:.
2370:.
2356:.
2350:.
2337:^
2329:56
2327:.
2321:.
2298:.
2288:.
2274:.
2268:.
2245:.
2233:.
2210:.
2190:30
2188:.
2184:.
2165:^
2155:.
2128:.
2099:.
2095:.
2065:^
2051:.
2041:.
2027:.
2021:.
2009:^
1490:.
975:,
892:.
867:.
841:,
780:.
750:.
738:,
726:,
677:.
669:,
634:,
598:,
582:,
484:,
439:.
405:,
212::
199::
110:Pg
54:Ma
48:,
44:-
4482:?
4474:?
4466:?
4458:?
4450:?
4442:?
4434:?
4383:?
4347:?
4339:?
4317:?
4295:?
4248:?
4240:?
4232:?
4224:?
4201:?
4193:?
4178:?
4170:?
4101:?
4051:?
3913:?
3905:?
3897:?
3889:?
3864:?
3815:?
3807:?
3773:?
3765:?
3750:?
3742:?
3720:?
3559:?
3544:?
3466:?
3444:?
3429:?
3322:?
3314:?
3306:?
3298:?
3283:?
3275:?
3260:?
3235:?
3227:?
3219:?
3211:?
3115:?
3100:?
3078:?
2947:?
2932:?
2924:?
2888:?
2873:?
2826:?
2790:?
2775:?
2582:e
2575:t
2568:v
2531:.
2499::
2493:4
2472:.
2448::
2416:.
2412::
2380:.
2364::
2306:.
2282::
2276:4
2253:.
2249::
2241::
2235:7
2216:8
2140:.
2136::
2113:.
2107::
2059:.
2035::
2029:9
1843:†
1782:†
1705:†
1685:†
1651:†
1624:†
1597:†
1570:†
1468:(
1193:†
1165:†
346:†
332:†
321:†
310:†
305:)
295:†
272:†
246:†
115:N
105:K
100:J
95:T
90:P
85:C
80:D
75:S
70:O
65:Ꞓ
20:)
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