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Beluga (sturgeon)

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780:, co-evolved with the formation of the Danube valley, resulting in very different survival strategies in its early life stages. This appears to explain why different individuals of the same long-migratory species spawn as far upstream as 1,700 km (1,100 mi) upstream, while others spawn just 100 km (62 mi). To make the long journey to very distant spawning grounds, the sturgeon adapted a two-stage migration strategy, beginning in autumn when they enter the Danube River overwinter in the river and the second stage is their spawning which takes place in the spring the river in fall and staying over winter in reaches of the river offering adequate substrate and water-flow resting conditions. Very few locations of existing wintering and spawning grounds for sturgeon are presently known in the lower Danube, and none are known to exist in the river's upper reaches. The same situation concerns nursery sites upon which young sturgeon depend during their journey to the 2640: 119: 788:
historic spawning routes. The female lays her eggs on gravel from 4 to 40 m (13 to 131 ft) underwater. Upon hatching, the embryo are 11 to 14 mm (0.43 to 0.55 in) long, and 10–14 days later when they absorb their yolk sack, the length is 18 to 20 mm (0.71 to 0.79 in). Thereafter, the larvae usually subsist on benthic invertebrates, but when reaching at least 10 cm (3.9 in) in length, they will switch to a fish-based diet. While swimming back to the ocean, the young sturgeon may cover up to 60 km (37 mi) a day.
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4 in)); and weight (2,000 kg (4,400 lb), 2,500 kg (5,500 lb), or even 3,000 kg (6,600 lb)) are disputed and unconfirmed; but they are not impossible. Several other records of aged sturgeon exceed 5 m (16 ft 5 in). Among sturgeons, only the closely related
1122:, became the first and only facility in the world to obtain a permit exemption for the sale of beluga sturgeon and its caviar in the U.S. Since 2017, the company has assisted in beluga sturgeon repopulation efforts across the world by providing over 160,000 fertilized eggs to the Caspian Sea region. 787:
Males attain sexual maturity at 12–16 years of age, whereas females do so at 16–22 years. They will spawn every four to seven years. At one time, beluga sturgeons could migrate up to 1,000 km (620 mi) upriver to spawn, but dams in almost every major tributary that they utilize have impeded
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Pollution from various sources, including oil, industries, sewage, and agriculture, is a critical threat to the beluga sturgeon. The species' long lifespan makes it vulnerable to pesticide contamination, resulting in reduced reproductive success and other health issues. Pollution negatively affects
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Adults are heavy-set, spindle-shaped, large and humpbacked. The head is massive with a very large protractile mouth that gradually moves in an almost frontal position during growth. The snout is quite short (one-third to one-quarter of the head), and scutes gradually undergo absorption and decrease
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The dorsal fin has 48 to 81 soft rays, and the anal fin, much shorter, has 22 to 41 soft rays. There are five in a series of longitudinal scutes: dorsal (one series, 9–17 scutes), lateral (two series, one per side, 28–60 scutes each) and ventral series (two series, one per side, 7–14 scutes each).
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estuary at 1,571 kg (3,463 lb) and 7.2 m (23 ft 7 in). Another specimen reportedly weighed 1,220 kg (2,690 lb) and measured 6.1 m (20 ft 0 in) in length. Claims about greater length (10 m (32 ft 10 in), or even 12 m (39 ft
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The beluga sturgeon faces a significant threat from illegal catches for its meat and caviar. The excessive harvesting and a sharp increase in poaching have removed the largest and most mature specimens from the population, almost eliminating natural reproduction. This exploitation has pushed the
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Beluga of such great sizes are very old (continuing to grow throughout life) and have become increasingly rare in recent decades because of heavy fishing of the species. Today, mature belugas that are caught are generally 142–328 cm (4 ft 8 in – 10 ft 9 in) long and
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The construction of dams, such as the Iron Gate in the Danube and the Volgograd Dam, has severely reduced the beluga sturgeon's available spawning grounds. These dams have blocked access to crucial river habitats, leading to a significant reduction in the species' ability to reproduce. Similar
745: 1018:, and all rivers interconnecting these waterways. Unfortunately, this range has been greatly reduced in modern times to the Caspian Sea, the Black Sea, and a few rivers such as the Danube, with attempts to reintroduce Belugas into various historic locations. 1117:
Management of sturgeon fisheries within the Caspian Sea began in the 1950s and while the initial regulations had honorable intentions, they achieved dwindling effects due to the ever present demand for the fish's caviar. In July 2016, Sturgeon Aquafarms in
832:. Different diets have been observed throughout the distribution range of beluga sturgeon, as well as according to spawners' migration stage. Adults mainly eat a great diversity of large fish (73% of the diet). Additional food items may include 1067:. Due to aforementioned poaching of the sturgeon, the Danube is the only river remaining with naturally reproducing sturgeon populations within the European Union. The sturgeon remains a protected species listed in Appendix III of the 759:
Due to rampant overfishing, the average lifespan of beluga sturgeon is unknown, with no specimens living past their 56th year. However, the species is reportedly quite long-lived, being capable of surviving over 100 years in the wild.
1011:, capable of moving freely between freshwater and estuaries, and thus can live in waters of varying saline content. Sturgeons are quite a durable species and can survive some of the most altered and polluted rivers in the world. 736:
weigh 19–264 kg (42–582 lb). The female beluga is typically 20% larger than the male. An exceptionally large beluga recently caught weighed 960 kg (2,120 lb) and measured 3.4 m (11 ft 2 in).
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is conical and contains numerous sensory pits on both ventral and dorsal surfaces. The mouth is large, crescent-shaped and protractile, with the upper lip continuous while the lower lip is interrupted by a large gap. The
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The beluga sturgeon is confronted by several critical threats that imperil its existence. Illegal harvesting, habitat disruption through dam construction, and pollution are among the most pressing challenges faced.
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Choudhury, Anindo; Dick, Terry A. (1998). "Special Paper: The Historical Biogeography of Sturgeons (Osteichthyes: Acipenseridae): A Synthesis of Phylogenetics, Palaeontology and Palaeogeography".
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on clean, hard substrate, which offers both support and cover to their sticky and adhesive eggs. Spawning biology and development of larval stages of the sturgeon, the most ancient fish of the
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Evolution of the Danube ichthyofauna, its recent state and an attempt to predict further changes after the construction of the planned hydroelectric power-stations and diversion schemes
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in number with age. Colouring is blue-grey or dark brown, with silver or grey flancs and white belly. The dark dorsum contrasts strongly against the rest of the body;
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Juveniles are slender, and the head is quite narrow with a mouth ventrally placed but projecting upward. The snout is thin and pointed (almost half of the head),
561: 4535: 3951: 3148: 747: 2688: 1628:. pp. 295–344. In: J. Holcík (ed.) The Freshwater Fishes of Europe, Vol. I/II: General Introduction of Fishes, Acipenseriformes, AULA-Verlag, Wiesbaden 3693: 1086:
has banned imports of beluga caviar and other beluga products from the Caspian Sea since 6 October 2005, after listing beluga sturgeon under the U.S.
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Suciu, M; Paraschiv, M; Ene, C; Suciu, R (2005). "Downstream migration of young of the year beluga sturgeons (Huso huso) in the lower Danube River".
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are laterally compressed with foliate appendages, arranged in two pairs, originating midway or closer to the mouth than to the tip of the snout.
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invertebrates in rivers and shallow coastal waters, where they grow quickly. At the length of 8–10 cm (3.1–3.9 in), they become largely
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Little is known about the diet of the extinct Adriatic population. It has been reported that in marine and brackish environments, adult Adriatic
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Khodorevskaya R.P., Polyaninova A.A., Geraskin P.P., Romanov, A.A. (1995). "A study on physiological and biochemical status of Beluga sturgeon,
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PARCO LOMBARDO DELLA VALLE DEL TICINO, Sviluppo sostenibile, tutela della biodiversità e dell’ambiente, qualità della vita (28 March 2019).
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Since 2015, an official captive breeding scheme has been established in Italy, with beluga from the Azov Sea. Then, after the building of a
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is considered a delicacy worldwide. The flesh of the beluga is not particularly renowned, but it is a hearty white meat similar to that of
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predator whose local distribution is not influenced by the nature of the substrates, unlike with most of the sturgeons that show
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population is strongly protected under Appendix II of the Bern Convention, prohibiting any intentional killing of these fish.
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habitat loss has occurred in other rivers due to dam construction, greatly impacting the species' survival.
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The piscivorous diet of beluga sturgeon tends to change with age: in the Caspian Sea, it mainly consists of
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the quality of the sturgeon's habitat, compounding the challenges faced by this endangered species.
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Dalia Onara, Radu Suciu, Marian Paraschiv, Marian Iani, Daniela Holostenco, Elena Tafla (2011).
4592: 4475: 2445: 2417: 2306: 1106:, on 2019 hundreds of young microchipped beluga and 60 tagged subadults were released into the 1030:
The caviar of a beluga sturgeon, the main reason for the massive overharvesting of this species
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Annex II of the Convention on the Conservation of European Wildlife and Natural Habitats
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crowdfunding appeal, over 7,000 three month-old beluga sturgeons were released into the
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were released in the Po river, attempting to resurrect the extinct Adriatic population.
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Historically, beluga sturgeon were found in the Caspian Sea, Black Sea, Adriatic Sea,
4650: 4509: 4243: 4146: 4060: 3880: 3855: 3850: 3806: 3801: 3756: 3751: 3731: 3603: 3382: 3301: 3063: 2950: 2890: 2854: 2791: 2591: 2497: 1818: 1250: 1034: 887: 862: 816: 725: 693: 685: 549: 518: 298: 154: 69: 1223: 4361: 4136: 3960: 3875: 3835: 3736: 3678: 3618: 3545: 3540: 3530: 3446: 3436: 3365: 3348: 3255: 3216: 3209: 2729: 2477: 2438: 2424: 2380: 1834:"Azioni preparatorie alla reintroduzione di Huso huso nel fiume Ticino (2015-2016)" 968: 841: 812: 732:) can attain similar size, with a maximum weight of 1,000 kg (2,200 lb). 619: 577: 537: 20: 4325: 1538: 1568: 4618: 4527: 4457: 4305: 4119: 4107: 4033: 3791: 3786: 3761: 3741: 3683: 3578: 3473: 3451: 3441: 3414: 3284: 3236: 2735: 2154: 2132: 2106: 2063: 2024: 1814:"Beluga Sturgeon Threatened With Extinction, Yet Caviar Quotas Remain Unchanged" 1099: 1045: 1015: 948: 853: 837: 829: 677: 646:
However, during growth, the beluga sturgeons show evident morphologic changes:
553: 529: 44: 37: 1137:, were caught trying to smuggle a 140 kilogram, 2.5-metre beluga sturgeon in a 1026: 4169: 3936: 3870: 3860: 3781: 3771: 3456: 3424: 3409: 3355: 3328: 3311: 2723: 2465: 2292: 2271: 1107: 1008: 996: 988: 820: 769: 700:). It is the largest freshwater fish in the world. The beluga also rivals the 504: 89: 54: 4296: 1788:"Thousands of Critically Endangered Beluga Sturgeon Released into the Danube" 3926: 3911: 3896: 3726: 3387: 3333: 3318: 3296: 3279: 3226: 3097: 2755: 2554: 2387: 1049: 1038: 992: 984: 904: 781: 692:
sp.) as the most massive fish and is the second-longest bony fish after the
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shows typical characteristics of other sturgeon, such as an elongated body,
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The Decline of the Beluga Sturgeon: A Case Study about Fisheries Management
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are evident, back and flancs are dark grey or black and the belly is white.
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Proceedings of the 5th International Symposium on Sturgeons, 9-13 May 2005
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Threatened fishes of the world: Huso huso (Linnaeus, 1758)(Acipenseridae)
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Very old specimens are stocky, with a large head and an enormous mouth.
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attitude. The prey is sucked into the mouth opening extremely quickly.
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The largest accepted record is of a female taken in 1827 in the
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Filippov, G.M. (1976). "Some data on the biology of the beluga
1475:(1962). (Israel Program for Scientific Translations). Jerusalem 4065: 3633: 1993: 1919:"WWF: Sturgeon poaching again on the rise in Romania, Ukraine" 545: 684:
Among all extant bony fishes, the beluga sturgeon rivals the
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Freshwater fishes of the USSR and adjacent countries. Vol. 1
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and rivers of the same area, migrating spawners eat various
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Pirogovskii, M.I., L.I. Sokolov & V.P. Vasil’ev. 1989.
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International Wildlife Encyclopedia: Index volume (Vol. 22)
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sp. for juveniles smaller than 40 cm, different species of
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Comparison between an adult (above) and a juvenile (below)
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The surface of the skin is covered by fine denticles. The
1849:"Huso Huso: riparte da Pavia il ripopolamento nel Ticino" 474: 3126: 576:
The common name for the sturgeon, as for the unrelated
1952:"Redlist - Huso Huso (Beluga) threats to the species" 591: 2597: 2587: 2570: 2560: 2530: 2520: 2510: 2493: 2483: 2473: 2405: 2368: 2353: 2342: 2252: 2236: 2225: 2214: 2204: 2193: 2182: 2171: 716:) for the title of largest actively predatory fish. 468: 459: 4280: 4157: 4090: 3997: 3959: 3950: 3889: 3820: 3707: 3659: 3569: 3494: 3254: 3164: 3095: 3071: 3062: 3016: 2972: 2939: 2753: 2744: 2544: 2464: 2455: 2404: 2367: 2327: 2251: 2152: 2105: 2078: 1711:
General Introduction to Fishes and Acipenseriformes
1537:Chakrabarty, Prosanta (2003). Fink, William (ed.). 1509:. 2011. Archived from the original on 21 July 2012. 1141:; the fish was later safely returned to the river. 755:
A sub-adult beluga sturgeon swimming in an aquarium
465: 1392:. Environmental Biology of Fishes, 65(3), 363-365. 1388:Vecsei, P., Sucui, R., & Peterson, D. (2002). 1773:. New York: P. F. Collier & Son Company. 1688:from the south-eastern part of the Caspian sea". 861:for fish ranging between 40 and 280 cm and then 626:skeleton, naked skin and longitudinal series of 552:, wild populations have been greatly reduced by 1265:10.2305/IUCN.UK.2022-1.RLTS.T10269A135087846.en 1240:Gessner, J.; Chebanov, M.; Freyhof, J. (2022). 1620: 1618: 1616: 542:third-most-massive living species of bony fish 3142: 2682: 2009: 1709:Freshwater fishes of Europe (Vol I part II). 1235: 1233: 680:(National Museum of Tatarstan, Kazan, Russia) 585: 8: 1869:: CS1 maint: multiple names: authors list ( 1669:: CS1 maint: multiple names: authors list ( 1605:: CS1 maint: multiple names: authors list ( 1573:. Ramsar, Iran: Blackwell. pp. 306–308. 676:A 1000-kg, 4.17-m-long beluga fish from the 19:For the whale species of the same name, see 4657:IUCN Red List critically endangered species 1639:The sturgeons and paddlefishes of the world 1637:Hochleithner, M. & J. Gessner. (2001). 1440:The Guinness Book of Animal Facts and Feats 544:. Heavily fished for the female's valuable 4268: 3956: 3149: 3135: 3127: 3068: 2750: 2689: 2675: 2667: 2461: 2335: 2164: 2160: 2149: 2084: 2075: 2032: 2016: 2002: 1994: 871:sp. and other sturgeons for the largest. 143: 117: 27: 3694:Tradeoffs for locomotion in air and water 1370:Fishbase.org. Accessed on 11 January 2008 1263: 1488:. Biologicke prace 13: 1–121, 24 plates. 1974:Integrated Taxonomic Information System 1434: 1432: 1430: 1186: 1084:United States Fish and Wildlife Service 1862: 1662: 1598: 1512: 16:Species of fish in the sturgeon family 1946: 1944: 1942: 1940: 1782: 1780: 1732:Larissa J. Graham and Brian R. Murphy 1723: 1721: 1532: 1530: 1406:. Calderini, Bologna. pp. 75–85. 971:) and fish belonging to the families 7: 4606:F0350B4A-28EF-D113-CF6E-7BDF2E822606 4424:dcdc2790-5aa4-49c9-b3c5-dec30867d1c9 1458:Burton, M., & Burton, R. (2002) 1415: 1413: 1384: 1382: 1380: 1378: 1376: 1159:species to the brink of extinction. 1071:, and its trade is restricted under 4250: 3521:Electroreception and electrogenesis 1707:Sokolov L.I., in Holcík J. (1989). 1251:IUCN Red List of Threatened Species 768:Like most sturgeons, the beluga is 540:. Based on maximum size, it is the 1761:Reynolds, Francis J., ed. (1921). 1208:10.1046/j.1365-2699.1998.2540623.x 804:eating a small fish in an aquarium 772:, migrating upstream in rivers to 14: 1657:Proc. Intern. Sturg. Symp. Moscow 840:, aquatic birds and young seals ( 4249: 4238: 4237: 4220: 4219: 3245: 2651: 2645: 2638: 1641:. Aquatech Publications. 202 pp. 886:In brackish environments of the 455: 185: 3205:Environmental impact of fishing 1655:(L.), and its feeding habits". 1442:. Sterling Pub Co Inc. (1983), 1404:Fauna d'Italia, X: Osteichthyes 1348:Deeplyfish- fishes of the world 1154:Illegal harvesting and poaching 1007:Beluga sturgeon are considered 967:are particularly common in the 947:foraged primarily on molluscs ( 528:. It is found primarily in the 412:Sal'nikov & Malyatskii 1934 400:Sal'nikov & Malyatskii 1934 388:Sal'nikov & Malyatskii 1934 380:Sal'nikov & Malyatskii 1934 1311:Froese, R.; Pauly, D. (2017). 1: 1750:. Food Fancy. September 2012. 3290:intramembranous ossification 536:basins, and formerly in the 4732:Taxa named by Carl Linnaeus 908:are the preferred prey. In 592: 564:to classify the species as 4748: 3669:Fin and flipper locomotion 3639:Sequential hermaphroditism 3526:Jamming avoidance response 3243: 1770:Collier's New Encyclopedia 1626:Huso huso (Linnaeus, 1758) 18: 4702:Fish of Georgia (country) 4233: 2706: 2635: 2338: 2167: 2163: 2148: 2129: 2087: 2074: 2031: 1714:. Wiesbaden: Aula Verlag. 1519:: CS1 maint: unfit URL ( 1063:classifies the beluga as 1048:is said to make the best 586: 317: 310: 287: 280: 182:Scientific classification 180: 163: 141: 132: 125: 116: 30: 4712:Fish of the Adriatic Sea 2866:A. oxyrinchus oxyrinchus 4722:Fish of the Caspian Sea 4185:Glossary of ichthyology 3747:Diel vertical migration 2918:European sea sturgeon ( 2699:sturgeon and paddlefish 1990:. Revised 1 March 2002. 1196:Journal of Biogeography 882:eating a small sturgeon 714:Somniosus microcephalus 4727:Fish described in 1758 3551:Surface wave detection 3516:Hydrodynamic reception 3190:Diseases and parasites 1507:caspianenvironment.org 1258:: e.T10269A135087846. 1088:Endangered Species Act 1031: 934:and, among sturgeons, 883: 805: 756: 706:Carcharodon carcharias 681: 612: 580:, is derived from the 4717:Fish of the Black Sea 4588:Paleobiology Database 3689:Undulatory locomotion 3506:Ampullae of Lorenzini 3083:American paddlefish ( 3037:Shovelnose sturgeon ( 2857:A. oxyrinchus desotoi 1462:. Marshall Cavendish. 1402:Tortonese E. (1970). 1065:critically endangered 1029: 877: 799: 754: 675: 607: 566:critically endangered 485:), also known as the 369:Acipenser vallisnerii 150:Critically Endangered 4692:Fish of Central Asia 4419:Fauna Europaea (new) 3917:Genetically modified 3107:Chinese paddlefish ( 3004:Amu Darya sturgeon ( 2986:Syr Darya sturgeon ( 2774:Shortnose sturgeon ( 2609:Pseudoscaphirhynchus 1547:Animal Diversity Web 1484:Balon, E.K. (1967). 1287:"Appendices | CITES" 1094:Repopulation efforts 437:Huso huso orientalis 3722:Aquatic respiration 3609:Life history theory 2864:Atlantic sturgeon ( 2837:Adriatic sturgeon ( 2828:Japanese sturgeon ( 2819:Sakhalin sturgeon ( 2765:Siberian sturgeon ( 2602:Protoscaphirhynchus 1887:. 20 December 2019. 1792:World Wildlife Fund 1163:Habitat impoundment 1127:World Wildlife Fund 377:Huso huso maeoticus 135:Conservation status 4682:Fish of Azerbaijan 3561:Weberian apparatus 3046:Alabama sturgeon ( 2900:Chinese sturgeon ( 2873:Persian sturgeon ( 2846:Bastard sturgeon ( 2803:A. gueldenstaedtii 2801:Russian sturgeon ( 2783:Dabry's sturgeon ( 2219:Bobasatraniiformes 1853:ticinobiosource.it 1104:Isola Serafini dam 1032: 957:Sepia officinalis, 940:is the main prey. 937:Acipenser ruthenus 890:basin, the genera 884: 824:Juveniles feed on 806: 757: 682: 613: 425:Huso huso caspicus 417:Huso huso caspicus 405:Huso huso ponticus 393:Huso huso ponticus 385:Huso huso ponticus 361:Huso ichthyocollus 4642: 4641: 4575:Open Tree of Life 4274:Taxon identifiers 4265: 4264: 4175:Fish common names 4086: 4085: 3717:Aquatic predation 3541:Capacity for pain 3270:Age determination 3124: 3123: 3120: 3119: 3058: 3057: 3028:Pallid sturgeon ( 2909:Starry sturgeon ( 2664: 2663: 2660: 2659: 2633: 2632: 2629: 2628: 2625: 2624: 2323: 2322: 2241:Saurichthyiformes 2230:Guildayichthyidae 2144: 2143: 2140: 2139: 2125: 2124: 1659:. VNIRO: 164–172. 1448:978-0-85112-235-9 1320:version (02/2017) 1075:Appendix II. The 953:common cuttlefish 931:Sander lucioperca 752: 702:great white shark 447: 446: 441: 433: 421: 413: 401: 389: 381: 373: 365: 357: 353:Acipenser albulus 349: 341: 333: 329:Acipenser belugus 325: 175: 158: 4739: 4635: 4634: 4622: 4621: 4609: 4608: 4596: 4595: 4583: 4582: 4570: 4569: 4557: 4556: 4544: 4543: 4531: 4530: 4528:NHMSYS0001691430 4518: 4517: 4505: 4504: 4492: 4491: 4479: 4478: 4466: 4465: 4453: 4452: 4440: 4439: 4427: 4426: 4414: 4413: 4401: 4400: 4388: 4387: 4378: 4377: 4365: 4364: 4352: 4351: 4339: 4338: 4329: 4328: 4316: 4315: 4314: 4301: 4300: 4299: 4269: 4253: 4252: 4241: 4240: 4223: 4222: 3957: 3249: 3180:Ethnoichthyology 3151: 3144: 3137: 3128: 3069: 2995:Dwarf sturgeon ( 2929:A. transmontanus 2927:White sturgeon ( 2830:A. multiscutatus 2810:Green sturgeon ( 2751: 2691: 2684: 2677: 2668: 2655: 2649: 2643: 2642: 2599: 2589: 2572: 2562: 2532: 2522: 2512: 2495: 2485: 2475: 2462: 2407: 2370: 2355: 2344: 2336: 2330:Acipenseriformes 2300:Plesiococcolepis 2254: 2238: 2227: 2216: 2206: 2195: 2184: 2173: 2165: 2161: 2150: 2085: 2076: 2069: 2068: 2033: 2018: 2011: 2004: 1995: 1984: 1982: 1980: 1956: 1955: 1948: 1935: 1934: 1932: 1930: 1915: 1909: 1908: 1907:. 19 April 2019. 1895: 1889: 1888: 1881: 1875: 1874: 1868: 1860: 1844: 1838: 1837: 1830: 1824: 1823: 1810: 1804: 1803: 1801: 1799: 1784: 1775: 1774: 1766: 1758: 1752: 1751: 1740: 1734: 1725: 1716: 1715: 1704: 1698: 1697: 1681: 1675: 1674: 1668: 1660: 1648: 1642: 1635: 1629: 1622: 1611: 1610: 1604: 1596: 1590: 1581: 1575: 1574: 1564: 1558: 1557: 1555: 1553: 1534: 1525: 1524: 1518: 1510: 1495: 1489: 1482: 1476: 1469: 1463: 1456: 1450: 1436: 1425: 1424: 1417: 1408: 1407: 1399: 1393: 1386: 1371: 1365: 1359: 1358: 1356: 1354: 1345: 1337: 1331: 1330: 1328: 1326: 1308: 1302: 1301: 1299: 1297: 1283: 1277: 1276: 1274: 1272: 1267: 1237: 1228: 1227: 1191: 1135:Grindu, Ialomita 965:Loligo vulgaris, 753: 698:Regalecus glesne 622:tail, partially 595: 589: 588: 526:Acipenseriformes 484: 483: 480: 479: 476: 473: 470: 467: 464: 461: 439: 431: 419: 411: 399: 387: 379: 371: 363: 355: 347: 345:Acipenser schypa 339: 331: 323: 293: 239:Acipenseriformes 190: 189: 169: 152: 147: 146: 121: 111: 41: 34:Temporal range: 28: 4747: 4746: 4742: 4741: 4740: 4738: 4737: 4736: 4662:Commercial fish 4647: 4646: 4643: 4638: 4630: 4625: 4617: 4612: 4604: 4599: 4591: 4586: 4578: 4573: 4565: 4560: 4552: 4549:Observation.org 4547: 4539: 4534: 4526: 4521: 4513: 4508: 4500: 4495: 4487: 4482: 4474: 4469: 4461: 4456: 4448: 4443: 4435: 4430: 4422: 4417: 4409: 4404: 4396: 4391: 4383: 4381: 4373: 4368: 4360: 4355: 4347: 4342: 4334: 4332: 4324: 4319: 4310: 4309: 4304: 4295: 4294: 4289: 4276: 4266: 4261: 4229: 4153: 4082: 3993: 3946: 3885: 3816: 3709: 3703: 3655: 3599:Ichthyoplankton 3565: 3497: 3490: 3486:Digital Library 3481:Teleost leptins 3420:Shark cartilage 3344:pharyngeal slit 3339:pharyngeal arch 3275:Anguilliformity 3260: 3258: 3250: 3241: 3160: 3155: 3125: 3116: 3091: 3054: 3039:S. platorynchus 3012: 2988:P. fedtschenkoi 2976: 2968: 2935: 2891:Amur sturgeon ( 2855:Gulf sturgeon ( 2792:Lake sturgeon ( 2776:A. brevirostrum 2740: 2702: 2695: 2665: 2656: 2637: 2621: 2540: 2451: 2400: 2363: 2347:Cylindracanthus 2332: 2319: 2247: 2157: 2136: 2121: 2101: 2070: 2038: 2037: 2027: 2022: 1978: 1976: 1968: 1965: 1960: 1959: 1950: 1949: 1938: 1928: 1926: 1925:. 31 March 2021 1923:Romania Insider 1917: 1916: 1912: 1897: 1896: 1892: 1883: 1882: 1878: 1861: 1846: 1845: 1841: 1832: 1831: 1827: 1822:. 4 March 2008. 1812: 1811: 1807: 1797: 1795: 1786: 1785: 1778: 1760: 1759: 1755: 1742: 1741: 1737: 1726: 1719: 1706: 1705: 1701: 1683: 1682: 1678: 1661: 1650: 1649: 1645: 1636: 1632: 1623: 1614: 1597: 1588: 1583: 1582: 1578: 1566: 1565: 1561: 1551: 1549: 1536: 1535: 1528: 1511: 1497: 1496: 1492: 1483: 1479: 1470: 1466: 1457: 1453: 1437: 1428: 1419: 1418: 1411: 1401: 1400: 1396: 1387: 1374: 1366: 1362: 1352: 1350: 1343: 1341:"Acipenseridae" 1339: 1338: 1334: 1324: 1322: 1313:"Acipenseridae" 1310: 1309: 1305: 1295: 1293: 1285: 1284: 1280: 1270: 1268: 1239: 1238: 1231: 1193: 1192: 1188: 1183: 1174: 1165: 1156: 1147: 1120:Bascom, Florida 1096: 1069:Bern Convention 1058: 1024: 1005: 925:Rutilus rutilus 919:Cyprinus carpio 794: 766: 744: 742: 710:greenland shark 670: 602: 574: 487:beluga sturgeon 458: 454: 306: 295: 289: 276: 184: 176: 159: 148: 144: 137: 112: 110: 109: 108: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 36: 35: 32: 24: 17: 12: 11: 5: 4745: 4743: 4735: 4734: 4729: 4724: 4719: 4714: 4709: 4707:Fish of Turkey 4704: 4699: 4697:Fish of Europe 4694: 4689: 4684: 4679: 4677:Fish of Russia 4674: 4669: 4664: 4659: 4649: 4648: 4640: 4639: 4637: 4636: 4623: 4610: 4597: 4584: 4571: 4558: 4545: 4532: 4519: 4506: 4493: 4480: 4467: 4454: 4441: 4428: 4415: 4406:Fauna Europaea 4402: 4389: 4379: 4366: 4353: 4340: 4330: 4317: 4302: 4286: 4284: 4278: 4277: 4272: 4263: 4262: 4260: 4259: 4247: 4234: 4231: 4230: 4228: 4227: 4217: 4212: 4211: 4210: 4205: 4197: 4192: 4187: 4182: 4177: 4172: 4167: 4161: 4159: 4155: 4154: 4152: 4151: 4150: 4149: 4144: 4134: 4133: 4132: 4127: 4122: 4112: 4111: 4110: 4105: 4094: 4092: 4088: 4087: 4084: 4083: 4081: 4080: 4079: 4078: 4073: 4068: 4058: 4057: 4056: 4051: 4046: 4041: 4031: 4030: 4029: 4024: 4019: 4014: 4003: 4001: 3999:Wild fisheries 3995: 3994: 3992: 3991: 3986: 3981: 3976: 3971: 3965: 3963: 3954: 3948: 3947: 3945: 3944: 3939: 3934: 3929: 3924: 3922:Hallucinogenic 3919: 3914: 3909: 3904: 3899: 3893: 3891: 3887: 3886: 3884: 3883: 3878: 3873: 3868: 3863: 3858: 3853: 3848: 3843: 3838: 3833: 3827: 3825: 3818: 3817: 3815: 3814: 3809: 3804: 3799: 3797:Schooling fish 3794: 3789: 3784: 3779: 3774: 3769: 3764: 3759: 3757:Filter feeders 3754: 3749: 3744: 3739: 3734: 3732:Bottom feeders 3729: 3724: 3719: 3713: 3711: 3705: 3704: 3702: 3701: 3696: 3691: 3686: 3681: 3676: 3671: 3665: 3663: 3657: 3656: 3654: 3653: 3652: 3651: 3641: 3636: 3631: 3626: 3621: 3616: 3611: 3606: 3601: 3596: 3591: 3586: 3581: 3575: 3573: 3567: 3566: 3564: 3563: 3558: 3553: 3548: 3543: 3538: 3533: 3528: 3523: 3518: 3513: 3508: 3502: 3500: 3492: 3491: 3489: 3488: 3483: 3478: 3477: 3476: 3471: 3461: 3460: 3459: 3454: 3444: 3439: 3434: 3433: 3432: 3422: 3417: 3412: 3407: 3402: 3401: 3400: 3390: 3385: 3380: 3378:Leydig's organ 3375: 3374: 3373: 3371:pharyngeal jaw 3368: 3358: 3353: 3352: 3351: 3346: 3341: 3336: 3331: 3326: 3324:branchial arch 3316: 3315: 3314: 3304: 3299: 3294: 3293: 3292: 3287: 3277: 3272: 3266: 3264: 3252: 3251: 3244: 3242: 3240: 3239: 3234: 3229: 3224: 3219: 3214: 3213: 3212: 3207: 3202: 3192: 3187: 3182: 3177: 3171: 3169: 3162: 3161: 3156: 3154: 3153: 3146: 3139: 3131: 3122: 3121: 3118: 3117: 3115: 3114: 3103: 3101: 3093: 3092: 3090: 3089: 3079: 3077: 3066: 3060: 3059: 3056: 3055: 3053: 3052: 3043: 3034: 3024: 3022: 3019:Scaphirhynchus 3014: 3013: 3011: 3010: 3001: 2992: 2982: 2980: 2977:scaphirhynchus 2970: 2969: 2967: 2966: 2957: 2947: 2945: 2937: 2936: 2934: 2933: 2924: 2915: 2906: 2897: 2888: 2879: 2870: 2861: 2852: 2848:A. nudiventris 2843: 2834: 2825: 2816: 2812:A. medirostris 2807: 2798: 2789: 2780: 2771: 2761: 2759: 2748: 2742: 2741: 2739: 2738: 2732: 2726: 2720: 2714: 2707: 2704: 2703: 2696: 2694: 2693: 2686: 2679: 2671: 2662: 2661: 2658: 2657: 2636: 2634: 2631: 2630: 2627: 2626: 2623: 2622: 2620: 2619: 2616:Scaphirhynchus 2612: 2605: 2595: 2585: 2578: 2575:Engdahlichthys 2568: 2565:Anchiacipenser 2558: 2550: 2548: 2542: 2541: 2539: 2538: 2535:Pugiopsephurus 2528: 2518: 2515:Protopsephurus 2508: 2501: 2491: 2488:Paleopsephurus 2481: 2470: 2468: 2459: 2457:Acipenseroidei 2453: 2452: 2450: 2449: 2442: 2435: 2428: 2421: 2413: 2411: 2409:Peipiaosteidae 2402: 2401: 2399: 2398: 2395:Strongylosteus 2391: 2384: 2376: 2374: 2372:Chondrosteidae 2365: 2364: 2362: 2361: 2358:Eochondrosteus 2351: 2339: 2333: 2328: 2325: 2324: 2321: 2320: 2318: 2317: 2314:Toarcocephalus 2310: 2303: 2296: 2289: 2282: 2275: 2268: 2260: 2258: 2256:Coccolepididae 2249: 2248: 2246: 2245: 2234: 2223: 2212: 2202: 2191: 2180: 2168: 2158: 2153: 2146: 2145: 2142: 2141: 2138: 2137: 2130: 2127: 2126: 2123: 2122: 2120: 2119: 2118: 2117: 2111: 2109: 2103: 2102: 2100: 2099: 2098: 2097: 2095:Actinopterygii 2088: 2082: 2080:Actinopterygii 2072: 2071: 2067: 2066: 2057: 2055:Actinopterygii 2051: 2045: 2036: 2029: 2028: 2023: 2021: 2020: 2013: 2006: 1998: 1992: 1991: 1985: 1964: 1963:External links 1961: 1958: 1957: 1936: 1910: 1890: 1876: 1839: 1825: 1805: 1794:. 29 June 2020 1776: 1753: 1735: 1717: 1699: 1676: 1643: 1630: 1612: 1576: 1559: 1526: 1490: 1477: 1464: 1451: 1426: 1409: 1394: 1372: 1360: 1332: 1303: 1278: 1229: 1202:(4): 623–640. 1185: 1184: 1182: 1179: 1173: 1170: 1164: 1161: 1155: 1152: 1146: 1143: 1095: 1092: 1057: 1054: 1023: 1020: 1004: 1001: 977:Pleuronectidae 961:European squid 800:Fingerling of 793: 790: 765: 762: 741: 738: 669: 666: 665: 664: 661: 656: 655: 601: 598: 573: 570: 491:great sturgeon 445: 444: 443: 442: 434: 432:Babushkin 1942 422: 420:Babushkin 1942 414: 402: 390: 382: 374: 366: 364:Bonaparte 1846 358: 350: 342: 337:Acipenser shyp 334: 326: 321:Acipenser huso 315: 314: 308: 307: 296: 285: 284: 278: 277: 270: 268: 264: 263: 256: 252: 251: 246: 242: 241: 236: 232: 231: 229:Actinopterygii 226: 222: 221: 216: 212: 211: 206: 202: 201: 196: 192: 191: 178: 177: 164: 161: 160: 142: 139: 138: 133: 130: 129: 123: 122: 114: 113: 105: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 42: 33: 15: 13: 10: 9: 6: 4: 3: 2: 4744: 4733: 4730: 4728: 4725: 4723: 4720: 4718: 4715: 4713: 4710: 4708: 4705: 4703: 4700: 4698: 4695: 4693: 4690: 4688: 4685: 4683: 4680: 4678: 4675: 4673: 4670: 4668: 4665: 4663: 4660: 4658: 4655: 4654: 4652: 4645: 4633: 4628: 4624: 4620: 4615: 4611: 4607: 4602: 4598: 4594: 4589: 4585: 4581: 4576: 4572: 4568: 4563: 4559: 4555: 4550: 4546: 4542: 4537: 4533: 4529: 4524: 4520: 4516: 4511: 4507: 4503: 4498: 4494: 4490: 4485: 4481: 4477: 4472: 4468: 4464: 4459: 4455: 4451: 4446: 4442: 4438: 4433: 4429: 4425: 4420: 4416: 4412: 4407: 4403: 4399: 4394: 4390: 4386: 4380: 4376: 4371: 4367: 4363: 4358: 4354: 4350: 4345: 4341: 4337: 4331: 4327: 4322: 4318: 4313: 4307: 4303: 4298: 4292: 4288: 4287: 4285: 4283: 4279: 4275: 4270: 4258: 4257: 4248: 4246: 4245: 4236: 4235: 4232: 4226: 4225:more lists... 4218: 4216: 4213: 4209: 4206: 4204: 4201: 4200: 4198: 4196: 4193: 4191: 4188: 4186: 4183: 4181: 4180:Fish families 4178: 4176: 4173: 4171: 4168: 4166: 4165:Aquarium life 4163: 4162: 4160: 4156: 4148: 4147:fleshy-finned 4145: 4143: 4140: 4139: 4138: 4135: 4131: 4128: 4126: 4123: 4121: 4118: 4117: 4116: 4115:Cartilaginous 4113: 4109: 4106: 4104: 4101: 4100: 4099: 4096: 4095: 4093: 4089: 4077: 4074: 4072: 4069: 4067: 4064: 4063: 4062: 4059: 4055: 4052: 4050: 4047: 4045: 4042: 4040: 4037: 4036: 4035: 4032: 4028: 4025: 4023: 4020: 4018: 4015: 4013: 4010: 4009: 4008: 4005: 4004: 4002: 4000: 3996: 3990: 3987: 3985: 3982: 3980: 3977: 3975: 3972: 3970: 3967: 3966: 3964: 3962: 3958: 3955: 3953: 3949: 3943: 3940: 3938: 3935: 3933: 3930: 3928: 3925: 3923: 3920: 3918: 3915: 3913: 3910: 3908: 3905: 3903: 3900: 3898: 3895: 3894: 3892: 3888: 3882: 3879: 3877: 3874: 3872: 3869: 3867: 3864: 3862: 3859: 3857: 3854: 3852: 3849: 3847: 3844: 3842: 3839: 3837: 3834: 3832: 3829: 3828: 3826: 3824: 3819: 3813: 3810: 3808: 3805: 3803: 3800: 3798: 3795: 3793: 3790: 3788: 3785: 3783: 3780: 3778: 3775: 3773: 3770: 3768: 3765: 3763: 3760: 3758: 3755: 3753: 3752:Electric fish 3750: 3748: 3745: 3743: 3740: 3738: 3735: 3733: 3730: 3728: 3725: 3723: 3720: 3718: 3715: 3714: 3712: 3706: 3700: 3697: 3695: 3692: 3690: 3687: 3685: 3682: 3680: 3677: 3675: 3672: 3670: 3667: 3666: 3664: 3662: 3658: 3650: 3647: 3646: 3645: 3642: 3640: 3637: 3635: 3632: 3630: 3627: 3625: 3622: 3620: 3617: 3615: 3612: 3610: 3607: 3605: 3602: 3600: 3597: 3595: 3592: 3590: 3587: 3585: 3582: 3580: 3577: 3576: 3574: 3572: 3568: 3562: 3559: 3557: 3554: 3552: 3549: 3547: 3544: 3542: 3539: 3537: 3534: 3532: 3529: 3527: 3524: 3522: 3519: 3517: 3514: 3512: 3509: 3507: 3504: 3503: 3501: 3499: 3493: 3487: 3484: 3482: 3479: 3475: 3472: 3470: 3467: 3466: 3465: 3462: 3458: 3455: 3453: 3450: 3449: 3448: 3445: 3443: 3440: 3438: 3435: 3431: 3428: 3427: 3426: 3423: 3421: 3418: 3416: 3413: 3411: 3408: 3406: 3403: 3399: 3396: 3395: 3394: 3391: 3389: 3386: 3384: 3383:Mauthner cell 3381: 3379: 3376: 3372: 3369: 3367: 3364: 3363: 3362: 3359: 3357: 3354: 3350: 3347: 3345: 3342: 3340: 3337: 3335: 3332: 3330: 3327: 3325: 3322: 3321: 3320: 3317: 3313: 3310: 3309: 3308: 3305: 3303: 3302:Chromatophore 3300: 3298: 3295: 3291: 3288: 3286: 3283: 3282: 3281: 3278: 3276: 3273: 3271: 3268: 3267: 3265: 3263: 3257: 3253: 3248: 3238: 3235: 3233: 3230: 3228: 3225: 3223: 3220: 3218: 3215: 3211: 3208: 3206: 3203: 3201: 3198: 3197: 3196: 3193: 3191: 3188: 3186: 3183: 3181: 3178: 3176: 3173: 3172: 3170: 3168: 3163: 3159: 3152: 3147: 3145: 3140: 3138: 3133: 3132: 3129: 3112: 3110: 3105: 3104: 3102: 3100: 3099: 3094: 3088: 3086: 3081: 3080: 3078: 3076: 3075: 3070: 3067: 3065: 3064:Polyodontidae 3061: 3051: 3049: 3044: 3042: 3040: 3035: 3033: 3031: 3026: 3025: 3023: 3021: 3020: 3015: 3009: 3007: 3002: 3000: 2998: 2993: 2991: 2989: 2984: 2983: 2981: 2979: 2978: 2971: 2965: 2963: 2958: 2956: 2954: 2949: 2948: 2946: 2944: 2943: 2938: 2932: 2930: 2925: 2923: 2921: 2916: 2914: 2912: 2907: 2905: 2903: 2898: 2896: 2894: 2893:A. schrenckii 2889: 2887: 2885: 2880: 2878: 2876: 2871: 2869: 2867: 2862: 2860: 2858: 2853: 2851: 2849: 2844: 2842: 2840: 2835: 2833: 2831: 2826: 2824: 2822: 2817: 2815: 2813: 2808: 2806: 2804: 2799: 2797: 2795: 2794:A. fulvescens 2790: 2788: 2786: 2781: 2779: 2777: 2772: 2770: 2768: 2763: 2762: 2760: 2758: 2757: 2752: 2749: 2747: 2746:Acipenseridae 2743: 2737: 2733: 2731: 2727: 2725: 2721: 2719: 2715: 2713: 2709: 2708: 2705: 2700: 2692: 2687: 2685: 2680: 2678: 2673: 2672: 2669: 2654: 2650: 2648: 2641: 2618: 2617: 2613: 2611: 2610: 2606: 2604: 2603: 2596: 2594: 2593: 2592:Priscosturion 2586: 2584: 2583: 2579: 2577: 2576: 2569: 2567: 2566: 2559: 2557: 2556: 2552: 2551: 2549: 2547: 2546:Acipenseridae 2543: 2537: 2536: 2529: 2527: 2526: 2519: 2517: 2516: 2509: 2507: 2506: 2502: 2500: 2499: 2498:Parapsephurus 2492: 2490: 2489: 2482: 2480: 2479: 2472: 2471: 2469: 2467: 2466:Polyodontidae 2463: 2460: 2458: 2454: 2448: 2447: 2443: 2441: 2440: 2436: 2434: 2433: 2429: 2427: 2426: 2422: 2420: 2419: 2415: 2414: 2412: 2410: 2403: 2397: 2396: 2392: 2390: 2389: 2385: 2383: 2382: 2378: 2377: 2375: 2373: 2366: 2360: 2359: 2352: 2349: 2348: 2341: 2340: 2337: 2334: 2331: 2326: 2316: 2315: 2311: 2309: 2308: 2304: 2302: 2301: 2297: 2295: 2294: 2290: 2288: 2287: 2283: 2281: 2280: 2276: 2274: 2273: 2269: 2267: 2266: 2262: 2261: 2259: 2257: 2250: 2243: 2242: 2235: 2232: 2231: 2224: 2221: 2220: 2213: 2211: 2210: 2203: 2200: 2199: 2192: 2189: 2188: 2181: 2178: 2177: 2170: 2169: 2166: 2162: 2159: 2156: 2151: 2147: 2135: 2134: 2128: 2115: 2114: 2113: 2112: 2110: 2108: 2104: 2096: 2092: 2091: 2090: 2089: 2086: 2083: 2081: 2077: 2073: 2065: 2061: 2058: 2056: 2052: 2050: 2046: 2044: 2040: 2039: 2034: 2030: 2026: 2019: 2014: 2012: 2007: 2005: 2000: 1999: 1996: 1989: 1986: 1975: 1971: 1967: 1966: 1962: 1953: 1947: 1945: 1943: 1941: 1937: 1924: 1920: 1914: 1911: 1906: 1905: 1900: 1894: 1891: 1886: 1880: 1877: 1872: 1866: 1858: 1854: 1850: 1843: 1840: 1835: 1829: 1826: 1821: 1820: 1819:Science Daily 1815: 1809: 1806: 1793: 1789: 1783: 1781: 1777: 1772: 1771: 1765: 1757: 1754: 1749: 1747: 1746:Beluga Caviar 1739: 1736: 1733: 1729: 1724: 1722: 1718: 1713: 1710: 1703: 1700: 1696:(4): 566–574. 1695: 1691: 1687: 1680: 1677: 1672: 1666: 1658: 1654: 1647: 1644: 1640: 1634: 1631: 1627: 1621: 1619: 1617: 1613: 1608: 1602: 1594: 1587: 1580: 1577: 1572: 1571: 1563: 1560: 1548: 1544: 1542: 1533: 1531: 1527: 1522: 1516: 1508: 1504: 1502: 1494: 1491: 1487: 1481: 1478: 1474: 1471:Berg, Leo S, 1468: 1465: 1461: 1455: 1452: 1449: 1445: 1441: 1435: 1433: 1431: 1427: 1422: 1416: 1414: 1410: 1405: 1398: 1395: 1391: 1385: 1383: 1381: 1379: 1377: 1373: 1369: 1364: 1361: 1349: 1342: 1336: 1333: 1321: 1319: 1314: 1307: 1304: 1292: 1288: 1282: 1279: 1266: 1261: 1257: 1253: 1252: 1247: 1245: 1236: 1234: 1230: 1225: 1221: 1217: 1213: 1209: 1205: 1201: 1197: 1190: 1187: 1180: 1178: 1171: 1169: 1162: 1160: 1153: 1151: 1144: 1142: 1140: 1136: 1132: 1128: 1123: 1121: 1115: 1113: 1109: 1105: 1101: 1093: 1091: 1089: 1085: 1080: 1078: 1077:Mediterranean 1074: 1070: 1066: 1062: 1055: 1053: 1051: 1047: 1044:The beluga's 1042: 1040: 1036: 1035:Beluga caviar 1028: 1021: 1019: 1017: 1012: 1010: 1002: 1000: 998: 994: 990: 986: 982: 978: 974: 970: 966: 962: 958: 954: 950: 946: 941: 939: 938: 933: 932: 927: 926: 921: 920: 915: 911: 907: 906: 901: 900: 895: 894: 889: 888:Ponto-Caspian 881: 876: 872: 870: 869: 864: 860: 856: 855: 849: 847: 843: 842:Caspian seals 839: 835: 831: 827: 822: 821: 818: 814: 810: 803: 798: 791: 789: 785: 783: 779: 775: 771: 763: 761: 739: 737: 733: 731: 730:Huso dauricus 727: 722: 717: 715: 711: 707: 703: 699: 695: 694:giant oarfish 691: 687: 686:ocean sunfish 679: 674: 667: 662: 658: 657: 653: 649: 648: 647: 644: 642: 637: 631: 629: 625: 624:cartilaginous 621: 617: 611: 606: 599: 597: 594: 583: 579: 571: 569: 567: 563: 559: 555: 551: 550:beluga caviar 547: 543: 539: 535: 531: 527: 524: 520: 519:Acipenseridae 516: 513: 509: 506: 502: 498: 497: 492: 488: 482: 452: 438: 435: 430: 426: 423: 418: 415: 410: 406: 403: 398: 394: 391: 386: 383: 378: 375: 370: 367: 362: 359: 354: 351: 346: 343: 338: 335: 330: 327: 324:Linnaeus 1758 322: 319: 318: 316: 313: 309: 304: 300: 294: 292: 286: 283: 282:Binomial name 279: 275: 274: 269: 266: 265: 262: 261: 257: 254: 253: 250: 249:Acipenseridae 247: 244: 243: 240: 237: 234: 233: 230: 227: 224: 223: 220: 217: 214: 213: 210: 207: 204: 203: 200: 197: 194: 193: 188: 183: 179: 173: 167: 162: 156: 151: 140: 136: 131: 128: 124: 120: 115: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 39: 29: 26: 22: 4687:Fish of Iran 4672:Fish of Asia 4644: 4281: 4254: 4242: 4142:spiny-finned 4091:Major groups 3812:Intelligence 3792:Scale eaters 3737:Cleaner fish 3619:Mouthbrooder 3571:Reproduction 3546:Schreckstoff 3531:Lateral line 3447:Swim bladder 3437:Spiral valve 3366:hyomandibula 3349:pseudobranch 3232:Hypoxia in - 3108: 3096: 3084: 3072: 3047: 3038: 3029: 3017: 3006:P. kaufmanni 3005: 2996: 2987: 2973: 2961: 2959: 2952: 2940: 2928: 2919: 2911:A. stellatus 2910: 2901: 2892: 2883: 2874: 2865: 2856: 2847: 2838: 2829: 2820: 2811: 2802: 2793: 2785:A. dabryanus 2784: 2775: 2766: 2754: 2730:Osteichthyes 2644: 2614: 2607: 2600: 2590: 2580: 2573: 2563: 2553: 2533: 2523: 2513: 2503: 2496: 2486: 2478:Crossopholis 2476: 2444: 2439:Stichopterus 2437: 2430: 2425:Peipiaosteus 2423: 2416: 2393: 2386: 2381:Chondrosteus 2379: 2356: 2345: 2312: 2305: 2298: 2291: 2284: 2277: 2270: 2263: 2239: 2228: 2217: 2207: 2196: 2187:Errolichthys 2185: 2174: 2131: 2059: 1977:. Retrieved 1927:. Retrieved 1922: 1913: 1902: 1893: 1879: 1852: 1842: 1828: 1817: 1808: 1796:. Retrieved 1791: 1768: 1764:"Huso"  1756: 1745: 1738: 1731: 1712: 1708: 1702: 1693: 1689: 1685: 1679: 1665:cite journal 1656: 1652: 1646: 1638: 1633: 1625: 1593:Danube Parks 1592: 1579: 1569: 1562: 1550:. Retrieved 1546: 1540: 1506: 1500: 1493: 1485: 1480: 1472: 1467: 1459: 1454: 1439: 1403: 1397: 1389: 1363: 1351:. Retrieved 1347: 1335: 1323:. Retrieved 1316: 1306: 1294:. Retrieved 1290: 1281: 1269:. Retrieved 1255: 1249: 1243: 1199: 1195: 1189: 1175: 1166: 1157: 1148: 1131:Danube River 1125:Following a 1124: 1116: 1112:H. huso 1111: 1097: 1081: 1059: 1043: 1033: 1013: 1006: 969:Adriatic Sea 964: 956: 945:H. huso 944: 942: 935: 929: 923: 917: 903: 897: 891: 885: 880:H. huso 879: 866: 852: 850: 846:Pusa caspica 845: 823: 808: 807: 801: 786: 767: 758: 734: 729: 718: 713: 705: 697: 689: 683: 645: 632: 620:heterocercal 615: 614: 609: 578:beluga whale 575: 538:Adriatic Sea 495: 494: 490: 486: 450: 448: 436: 428: 424: 416: 408: 404: 397:occidentalis 396: 392: 384: 376: 368: 360: 356:Forster 1767 352: 344: 340:Forster 1767 336: 332:Forster 1767 328: 320: 290: 288: 273:H. huso 272: 271: 259: 126: 25: 21:Beluga whale 4458:iNaturalist 4306:Wikispecies 4256:WikiProject 4215:Prehistoric 4199:Threatened 3890:Other types 3787:Sardine run 3762:Forage fish 3742:Corallivory 3594:Development 3579:Bubble nest 3452:physoclisti 3442:Suckermouth 3415:Root effect 3237:Ichthyology 3085:P. spathula 3048:S. suttkusi 2997:P. hermanni 2953:H. dauricus 2902:A. sinensis 2884:A. ruthenus 2875:A. persicus 2839:A. naccarii 2736:Chondrostei 2728:Superclass 2432:Spherosteus 2279:Condorlepis 2209:Neochallaia 2155:Chondrostei 2133:Chondrostei 2107:Chondrostei 2064:Actinopteri 2025:Chondrostei 1970:"Huso huso" 1690:J. Ichthyol 1100:fish ladder 1046:air bladder 1016:Sea of Azov 951:, of which 949:Cephalopoda 854:Clupeonella 838:crustaceans 830:piscivorous 678:Volga River 600:Description 554:overfishing 548:, known as 348:Gmelin 1789 168:Appendix II 38:Pleistocene 4651:Categories 3871:Groundfish 3866:Freshwater 3861:Euryhaline 3846:Coral reef 3782:Salmon run 3772:Paedophagy 3674:Amphibious 3661:Locomotion 3469:pharyngeal 3457:physostome 3410:Photophore 3356:Glossohyal 3329:gill raker 3312:dorsal fin 3262:physiology 3109:P. gladius 2821:A. mikadoi 2724:Vertebrata 2722:Subphylum 2293:Morrolepis 2272:Coccolepis 2265:Barbalepis 2198:Gualolepis 2116:see below↓ 1979:24 January 1368:Huso huso. 1296:14 January 1181:References 1009:euryhaline 997:Cyprinidae 989:Scombridae 770:anadromous 708:) and the 560:, leading 505:anadromous 440:Lelek 1987 409:orientalis 372:Molin 1853 4667:Sturgeons 4375:huso-huso 4326:Huso_huso 4312:Huso huso 4282:Huso huso 4120:chimaeras 4007:Predatory 3984:Salmonids 3942:Whitefish 3932:Poisonous 3907:Diversity 3841:Coldwater 3777:Predatory 3767:Migratory 3727:Bait ball 3710:behaviour 3629:Pregnancy 3624:Polyandry 3398:papillare 3393:Operculum 3388:Meristics 3334:gill slit 3297:Cleithrum 3227:Fish kill 3217:Fear of - 3210:- as food 3200:Fisheries 3185:Evolution 3175:Diversity 3098:Psephurus 2920:A. sturio 2882:Sterlet ( 2767:A. baerii 2756:Acipenser 2734:Subclass 2555:Acipenser 2525:Psephurus 2446:Yanosteus 2418:Liaosteus 2388:Gyrosteus 2307:Sunolepis 2041:Kingdom: 1686:Huso huso 1653:Huso huso 1543:(Beluga)" 1541:Huso huso 1501:Huso huso 1291:cites.org 1244:Huso huso 1172:Pollution 1050:isinglass 1039:swordfish 993:Mugilidae 985:Clupeidae 916:, mainly 914:cyprinids 910:estuaries 905:Engraulis 878:An adult 809:Huso huso 802:Huso huso 782:Black Sea 616:Huso huso 610:Huso huso 572:Etymology 534:Black Sea 521:) of the 496:Huso huso 291:Huso huso 267:Species: 205:Kingdom: 199:Eukaryota 127:Huso huso 4614:Species+ 4476:10145452 4432:FishBase 4398:46561184 4333:BioLib: 4291:Wikidata 4244:Category 4195:Smallest 4108:lampreys 4071:flatfish 4061:Demersal 4017:mackerel 4012:billfish 3952:Commerce 3881:Tropical 3856:Demersal 3851:Deep-sea 3807:Venomous 3699:RoboTuna 3649:triggers 3644:Spawning 3604:Juvenile 3589:Egg case 3222:FishBase 3074:Polyodon 3030:S. albus 2975:Pseudo- 2960:Beluga ( 2951:Kaluga ( 2718:Chordata 2712:Animalia 2710:Kingdom 2505:Polyodon 2286:Iyalepis 2176:Birgeria 2049:Chordata 2047:Phylum: 2043:Animalia 1865:cite web 1601:cite web 1515:cite web 1318:FishBase 1224:84869858 1108:Po river 981:Gobiidae 859:Gobiidae 834:molluscs 817:demersal 764:Spawning 558:poaching 512:sturgeon 499:), is a 429:curensis 312:Synonyms 299:Linnaeus 245:Family: 219:Chordata 215:Phylum: 209:Animalia 195:Domain: 155:IUCN 3.1 4450:2402105 4297:Q191816 4190:Largest 4103:hagfish 4098:Jawless 4076:pollock 4049:sardine 4044:herring 4039:anchovy 3989:Tilapia 3979:Octopus 3974:Catfish 3961:Farming 3876:Pelagic 3836:Coastal 3823:habitat 3679:Walking 3584:Clasper 3536:Otolith 3498:systems 3496:Sensory 3430:ganoine 3405:Papilla 3256:Anatomy 3195:Fishing 2962:H. huso 2716:Phylum 2701:species 2697:Recent 2053:Class: 1216:2846137 1145:Threats 1003:Habitat 973:Gadidae 863:mullets 826:benthic 813:pelagic 740:Biology 641:barbels 636:rostrum 582:Russian 530:Caspian 510:in the 501:species 255:Genus: 235:Order: 225:Class: 170: ( 153: ( 106:↓ 40:–Recent 4632:126280 4580:751886 4567:126280 4554:208243 4502:161084 4463:102975 4411:304329 4382:ECOS: 4208:sharks 4125:sharks 4054:sprats 4034:Forage 4022:salmon 3902:Coarse 3684:Flying 3556:Vision 3511:Barbel 3425:Scales 3285:dermal 3165:About 1929:28 May 1904:Forbes 1798:28 May 1730:(PDF) 1552:24 May 1446:  1438:Wood, 1353:18 May 1325:18 May 1271:24 May 1222:  1214:  1056:Status 899:Aspius 778:Danube 726:Kaluga 652:scutes 628:scutes 515:family 451:beluga 31:Beluga 4627:WoRMS 4601:Plazi 4593:94895 4541:61971 4515:10269 4489:79864 4471:IRMNG 4362:3MV27 4349:29692 4336:15453 4170:Blind 4158:Lists 3937:Rough 3802:Sleep 3708:Other 3474:shark 3464:Teeth 2060:Clade 1589:(PDF) 1344:(PDF) 1220:S2CID 1212:JSTOR 1139:wagon 1073:CITES 893:Alosa 868:Alosa 811:is a 774:spawn 721:Volga 593:belyj 587:белый 584:word 523:order 172:CITES 166:CITES 4619:6061 4562:OBIS 4536:NCBI 4510:IUCN 4497:ITIS 4445:GBIF 4437:2067 4385:5279 4344:BOLD 4203:rays 4137:Bony 4130:rays 4027:tuna 3969:Carp 3927:Oily 3912:Game 3897:Bait 3831:Cave 3614:Milt 3319:Gill 3307:Fins 3280:Bone 3167:fish 3158:Fish 2942:Huso 2582:Huso 2093:see 1981:2006 1931:2022 1871:link 1800:2022 1671:link 1607:link 1554:2023 1521:link 1444:ISBN 1355:2017 1327:2017 1298:2022 1273:2023 1256:2022 1082:The 1061:IUCN 1022:Uses 991:and 959:and 922:and 902:and 848:). 836:and 792:Diet 690:Mola 668:Size 562:IUCN 556:and 532:and 508:fish 449:The 303:1758 260:Huso 45:PreꞒ 4523:NBN 4484:ISC 4393:EoL 4370:CMS 4357:CoL 4321:ADW 4066:cod 3821:By 3634:Roe 3361:Jaw 3259:and 1857:PDF 1260:doi 1204:doi 1102:on 546:roe 503:of 489:or 427:n. 407:n. 395:n. 4653:: 4629:: 4616:: 4603:: 4590:: 4577:: 4564:: 4551:: 4538:: 4525:: 4512:: 4499:: 4486:: 4473:: 4460:: 4447:: 4434:: 4421:: 4408:: 4395:: 4372:: 4359:: 4346:: 4323:: 4308:: 4293:: 2062:: 1972:. 1939:^ 1921:. 1901:. 1867:}} 1863:{{ 1855:. 1851:. 1816:. 1790:. 1779:^ 1767:. 1720:^ 1694:16 1692:. 1667:}} 1663:{{ 1615:^ 1603:}} 1599:{{ 1591:. 1545:. 1529:^ 1517:}} 1513:{{ 1505:. 1429:^ 1412:^ 1375:^ 1346:. 1315:. 1289:. 1254:. 1248:. 1232:^ 1218:. 1210:. 1200:25 1198:. 1090:. 1052:. 999:. 987:, 983:, 979:, 975:, 963:, 955:, 928:, 896:, 865:, 844:, 784:. 630:. 568:. 472:uː 301:, 95:Pg 3150:e 3143:t 3136:v 3113:† 3111:) 3087:) 3050:) 3041:) 3032:) 3008:) 2999:) 2990:) 2964:) 2955:) 2931:) 2922:) 2913:) 2904:) 2895:) 2886:) 2877:) 2868:) 2859:) 2850:) 2841:) 2832:) 2823:) 2814:) 2805:) 2796:) 2787:) 2778:) 2769:) 2690:e 2683:t 2676:v 2598:† 2588:† 2571:† 2561:† 2531:† 2521:† 2511:† 2494:† 2484:† 2474:† 2406:† 2369:† 2354:† 2350:? 2343:† 2253:† 2244:? 2237:† 2233:? 2226:† 2222:? 2215:† 2205:† 2201:? 2194:† 2190:? 2183:† 2179:? 2172:† 2017:e 2010:t 2003:v 1983:. 1954:. 1933:. 1873:) 1859:. 1836:. 1802:. 1748:" 1744:" 1673:) 1609:) 1595:. 1556:. 1539:" 1523:) 1503:" 1499:" 1423:. 1357:. 1329:. 1300:. 1275:. 1262:: 1246:" 1242:" 1226:. 1206:: 728:( 712:( 704:( 696:( 688:( 590:( 517:( 493:( 481:/ 478:ə 475:ɡ 469:l 466:ˈ 463:ə 460:b 457:/ 453:( 305:) 297:( 174:) 157:) 100:N 90:K 85:J 80:T 75:P 70:C 65:D 60:S 55:O 50:Ꞓ 23:.

Index

Beluga whale
Pleistocene
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Conservation status
Critically Endangered
IUCN 3.1
CITES
CITES
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Actinopterygii
Acipenseriformes
Acipenseridae
Huso
Binomial name

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