31:
75:
50:
322:
brown above, yellow or cream below; the area of demarcation is indistinct. The tympanum and posterior part of the lower jaw are yellow with a few gray blotches and orange spots. Jaws are grayish yellow; the iris is brown tan. Dorsally, the head is covered with large distinct scales. There are two chin barbels. The neck is grayish brown
321:
is about half as long as the length of the fore lobe. The plastral formula is: intergul > fem > abd > hum > an > gul > pect. The head is extremely broad; the carapace length averages only 4.4 times the tympanic head width, and older females may have massive heads. It is dark grayish
304:
and pleural scutes may be rugose with growth annuli. Marginals 1, 2, and 8-10 are slightly expanded but not flared, and 3-7 are often slightly upturned. The carapace is highest just behind the center and broadest at the level of the anterior part of the eighth marginals; its posterior rim may be
30:
508:. 2014. Turtles of the world, 7th edition: annotated checklist of taxonomy, synonymy, distribution with maps, and conservation status. In: Rhodin, A.G.J., Pritchard, P.C.H., van Dijk, P.P., Saumure, R.A., Buhlmann, K.A., Iverson, J.B., and Mittermeier, R.A. (Eds.).
396:
probably emerge between
December and March. Clutch size ranges from 4-8 white, rounded (28-32.5 x 25.8–31 mm; 11-20 g), hard-shelled eggs (Cintra and Yamashita, 1989). Incubation takes over six months; hatchlings (about 3.8 mm) have orange-red spots on
299:
with a shallow dorsal groove extending along the second to fourth vertebrals. The first and fifth vertebral scutes are very broad, the second through fourth may be slightly longer than broad, and the fifth is laterally expanded.
330:, and has a few scattered blunt, conical tubercles on the dorsal surface. Limbs are gray on the outside, yellow beneath, and covered with large scales. Large conical tubercles are present on the inside of each thigh. The
484:
Rhodin, A. G. J.;Mittermeier, R. A.;McMorris, J. R. 1984. Platemys macrocephala, a new species of chelid turtle from central
Bolivia and the pantanal region of Brazil. Herpetologica 40 (1): 38-46
2312:
2380:
2229:
2335:
381:, and slow-flowing streams.They also live in Venezuela also the yellow footed turtles.. In Venezuela Chaco it's called Galapago they have big neck and big tail.
2268:
496:
Fritz, U. and Pauler, I. 1992. Phrynops chacoensis spec. nov. (Reptilia, Chelidae), eine neue Krötenkopfschildkröte. Mitt. Zool. Mus. Berl. 68(2):299–307.
2203:
2242:
520:
305:
weakly serrated. The carapace is dark to blackish brown, but may be light brown in some. Juveniles often have lighter brown radiations on their
510:
Conservation
Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group
313:
and bridge are yellow with some dark pigment extending along the seams (sometimes covering most of a scute, but usually not the areola); this
547:
2385:
865:
1134:
2345:
2043:
721:
365:
2375:
1249:
1141:
1543:
1489:
1213:
1112:
2365:
2350:
2022:
1989:
1972:
1127:
2247:
2086:
Turtles of the World 2021 Update: Annotated
Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status
413:
The status of this species is listed as lower risk: near threatened by the IUCN. The Chaco population, listed separately as
2142:
2370:
1885:
1651:
1644:
1557:
728:
505:
2360:
1956:
1939:
1220:
1008:
991:
809:
74:
2164:
2050:
1536:
2156:
2355:
2273:
2057:
1829:
1381:
1105:
880:
2084:
1864:
1822:
1808:
851:
1878:
1857:
1206:
714:
540:
521:
https://web.archive.org/web/20110608014335/http://nlbif.eti.uva.nl/bis/turtles.php?menuentry=soorten&id=183
873:
823:
1899:
1815:
1787:
1754:
1737:
1178:
1148:
858:
2340:
2104:
2036:
1836:
1227:
1041:
1024:
2169:
2294:
1843:
1300:
317:
fades with age. The fore lobe is broader than the hind lobe, which contains a deep posterior notch. The
179:
776:
458:
2190:
1697:
1680:
1670:
1336:
1293:
928:
913:
844:
837:
1871:
1794:
1199:
1192:
975:
958:
533:
357:
39:
337:
Females are larger and more domed; males have slightly concave plastra and longer, thicker tails.
1801:
1550:
1496:
1171:
1098:
942:
935:
830:
768:
201:
69:
2299:
2234:
2281:
2260:
2177:
1892:
1503:
1482:
1468:
1435:
1418:
1344:
1329:
1185:
1120:
816:
2307:
1598:
1475:
1402:
920:
610:
453:
2286:
512:. Chelonian Research Monographs 5(7):000.329–479, doi:10.3854/ crm.5.000.checklist.v7.2014.
1850:
1388:
1374:
1352:
1322:
1091:
54:
435:
1922:
1395:
1286:
2329:
2255:
1720:
690:
444:
287:
156:
59:
2182:
2029:
353:
318:
310:
306:
2216:
2136:
1627:
792:
434:
Rhodin, A.G.J.; Vinke, T.; Vinke, S.; MĂ©trailler, S.; Mittermeier, R.A. (2018).
2005:
600:
393:
136:
2127:
525:
1590:
1573:
1519:
1239:
1160:
1081:
1070:
761:
619:
331:
301:
270:
86:
2121:
1770:
1617:
1060:
896:
680:
671:
637:
576:
405:, and sides of the neck. Snails form a large part of this turtle's diet.
402:
398:
349:
296:
292:
274:
258:
146:
106:
2208:
1451:
647:
588:
582:
374:
360:, to the Chaco of Paraguay. Its range in Bolivia may be more extended.
327:
323:
314:
266:
116:
2221:
2151:
1364:
1312:
1276:
1265:
744:
594:
570:
557:
378:
262:
254:
126:
96:
2098:
348:
is known from the upper Rio Mamoré drainage of central
Bolivia, the
628:
389:
2195:
2102:
529:
295:
length. It has a broad, oval to moderately elongated, deep
392:
starts near the end of the wet season, in April–May, and
2111:
2014:
2003:
1981:
1970:
1948:
1937:
1930:
1921:
1779:
1768:
1746:
1735:
1728:
1719:
1689:
1678:
1669:
1636:
1625:
1616:
1582:
1571:
1528:
1517:
1460:
1449:
1427:
1416:
1362:
1310:
1274:
1263:
1237:
1158:
1079:
1068:
1059:
1033:
1022:
1000:
989:
967:
956:
905:
894:
801:
790:
753:
742:
699:
688:
679:
670:
636:
627:
618:
609:
459:10.2305/IUCN.UK.2018-2.RLTS.T97259978A144765482.en
291:species, growing to 23.5 cm (9.3 in) in
2083:Phylogenetic arrangement of turtles based on
541:
214:Fitzinger in Siebenrock 1904:27 (nomen nudum)
8:
2099:
2011:
1978:
1945:
1934:
1927:
1776:
1743:
1732:
1725:
1686:
1675:
1633:
1622:
1579:
1525:
1457:
1424:
1271:
1076:
1065:
1030:
997:
964:
902:
798:
750:
696:
685:
676:
642:
633:
624:
615:
548:
534:
526:
285:This is the largest of the South American
195:(Rhodin, Mittermeier & McMorris, 1984)
48:
29:
20:
457:
352:region and other swamplands of the upper
222:Rhodin, Mittermeier, and McMorris 1984:38
492:
490:
480:
478:
476:
363:This species is most closely related to
426:
2381:Near threatened biota of South America
2336:IUCN Red List near threatened species
7:
445:IUCN Red List of Threatened Species
14:
24:Big-headed pantanal swamp turtle
241:big-headed pantanal swamp turtle
73:
1:
1957:Erymnochelys madagascariensis
715:Acanthochelys pallidipectoris
506:Turtle Taxonomy Working Group
417:, is considered vulnerable.
852:Mesoclemmys sabiniparaensis
2402:
2386:Reptiles described in 1984
2157:acanthochelys-macrocephala
2143:Acanthochelys macrocephala
2113:Acanthochelys macrocephala
1990:Peltocephalus dumerilianus
708:Acanthochelys macrocephala
438:Acanthochelys macrocephala
250:Acanthochelys macrocephala
189:Acanthochelys macrocephala
2079:
2044:Podocnemis sextuberculata
2023:Podocnemis erythrocephala
645:
565:
452:: e.T97259978A144765482.
356:drainage in southwestern
230:Fritz and Pauler 1992:299
207:
200:
185:
178:
70:Scientific classification
68:
46:
37:
28:
23:
2346:Turtles of South America
1207:Chelodina kurrichalpongo
866:Mesoclemmys vanderhaegei
415:Acanthochelys chacoensis
2376:Near threatened animals
1645:Hydromedusa maximiliani
1179:Chelodina burrungandjii
1149:Chelodina steindachneri
874:Mesoclemmys vanegasorum
859:Mesoclemmys tuberculata
824:Mesoclemmys jurutiensis
722:Acanthochelys radiolata
366:Acanthochelys radiolata
1228:Chelodina walloyarrina
1128:Chelodina novaeguineae
2366:Reptiles of Argentina
2351:Fauna of the Pantanal
2295:Paleobiology Database
1652:Hydromedusa tectifera
1551:Myuchelys latisternum
1301:Elseya flaviventralis
1106:Chelodina longicollis
976:Platemys platycephala
245:pantanal swamp turtle
220:Platemys macrocephala
2371:Reptiles of Paraguay
1872:Pelusios rhodesianus
1823:Pelusios castanoides
1795:Pelusios bechuanicus
1698:Pseudemydura umbrina
1294:Elseya branderhorsti
1135:Chelodina pritchardi
929:Phrynops paranaensis
914:Phrynops geoffroanus
845:Mesoclemmys raniceps
838:Mesoclemmys perplexa
729:Acanthochelys spixii
171:A. macrocephala
2361:Reptiles of Bolivia
2051:Podocnemis unifilis
1844:Pelusios gabonensis
1382:Elseya novaeguineae
1337:Elseya lavarackorum
1200:Chelodina kuchlingi
1193:Chelodina insculpta
371:Phrynops chacoensis
358:Mato Grosso, Brazil
228:Phrynops chacoensis
212:Phrynops schoepffii
40:Conservation status
2037:Podocnemis lewyana
2030:Podocnemis expansa
1900:Pelusios williamsi
1837:Pelusios cupulatta
1816:Pelusios castaneus
1809:Pelusios carinatus
1802:Pelusios broadleyi
1788:Pelusios adansonii
1755:Pelomedusa subrufa
1544:Myuchelys georgesi
1497:Emydura tanybaraga
1490:Emydura subglobosa
1172:Chelodina alanrixi
1142:Chelodina reimanni
1099:Chelodina gunaleni
943:Phrynops williamsi
936:Phrynops tuberosus
881:Mesoclemmys zuliae
831:Mesoclemmys nasuta
769:Chelus orinocensis
253:) is a species of
2356:Turtles of Brazil
2323:
2322:
2282:Open Tree of Life
2105:Taxon identifiers
2096:
2095:
2089:. Key: †=extinct.
2075:
2074:
2071:
2070:
2067:
2066:
1999:
1998:
1966:
1965:
1917:
1916:
1913:
1912:
1909:
1908:
1886:Pelusios subniger
1879:Pelusios sinuatus
1764:
1763:
1715:
1714:
1711:
1710:
1707:
1706:
1665:
1664:
1661:
1660:
1612:
1611:
1608:
1607:
1591:Rheodytes leukops
1567:
1566:
1558:Myuchelys purvisi
1513:
1512:
1504:Emydura victoriae
1483:Emydura macquarii
1469:Emydura australis
1445:
1444:
1412:
1411:
1345:Elseya nadibajagu
1259:
1258:
1250:Chelodina oblonga
1214:Chelodina parkeri
1186:Chelodina expansa
1121:Chelodina murrayi
1113:Chelodina mccordi
1055:
1054:
1051:
1050:
1018:
1017:
1009:Ranacephala hogei
985:
984:
952:
951:
890:
889:
817:Mesoclemmys gibba
810:Mesoclemmys dahli
786:
785:
777:Chelus colombiana
738:
737:
666:
665:
662:
661:
658:
657:
654:
653:
307:carapacial scutes
237:
236:
231:
223:
215:
63:
16:Species of turtle
2393:
2316:
2315:
2303:
2302:
2290:
2289:
2277:
2276:
2264:
2263:
2251:
2250:
2238:
2237:
2225:
2224:
2212:
2211:
2199:
2198:
2186:
2185:
2173:
2172:
2160:
2159:
2147:
2146:
2145:
2132:
2131:
2130:
2100:
2058:Podocnemis vogli
2012:
1979:
1946:
1935:
1928:
1893:Pelusios upembae
1830:Pelusios chapini
1777:
1744:
1733:
1726:
1687:
1676:
1634:
1623:
1599:Rheodytes devisi
1580:
1526:
1476:Emydura gunaleni
1458:
1425:
1403:Elseya schultzei
1272:
1221:Chelodina rugosa
1077:
1066:
1042:Rhinemys rufipes
1031:
998:
965:
921:Phrynops hilarii
903:
799:
762:Chelus fimbriata
751:
697:
686:
677:
643:
634:
625:
616:
550:
543:
536:
527:
513:
503:
497:
494:
485:
482:
471:
470:
468:
466:
461:
431:
319:intergular scute
226:
218:
210:
191:
78:
77:
57:
52:
51:
33:
21:
2401:
2400:
2396:
2395:
2394:
2392:
2391:
2390:
2326:
2325:
2324:
2319:
2311:
2306:
2298:
2293:
2285:
2280:
2272:
2267:
2259:
2254:
2246:
2241:
2233:
2228:
2220:
2215:
2207:
2202:
2194:
2189:
2181:
2176:
2168:
2163:
2155:
2150:
2141:
2140:
2135:
2126:
2125:
2120:
2107:
2097:
2092:
2063:
1995:
1962:
1905:
1851:Pelusios marani
1760:
1703:
1671:Pseudemydurinae
1657:
1604:
1563:
1537:Myuchelys belli
1509:
1441:
1436:Elusor macrurus
1408:
1389:Elseya orestiad
1375:Elseya caelatus
1358:
1353:Elseya uberrima
1323:Elseya albagula
1306:
1255:
1233:
1154:
1092:Chelodina canni
1047:
1014:
981:
948:
886:
782:
734:
650:
605:
561:
554:
517:
516:
504:
500:
495:
488:
483:
474:
464:
462:
433:
432:
428:
423:
411:
387:
385:Natural history
346:A. macrocephala
343:
283:
196:
193:
187:
174:
72:
64:
55:Near Threatened
53:
49:
42:
17:
12:
11:
5:
2399:
2397:
2389:
2388:
2383:
2378:
2373:
2368:
2363:
2358:
2353:
2348:
2343:
2338:
2328:
2327:
2321:
2320:
2318:
2317:
2304:
2291:
2278:
2265:
2252:
2239:
2226:
2213:
2200:
2187:
2174:
2161:
2148:
2133:
2117:
2115:
2109:
2108:
2103:
2094:
2093:
2091:
2090:
2080:
2077:
2076:
2073:
2072:
2069:
2068:
2065:
2064:
2062:
2061:
2054:
2047:
2040:
2033:
2026:
2018:
2016:
2009:
2001:
2000:
1997:
1996:
1994:
1993:
1985:
1983:
1976:
1968:
1967:
1964:
1963:
1961:
1960:
1952:
1950:
1943:
1932:
1925:
1923:Podocnemididae
1919:
1918:
1915:
1914:
1911:
1910:
1907:
1906:
1904:
1903:
1896:
1889:
1882:
1875:
1868:
1865:Pelusios niger
1861:
1858:Pelusios nanus
1854:
1847:
1840:
1833:
1826:
1819:
1812:
1805:
1798:
1791:
1783:
1781:
1774:
1766:
1765:
1762:
1761:
1759:
1758:
1750:
1748:
1741:
1730:
1723:
1717:
1716:
1713:
1712:
1709:
1708:
1705:
1704:
1702:
1701:
1693:
1691:
1684:
1673:
1667:
1666:
1663:
1662:
1659:
1658:
1656:
1655:
1648:
1640:
1638:
1631:
1620:
1618:Hydromedusinae
1614:
1613:
1610:
1609:
1606:
1605:
1603:
1602:
1594:
1586:
1584:
1577:
1569:
1568:
1565:
1564:
1562:
1561:
1554:
1547:
1540:
1532:
1530:
1523:
1515:
1514:
1511:
1510:
1508:
1507:
1500:
1493:
1486:
1479:
1472:
1464:
1462:
1455:
1447:
1446:
1443:
1442:
1440:
1439:
1431:
1429:
1422:
1414:
1413:
1410:
1409:
1407:
1406:
1399:
1396:Elseya rhodini
1392:
1385:
1378:
1370:
1368:
1360:
1359:
1357:
1356:
1348:
1340:
1333:
1326:
1318:
1316:
1308:
1307:
1305:
1304:
1297:
1290:
1287:Elseya dentata
1282:
1280:
1269:
1261:
1260:
1257:
1256:
1254:
1253:
1245:
1243:
1240:Macrochelodina
1235:
1234:
1232:
1231:
1224:
1217:
1210:
1203:
1196:
1189:
1182:
1175:
1166:
1164:
1156:
1155:
1153:
1152:
1145:
1138:
1131:
1124:
1116:
1109:
1102:
1095:
1087:
1085:
1074:
1063:
1057:
1056:
1053:
1052:
1049:
1048:
1046:
1045:
1037:
1035:
1028:
1020:
1019:
1016:
1015:
1013:
1012:
1004:
1002:
995:
987:
986:
983:
982:
980:
979:
971:
969:
962:
954:
953:
950:
949:
947:
946:
939:
932:
924:
917:
909:
907:
900:
892:
891:
888:
887:
885:
884:
877:
869:
862:
855:
848:
841:
834:
827:
820:
813:
805:
803:
796:
788:
787:
784:
783:
781:
780:
772:
765:
757:
755:
748:
740:
739:
736:
735:
733:
732:
725:
718:
711:
703:
701:
694:
683:
674:
668:
667:
664:
663:
660:
659:
656:
655:
652:
651:
646:
640:
631:
622:
613:
607:
606:
604:
603:
597:
591:
585:
579:
573:
566:
563:
562:
555:
553:
552:
545:
538:
530:
524:
523:
515:
514:
498:
486:
472:
425:
424:
422:
419:
410:
407:
386:
383:
373:. It inhabits
369:; it includes
342:
339:
282:
279:
257:in the family
235:
234:
233:
232:
224:
216:
205:
204:
198:
197:
194:
183:
182:
176:
175:
168:
166:
162:
161:
154:
150:
149:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
2398:
2387:
2384:
2382:
2379:
2377:
2374:
2372:
2369:
2367:
2364:
2362:
2359:
2357:
2354:
2352:
2349:
2347:
2344:
2342:
2341:Acanthochelys
2339:
2337:
2334:
2333:
2331:
2314:
2309:
2305:
2301:
2296:
2292:
2288:
2283:
2279:
2275:
2270:
2266:
2262:
2257:
2253:
2249:
2244:
2240:
2236:
2231:
2227:
2223:
2218:
2214:
2210:
2205:
2201:
2197:
2192:
2188:
2184:
2179:
2175:
2171:
2166:
2162:
2158:
2153:
2149:
2144:
2138:
2134:
2129:
2123:
2119:
2118:
2116:
2114:
2110:
2106:
2101:
2088:
2087:
2082:
2081:
2078:
2060:
2059:
2055:
2053:
2052:
2048:
2046:
2045:
2041:
2039:
2038:
2034:
2032:
2031:
2027:
2025:
2024:
2020:
2019:
2017:
2013:
2010:
2008:
2007:
2002:
1992:
1991:
1987:
1986:
1984:
1980:
1977:
1975:
1974:
1973:Peltocephalus
1969:
1959:
1958:
1954:
1953:
1951:
1947:
1944:
1942:
1941:
1936:
1933:
1929:
1926:
1924:
1920:
1902:
1901:
1897:
1895:
1894:
1890:
1888:
1887:
1883:
1881:
1880:
1876:
1874:
1873:
1869:
1867:
1866:
1862:
1860:
1859:
1855:
1853:
1852:
1848:
1846:
1845:
1841:
1839:
1838:
1834:
1832:
1831:
1827:
1825:
1824:
1820:
1818:
1817:
1813:
1811:
1810:
1806:
1804:
1803:
1799:
1797:
1796:
1792:
1790:
1789:
1785:
1784:
1782:
1778:
1775:
1773:
1772:
1767:
1757:
1756:
1752:
1751:
1749:
1745:
1742:
1740:
1739:
1734:
1731:
1727:
1724:
1722:
1721:Pelomedusidae
1718:
1700:
1699:
1695:
1694:
1692:
1688:
1685:
1683:
1682:
1677:
1674:
1672:
1668:
1654:
1653:
1649:
1647:
1646:
1642:
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1639:
1635:
1632:
1630:
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1624:
1621:
1619:
1615:
1601:
1600:
1595:
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1588:
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1578:
1576:
1575:
1570:
1560:
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1548:
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1541:
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1534:
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1527:
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1516:
1506:
1505:
1501:
1499:
1498:
1494:
1492:
1491:
1487:
1485:
1484:
1480:
1478:
1477:
1473:
1471:
1470:
1466:
1465:
1463:
1459:
1456:
1454:
1453:
1448:
1438:
1437:
1433:
1432:
1430:
1426:
1423:
1421:
1420:
1415:
1405:
1404:
1400:
1398:
1397:
1393:
1391:
1390:
1386:
1384:
1383:
1379:
1377:
1376:
1372:
1371:
1369:
1367:
1366:
1365:Hanwarachelys
1361:
1355:
1354:
1349:
1347:
1346:
1341:
1339:
1338:
1334:
1332:
1331:
1330:Elseya irwini
1327:
1325:
1324:
1320:
1319:
1317:
1315:
1314:
1309:
1303:
1302:
1298:
1296:
1295:
1291:
1289:
1288:
1284:
1283:
1281:
1279:
1278:
1273:
1270:
1268:
1267:
1262:
1252:
1251:
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1244:
1242:
1241:
1236:
1230:
1229:
1225:
1223:
1222:
1218:
1216:
1215:
1211:
1209:
1208:
1204:
1202:
1201:
1197:
1195:
1194:
1190:
1188:
1187:
1183:
1181:
1180:
1176:
1174:
1173:
1168:
1167:
1165:
1163:
1162:
1157:
1151:
1150:
1146:
1144:
1143:
1139:
1137:
1136:
1132:
1130:
1129:
1125:
1123:
1122:
1117:
1115:
1114:
1110:
1108:
1107:
1103:
1101:
1100:
1096:
1094:
1093:
1089:
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1086:
1084:
1083:
1078:
1075:
1073:
1072:
1067:
1064:
1062:
1058:
1044:
1043:
1039:
1038:
1036:
1032:
1029:
1027:
1026:
1021:
1011:
1010:
1006:
1005:
1003:
999:
996:
994:
993:
988:
978:
977:
973:
972:
970:
966:
963:
961:
960:
955:
945:
944:
940:
938:
937:
933:
931:
930:
925:
923:
922:
918:
916:
915:
911:
910:
908:
904:
901:
899:
898:
893:
883:
882:
878:
876:
875:
870:
868:
867:
863:
861:
860:
856:
854:
853:
849:
847:
846:
842:
840:
839:
835:
833:
832:
828:
826:
825:
821:
819:
818:
814:
812:
811:
807:
806:
804:
800:
797:
795:
794:
789:
779:
778:
773:
771:
770:
766:
764:
763:
759:
758:
756:
752:
749:
747:
746:
741:
731:
730:
726:
724:
723:
719:
717:
716:
712:
710:
709:
705:
704:
702:
698:
695:
693:
692:
691:Acanthochelys
687:
684:
682:
678:
675:
673:
669:
649:
644:
641:
639:
635:
632:
630:
626:
623:
621:
617:
614:
612:
608:
602:
598:
596:
592:
590:
586:
584:
580:
578:
574:
572:
568:
567:
564:
559:
551:
546:
544:
539:
537:
532:
531:
528:
522:
519:
518:
511:
507:
502:
499:
493:
491:
487:
481:
479:
477:
473:
460:
455:
451:
447:
446:
441:
439:
430:
427:
420:
418:
416:
408:
406:
404:
400:
395:
391:
384:
382:
380:
376:
372:
368:
367:
361:
359:
355:
351:
347:
340:
338:
335:
333:
329:
325:
320:
316:
312:
308:
303:
298:
294:
290:
289:
288:Acanthochelys
280:
278:
276:
272:
268:
264:
260:
256:
252:
251:
246:
242:
229:
225:
221:
217:
213:
209:
208:
206:
203:
199:
192:
190:
184:
181:
180:Binomial name
177:
173:
172:
167:
164:
163:
160:
159:
158:Acanthochelys
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
76:
71:
67:
61:
56:
45:
41:
36:
32:
27:
22:
19:
2313:macrocephala
2112:
2085:
2056:
2049:
2042:
2035:
2028:
2021:
2004:
1988:
1971:
1955:
1940:Erymnochelys
1938:
1898:
1891:
1884:
1877:
1870:
1863:
1856:
1849:
1842:
1835:
1828:
1821:
1814:
1807:
1800:
1793:
1786:
1769:
1753:
1736:
1696:
1681:Pseudemydura
1679:
1650:
1643:
1626:
1597:
1589:
1572:
1556:
1549:
1542:
1535:
1518:
1502:
1495:
1488:
1481:
1474:
1467:
1450:
1434:
1417:
1401:
1394:
1387:
1380:
1373:
1363:
1351:
1343:
1335:
1328:
1321:
1313:Pelocomastes
1311:
1299:
1292:
1285:
1275:
1264:
1248:
1238:
1226:
1219:
1212:
1205:
1198:
1191:
1184:
1177:
1170:
1159:
1147:
1140:
1133:
1126:
1119:
1111:
1104:
1097:
1090:
1080:
1069:
1061:Chelodininae
1040:
1023:
1007:
990:
974:
957:
941:
934:
927:
919:
912:
895:
879:
872:
864:
857:
850:
843:
836:
829:
822:
815:
808:
791:
775:
767:
760:
743:
727:
720:
713:
707:
706:
689:
556:Side-necked
509:
501:
463:. Retrieved
449:
443:
437:
429:
414:
412:
388:
370:
364:
362:
354:Rio Paraguay
345:
344:
341:Distribution
336:
334:is 2n = 48.
309:. The broad
286:
284:
249:
248:
244:
240:
238:
227:
219:
211:
188:
186:
170:
169:
157:
18:
2217:iNaturalist
2137:Wikispecies
1628:Hydromedusa
992:Ranacephala
793:Mesoclemmys
465:17 November
281:Recognition
2330:Categories
2006:Podocnemis
1738:Pelomedusa
601:Pleurodira
599:Suborder:
595:Testudines
587:Subclass:
421:References
394:hatchlings
137:Pleurodira
133:Suborder:
127:Testudines
1574:Rheodytes
1520:Myuchelys
1161:Chelydera
1082:Chelodina
1071:Chelodina
620:Subfamily
569:Kingdom:
332:karyotype
328:ventrally
326:, yellow
302:Vertebral
271:Argentina
261:found in
165:Species:
93:Kingdom:
87:Eukaryota
2261:97259978
2235:10362106
2122:Wikidata
1771:Pelusios
1025:Rhinemys
959:Platemys
897:Phrynops
681:Chelinae
672:Chelidae
638:Subgenus
589:Anapsida
583:Reptilia
577:Chordata
575:Phylum:
571:Animalia
403:plastron
399:carapace
350:Pantanal
324:dorsally
311:plastron
297:carapace
293:carapace
275:Paraguay
259:Chelidae
202:Synonyms
147:Chelidae
143:Family:
117:Reptilia
107:Chordata
103:Phylum:
97:Animalia
83:Domain:
60:IUCN 3.1
2209:2441991
2128:Q304406
1452:Emydura
648:Species
593:Order:
581:Class:
560:species
390:Nesting
375:marshes
315:pigment
267:Bolivia
153:Genus:
123:Order:
113:Class:
58: (
2300:375069
2274:440077
2248:551877
2196:791049
2170:190955
2152:ARKive
2015:
1982:
1949:
1931:
1780:
1747:
1729:
1690:
1637:
1583:
1529:
1461:
1428:
1419:Elusor
1277:Elseya
1266:Elseya
1034:
1001:
968:
906:
802:
754:
745:Chelus
700:
611:Family
558:turtle
409:Status
379:swamps
273:, and
263:Brazil
255:turtle
2287:19159
2230:IRMNG
2222:39610
2183:64CCD
629:Genus
2269:NCBI
2256:IUCN
2243:ITIS
2204:GBIF
2165:BOLD
467:2021
450:2018
239:The
2191:EoL
2178:CoL
454:doi
243:or
2332::
2310::
2308:RD
2297::
2284::
2271::
2258::
2245::
2232::
2219::
2206::
2193::
2180::
2167::
2154::
2139::
2124::
489:^
475:^
448:.
442:.
401:,
377:,
277:.
269:,
265:,
1596:â€
1350:â€
1342:â€
1169:â€
1118:â€
926:â€
871:â€
774:â€
549:e
542:t
535:v
469:.
456::
440:"
436:"
247:(
62:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
