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analyzed independently by a panel of five forensic anthropologists, including Dr. Tim White, Dr. Alison
Galloway, Dr. Mark Griffin, Dr. P. Willey and Dr. Eric Bartelink. All five concluded that two of the larger fragments, erroneously identified in 1986 by the Viennese physicians Hans Bankl and Hans Jesserer as fragments of a human parietal, were fragments of a human frontal bone. As these fragments lacked Beethoven’s diagnostic craniotomy cut from his autopsy, it was unanimously agreed that the frontal bone fragments could not have originated from Beethoven. The portion of Beethoven’s frontal bone corresponding to the cranial fragments then under examination were furthermore described in considerable detail, and measured three times, during Beethoven’s second exhumation in 1888, twenty-five years after their purported theft in 1863. The authenticity status of the third large cranial element, the squamous of a human occipital, was not conclusively arrived at by this panel of forensic anthropologists. However, the squamous of Beethoven’s occipital is also listed among the cranial elements recovered during Beethoven’s second exhumation in 1888, and salient features of the squamous of Beethoven’s occipital (e.g. the curvature of the squamous on the horizontal plane; the overhang of the occipital) are likewise described in this same 1888 exhumation report.
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ongoing HBV infection at the time those hairs were forming, estimated to have been most likely in ca. 1826. Owing to the limited sensitivity of the HBV analyses, the authors could not determine the nature, period of onset, or origin of
Beethoven’s HBV infection, but suggest that a chronic infection was nonetheless more probable than an acute infection. They concluded that a lifelong infection would have been a strong driver of liver disease, no doubt exacerbated by alcohol consumption and genetic risk, while an infection later in life would have been of debatable relevance.
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164:, in his notebook, recorded HĂĽttenbrenner's account of Beethoven's death. HĂĽttenbrenner's eyewitness report is sometimes recast to imply that Beethoven "shook his fist at the heavens" in the moment before death. Since any imputations as to the dying man's emotional state are impossible to verify, they tend to be glossed over or ignored as irrelevant by modern Beethoven scholars.
285:. Beethoven's brain was described as possessing "exaggerated folds", an excess of fluid in the skull, and some thickening of the membranes inside the left ventricle. Scholars believe he may have had a degree of cerebral atrophy, although he showed no sign of cognitive impairment to the end. The skull was described as "possessing unusual thickness".
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toxicological analyses of the Hiller Lock, which had been taken to conclude that plumbism caused or contributed to
Beethoven’s health complaints, and that he had neither been administered opiates during the course of his final illness, nor mercury for a hypothesized infection with syphilis, no longer applied to Beethoven.
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into wine was a very common practice to sweeten cheap wines, and despite being outlawed in most
European countries during the 18th century, the prohibition was difficult to enforce and production of lead-fortified wine (which originated in Roman times) continued unabated. There is no indication the
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Among lines of evidence considered by some to be consistent with lead poisoning, the finding of shrunken cochlear nerves at his autopsy is consistent with axonal degeneration due to heavy metals such as lead. Chronic low-level lead exposure causes a slowly progressive hearing loss with sensory and
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In 2010, Dr. Andrew C. Todd tested two skull fragments purportedly to have been stolen during
Beethoven’s first exhumation in 1863 for lead, and determined that the concentration of lead was no greater than would be expected for a normal man of 56 at that time. In 2012, these skull fragments were
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as "shrunken and fibrous", with the exit duct being very thin and narrowed. Large amounts of reddish fluid had accumulated in
Beethoven's abdomen, likely from spontaneous bacterial infections mixed with some blood. This was possibly a result of draining fluid from his abdomen in his last days, a
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was sequenced using a targeted hybridization capture technique, and was assigned to the D2 subgenotype. Only the
Stumpff Lock, cut after Beethoven’s death and plausibly representing a growth interval many months before his death, was positive for HBV. The authors concluded that Beethoven had an
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cemetery, northwest of Vienna. Many thousands of citizens lined the streets for the funeral procession. As with all crowds, estimates vary, with witnesses reporting anywhere from 10,000 to 30,000 onlookers. Theaters were closed, and many notable artists participated in the funeral procession as
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mitochondrial haplogroup, common among individuals of
Ashkenazi Jewish matrilineal ancestry. The authors reported the lowest possible final mitochondrial contamination estimate of 0-2% for the Hiller Lock, using the algorithm Schmutzi. The study authors therefore concluded that the results of
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The authors concluded that, had
Beethoven’s alcohol consumption been sufficiently heavy over a long enough period of time, its interaction with his heritable risk factors may constitute a plausible causal explanation for his liver disease. The authors cautioned, however, that the extent of
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and killed him. Reiter's hypothesis however is at odds with Dr. Wawruch's written instruction "that the wound was kept dry all the time". Furthermore, human hair is a very bad biomarker for lead contamination, and Reiter's hypothesis must be considered dubious as long as proper scholarly
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Kocher, Arthur; Papac, Luka; Barquera, Rodrigo; Key, Felix M.; Spyrou, Maria A.; Hübler, Ron; Rohrlach, Adam B.; Aron, Franziska; Stahl, Raphaela; Wissgott, Antje; van Bömmel, Florian; Pfefferkorn, Maria; Mittnik, Alissa; Villalba-Mouco, Vanessa; Neumann, Gunnar U. (8 October 2021).
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Begg, Tristan James
Alexander; Schmidt, Axel; Kocher, Arthur; Larmuseau, Maarten H.D.; Runfeldt, Göran; Maier, Paul Andrew; Wilson, John D.; Barquera, Rodrigo; Maj, Carlo; Szolek, András; Sager, Michael; Clayton, Stephen; Peltzer, Alexander; Hui, Ruoyun; Ronge, Julia (April 2023).
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At this startling, awful peal of thunder, the dying man suddenly raised his head from Hüttenbrenner's arm, stretched out his own right arm majestically—like a general giving orders to an army. This was but for an instant; the arm sunk back; he fell back; Beethoven was dead.
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Beethoven’s alcohol consumption, and thus the extent of this harmful interaction, remain uncertain. The authors also concluded that, despite Beethoven being compound heterozygous for type-1 hereditary haemochromatosis, he remained at a low overall risk for an
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Bycroft, Clare; Freeman, Colin; Petkova, Desislava; Band, Gavin; Elliott, Lloyd T.; Sharp, Kevin; Motyer, Allan; Vukcevic, Damjan; Delaneau, Olivier; O’Connell, Jared; Cortes, Adrian; Welsh, Samantha; Young, Alan; Effingham, Mark; McVean, Gil (October 2018).
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Christian Reiter asserted that Beethoven's doctor, Andreas Wawruch, accidentally killed him by giving him an overdose of a lead-based cure. According to Reiter, Wawruch used the cure to alleviate fluid in the abdomen; the lead penetrated Beethoven's
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HĂĽttenbrenner's account has been used to ascribe motivations of resistance and anger to Beethoven in his final moments. Beethoven's last words and the exact cause of his death have also been the subject of some historical debate with causes such as
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Emdin, Connor A.; Haas, Mary; Ajmera, Veeral; Simon, Tracey G.; Homburger, Julian; Neben, Cynthia; Jiang, Lan; Wei, Wei-Qi; Feng, Qiping; Zhou, Alicia; Denny, Joshua; Corey, Kathleen; Loomba, Rohit; Kathiresan, Sekar; Khera, Amit V. (April 2021).
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Heavy metal contamination is thought to be a contributing factor in Beethoven's death as these were commonly used in medicines of the time. It has also been theorized that he consumed large amounts of lead from illegally fortified wine. Putting
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Beethoven suffered declining health throughout the last years of his life, including the so-called "Late period" when he produced some of his most admired work. The last work he was able to complete was the substitute final movement of the
328:. One of Beethoven's friends incorrectly thought that "strangers had cut all of his hair off"; in fact, the apparent lack of hair was due to a cloth cap that covered most of the hair while the body was lying in state.
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In the days following the funeral, one of the grave-diggers was reportedly offered a substantial sum of money to remove the head from the grave. As a result, Beethoven's friends had a watch put on the grave.
589:, with approximately 20% of these men having a primary or secondary diagnosis of some form of cirrhosis at the time they were recruited as UK BioBank participants (aged 40 to 69, average 56.74 for men).
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genome sequenced from a lock of hair which the study’s authors considered “almost certainly authentic” revealed several genetic risk factors for liver disease, as well as evidence for an infection with
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The autopsy indicated damage to his aural nerves as well as hardening of their accompanying arteries, although the latter appears to be consistent with natural aging and not inflammatory damage from
120:, Karl Holz and some friends were probably in the house, HĂĽttenbrenner reports in his account given in 1860, that only he and Beethoven's sister-in-law were present in the room at the time of death.
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on the other hand can be contracted from food and water that were not handled properly and was very common in the 19th century, although it does not cause liver cirrhosis or permanent organ damage.
56:, who provided a vivid description of the event. Beethoven's funeral was held three days later, and the procession was witnessed by a large crowd. He was originally buried in the cemetery at
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of progressive liver disease or cirrhosis among those men in the general population, a substantial interaction effect was observed among those men who had also been diagnosed with an
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As it became apparent that Beethoven would not recover, his friends gathered to help and to pay their final respects. Beethoven's doctors conducted four minor operations to relieve
1326:"Systematic review with meta-analysis: the I148M variant of patatin-like phospholipase domain-containing 3 gene (PNPLA3) is significantly associated with alcoholic liver cirrhosis"
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1261:"Association Between Patatin-Like Phospholipase Domain Containing 3 Gene (PNPLA3) Polymorphisms and Nonalcoholic Fatty Liver Disease: A HuGE Review and Meta-Analysis"
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were taken. The body was washed, clothed and placed in an oaken coffin, with the head given a wreath of white roses. Beethoven's hands held a wax cross and a lily.
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is a common consequence. Scholars disagree over whether Beethoven's liver damage was the result of heavy alcohol consumption, hepatic infection, or both.
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157:; this was specifically denied by HĂĽttenbrenner in 1860. Another invention is that his last words were, "I shall hear in heaven," apropos his deafness.
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Beethoven's last recorded words were "Pity, pity—too late!", as the dying composer was told of a gift of twelve bottles of wine from his publisher,
48:, Vienna, on 26 March 1827 at the age of 56, following a prolonged illness. His death was witnessed by his sister-in-law, possibly by his secretary
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In the same 2023 study, the Hiller Lock, elsewhere referred to as the Guevara Lock, was revealed to have originated from a woman, assigned to the
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In 1863 Beethoven's body (and also that of Schubert, who was buried nearby) was exhumed, studied and reburied, in proceedings paid for by the
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practice that frequently caused infection and often death of the patient in a time before antibiotics and bacterial pathology were known.
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was performed on 27 March 1827 by Dr. Johann Wagner. While it is unclear who ordered the autopsy, a specific request by Beethoven in his
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416:. At that time, fragments from the back of his skull, which had been separated during the autopsy, were acquired by the Austrian doctor
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composer had syphilis beyond a mercury treatment prescribed to him around 1815, but these were used for various other ailments as well.
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101:, which was published separately as Opus 133. Shortly thereafter, in late 1826, illness struck again, with episodes of vomiting and
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are causes of cirrhosis, but they spread from contact with contaminated body fluids and were extremely rare in Beethoven's day .
344:, wrote a post-mortem analysis. This post-mortem became well known by being referenced by a short comical essay by the humorist
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1378:"Association of Genetic Variation With Cirrhosis: A Multi-Trait Genome-Wide Association and Gene–Environment Interaction Study"
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116:, and on 26 March he slipped into unconsciousness and died early that evening. While others, including Beethoven's brother
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for cirrhosis was in agreement with these findings, placing him in the 96th polygenic risk percentile, and conferring an
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Chamorro, A.-J.; Torres, J.-L.; Mirón-Canelo, J.-A.; González-Sarmiento, R.; Laso, F.-J.; Marcos, M. (September 2014).
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292:(RPN). Diabetes is also a cause of RPN, and scholars have not ruled out the possibility that the composer had
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for the most robustly associated common risk genotype for the full spectrum of progressive liver disease, at
112:(abdominal swelling), of which the first resulted in infection, the others not. On 24 March he was given his
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autonomic findings, rather than the classic wrist drop due to motor neuropathy from sub-acute poisoning.
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may have played a role in the decision. The autopsy revealed a severely cirrhotic and shrunken
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Beethoven's kidneys had calcareous growths in them, indicating that he was likely developing
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In the days immediately preceding and following his death, a number of people, including
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Lewis, Dyani (22 March 2023). "Beethoven's cause of death revealed from locks of hair".
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Please help update this article to reflect recent events or newly available information.
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Xu, Renfan; Tao, Anyu; Zhang, Shasha; Deng, Youbin; Chen, Guangzhi (20 March 2015).
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A lock of Beethoven's hair mounted in a jewel-encrusted case with a glass covering
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Renaud, Gabriel; Slon, Viviane; Duggan, Ana T.; Kelso, Janet (12 October 2015).
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Meredith, William Rhea (2005). "The History of Beethoven's Skull Fragments".
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who shared Beethoven’s genetic risk factors, did not reveal a high
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Eisinger, Josef (1 January 2008). "The lead in Beethoven's hair".
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1170:(1st ed.). Westport, CT: Greenwood Press. pp. 112–115.
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There is dispute about the cause of Beethoven's death; alcoholic
1436:"The UK Biobank resource with deep phenotyping and genomic data"
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875:"Review: Chocolate and Cuckoo clocks: The Essential Alan Coren"
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The funeral was held on 29 March 1827 at the parish church in
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1068:"Beethoven May Not Have Died of Lead Poisoning, After All"
1698:. Vol. 3 (2nd ed.). The Beethoven Association.
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Laryngologie, Rhinologie, Otologie und Ihre Grenzgebiete
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entitled "Careful, Mr. Beethoven, that was your fifth!"
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In 1970, Dr. John Spencer Madden, editor of the journal
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Journal of the Royal College of Physicians of Edinburgh
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Beethoven in person: his deafness, illnesses and death
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Madden, John Spencer (1970). "Ludwig van Beethoven".
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In the same 2023 genomic study, a 1.26-fold coverage
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to that conferred by his PNPLA3 risk genotype. While
1194:"Genomic analyses of hair from Ludwig van Beethoven"
393:. At a memorial mass in a Vienna church on 3 April,
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Beethoven with the Manuscript of the Missa Solemnis
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1092:"Ertaubte Beethoven an einer Pagetschen Krankheit?"
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44:, a German composer, died in his apartment in the
145:. One common belief was that his last words were
60:, although his remains were moved in 1888 to the
300:was swollen to twice the normal size and he had
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1502:"Ten millennia of hepatitis B virus evolution"
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449:. The reason given is: Article should mention
2109:Ira F. Brilliant Center for Beethoven Studies
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577:performed in that study, using men from the
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218:Learn how and when to remove this message
1090:Jesserer, H.; Bankl, H. (October 1986).
966:"Beethoven's terminal illness and death"
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939:
928:"Ludwig van Beethoven's Funeral Oration"
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520:In March of 2023, genomic analyses of a
373:pallbearers or torch bearers, including
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999:"Pathologist: Doctor Killed Beethoven"
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857:10.1093/oxfordjournals.alcalc.a045900
148:"Plaudite, amici, comedia finita est"
136:Thayer's summary of Beethoven's death
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200:adding citations to reliable sources
97:, composed to replace the difficult
509:documentation remains unpublished.
33:Beethoven's funeral as depicted by
2099:British première of Symphony No. 9
673:Strickland, Ashley (11 May 2024).
25:
1137:William Meredith (24 July 2015).
496:have all been proposed. In 2008,
331:On 28 March 1827, castings for a
88:Plaque at SchwarzspanierstraĂźe 15
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2060:Birthplace of Beethoven's mother
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1787:Relationship with contemporaries
1696:The Life of Ludwig van Beethoven
873:Moss, Peter (27 November 2008).
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168:Autopsy and post-mortem findings
997:Jahn, George (28 August 2007).
308:, as the doctors described his
187:needs additional citations for
1643:. Cambridge University Press.
806:Ritson, Bruce (12 June 2012).
95:String Quartet No. 13, Op. 130
1:
964:Mai, F. M. (1 October 2006).
575:retrospective cohort analyses
451:2024 results of hair analysis
414:Gesellschaft der Musikfreunde
1718:Center for Beethoven Studies
1394:10.1053/j.gastro.2020.12.011
397:was sung with an additional
105:that nearly ended his life.
2187:Deaths by person in Austria
1814:Concert of 22 December 1808
1635:Gibbs, Christopher (2000).
368:, and he was buried in the
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1924:New York and San Francisco
1680:Thayer, Alexander Wheelock
652:10.1038/d41586-023-00846-z
52:, and by his close friend
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2094:Beethoven Quartet Society
1839:Relationship with C minor
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1722:San Jose State University
1592:10.1186/s13059-015-0776-0
1460:10.1038/s41586-018-0579-z
1229:10067/1974860151162165141
1219:10.1016/j.cub.2023.02.041
1166:Davies, Peter J. (2001).
1143:San Jose State University
1047:10.1080/02772240701630588
738:History Channel on Foxtel
445:This article needs to be
326:San Jose State University
162:Alexander Wheelock Thayer
911:" Johann Nepomuk Hummel"
808:"Obituary: J. S. Madden"
602:hepatitis B virus genome
290:renal papillary necrosis
1804:Heiligenstadt Testament
1526:10.1126/science.abi5658
1139:"Update on skull bones"
916:Encyclopædia Britannica
862:(subscription required)
422:Vienna Central Cemetery
247:Heiligenstadt Testament
62:Vienna Central Cemetery
2089:Beethoven Project Trio
1684:Krehbiel, Henry Edward
1108:10.1055/s-2007-1008044
844:Alcohol and Alcoholism
813:Alcohol and Alcoholism
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595:iron overload disorder
387:Klemens von Metternich
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341:Alcohol and Alcoholism
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18:Beethoven's liver
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543:compound heterozygous
375:Johann Nepomuk Hummel
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160:Beethoven biographer
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2202:Ludwig van Beethoven
2070:Beethoven quadrangle
1854:Compositional method
1768:List of compositions
1759:Ludwig van Beethoven
1639:The Life of Schubert
1388:(5): 1620–1633.e13.
1204:(8): 1431–1447.e22.
1141:. Beethoven Center,
880:The Jewish Chronicle
587:alcohol use disorder
482:infectious hepatitis
381:who wrote a eulogy,
196:improve this article
54:Anselm HĂĽttenbrenner
42:Ludwig van Beethoven
2197:Funerals in Austria
2065:Beethoven Peninsula
1518:2021Sci...374..182K
1452:2018Natur.562..203B
1277:2015NatSR...5E9284X
1210:2023CBio...33E1431B
1039:2008TxEC...90....1E
1004:The Washington Post
954:, pp. 5–6, 17.
302:portal hypertension
2192:Funerals by person
1942:Mähler's portraits
1904:List of sculptures
1871:Biamonti Catalogue
1265:Scientific Reports
1077:The New York Times
622:
403:Ignaz von Seyfried
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352:Funeral and burial
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154:Commedia dell'arte
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46:Schwarzspanierhaus
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35:Franz Xaver Stöber
2212:March 1827 events
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1979:Copying Beethoven
1844:Collected edition
1660:Beethoven Journal
1512:(6564): 182–188.
1446:(7726): 203–209.
1343:10.1111/apt.12890
1285:10.1038/srep09284
930:, lvbeethoven.com
527:hepatitis B virus
494:Whipple's disease
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379:Franz Grillparzer
360:Beethoven's grave
294:diabetes mellitus
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1963:Immortal Beloved
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749:
747:
745:
734:"Beethoven Dies"
730:
724:
718:
712:
706:
700:
694:
683:
682:
670:
664:
663:
639:
529:. Beethoven was
465:
462:
456:
440:
439:
432:
395:Mozart's Requiem
322:Ferdinand Hiller
223:
216:
212:
209:
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150:
137:
76:being theories.
21:
2227:
2226:
2222:
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2216:
2177:
2176:
2175:
2171:Classical music
2165:
2163:
2153:
2151:
2143:
2141:
2136:
2118:
2079:Beethoven Burst
2043:
2034:(sister-in-law)
1990:
1930:
1899:Beethoven House
1877:
1827:
1818:
1770:
1761:
1756:
1714:
1678:
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1258:
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1198:Current Biology
1190:
1189:
1185:
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1165:
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1147:
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1136:
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1131:
1102:(10): 592–597.
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719:
715:
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695:
686:
672:
671:
667:
641:
640:
636:
632:
627:
563:polygenic score
466:
460:
457:
454:
441:
437:
430:
418:Romeo Seligmann
354:
318:Anton Schindler
236:Josef Danhauser
224:
213:
207:
204:
193:
181:
170:
139:
135:
126:
82:
23:
22:
15:
12:
11:
5:
2225:
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2215:
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2207:1827 in Europe
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2106:
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2062:
2057:
2055:1815 Beethoven
2051:
2049:
2045:
2044:
2042:
2041:
2035:
2029:
2023:
2017:
2011:
2005:
1998:
1996:
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1967:
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1932:
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1928:
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1926:
1921:
1916:
1911:
1901:
1896:
1894:Beethovenhalle
1891:
1889:Beethoven Hall
1885:
1883:
1879:
1878:
1876:
1875:
1874:
1873:
1868:
1861:Works catalogs
1858:
1857:
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1713:
1712:External links
1710:
1709:
1708:
1676:
1655:
1649:
1630:
1627:
1625:
1624:
1579:Genome Biology
1565:
1491:
1425:
1367:
1336:(6): 571–581.
1316:
1251:
1183:
1177:978-0313315879
1176:
1155:
1129:
1082:
1060:
1017:
989:
978:(3): 258–263.
956:
944:
932:
920:
903:
901:, p. 139.
886:
865:
851:(3): 101–103.
833:
795:
778:
763:
751:
740:. 19 June 2016
725:
723:, p. 306.
713:
711:, p. 307.
701:
684:
665:
633:
631:
628:
626:
623:
561:. Beethoven’s
486:lead poisoning
468:
467:
444:
442:
435:
429:
426:
391:Franz Schubert
353:
350:
234:Death mask by
226:
225:
184:
182:
175:
169:
166:
127:
125:
122:
81:
78:
74:lead poisoning
24:
14:
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2050:
2046:
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2036:
2033:
2030:
2028:(grandfather)
2027:
2024:
2021:
2018:
2015:
2012:
2009:
2006:
2003:
2000:
1999:
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1849:Musical style
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1692:Riemann, Hugo
1689:
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1666:(1–2): 3–46.
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1650:0-521-59426-X
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1101:
1098:(in German).
1097:
1093:
1086:
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1080:, 28 May 2010
1079:
1078:
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952:Meredith 2005
948:
945:
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940:Meredith 2005
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933:
929:
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823:
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791:Meredith 2005
787:
785:
783:
779:
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774:Meredith 2005
770:
768:
764:
760:
759:Meredith 2005
755:
752:
739:
735:
729:
726:
722:
717:
714:
710:
705:
702:
699:, p. 308
698:
693:
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689:
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676:
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556:
552:
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544:
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532:
528:
523:
522:high-coverage
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307:
303:
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284:
279:
276:
275:sugar of lead
270:
268:
264:
260:
256:
252:
248:
244:
237:
232:
222:
219:
211:
201:
197:
191:
190:
185:This section
183:
179:
174:
173:
167:
165:
163:
158:
156:
155:
149:
144:
138:
132:
123:
121:
119:
115:
111:
106:
104:
100:
96:
86:
80:Final illness
79:
77:
75:
71:
70:liver disease
65:
63:
59:
55:
51:
47:
43:
36:
31:
27:
19:
1977:
1969:
1961:
1953:
1946:
1798:
1695:
1663:
1659:
1638:
1582:
1578:
1568:
1535:10261/268494
1509:
1505:
1494:
1443:
1439:
1428:
1385:
1381:
1370:
1353:10366/154068
1333:
1329:
1319:
1268:
1264:
1254:
1201:
1197:
1186:
1167:
1146:. Retrieved
1132:
1099:
1095:
1085:
1075:
1072:James Barron
1063:
1030:
1026:
1020:
1008:. Retrieved
1002:
992:
975:
969:
959:
947:
942:, p. 4.
935:
923:
914:
906:
878:
868:
848:
842:
836:
817:
811:
761:, p. 1.
754:
742:. Retrieved
737:
728:
716:
704:
678:
668:
643:
637:
607:
599:
591:
519:
515:
511:
471:
458:
446:
411:
407:
363:
339:
337:
330:
315:
306:pancreatitis
287:
280:
271:
240:
214:
205:
194:Please help
189:verification
186:
159:
152:
143:Schott Music
140:
134:
129:
107:
91:
66:
40:
26:
1986:Other films
1982:(2006 film)
1974:(2003 film)
1966:(1994 film)
1958:(1949 film)
1919:Mexico City
1271:(1): 9284.
1010:29 December
793:, p. 3
776:, p. 2
721:Thayer 1921
709:Thayer 1921
697:Thayer 1921
571:effect-size
569:similar in
549:mutations,
539:PNPLA3 gene
501:pathologist
490:sarcoidosis
383:Carl Czerny
267:Hepatitis A
259:Hepatitis B
253:, of which
208:August 2021
124:Final words
37:(1795–1858)
2181:Categories
1935:Depictions
1782:Birthplace
1585:(1): 224.
1033:(1): 1–5.
899:Gibbs 2000
820:(4): 369.
744:16 October
625:References
583:prevalence
579:UK BioBank
567:odds ratio
541:, and was
531:homozygous
366:Alsergrund
346:Alan Coren
333:death mask
114:last rites
99:GroĂźe Fuge
2159:Biography
2022:(brother)
2016:(brother)
1882:Memorials
1601:1474-760X
1560:238475573
1544:0036-8075
1468:1476-4687
1402:0016-5085
1293:2045-2322
1238:0960-9822
1116:0935-8943
557:, in the
555:rs1800562
551:rs1799945
474:cirrhosis
399:Libera me
50:Karl Holz
2132:Category
2114:Three Bs
2040:(nephew)
2010:(mother)
2004:(father)
1694:(1921).
1672:64392567
1619:26458810
1552:34618559
1486:30305743
1420:33310085
1362:25060292
1311:25791171
1246:36958333
1124:57055511
1055:95824344
984:17214130
660:36949134
559:HFE gene
545:for two
535:rs738409
498:Austrian
478:syphilis
461:May 2024
428:Theories
310:pancreas
283:syphilis
103:diarrhea
2145:Portals
2048:Related
2032:Johanna
1629:Sources
1610:4601135
1514:Bibcode
1506:Science
1477:6786975
1448:Bibcode
1411:8035329
1302:4366950
1273:Bibcode
1206:Bibcode
1035:Bibcode
610:K1a1b1a
537:in the
447:updated
370:Währing
255:ascites
243:autopsy
110:ascites
58:Währing
2026:Ludwig
2020:Johann
2014:Kaspar
2002:Johann
1995:Family
1971:Eroica
1955:Eroica
1909:Berlin
1792:Mozart
1704:422583
1702:
1670:
1647:
1617:
1607:
1599:
1558:
1550:
1542:
1484:
1474:
1466:
1440:Nature
1418:
1408:
1400:
1360:
1309:
1299:
1291:
1244:
1236:
1174:
1122:
1114:
1053:
982:
658:
644:Nature
492:, and
298:spleen
296:. His
118:Johann
2008:Maria
1832:Music
1799:Death
1556:S2CID
1148:7 May
1120:S2CID
1051:S2CID
630:Notes
506:liver
251:liver
2038:Karl
1914:Bonn
1775:Life
1700:OCLC
1668:OCLC
1645:ISBN
1615:PMID
1597:ISSN
1548:PMID
1540:ISSN
1482:PMID
1464:ISSN
1416:PMID
1398:ISSN
1358:PMID
1307:PMID
1289:ISSN
1242:PMID
1234:ISSN
1172:ISBN
1150:2020
1112:ISSN
1012:2008
980:PMID
746:2018
656:PMID
553:and
389:and
320:and
261:and
72:and
1866:WoO
1720:at
1605:PMC
1587:doi
1530:hdl
1522:doi
1510:374
1472:PMC
1456:doi
1444:562
1406:PMC
1390:doi
1386:160
1348:hdl
1338:doi
1297:PMC
1281:doi
1224:hdl
1214:doi
1104:doi
1070:by
1043:doi
853:doi
822:doi
679:CNN
648:doi
401:by
385:,
241:An
198:by
2183::
2072:/
1690:;
1686:;
1682:;
1664:20
1662:.
1613:.
1603:.
1595:.
1583:16
1581:.
1577:.
1554:.
1546:.
1538:.
1528:.
1520:.
1508:.
1504:.
1480:.
1470:.
1462:.
1454:.
1442:.
1438:.
1414:.
1404:.
1396:.
1384:.
1380:.
1356:.
1346:.
1334:40
1332:.
1328:.
1305:.
1295:.
1287:.
1279:.
1267:.
1263:.
1240:.
1232:.
1222:.
1212:.
1202:33
1200:.
1196:.
1158:^
1118:.
1110:.
1100:65
1094:.
1074:,
1049:.
1041:.
1031:90
1029:.
1001:.
976:36
974:.
968:.
913:,
889:^
877:.
847:.
818:47
816:.
810:.
798:^
781:^
766:^
736:.
687:^
677:.
654:.
646:.
488:,
484:,
480:,
476:,
424:.
405:.
377:,
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2147::
1751:e
1744:t
1737:v
1706:.
1674:.
1653:.
1621:.
1589::
1562:.
1532::
1524::
1516::
1488:.
1458::
1450::
1422:.
1392::
1364:.
1350::
1340::
1313:.
1283::
1275::
1269:5
1248:.
1226::
1216::
1208::
1180:.
1152:.
1126:.
1106::
1057:.
1045::
1037::
1014:.
986:.
883:.
859:.
855::
849:5
830:.
824::
748:.
681:.
662:.
650::
463:)
459:(
453:.
263:C
221:)
215:(
210:)
206:(
192:.
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
