Specimens of Tyrannosaurus

2745:"Scotty" was discovered by Robert Gebhardt, a high school principal from Eastend, SK who accompanied palaeontologists from the Royal Saskatchewan Museum on a prospective expedition into the Frenchman Formation in southwestern Saskatchewan on 16 August 1991. It wasn't until June 1994 that the Royal Saskatchewan Museum was able to begin the excavation, which was led and overseen by the Museum's Ron Borden, as well as resident paleontologists Tim Tokaryk and John Storer who were with Gebhardt when he uncovered the first fossils. The bones were deeply packed in dense, iron-laden sandstone, which took more than twenty years for the team to fully remove, excavate, and assemble the majority of the skeleton, with additional trips being made to the site to retrieve smaller bones and teeth. The entire process of excavating the skeleton was also slowed down by its considerable size. 2748:"Scotty" is reported to be 13.1 metres (43 feet) long and weighed an estimated 8.8 tonnes (8,800 kg). Despite it not being a complete fossil, paleontologists were able to create the estimation for the weight and length through measurements of important weight bearing bones such as the femur, hip, and shoulder bones that have all been measured to be larger and thicker with "Scotty" than the corresponding bones with "Sue". Going from the latest study "Scotty" exceeds "Sue" in 84.6% of the published measurements. While the reported measurements and weight for "Scotty" are larger than those of "Sue", some scientists posit that the two fossils are too close in size to officially declare "Scotty" the largest. 2980: 3239: 3173:"Barbara's" circumstances are particularly rare, more so when taken into account that she suffered and survived long after a debilitating foot injury. While she was no longer able to capture her prey, it is suggested that she got by with the help of a mate or cohort feeding her, as the injury would have rendered her immobile for upwards of 6 months. This is vaguely supported by trackway evidence that has been used to imply tyrannosaur group hunting. It is doubtful that "Barbara" ever successfully hunted again as a predator unless its prey was nearby. 2769: 1907: 1698: 3106:. The specimen was found in Late Cretaceous deposits and it is estimated to be 66.3 million years old. The Tufts-Love rex is undergoing preparation by Michael Holland and his team at the Burke Museum. The skeleton is estimated to be 30% complete, but it includes a complete (all of the bones of the skull and jaws are preserved) and mostly articulated skull. Holland describes the skull as minimally distorted and in an "exquisite" state of preservation. 2324: 1792: 1527: 2809: 74: 2845: 1403: 1579:

balancing with tail outstretched. Along with a more lifelike posture, the specimen also now includes a composite reconstruction of the skull by Michael Holland. It has been reconstructed in recent years, it measured an estimated 11.9 meters in length and an estimated weight of 7.4–14.6 metric tonnes, 9.1 metric tonnes being the average estimate in that study, although most earlier studies have suggested lower weight figures.

2587: 2428: 1460: 2718: 2647: 3155: 3115: 2911: 43: 2869: 2122: 3079: 1643: 1588: 2209: 3296: 2524: 2000: 2958:), due to the fact that the museum was undergoing restoration. When the Netherlands exhibition ended, it continued travelling through other European countries in 2017, 2018 and 2019. As of August 2019, Trix was returned to display at the Naturalis museum where it is installed in a special room that was under construction during Trix's European tour. 3227:". Given the injuries present on both fossils, it has been theorized that both died while fighting one another. The specimen is presently undergoing study. Important biological data is likely preserved within the specimen, including body outlines, skin impressions, soft tissues, injuries, stomach contents, and even original proteins. 2503:(2011) also has a T-shaped postorbital. Paleontologists who support the theory that Jane represents a juvenile believe the tyrannosaur was approximately 11 years old at its time of death, and its fully restored skeleton measured 6.5 meters (21 ft) long, a bit more than half as long as the largest-known complete 1555:, assistant curator of the American Museum of Natural History and a famous paleontologist in his own right. He forwarded news of it to Osborn; it would be three years before they found the rest of it. In 1905 when the type was described by Osborn, previous knowledge of dinosaur predators at the time were based on 2855:

Victoria is a specimen found near Faith, South Dakota in 2013. Victoria is estimated to be around 12 ft tall and 40 ft long, and she is thought to have died in her subadult stage, between 15 and 25 years of age. Victoria has also been the subject of a traveling exhibition being displayed in

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specimen discovered in South Dakota by Bucky Derflinger in 1999 at the same site as AMNH 3982. Excavations of this 10% complete skeleton began in 2000. The known material includes a partial skull, several vertebrae, and ribs. A very wide femur with a length of 1300 mm and a circumference of 630 mm is

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casts (replicas) or parts, but Tristan is one of only two original skeletons on display in the continent (the other is "Trix" in the Netherlands). The matte-black fossilised skeleton is about 12 m (39 ft) long and 3.4 m (11 ft) tall at the hips. Tristan is among the most complete

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fossils, "Scotty" shows signs of trichomoniasis, a parasitic infection in the jaw that left visible holes in the bone and was unique to this specific species of dinosaur. Additionally, a broken and healed rib on its right side, broken tail vertebra, as well as a hole near the eye socket are possibly

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Sue has a length of 12.3–12.4 meters (40.4–40.7 ft), stands 3.66–3.96 m (12–13 ft) tall at the hips, and according to the most recent studies estimated to have weighed between 8.4 and 14 metric tons when alive. It has been hypothesised that Sue's impressive size may have been achieved

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for its apparently small adult size. The specimen is estimated to have been around 5.2 meters (17 ft) long when it died. However, a detailed analysis of the specimen by Thomas Carr in 1999 showed that the specimen was, in fact, a juvenile, leading Carr and many other paleontologists to consider

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specimen was discovered on private land in Harding County, South Dakota, once in 1981 by Michael and Dee Zimmerschied, and again on 4 October 1992 (Alan and Robert Detrich re-discovered Samson after it was originally found and deemed by paleontologists that several bones had washed in and there was

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for protection against possible bombing raids. The specimen, now labeled CM 9380, is still mounted in Pittsburgh, at first with the tail acting as a tripod in the old-fashioned kangaroo pose. It has since received a modernization of its posture (mounted by Phil Fraley and crew) and can now be found

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specimen at more than 98% preservation, was discovered in Montana in 2006. Following years of unsuccessful attempts to sell it to museums or auction it off, the NCMNS started negotiations in 2016, which were prolonged due to a legal battle over the rights to the fossil, which was resolved in 2020.

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skeletons: It was re-assembled from about 300 separate parts, 170 of which are original (including 98% of the skull and all the teeth), the rest reproductions. It is estimated to have died when about 20 years old and it was in poor health, having several bone fractures, bite marks to the skull and

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in 2001. Despite having a typically female name, Jane's sex is unknown—the specimen was named after Burpee Museum benefactor Jane Solem. The specimen was found in the summer of 2001 by Carol Tuck and Bill Harrison on an expedition led by Burpee Museum curator Michael Henderson. After four years of

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forelimb. It has an estimated length of around 11.6 meters (38 ft) and a weight between 5.8 metric tons (6.4 short tons) and 10.8 metric tons (11.9 short tons) in newer figures. It is estimated that the "Wankel rex" was 18 years old when it died, an adult but not completely grown. The "Wankel

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ever discovered in the world, the largest of any dinosaur discovered in Canada, and as one of the oldest and most complete fossils of its kind at more than 70% bulk. "Scotty" resides at the Royal Saskatchewan Museum's T. rex Discovery Centre in Eastend, SK, Canada. In May 2019, a second mount was

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in the Charles M. Russell National Wildlife Refuge in Garfield County, Montana. Its discoverer was Bob Harmon, a fossil preparator for the Museum of the Rockies, and was nicknamed the "B-rex" (or "Bob-rex") in honor of Harmon. The specimen was discovered in 2000, and excavated by MOR from 2001 to

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have been sold worldwide, each for a price of about $ 100,000. Stan's skeleton was auctioned for $ 31.8 million in a 2020 Christie's New York sale, making it a record-breaking dinosaur sale, with the buyer eventually being revealed as the under construction Natural History Museum Abu Dhabi in the

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The mount retained a rearing pose similar to the initial proposal. By the 1980s it was generally accepted that such a pose would have been anatomically impossible in life, and the skeleton was re-mounted in a more accurate, horizontal pose during a renovation of the museum's dinosaur halls in the

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blood cells and vessels. However, since an unknown process distinct from normal fossilization seems to have preserved the material, the researchers are being careful not to claim that it is original material from the dinosaur. Paleontologist Thomas Kaye of the University of Washington in Seattle

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museum in Washington, D.C., for 50 years. (The Museum of the Rockies continues to display a cast reconstruction of the skull by Michael Holland). The specimen went on temporary display on National Fossil Day, 16 October 2013, and was exhibited until the museum's dinosaur hall exhibit closed for

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collaborated with the team in the excavation. The bones were cleaned and assembled in a mount at Black Hills Institute's installations, with the help of both Chicago's Field Museum of Natural History and the Naturalis Museum in Leiden. Chicago's Field museum sent digital models of their famous

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specimens ever found. Baby Bob has been fully excavated, although it will take another year to clean. Detrich said the skull, which is about 75 percent complete, and most of the major skeletal elements were found strewn across a flood plain, although very few vertebra and ribs were found.

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from Montana. The partial skeleton was found by two Burke Museum volunteers, Jason Love and Luke Tufts, and was named the "Tufts-Love" rex. Paleontologists at the Burke Museum believe that the Tufts-Love rex was around 15 years old when it died. The skull is of average size for an adult

1668:. However, technical difficulties prevented the mount from being executed. One obvious problem was that the Cretaceous Dinosaur Hall was too small to accommodate this dramatic display, and AMNH 5027 was already mounted by itself as the central attraction of the hall. The forearms of 2776:

Commercial paleontologist Craig Pfister discovered the specimen in the lower Hell Creek Formation in Carter County, Montana, in 2010. Its excavation and preservation lasted four years. It was later sold to Danish-born investment banker Niels Nielsen, who loaned the specimen to the

1750:, where the original specimen was housed and is now on display. Their initial research indicated that the skull bones were fused, and that it therefore represented an adult specimen. In light of this, Bakker and colleagues assigned the skull to a new genus, named 3069:

for use in the exhibition, alongside the display of the real fossil skeleton, and which remained at the museum after the end of the exhibition. For the exhibit at the Nottingham Natural History Museum at Wollaton Hall, Titus was reconstructed in a walking pose.

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2003. Although only 37 percent of the skeleton was present, this included almost all of the skull (although the skull was nearly completely disarticulated). The specimen also includes several cervical, dorsal, sacral, and caudal vertebrae; several

3038:. External bone inspection has revealed injuries to Titus' right tibia (possibly a claw or bite wound); a deformed toe on the right foot; and a bitten and healed tail. The bite wound near the end of the tail indicates a possible attack by another 2785:

for one year, and expected back in 2021. Nielsen and his friend Jens Jensen named the specimen Tristan-Otto (short: Tristan) for their sons. The Museum für Naturkunde Berlin lists it under specimen number MB.R.91216. Several European museums have

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in 2000 under the name of "Z-rex", with an asking price of over US$ 8 million. It failed to sell online but was purchased for an undisclosed price in 2001 by British millionaire Graham Ferguson Lacey, who renamed the skeleton "Samson" after the

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in the Latest Cretaceous of North America: A Comment on "The Tyrant Lizard King, Queen and Emperor: Multiple Lines of Morphological and Stratigraphic Evidence Support Subtle Evolution and Probable Speciation Within the North American Genus

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as entire sections of bone were damaged, and some were split open by huge crushing bite-forces. The nature of the crushing on the femur and tibia, along with size of the bite marks, indicates that these bones were bitten through by another

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paleontologists in southeastern Montana. At 17 years old, 34 ft (10 m) long and nearly 7,000 lb (3,200 kg), it is estimated to be a 70% complete specimen. Thomas is mounted in a "growth series" with the youngest-known

2679:. It was prepared by the Carnegie Museum starting in May 2004. After preparation was complete in March 2006, the specimen was returned to Lacey. It, along with some other dinosaur skeletons, was sold again at auction on 3 October 2009. 2629:

biofilm created by bacteria while digesting and breaking down the original specimen. He has discovered this to be true in many specimens from the same area. In 2016, it was finally confirmed by Mary Higby Schweitzer and Lindsay Zanno

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he belonged to, claimed ownership, the Institute had considered itself to have. In 1997, the suit was settled in favor of Williams and the fossil was returned to Williams' ownership. Williams quickly offered up "Sue" for auction by

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and colleagues illustrated the concept of parasitic infections in dinosaurs by analysing the lesions found on the cranial bones of Black Beauty. The specimen has been used to study comparative morphology between tyrannosaurids and

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and shown to their teacher. Soon afterward, the Royal Tyrrell Museum was contacted, and excavation of the sandstone matrix surrounding the fossils began in 1982. The dig site where the fossil was found is located at approximately

2698:. Detrich unearthed the fossil dubbed "Baby Bob" in a fossil-rich area near the Eastern Montana town of Jordan. Its femur measures about 25 inches, and if all the preliminary data pans out, that would make it among the smallest 3126:

by an anonymous owner, currently on display alongside “Barbara” until the end of 2023. The specimen is estimated to be 66.8 million years old and almost of adult size. He was recovered from Niobrara County, Wyoming.

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Larson, P. L. 2013. The case for Nanotyrannus in J. M. Parrish, R. A. Molnar, P. J. Currie., and E. B. Koppelhus (eds.) Tyrannosaurid Paleobiology, University of Indiana Press, Bloomington and Indianapolis, pp.

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in having a higher tooth count, large pneumatic foramen on the center of the quadratojugal, T-shaped postorbital, and fused shoulder blade and pelvis, Yun (2015) concurred with the opinion of most workers that

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early 1990s. The mount can still be seen on display on the fourth floor of the American Museum. The American Museum of Natural History features AMNH 5027 in its famed Hall of Saurischian Dinosaurs to this day.

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Considered the 14th T-rex specimen to be found. The first in-situ t-rex to be known from the state of North Dakota. Discovered and collected by amateurs and shows tooth marks on bones, indicating scavenging.

2031:. The specimen, given the number MOR 555 but informally called the "Wankel rex," includes approximately 80-85 percent of the skeleton, including the skull, as well as what at the time was the first complete 4216: 800:

Proposed to be largest known specimen, scientific consensus not yet reached. Exceeds "Sue" in 84.6% of the published bone measurements. Has been estimated to be larger than "Sue" in two published studies.

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are exhibited at Museum of the Rockies as part of a full skeletal mount completed with cast elements replacing the missing bones. This mount was installed after the Wankel Rex (now nicknamed The Nation's

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Dalman, Sebastian G.; Loewen, Mark A.; Pyron, R. Alexander; Jasinski, Steven E.; Malinzak, D. Edward; Lucas, Spencer G.; Fiorillo, Anthony R.; Currie, Philip J.; Longrich, Nicholas R. (11 January 2024).

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skeleton. This specimen, named "Sue" in honor of its discoverer, soon became embroiled in a legal battle over its ownership. The owner of the land the fossil was found, Maurice Williams, as well the

4515:"The Tyrant Lizard King, Queen and Emperor: Multiple Lines of Morphological and Stratigraphic Evidence Support Subtle Evolution and Probable Speciation Within the North American Genus Tyrannosaurus" 7096: 6030: 4613: 3826:

Tsuihiji, T.; Watabe, M.; Tsogtbaatar, K.; Tsubamoto, T.; Barsbold, R.; Suzuki, S.; Lee, A.H.; Ridgely, R.C.; Kawahara, Y.; Witmer, L.M. (2011). "Cranial osteology of a juvenile specimen of

2515:(2011), the weight of the Jane specimen in life was probably between 639 kg (1,409 lb) and 1,269 kg (2,798 lb), 954 kg (2,103 lb) being the average estimate. 3053:. Excavation of the specimen began in 2018, and took 18 months. The bones of "Titus" were shipped to conservationist Nigel Larkin in the UK, who constructed the mount using a cast of the 1892:

Replicas of Black Beauty have been shown in some exhibitions and museums, including both simple skull montages and complete skeletons. Casts are on display in museums around the world.

1863: 6924: 1811:. The nickname stems from the apparent shiny dark color of the fossil bones, which occurred during fossilization by the presence of minerals in the surrounding rock; it was the first 86:

is one of the most iconic dinosaurs and is known from numerous specimens, some of which have individually acquired notability due to their scientific significance and media coverage.

6295: 6251: 2954:. The first exhibit spanned from 10 September 2016 to 5 June 2017 and was set at the only room of the Naturalis museum open to public at the time (the 17th-century building known as 2480:. However, the Jane material has yet to be properly studied and described by scientists. Although Larson (2013) saw Jane as more identical to CMNH 7541 and LACM 28471 than to adult 512:

One of the best-known specimens, with casts present in numerous institutions. Sold to a private owner in 2020 for US$ 31 million, formerly in the Black Hills Institute as BHI 3033.

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in having smaller postorbital crests, a proportionately longer and shallower lower jaw with a less prominent chin suggestive of a weaker bite, and more laterally compressed teeth.

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rex" was also one of the first fossil dinosaur skeletons studied to see if biological molecules still existed within the fossilized bones. Doctoral candidate Mary Schweitzer found

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Burnham, M.; Mclain, M.; Nelson, D.R.; Snyder, K. (2018). "Tyrannosaur cannibalism: a case of a tooth-traced tyrannosaurid bone in the Lance Formation (Maastrichtian), Wyoming".

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Peter the T. rex, a Paleontological Study on a New Specimen of Tyrannosaurus Rex Named 'Peter', from the Lance Formation (Maastrichtian) of Wyoming specimen number AWMM-IL 2-22.9

1971:. One of the most prominent injuries are in the neck and the skull. A piece of bone is missing at the rear, and the skull also bears a hole 1 inch wide, probably made by another 6761: 6529: 6440: 6008: 4694: 2398:, or belly ribs, and an ulna, or lower arm bone. As of now, 101 bones, or about 34% of Bucky's skeleton, has been discovered and verified. Bucky is the sixth-most complete 7111: 469: 5978: 4450: 4237: 3255: 1355: 2073: 1623:

PR2081, for instance) the cervical series of AMNH 5027 is much more gracile, so with later discoveries the distinction between tyrannosaurid necks and the necks of

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carcass. It has been stated by Pete Makovicky, the Chicago museum's lead curator of dinosaurs, that this specimen is in the same size range as "Sue" and "Scotty".

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were not well documented and the hands were unknown, so for the sake of the display, the forearms of AMNH 5027 were given three fingers, based on the forelimbs of

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known. Sue's age at the time of death was estimated by Peter Mackovicky and the University of Florida to be 28 years old, over 6–10 years older than most big

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Osborn planned to mount the similarly sized AMNH 5027 and AMNH 973 together in dynamic poses. Designed by E.S. Christman, the scene was to depict a rearing

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Carr, Thomas D.; Napoli, James G.; Brusatte, Stephen L.; Holtz, Thomas R.; Hone, David W. E.; Williamson, Thomas E.; Zanno, Lindsay E. (1 September 2022).

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nothing left). It was shortly after that when Fred Nuss and Candace Nuss of Nuss Fossils with the Detrich brothers found the most complete and undistorted

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had become jumbled with the remains of the bigger fossil. Following the 1941 entry of the United States into World War II, the holotype was sold to the

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in 1902 and excavated over the next three years. Another specimen (AMNH 5866), found in Wyoming in 1900, was described in the same paper under the name

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have also been studied as they show evidence of use. This evidence includes the construction of metacarpal III, as well as repeated fractures in the

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A juvenile or baby specimen. The discoverers have stated its age to be 4 years based on histology. An attempt was made to sell the specimen on eBay.

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places such as the Arizona Science Center. Her cause of death is unknown; however, she was believed to have been bitten in the lower jaw by another

2332: 1779:(LACM 23844) which included most of the skull of a very large, mature animal. When it was put on display in Los Angeles, LACM 23844 was the largest 860: 3166:

by the same anonymous owner as "Peter", currently on display alongside him until the end of 2023. The pair will be the first adult male and female

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that the soft tissue was medullary bone tissue, like that in modern birds when they are readying to lay eggs. This confirmed the identity of the

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One of the largest known specimens that is also well preserved. Has been stated to be the oldest known specimen but this has not been confirmed.

6129: 5140: 5100: 3408: 7080: 4198: 6932: 5588: 5210: 3947: 1870: 1800: 421: 2263:. The left hindleg is relatively complete with a 1.2-meter-long (3.9 ft) femur, missing only some toe bones. The forelimbs include the 7106: 5714: 2069: 961:

An adolescent specimen, approximately 14 years old. Considered by a minority of paleontologists to be a separate taxon from Tyrannosaurus.

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and colleagues announced the recovery of soft tissue from the marrow cavity of a fossilized femur belong to B-Rex. Flexible, bifurcating

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was not previously known, so it was this specimen that brought the short, stocky tyrannosaur neck to light. Compared to later specimens (

4097: 1759:. The current classification of CMNH 7541 is not universal, with some research suggesting the specimen belongs to a taxon distinct from 1747: 1717: 329: 5839: 5477: 5154: 3146:. Explanations for this behavior range from response to over-crowded populations, limited food supply, sexual dominance, or even play. 1819:

finds. Black Beauty was found in 1980 by a high school student, Jeff Baker, while on a fishing trip with a friend in the region of the

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Crosslin, Rick; Mary Fortney; Dinosphere Exhibit Development Team (2004). "Tyrannosaurus rex Attack Scene-What will be the outcome?".

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in Berlin, Germany, for research and exhibition. It has been on display at Museum für Naturkunde between 2015 and 2020, moving to the

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Also known as The Nation's Rex. Specimen on loan to the Smithsonian Institution, with a cast on display at the Museum of the Rockies.

284: 241: 143: 6409: 2833: 7116: 6451: 3690: 3595: 2849: 2782: 1616: 1217: 1065: 6339: 1947:, Inc. center. It is a well known specimen, and one of the most complete, with 199 bones recovered. About 30 casts of the original 5048: 4701: 4351:"A Computational Analysis of Limb and Body Dimensions in Tyrannosaurus rex with Implications for Locomotion, Ontogeny, and Growth" 3613:"A Computational Analysis of Limb and Body Dimensions in Tyrannosaurus rex with Implications for Locomotion, Ontogeny, and Growth" 5902: 3311: 2468: 2436: 2107:

when they were approached by various media outlets for comment. Their criticism was subsequently published in a technical paper.

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remains were later identified as those of a theropod rather than a ceratopsid, and H.F. Osborn recognized the similarity between

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found. Between 75% and 80% of its skeletal volume was recovered. They are thought to have been at least 30 years old at death.

2609: 2162: 2126: 1620: 731: 2307:) was loaned to the Smithsonian to occupy a central part in the museum's dinosaur hall, featuring a dynamic mount showing the 6503: 5982: 3350:"A giant tyrannosaur from the Campanian–Maastrichtian of southern North America and the evolution of tyrannosaurid gigantism" 3035: 1089: 190: 4244: 2825:

fossil, a two-year-old, 11-foot-long (3.4 m) specimen, and a 13-year-old, 20-foot-long (6.1 m) juvenile specimen.

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attack. Other abnormalities, such as impacted teeth, suggest that "Scotty" was not only bitten, but also bit other animals.

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Preparation of "Sue" (FMNH PR2081) was completed at the Field Museum and the skeleton was placed on exhibit on 17 May 2000.

1498:. At the time of their initial description and naming, these specimens had not been fully prepared and the type specimen of 7066: 6371: 5421:

Schweitzer, M.H.; Wittmeyer, J.L.; Horner, J.R.; Toporski, J.B. (2005). "Soft Tissue Vessels and Cellular Preservation in

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specimen, FMNH PR 2081 (Sue) to complete the cast and Naturalis museum replicated the bones using 3D-printing technology.

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Crosslin, Rick; Mary Fortney; Dinosphere Exhibit Development Team (2004). "Resource Materials: Bucky-Tyrannosaurus rex".

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erected at the Royal Saskatchewan Museum in Regina, where the exhibit reflects the recent discoveries about the fossil.

2092: 2028: 2024: 1881: 1315: 1276: 5070: 5870: 2065: 833: 50: 1432:, meaning "giant porous vertebra" in reference to the numerous openings for blood vessels he found in the bone. The 7075: 6296:"Victoria, the 66-million-year-old Tyrannosaurus rex, will have her exhibit extended at the Arizona Science Center" 6252:"Victoria, the 66-million-year-old Tyrannosaurus rex, will have her exhibit extended at the Arizona Science Center" 4957: 2947: 5233:

Larson, Neal L.; Peter Larson; Kenneth Carpenter (2008). "One Hundred Years of Tyrannosaurus Rex: The Skeletons".

3142:. There is also a set of smaller, parallel tooth marks nearby on the shaft that are not attributable to an adult 2885: 2375:. Part of The Children's Museum of Indianapolis Dinosphere exhibit, Bucky is displayed along with Stan, an adult 1985:

bite. The bite marks are healed, indicating that Stan survived the wounds. Stan could also have been infected by

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in Osborn's 1905 work, it was considered the older name and has been used since. Had it not been for page order,

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Larkin, N.R., Dey, S., Smith, A.S. and Evans, R. 2022. 21st Century Rex: maximising access to a privately owned

2367:. It was discovered by rancher and cowboy Bucky Derflinger. Bucky was well preserved and easily prepared by the 1831: 7005: 5602:"Bonhams : The T. rex known as Samson - One of the Most Complete Tyrannosaurus rex Specimens in Existence" 3094: 2797: 2064:" became the centerpiece of the dinosaur hall when it reopened in 2019. Casts of MOR 555 are on display at the 6955: 6102: 4929: 3258:, the fossil material (NMMNH P-3698) consists of skull and lower jaw bones, in addition to isolated teeth and 2778: 6904:"North Carolina Museum of Natural Sciences to receive the "Dueling Dinosaurs" | Programs and Events Calendar" 4767:"Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Winter 2011 Appendix" 4562: 4162: 3931:

Larson, Peter H. (2008). "One Hundred Years of Tyrannosaurus Rex: The Skeletons.". In Larson, Neal L. (ed.).

3130:“Peter” is one of only four incredibly rare and visually stunning obisidian black colored tyrannosaurus rex. 6277: 6177: 4730:: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull". 2474:

The Jane specimen has been central to the debate regarding the validity of the proposed tyrannosaurid genus

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from the McRae Formation (Lancian, Upper Cretaceous), Elephant Butte reservoir, Sierra County, New Mexico"

5887: 5840:"Scotty vs. Sue: Is the Canadian T. rex really bigger than Chicago's? The Field Museum disputes new study" 5155:"Scotty vs. Sue: Is the Canadian T. Rex really bigger than Chicago's? The Field Museum disputes new study" 4488: 3611:

Makovicky, Peter J.; Allen, Vivian; Molnar, Julia; Bates, Karl T.; Hutchinson, John R. (12 October 2011).

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in 1991. Since its discovery and extensive subsequent study, "Scotty" has been referred to as the largest

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includes a relatively complete skull with jaws, multiple vertebrae of the back and tail, a well preserved

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Schweitzer, Mary Higby; Zheng, Wenxia; Zanno, Lindsay; Werning, Sarah; Sugiyama, Toshie (15 March 2016).

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The specimen belongs to a private collector, but 3D-printed replicas of the bones are accessioned in the

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Named 'Barbara', from the Hell Creek Formation (Maastrichtian) of Montana specimen number AWMM-IL2022.21

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in 2014 and excavated in 2018. It is 20% complete, and was named after the protagonist in Shakespeare's

2950:. At arrival in the Netherlands, it started touring on public display in an itinerant exhibition titled 2605: 2368: 2237: 2020: 1930: 688: 649: 609: 567: 529: 507: 6397:

http://www.thefossilforum.com/uploads/monthly_2016_09/10.thumb.jpg.592a065fd0a51cd2c23fa6d1a2063c29.jpg

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by Neal Larson. Black Hills Institute of Geological Research, Inc. 19 May 2000. Retrieved 16 July 2006.

3518:

Osborn, Henry Fairfield; Brown, Barnum (1906). "Tyrannosaurus, Upper Cretaceous carnivorous dinosaur".

2768: 1906: 5805: 6701: 5516: 5434: 4894: 4657: 4526: 4514: 4362: 4273: 4046: 3792: 3624: 3009: 2455: 2348: 2236:

in 1997. The discovery was made by Louis E. Tremblay on 4 July 1997 working under the supervision of

1922: 1328: 1252: 1208: 1169: 1119: 1078: 1034: 993: 953: 912: 873: 828: 820: 746: 701: 662: 622: 580: 542: 502: 395: 344: 299: 260: 158: 6585: 5359: 5337: 4260:

Wolff, Ewan D. S.; Salisbury, Steven W.; Horner, John R.; Varricchio, David J. (30 September 2009).

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and fibrous but elastic bone matrix tissue were recognized. In addition, microstructures resembling

6611: 6556: 6534: 6133: 6034: 5505:"Chemistry supports the identification of gender-specific reproductive tissue in Tyrannosaurus rex" 3267: 3251: 2923: 2411: 2352: 2145: 2100: 1421: 1371: 917: 878: 790: 751: 163: 153: 5481: 3955: 2165:

in Chicago for US$ 8.4 million—the highest price ever paid for a fossil before being surpassed by

632:

One of the largest specimens,although size estimation is difficult due to its fragmentary nature.

447: 7032: 6986: 6717: 5693: 5458: 4747: 4542: 3892: 3847: 3808: 2464: 1978: 1731: 1608: 943: 255: 6925:"'Dueling Dinosaurs' fossil, hidden from science for 14 years, could finally reveal its secrets" 5722: 5591:

by Dan Lagiovane. Carnegie Museum of Natural History . 2 March 2006. Retrieved 30 December 2010.

3587: 3580: 2796:

signs of disease in the jaw. The disease present in Tristan's jaw was suggested to be a case of

2355:. The skeleton, transported by water, ended up in a low shallow valley along with bones from an 2076:. A bronze cast of the specimen, known as "Big Mike", stands outside the Museum of the Rockies. 5296: 4119: 2961:

According to Peter Larson (director of Black Hills Institute), Trix is among the most complete

2287:

has been the subject of research regarding parasitic infections in dinosaurs. The forelimbs of

1865:

near the confluence of the Crowsnest and Willow Rivers, and consisted of rock belonging to the

1697: 1042:

Currently the most complete specimen known to science,an adolescent preserved entangled with a

6978: 5903:

https://docs.google.com/spreadsheets/d/1xbuGjf_PaBcJNKJSu4sEixEqZzEc-wyvfrg1iTnCSno/edit#gid=0

5813: 5685: 5552: 5534: 5450: 5206: 4961: 4675: 4444: 4390: 4301: 4141: 4074: 3686: 3670: 3652: 3591: 3389: 3371: 3224: 3182: 2730: 2712: 2626: 2600: 2537: 1911: 1658:(AMNH 5027) snapping at another cowering one (AMNH 973), as they fought over the remains of a 1009: 5389: 5248: 4101: 2347:

ever placed on permanent exhibit in a museum. The dinosaur remains were found in 1998 in the

7024: 6970: 6709: 6155: 5675: 5542: 5524: 5442: 5401: 5022: 4822: 4739: 4665: 4534: 4380: 4370: 4291: 4281: 4064: 4054: 3882: 3839: 3800: 3719: 3674: 3642: 3632: 3560: 3527: 3500: 3492: 3458: 3379: 3361: 3301: 3014: 2988: 2974: 2323: 2088: 2048: 1824: 1791: 1739: 1735: 1526: 1186: 693: 654: 614: 572: 534: 6357: 3564: 2808: 1607:

in Montana, and described by Osborn in 1912 and 1916. At the time of discovery, a complete

7070: 5747: 5159: 5009: 4990: 4972: 4248: 4169: 3250:

The remains of a tyrannosaur were discovered in 1983 in the Campanian-early Maastrichtian

3058: 2905: 2889: 2691: 2190: 2166: 2133: 1981:

are fused, and another has additional bone growth. This could have been caused by another

1953: 1901: 1289: 1138: 1124: 1111: 988: 741: 480: 205: 6956:"Late Cretaceous (Lancian) dinosaurs from the McRae Formation, Sierra County, New Mexico" 3737:

Gilmore, C. W. (1946). "A new carnivorous dinosaur from the Lance Formation of Montana".

756:

The largest known specimen at the time of discovery as well as one of the most complete.

6705: 5520: 5438: 4962:"Oldest T. rex Trix on view in Leiden, VUB researchers established the dinosaur's age", 4661: 4530: 4366: 4277: 4050: 3977: 3796: 3628: 3384: 3349: 3191:) is a nickname for a nearly complete tyrannosaurid (generally considered an adolescent 1402: 73: 6223: 5547: 5504: 4385: 4350: 4296: 4261: 4092: 4069: 4034: 3647: 3612: 3134: 3062: 3027: 3022:

for 13 months beginning July 2021, it was during that time only the second specimen of

2586: 2532: 2508: 2427: 2116: 1820: 1542: 1502:

had not even been fully recovered. In 1906, after further preparation and examination,

1463: 719: 589: 439: 17: 2079:

In 2022, Gregory S. Paul and colleagues argued that the Wankel rex was not actually a

1678:(the more allosaur-like arms were replaced several years later when better fossils of 1459: 7090: 7036: 6990: 6954:

Lozinsky, Richard P.; Hunt, Adrian P.; Wolberg, Donald L.; Lucas, Spencer G. (1984).

6721: 5366: 4546: 4011: 3019: 3003: 2992: 2577: 2448: 2357: 2308: 2104: 1973: 1935: 1679: 1659: 1603:

With a length of 12.1-12.2 meters, AMNH 5027 was discovered and excavated in 1908 by

1592: 1547: 289: 148: 82: 6639:"Peter the T Rex: Auckland Museum hosts world debut of dinosaur bones found in 2018" 5697: 5462: 4145: 3851: 3812: 2896:, only missing around 3 vertebrae. Ivan is around 40 ft long and 12 feet high. 2717: 2646: 2216:(formerly nicknamed "Peck's Rex) real fossils displayed at the Museum of the Rockies 6182: 5334:"NIU teams up with Burpee Museum to bring world's top dinosaur hunters to Rockford" 3804: 3197: 3188: 3154: 3114: 3046: 2910: 2734: 2613: 2580: 2476: 2372: 2336: 2096: 1926: 1743: 1604: 1552: 1425: 904: 865: 837: 795: 339: 294: 251: 200: 5572: 5406: 2844: 2158: 2087:. This was heavily criticized by several other leading paleontologists, including 5601: 5360:

NIU teams up with Burpee Museum to bring world's top dinosaur hunters to Rockford

5200: 4375: 4349:

Hutchinson, J. R.; Bates, K. T.; Molnar, J.; Allen, V.; Makovicky, P. J. (2011).

4286: 4241: 4217:"Stan the T. rex found! World's most expensive fossil finds home in a new museum" 4059: 3843: 3637: 3170:

to be displayed together. She is one of a few specimens believed to be pregnant.

1815:

specimen to receive a nickname, beginning a trend that continues with most major

7063: 6530:"'Titus' the T. rex is coming to the UK this summer. Here's why it's a big deal" 4563:"They Want to Break T. Rex Into 3 Species. Other Paleontologists Aren't Pleased" 3259: 3219: 3066: 2495: 2363: 1987: 1963:, the skeleton of Stan shows many broken and healed bones. These include broken 1722: 1624: 1559: 1044: 983: 387: 307:

The base skeleton that was used as the iconic symbol for the famous logo of the

4983: 4670: 4636: 4538: 4091:

Many sources says that "Stan" is the second most complete specimen, except for

3551:, a Cretaceous carnivorous dinosaur, the largest flesh-eater that ever lived". 3445:(1917). "Skeletal adaptations of Ornitholestes, Struthiomimus, Tyrannosaurus". 3366: 1635:

skeletons. The skeleton of this specimen was used as the iconic symbol for the

7081:

Comparison of known skeletal material and sizes for five tyrannosaur specimens

3291: 2996: 2868: 2617: 2573: 2531:

This specimen was found in the lower portion of the Hell Creek Formation near

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Peter (Black, Left) & Barbara (Right, Beige) Pose at Auckland Museum 2022

2620:

were found inside the matrix and vessels. The structures bear resemblance to

1627:

became more obvious. This specimen also provided the first complete skull of

1506:

recognized both skeletons as belonging to the same species. Because the name

5446: 5078: 4158: 3871:(Dinosauria: Tyrannosauroidea)—A Distinct Taxon of Small-Bodied Tyrannosaur" 3760:, a new genus of pygmy tyrannosaur, from the latest Cretaceous of Montana". 3078: 2569: 2276: 1664: 1642: 1587: 334: 6854:"Montana's Dueling Dinosaurs To Fetch Up To $ 9 Million At Bonhams Auction" 5689: 5556: 5454: 4394: 4305: 4078: 3887: 3866: 3656: 3496: 3417:

Geological Society of America Annual Meeting — Abstracts with Programs

3393: 3266:, and were described in 1986. In 2024, Sebastian G. Dalman and colleagues 2208: 2185:

specimens, like MOR 555, AMNH 5027 or BHI 3033. The only known specimen of

2121: 1482:, a partial skull and skeleton originally called AMNH 973 (AMNH stands for 1048:. Like Jane, a minority of paleontologists consider it a different taxon. 6713: 5012:

Field Museum of Natural History. 9 September 2005. Retrieved 16 July 2006.

2682:

Samson measured 11.9 m (39 ft), only slightly shorter than Sue.

2295:—possibly caused by heavy loads or pressure (Carpenter and Lipkin, 2005). 1566:

were treated with extreme caution, with suspicion that bones of a smaller

6974: 5748:"World's Largest T. Rex Skeleton nicknamed "Scotty" Discovered in Canada" 5658:

Persons, W. Scott; Currie, Philip J.; Erickson, Gregory M. (April 2020).

5390:"Evidence points out that "Nanotyrannus" is a juvenile Tyrannosaurus rex" 3061:

to supplement the known bones of "Titus", after scanning the bones using

2591: 2545: 2511:," which measures 12.4 m (41 ft) long. According to Hutchinson 2451: 2174: 1567: 1556: 1475: 1406: 1320: 1281: 230: 30: 7028: 2523: 1999: 1420:

consists of two partial vertebrae (one of which has been lost) found by

471:

W longitude. Noted for black colouration taken on during fossilisation.

4751: 3830:

from the Nemegt Formation (Upper Cretaceous) of Bugin Tsav, Mongolia".

3045:

The remains of "Titus" were discovered in September 2014 by commercial

2927: 2621: 2549: 2541: 2459: 2387: 2340: 2292: 2272: 2268: 2264: 2256: 2252: 2233: 1491: 1333: 1294: 1242: 778: 736: 706: 667: 627: 400: 349: 6445:

Named "Titus" Nottingham Natural History Museum Specimen #NGMG 2022-7"

5529: 5249:"Meet the Dinos of Dinosphere | The Children's Museum of Indianapolis" 5141:"New exhibit at Museum of the Rockies features real fossilized T. Rex" 5101:"New exhibit at Museum of the Rockies features real fossilized T. Rex" 3781:"Craniofacial ontogeny in Tyrannosauridae (Dinosauria, Coelurosauria)" 2860:. The bite may have become infected, spreading and leading to sepsis. 6278:"Meet Victoria, one of the most complete T. rex fossils in the world" 5680: 5659: 3723: 3532: 3504: 3462: 2676: 2471:

as the centerpiece of an exhibit called "Jane: Diary of a Dinosaur."

2444: 2229: 1948: 1940: 1918: 1804: 1155: 195: 5573:"Carnegie Museum digs into controversial, but promising T-rex skull" 4743: 4184: 3926: 3924: 3922: 3780: 3407:

Quinlan, Elizibeth D.; Derstler, Kraig; Miller, Mercedes M. (2007).

2930:

where they discovered and unearthed a large and remarkably complete

2060:

renovation in the spring of 2014. The skeleton, named "The Nation's

3262:. Some of the bones were briefly mentioned in 1984 as belonging to 3089:

In 2016 Greg Wilson, David DeMar, and a paleontology team from the

5002: 3237: 3153: 3113: 3077: 2978: 2909: 2867: 2843: 2807: 2767: 2716: 2673: 2645: 2585: 2561: 2557: 2522: 2426: 2322: 2260: 2207: 2153: 2120: 1998: 1968: 1905: 1790: 1696: 1641: 1586: 1525: 1458: 1401: 219:, analysis of the specimen suggests a large individual similar to 72: 6795:

Barbara the T. rex, a Paleontological Study on a New Specimen of

5627:"New Arrivals – T-Rex & Nanotyrannus Teeth From South Dakota" 4433:"Homecoming King: The Nation's T. rex Returns to the Smithsonian" 3254:

in New Mexico by Donald Staton and Joe LaPoint. Reposited at the

1650:

exhibit Osborn planned for the American Museum of Natural History

7004:

Gillette, David D.; Wolberg, Donald L.; Hunt, Adrian P. (1986).

6878: 6793:

Burnham, David A.; Rothschild, Bruce M.; Nudds, John R. (2022).

6663:

Burnham, David A.; Rothschild, Bruce M.; Nudds, John R. (2022).

6439:

Burnham, David A.; Rothschild, Bruce M.; Nudds, John R. (2021).

5806:"'Scotty' the T. Rex Is the Heaviest Ever Found, Scientists Say" 5336:. Northern Illinois University. 7 September 2007. Archived from 4513:

Paul, Gregory S.; Persons IV, W. Scott; van Raalte, Jay (2022).

4321:"Smithsonian Welcomes "The Nation's T. rex" to Washington, D.C." 3334:

skeleton in the digital age. Geological Curator, 11(6), 341–354.

2668: 2565: 2553: 2037: 1444:

as early as 1917. However, due to the fragmentary nature of the

6879:"Dueling Dinosaurs | North Carolina Museum of Natural Sciences" 4956:

Koen Stein, paleontologist at the Free University of Brussels (

4468:

Gets Long-Term Lease at Smithsonian's Natural History Museum."

3582:

Discovering Dinosaurs in the American Museum of Natural History

2694:, unearthed the remains of what is believed to be a 4-year-old 6178:"Famous T. Rex had a bone infection, new medical scans reveal" 5775:"Meet 'Scotty,' the largest Tyrannosaurus rex ever discovered" 5049:"Parasite may have felled a mighty T. rex | Paleontology" 3097:, and the Dig Field school excavated the partial remains of a 1964: 1424:

in 1892. Cope believed that they belonged to an "agathaumid" (

37: 6883:

Dueling Dinosaurs | North Carolina Museum of Natural Sciences

4199:"T. Rex Skeleton Brings $ 31.8 Million at Christie's Auction" 4033:

Wolff, ED; Salisbury, SW; Horner, JR; Varricchio, DJ (2009).

3002:"Titus" is the name given to an obsidian-black skeleton of a 2663:

skull ever discovered. Following the sale of "Sue," another

29:"B-rex" redirects here. For the bird Shoebill (B. rex), see 6372:"Trix the T Rex makes her mark on Leiden in new exhibition" 3162:

Barbara is the nickname given to a specimen on loan to the

2019:

of Montana. This specimen was excavated by a team from the

1551:. Fragments of (then) AMNH 973 were first found in 1902 by 1416:

The first-named fossil specimen which can be attributed to

77:"Sue", AMNH 5027, "Stan", and "Jane", to scale with a human 6314:"Victoria: The T. rex | Houston Museum Of Natural Science" 5871:"Sask. T-rex Scotty is officially biggest ever discovered" 4614:"Call to split T. rex into 3 species sparks fierce debate" 2224:(also known as "Peck's rex", "Peckrex", "Rigby's rex" and 6504:"Real T. Rex descends on Nottingham in a world exclusive" 3122:

Peter is the nickname given to a specimen on loan to the

1775:

under the direction of Harley Garbani discovered another

5916:"Mighty T. rex Killed by Lowly Parasite, Study Suggests" 2390:; Bucky's furcula was the first one found for the genus 1092:, although the specimen belongs to a private collector. 4344: 4342: 4340: 3865:

Longrich, Nicholas R.; Saitta, Evan T. (1 March 2024).

2987:

skeleton incorporating authentic fossil elements from "

2454:, officially known as BMRP 2002.4.1, discovered in the 2331:

Bucky is a fossil of a juvenile specimen on display at

2047:

It was long on exhibit at the Museum of the Rockies in

2011:

In 1988, local rancher Kathy Wankel discovered another

54: 6340:"Wichita gets to keep its T. rex skeleton in Old Town" 6031:"Nach drei Jahren verlässt T. rex Tristan Otto Berlin" 2721:

Scotty is currently thought by many to be the largest

2590:

Femur of MOR 1125 from which demineralized matrix and

2386:

Bucky is one of the few dinosaur fossils found with a

6002: 6000: 5653: 5651: 5649: 5647: 4952: 4950: 4262:"Common Avian Infection Plagued the Tyrant Dinosaurs" 4098:"Yesterday.sg: SUE, STAN & Friends were in town…" 4035:"Common Avian Infection Plagued the Tyrant Dinosaurs" 3217:

is an adolescent that was preserved entangled with a

3205:. This specimen, considered the most complete of any 2667:

skeleton was, the specimen was put up for auction on

2379:, in a hunting scene. Both dinosaurs are attacking a 2051:. In June 2013, the Corps loaned the specimen to the 1834: 1448:

vertebrae, Osborn did not synonymize the two genera.

450: 7097:

Collection of the Carnegie Museum of Natural History

6130:"Dansk velhaver investerer i ægte Tyrannosaurus rex" 5660:"An Older and Exceptionally Large Adult Specimen of 1734:in 1946. In 1988, the specimen was re-described by 6766:Barbara joins Auckland Museums growing collection" 5320:The First Recognized Furcula For Tyrannosaurus Rex 5194: 5192: 5190: 5188: 4561: 3756:Bakker, R.T.; Williams, M.; Currie, P.J. (1988). " 3712:Bulletin of the American Museum of Natural History 3579: 3578:Norell, M. A.; Gaffney, E. S.; Dingus, L. (1995). 3520:Bulletin of the American Museum of Natural History 3485:Bulletin of the American Museum of Natural History 3447:Bulletin of the American Museum of Natural History 2144:currently known, in the Hell Creek Formation near 1933:in 1987 by Stan Sacrison, who also discovered the 1857: 1716:, was recovered from Montana in 1942. This skull, 463: 6410:"Don't worry, Sue – Trix is just passing through" 4825:. Field Museum of Natural History. 11 August 2016 4732:Memoirs of the Society of Vertebrate Paleontology 4700:(Second ed.). pp. 3, 52. Archived from 3908: 3906: 3030:, the other being the type of the junior synonym 2892:. The specimen has the most complete tail of any 1031:Clayton Phipps, Mark Eatman, & Chad O’Connor 4238:New Mexico Museum of Natural History and Science 3256:New Mexico Museum of Natural History and Science 3210:The fossils were officially displayed in 2024. 2828:This fossil is one of the geologically youngest 2690:On 7 July 2013, fossil hunter Robert Detrich of 1356:New Mexico Museum of Natural History and Science 7076:AMNH Article on the First Tyrannosaur specimens 6788: 6786: 5478:"Scientists question dinosaur soft tissue find" 5322:. Black Hills Institute of Geological Research. 4917: 4915: 4695:"Unearthing the Secrets of Sue: Educator Guide" 4637:"Insufficient Evidence for Multiple Species of 4595:"Tyrannosaurus rex may have been misunderstood" 4485:"T. rex arrives at National Museum of Scotland" 4163:"New Mexico museum has funds for half a T. rex" 3223:; their combined fossil is referred to as the " 2733:", cataloged as RSM P2523.8, was discovered in 2402:out of more than 40 that have been discovered. 2074:University of California Museum of Paleontology 6828:"Nanotyrannus: Up Hell Creek Without a Paddle" 6441:"A Paleontological Study of a New Specimen of 5715:"The Cretaceous « T.rex Discovery Centre" 5205:. Life of the past. Indiana University Press. 3342: 3340: 6123: 6121: 6119: 6077:"Schädel eines T. Rex in Berlin eingetroffen" 5115:"What's the Army doing with dinosaurs? Redux" 4406: 4404: 3483:and other Cretaceous carnivorous dinosaurs". 3118:Peter (L) & Barbara (R) side by side 2022 2240:who led the excavation and bone preparation. 2083:, but rather the holotype for a new species: 8: 6477:"Titus T. rex Scientific Report: Discussion" 6202:Natural History Museum of Los Angeles County 5973: 5971: 5969: 5967: 5077:. Dml.cmnh.org. 13 June 2005. Archived from 4449:: CS1 maint: multiple names: authors list ( 3474: 3472: 2918:In 2013, a team of paleontologists from the 2015:in Hell Creek sediments on an island in the 1945:Black Hills Institute of Geological Research 1773:Natural History Museum of Los Angeles County 1742:, and Michael Williams, then the curator of 1385:, dating to around 5–7 million years before 979:Natural History Museum of Los Angeles County 588:Confirmed as female, due to the presence of 383:Natural History Museum of Los Angeles County 7064:The Black Hills Institute's article on Stan 5172: 5170: 4508: 4506: 3685:. Cambridge University Press. p. 143. 3091:Burke Museum of Natural History and Culture 2816:From 2003 to 2005, Thomas was excavated by 2017:Charles M. Russell National Wildlife Refuge 1238:Burke Museum of Natural History and Culture 444:One of the westernmost specimens, found at 6009:"T-Rex Tristan: Entzauberung eines Mythos" 5568: 5566: 5295:. University of California. Archived from 3710:: restoration and model of the skeleton". 3409:"Anatomy and function of digit III of the 2934:specimen that lived 67 million years ago. 2832:specimens known, discovered very near the 2812:Thomas, mounted with two younger specimens 2177:development, since it is the third oldest 2040:, a biological form of iron that makes up 1858:{\displaystyle 114^{\circ }~{\textrm {W}}} 93: 7112:Collection of the Smithsonian Institution 5940: 5938: 5936: 5679: 5546: 5528: 5405: 4669: 4384: 4374: 4295: 4285: 4068: 4058: 3886: 3646: 3636: 3531: 3383: 3365: 3246:indicating the holotype material in white 3203:North Carolina Museum of Natural Sciences 1939:specimen nicknamed "Duffy". The original 1849: 1848: 1839: 1833: 1712:, frequently considered to be a juvenile 1022:North Carolina Museum of Natural Sciences 455: 449: 5589:"It's "Bone Voyage" for a famous fossil" 2463:preparation, Jane was put on display at 2394:Bucky also has a nearly complete set of 2228:"imperator") is the nickname given to a 1510:had appeared just one page earlier than 1220:collection under the number NCMG 2021-7 362:. Now considered a juvenile specimen of 7017:New Mexico Geological Society Guidebook 6434: 6432: 6430: 6246: 6244: 5981:. Museum für Naturkunde. Archived from 3323: 3270:this specimen as the holotype of a new 123: 6586:"Burke Museum team discovers a T. rex" 6557:"Mounting the skeleton of 'Titus' the 5914:Choi, Charles Q. (29 September 2009). 5709: 5707: 5579:. 12 May 2004. Retrieved 16 July 2006. 4874:. Scott Hartman's Skeletal Drawing.com 4872:"Mass estimates: North vs South redux" 4442: 4095:. 65% of his total skeleton was found 3916:New York: Simon & Schuster. 238pp. 3565:10.1038/scientificamerican10091915-322 2540:; some cervical and dorsal ribs; left 2161:in New York, where it was sold to the 6612:"Tyrannosaurus rex: A gigantic beast" 5864: 5862: 5860: 5799: 5797: 5795: 5742: 5740: 4116:"Stan (The University of Manchester)" 3739:Smithsonian Miscellaneous Collections 2625:hypothesized that the soft-tissue is 2493:, noting that a juvenile specimen of 2333:The Children's Museum of Indianapolis 1708:A small but nearly complete skull of 1631:. In total, Brown found five partial 1518:would have become the official name. 861:The Children's Museum of Indianapolis 422:Royal Tyrrell Museum of Palaeontology 7: 6637:Basagre, Bernadette (1 April 2022). 6007:Gennies, Sidney (17 December 2015). 5945:Cookson, Clive (25 September 2015). 5199:Larson, P.L.; Carpenter, K. (2008). 3912:Horner, J.R. & Lessem, D. 1993. 3034:which has the jaw on display in the 2199: 2070:Australian Fossil and Mineral Museum 1921:about 11.78 m (38 ft) long found in 6760:Anderson, Ryan (24 November 2022). 5869:Francis, Jennifer (22 March 2019). 5773:Ashley Strickland (26 March 2019). 4984:"The Story of a Dinosaur Named Sue" 4726:Brochu, C.R. (2003). "Osteology of 3954:. Dsc.discovery.com. Archived from 3065:to create digital models that were 1748:Cleveland Museum of Natural History 1718:Cleveland Museum of Natural History 1646:Scale model of the never-completed 1088:Inventory number attributed by the 330:Cleveland Museum of Natural History 6931:. 17 November 2020. Archived from 6037:. 26 November 2018. Archived from 5804:Holson, Laura M. (28 March 2019). 5235:Tyrannosaurus Rex, the Tyrant King 5202:Tyrannosaurus Rex, the Tyrant King 4560:Elbein, Asher (28 February 2022). 4431:Wu, Katherine, J. (17 July 2018). 4185:"T-Rex Replica Skeletons for Sale" 3984:. Unearthingtrex.com. 26 June 2002 3832:Journal of Vertebrate Paleontology 3785:Journal of Vertebrate Paleontology 3455:American Museum of Natural History 2053:National Museum of Natural History 1598:American Museum of Natural History 1572:Carnegie Museum of Natural History 1484:American Museum of Natural History 285:American Museum of Natural History 242:Carnegie Museum of Natural History 144:American Museum of Natural History 25: 6502:Pinchess, Lynette (1 July 2021). 6408:Johnson, Steve (24 August 2016). 5029:. Article-niche.com. 4 March 2009 4612:Greshko, Michael (1 March 2022). 4574:from the original on 1 March 2022 4197:Small, Zachary (7 October 2020). 3933:Tyrannosaurus rex the Tyrant King 3020:Nottingham Natural History Museum 2783:Natural History Museum of Denmark 2007:" mount in the Smithsonian Museum 1218:Nottingham Natural History Museum 1066:Natural History Museum of Denmark 6565:Natural-History-Conservation.com 6338:Reidl, Matt (15 February 2018). 6075:Karberg, Sascha (13 July 2015). 5947:"How to buy a Tyrannosaurus Rex" 5838:Johnson, Steve (29 March 2019). 3312:Timeline of tyrannosaur research 3294: 2469:Burpee Museum of Natural History 2437:Burpee Museum of Natural History 1917:Stan is the nickname given to a 1869:. The specimen is housed in the 1823:. A large bone was found in the 1799:"Black Beauty" (specimen number 1771:In 1966, a crew working for the 1562:, so the short fore-arms of the 939:Burpee Museum of Natural History 41: 6450:. Wollaton Hall. Archived from 6154:Ring, Caroline (25 July 2015). 5275:Dinosphere: A 3–5 Unit of Study 5179:Dinosphere: A 3–5 Unit of Study 4923:"T. rex is coming to Naturalis" 4802:Field Museum of Natural History 4223:. 23 March 2022. Archived from 2610:North Carolina State University 2412:paleontologist of the same name 2163:Field Museum of Natural History 2127:Field Museum of Natural History 817:Pioneer Trails Regional Museum 732:Field Museum of Natural History 358:and later given its own genus, 6832:A Central Coast Paleontologist 6128:Okke, Tom (17 December 2015). 5979:"Tristan – Berlin bares teeth" 4870:Hartman, Scott (7 July 2013). 3805:10.1080/02724634.1999.10011161 3036:Natural History Museum, London 2140:discovered the best-preserved 1090:Natural History Museum, Berlin 191:Natural History Museum, London 1: 6528:Ingram, Simon (12 May 2021). 5407:10.7287/peerj.preprints.852v1 2920:Naturalis Biodiversity Center 1161:Naturalis Biodiversity Center 1151:Naturalis Biodiversity Center 6806:Auckland War Memorial Museum 6672:Auckland War Memorial Museum 5476:Fox, Maggie (30 July 2008). 4593:Hunt, Katie (1 March 2022). 4376:10.1371/journal.pone.0026037 4287:10.1371/journal.pone.0007288 4146:"Stan the Tyrannosaurus rex" 4060:10.1371/journal.pone.0007288 3844:10.1080/02724634.2011.557116 3638:10.1371/journal.pone.0026037 3278:. This species differs from 3164:Auckland War Memorial Museum 3124:Auckland War Memorial Museum 3049:Craig Pfister near Ekalaka, 2834:Cretaceous–Tertiary boundary 2527:Reconstructed skull of B-rex 2410:E. D. Cope (named after the 2383:specimen known as "Kelsey". 2327:Cast of the "Bucky" specimen 2044:(the red pigment in blood). 2029:U.S. Army Corps of Engineers 1803:81.6.1) is a well-preserved 1611:(neck vertebrae) series for 1316:Auckland War Memorial Museum 1277:Auckland War Memorial Museum 464:{\displaystyle 114^{\circ }} 7107:Lists of dinosaur specimens 6065:, accessed 13 February 2020 4823:"How well do you know SUE?" 4411:Schontzler, Gail. "Montana 3586:. New York: Knopf. p. 3187:"Bloody Mary" (as named by 2435:fossil "Jane" displayed at 2343:. It is the first juvenile 2066:National Museum of Scotland 2027:, with assistance from the 1959:Like many other fossils of 1787:"Black Beauty": RTMP 81.6.1 1783:skull on exhibit anywhere. 1662:, described at the time as 1428:) dinosaur, and named them 7133: 6826:accpaleo (11 March 2018). 4958:Vrije Universiteit Brussel 4671:10.1007/s11692-022-09573-1 4539:10.1007/s11692-022-09561-5 4319:Institution, Smithsonian. 4010:. Uleth.ca. Archived from 3367:10.1038/s41598-023-47011-0 3180: 3150:"Barbara": AWMM-IL 2022.21 2972: 2946:after former Netherlands' 2903: 2710: 2243:The skeleton of Montana's 2189:that is older than Sue is 2114: 1954:Saadiyat Cultural District 1899: 1755:it a juvenile specimen of 1596:specimen AMNH 5027 at the 1545:specimen used to describe 210:Seven Mile Creek, Wyoming 28: 7060:at The Theropod Database. 6736:"Peter Scientific Report" 5480:. Reuters. Archived from 5003:"Sue at the Field Museum" 4415:Heading to Smithsonian." 2886:Museum of World Treasures 2798:tumefactive osteomyelitis 2725:specimen ever discovered. 2298:The fossils of Montana's 1637:Jurassic Park film series 1486:), was discovered in the 774:Royal Saskatchewan Museum 309:Jurassic Park film series 7117:Cretaceous fossil record 5888:Article about Scotty on 4989:27 December 2014 at the 4417:Bozeman Daily Chronicle. 3232:Tyrannosaurus mcraeensis 3133:He was likely killed by 3095:University of Washington 3074:"Tufts-Love": UWBM 99000 3008:discovered in Montana's 2914:Mounted skeleton of Trix 1409:specimen (AMNH 3982) of 1346:Tyrannosaurus mcraeensis 7069:12 October 2007 at the 6224:"Collection Highlights" 6156:"Der T. rex von Berlin" 5577:Pittsburgh Post-Gazette 5447:10.1126/science.1108397 5253:www.childrensmuseum.org 4971:22 October 2019 at the 4798:Sue at the Field Museum 4324:Smithsonian Institution 3242:Reconstructed skull of 3177:"Bloody Mary": BHI 6437 3110:"Peter": AWMM-IL 2022.9 2942:The specimen was named 2880:Ivan is a 65% complete 2848:Victoria on display at 2677:figure of the same name 2057:Smithsonian Institution 1821:Crowsnest Pass, Alberta 1530:Reconstructed mount of 1496:Dynamosaurus imperiosus 1468:Dynamosaurus imperiosus 440:Crowsnest Pass, Alberta 217:Dynamosaurus imperiosus 18:Black Beauty (dinosaur) 6762:"Skeleton of pregnant 6062:Back in 2021 - Tristan 4893:kcook (30 June 2014). 4474:Accessed 28 June 2013. 4421:Accessed 28 June 2013. 3980:www.unearthingtrex.com 3888:10.3390/fossils2010001 3869:Nanotyrannus lancensis 3706:Osborn, H.F. (1913). " 3677:(1990). "Variation in 3497:10.5281/zenodo.1038151 3479:Osborn, H.F. (1905). " 3247: 3213:The Dueling Dinosaurs 3159: 3119: 3086: 3051:Carter County, Montana 3032:Dynamosaurus imperosus 2999: 2915: 2877: 2852: 2813: 2773: 2756:the result of another 2726: 2651: 2638:MOR 1125 as a female. 2595: 2594:(insets) were obtained 2528: 2499:described by Tsuihiji 2440: 2406:"E. D. Cope": BHI 6248 2328: 2319:"Bucky": TCM 2001.90.1 2217: 2130: 2023:led by paleontologist 2008: 1914: 1867:Willow Creek Formation 1859: 1796: 1710:Nanotyrannus lancensis 1705: 1651: 1600: 1535: 1504:Henry Fairfield Osborn 1471: 1413: 1084:Carter County, Montana 465: 435:Willow Creek Formation 78: 6714:10.2110/palo.2017.076 6103:Museum für Naturkunde 5668:The Anatomical Record 5289:"The Origin of Birds" 4765:Holtz, T. R. (2011). 3867:"Taxonomic Status of 3453:(43). New York City: 3241: 3157: 3117: 3081: 2982: 2936:Black Hills institute 2913: 2876:, during installation 2871: 2847: 2811: 2779:Museum für Naturkunde 2771: 2720: 2707:"Scotty": RSM P2523.8 2649: 2606:Mary Higby Schweitzer 2589: 2526: 2507:specimen, nicknamed " 2439:at Rockford, Illinois 2430: 2423:"Jane": BMRP 2002.4.1 2369:Black Hills Institute 2326: 2211: 2138:Black Hills Institute 2124: 2021:Museum of the Rockies 2002: 1995:"Wankel Rex": MOR 555 1909: 1860: 1794: 1700: 1645: 1590: 1529: 1462: 1405: 1165:Black Hills Institute 1116:Robert (Bob) Detrich 900:Black Hills Institute 689:Museum of the Rockies 650:Museum of the Rockies 610:Museum of the Rockies 568:Museum of the Rockies 530:Museum of the Rockies 508:Buffalo, South Dakota 466: 366:by most researchers. 356:Gorgosaurus lancensis 76: 7083:at Skeletal Drawing. 6975:10.58799/NMG-v6n4.72 6284:. 12 September 2019. 5008:4 April 2014 at the 4895:"SUE Educator Guide" 4650:Evolutionary Biology 4519:Evolutionary Biology 4491:on 20 September 2011 4464:du Lac, J. Freedom " 4437:Smithsonian Magazine 4168:16 July 2011 at the 4122:on 18 September 2010 4093:"Sue" (FMNH PR 2081) 3683:Dinosaur Systematics 3419:: 77. Archived from 3026:to be on exhibit in 3010:Hell Creek Formation 2735:Saskatchewan, Canada 2456:Hell Creek Formation 2349:Hell Creek Formation 2085:Tyrannosaurus regina 1923:Hell Creek Formation 1880:In 2009, a paper by 1871:Royal Tyrrell Museum 1832: 1329:Hell Creek Formation 1253:Hell Creek Formation 1209:Hell Creek Formation 1170:Hell Creek Formation 1120:Hell Creek Formation 1079:Hell Creek Formation 1035:Hell Creek Formation 994:Hell Creek Formation 954:Hell Creek Formation 913:Hell Creek Formation 874:Hell Creek Formation 829:Hell Creek Formation 747:Hell Creek Formation 702:Hell Creek Formation 663:Hell Creek Formation 623:Hell Creek Formation 581:Hell Creek Formation 543:Hell Creek Formation 503:Hell Creek Formation 448: 396:Hell Creek Formation 345:Hell Creek Formation 300:Hell Creek Formation 268:Originally AMNH 973 261:Hell Creek Formation 213:Originally AMNH 5866 159:Hell Creek Formation 55:adding missing items 7029:10.56577/FFC-37.235 6935:on 17 November 2020 6706:2018Palai..33..164M 6535:National Geographic 6508:NottinghamshireLive 6035:Berliner Morgenpost 5521:2016NatSR...623099S 5439:2005Sci...307.1952S 5433:(5717): 1952–1955. 5081:on 13 November 2007 4935:on 15 December 2016 4662:2022EvBio..49..327C 4618:National Geographic 4531:2022EvBio..49..156P 4367:2011PLoSO...626037H 4278:2009PLoSO...4.7288W 4247:5 June 2010 at the 4221:National Geographic 4172:, 26 February 2006. 4051:2009PLoSO...4.7288W 4014:on 23 February 2012 3950:Discovery Channel's 3914:The Complete T. rex 3797:1999JVPal..19..497C 3779:Carr, T.D. (1999). 3629:2011PLoSO...626037H 3553:Scientific American 3547:Brown, B. (1915). " 3423:on 24 February 2008 3252:Hall Lake Formation 3018:. Exhibited in the 2924:Leiden, Netherlands 2900:"Trix": RGM 792.000 2353:Faith, South Dakota 2173:due to a prolonged 2146:Faith, South Dakota 1977:. Also, two of the 1967:and damages in the 1877:, Alberta, Canada. 1430:Manospondylus gigas 1422:Edward Drinker Cope 1411:Manospondylus gigas 1372:Hall Lake Formation 918:Faith, South Dakota 879:Faith, South Dakota 791:Frenchman Formation 752:Faith, South Dakota 170:Manospondylus gigas 164:Faith, South Dakota 6963:New Mexico Geology 6378:. 6 September 2019 6302:. 16 October 2020. 6204:. 15 December 2010 6079:. Der Tagesspiegel 6011:. Der Tagesspiegel 5810:The New York Times 5719:royalsaskmuseum.ca 5509:Scientific Reports 4811:on 18 August 2016. 4693:The Field Museum. 4568:The New York Times 4203:The New York Times 3958:on 10 October 2012 3828:Tarbosaurus bataar 3354:Scientific Reports 3248: 3160: 3120: 3087: 3000: 2916: 2878: 2853: 2814: 2774: 2727: 2652: 2598:In the March 2005 2596: 2576:; and a number of 2529: 2465:Rockford, Illinois 2447:specimen of small 2441: 2329: 2238:J. Keith Rigby Jr. 2232:specimen found in 2218: 2131: 2111:"Sue": FMNH PR2081 2009: 1979:cervical vertebrae 1943:are now housed at 1915: 1910:Cast of "Stan" at 1855: 1797: 1732:Charles W. Gilmore 1706: 1682:arms were found). 1652: 1601: 1536: 1472: 1414: 909:Bucky Derfilinger 461: 79: 53:; you can help by 7056:Tyrannosaurus rex 7008:Tyrannosaurus rex 6852:Keats, Jonathon. 6797:Tyrannosaurus Rex 6559:Tyrannosaurus rex 6483:. 2 February 2022 6443:Tyrannosaurus rex 6360:. September 2016. 6344:The Wichita Eagle 6258:. 16 October 2020 6228:dinosaurs.nhm.org 5662:Tyrannosaurus rex 5530:10.1038/srep23099 5423:Tyrannosaurus rex 5365:5 August 2012 at 5293:The Macro Library 5212:978-0-253-35087-9 5025:article-niche.com 4966:, 23 January 2017 4851:. 5 February 2018 4728:Tyrannosaurus rex 4707:on 3 October 2018 4466:Tyrannosaurus rex 4227:on 23 March 2022. 4142:Manchester Museum 4104:on 1 October 2011 3935:. pp. 16–17. 3679:Tyrannosaurus rex 3671:Kenneth Carpenter 3411:Tyrannosaurus rex 3332:Tyrannosaurus rex 3225:Dueling Dinosaurs 3183:Dueling Dinosaurs 3140:Tyrannosaurus rex 3099:Tyrannosaurus rex 3040:Tyrannosaurus rex 3005:Tyrannosaurus rex 2932:Tyrannosaurus rex 2884:displayed at the 2823:Tyrannosaurus rex 2772:Tristan in Berlin 2713:Scotty (dinosaur) 2696:Tyrannosaurus rex 2665:Tyrannosaurus rex 2661:Tyrannosaurus rex 2519:"B-rex": MOR 1125 2400:Tyrannosaurus rex 2351:near the town of 2150:Tyrannosaurus rex 2134:Susan Hendrickson 2013:Tyrannosaurus rex 1991:-like parasites. 1961:Tyrannosaurus rex 1912:Manchester Museum 1852: 1847: 1813:Tyrannosaurus rex 1809:Tyrannosaurus rex 1648:Tyrannosaurus rex 1629:Tyrannosaurus rex 1548:Tyrannosaurus rex 1522:Holotype: CM 9380 1508:Tyrannosaurus rex 1480:Tyrannosaurus rex 1442:Tyrannosaurus rex 1418:Tyrannosaurus rex 1392: 1391: 950:William Harrison 870:Bucky Derflinger 364:Tyrannosaurus rex 71: 70: 16:(Redirected from 7124: 7041: 7040: 7014: 7001: 6995: 6994: 6960: 6951: 6945: 6944: 6942: 6940: 6921: 6915: 6914: 6912: 6910: 6900: 6894: 6893: 6891: 6889: 6875: 6869: 6868: 6866: 6864: 6849: 6843: 6842: 6840: 6838: 6823: 6817: 6816: 6814: 6812: 6803: 6790: 6781: 6780: 6778: 6776: 6757: 6751: 6750: 6748: 6746: 6740: 6732: 6726: 6725: 6689: 6683: 6682: 6680: 6678: 6669: 6660: 6654: 6653: 6651: 6649: 6634: 6628: 6627: 6625: 6623: 6608: 6602: 6601: 6599: 6597: 6592:. 17 August 2016 6582: 6576: 6575: 6573: 6571: 6553: 6547: 6546: 6544: 6542: 6525: 6519: 6518: 6516: 6514: 6499: 6493: 6492: 6490: 6488: 6473: 6467: 6466: 6464: 6462: 6456: 6449: 6436: 6425: 6424: 6422: 6420: 6405: 6399: 6394: 6388: 6387: 6385: 6383: 6368: 6362: 6361: 6354: 6348: 6347: 6335: 6329: 6328: 6326: 6324: 6310: 6304: 6303: 6292: 6286: 6285: 6274: 6268: 6267: 6265: 6263: 6248: 6239: 6238: 6236: 6234: 6220: 6214: 6213: 6211: 6209: 6194: 6188: 6187: 6186:. December 2021. 6174: 6168: 6167: 6165: 6163: 6151: 6145: 6144: 6142: 6140: 6125: 6114: 6113: 6111: 6109: 6095: 6089: 6088: 6086: 6084: 6072: 6066: 6057: 6051: 6050: 6048: 6046: 6027: 6021: 6020: 6018: 6016: 6004: 5995: 5994: 5992: 5990: 5975: 5962: 5961: 5959: 5957: 5942: 5931: 5930: 5928: 5926: 5911: 5905: 5900: 5894: 5885: 5879: 5878: 5866: 5855: 5854: 5852: 5850: 5835: 5829: 5828: 5826: 5824: 5801: 5790: 5789: 5787: 5785: 5770: 5764: 5763: 5761: 5759: 5752:The Vintage News 5744: 5735: 5734: 5732: 5730: 5725:on 11 April 2019 5721:. Archived from 5711: 5702: 5701: 5683: 5681:10.1002/ar.24118 5655: 5642: 5641: 5639: 5637: 5623: 5617: 5616: 5614: 5612: 5598: 5592: 5586: 5580: 5575:by Byron Spice. 5570: 5561: 5560: 5550: 5532: 5500: 5494: 5493: 5491: 5489: 5473: 5467: 5466: 5418: 5412: 5411: 5409: 5385: 5379: 5375: 5369: 5356: 5350: 5349: 5347: 5345: 5340:on 5 August 2012 5330: 5324: 5323: 5315: 5309: 5308: 5306: 5304: 5285: 5279: 5278: 5270: 5264: 5263: 5261: 5259: 5245: 5239: 5238: 5230: 5224: 5223: 5221: 5219: 5196: 5183: 5182: 5174: 5165: 5164: 5163:. 29 March 2019. 5151: 5145: 5144: 5137: 5131: 5130: 5128: 5126: 5111: 5105: 5104: 5097: 5091: 5090: 5088: 5086: 5067: 5061: 5060: 5058: 5056: 5045: 5039: 5038: 5036: 5034: 5019: 5013: 5000: 4994: 4981: 4975: 4954: 4945: 4944: 4942: 4940: 4934: 4928:. Archived from 4927: 4919: 4910: 4909: 4907: 4905: 4890: 4884: 4883: 4881: 4879: 4867: 4861: 4860: 4858: 4856: 4845:"Sue the T. Rex" 4841: 4835: 4834: 4832: 4830: 4819: 4813: 4812: 4810: 4804:. Archived from 4795: 4791:"Sue Fact Sheet" 4787: 4781: 4780: 4778: 4776: 4771: 4762: 4756: 4755: 4723: 4717: 4716: 4714: 4712: 4706: 4699: 4690: 4684: 4683: 4673: 4632: 4626: 4625: 4624:on 1 March 2022. 4620:. Archived from 4609: 4603: 4602: 4590: 4584: 4583: 4581: 4579: 4565: 4557: 4551: 4550: 4510: 4501: 4500: 4498: 4496: 4487:. Archived from 4481: 4475: 4470:Washington Post. 4461: 4455: 4454: 4448: 4440: 4428: 4422: 4408: 4399: 4398: 4388: 4378: 4346: 4335: 4334: 4332: 4330: 4316: 4310: 4309: 4299: 4289: 4257: 4251: 4242:"T. REX ATTACK!" 4235: 4229: 4228: 4213: 4207: 4206: 4194: 4188: 4179: 4173: 4154: 4148: 4138: 4132: 4131: 4129: 4127: 4118:. Archived from 4113: 4111: 4109: 4100:. Archived from 4089: 4083: 4082: 4072: 4062: 4030: 4024: 4023: 4021: 4019: 4000: 3994: 3993: 3991: 3989: 3974: 3968: 3967: 3965: 3963: 3943: 3937: 3936: 3928: 3917: 3910: 3901: 3900: 3890: 3862: 3856: 3855: 3823: 3817: 3816: 3776: 3770: 3769: 3753: 3747: 3746: 3734: 3728: 3727: 3703: 3697: 3696: 3675:Philip J. Currie 3667: 3661: 3660: 3650: 3640: 3608: 3602: 3601: 3585: 3575: 3569: 3568: 3544: 3538: 3537: 3535: 3515: 3509: 3508: 3476: 3467: 3466: 3439: 3433: 3432: 3430: 3428: 3404: 3398: 3397: 3387: 3369: 3344: 3335: 3328: 3304: 3302:Dinosaurs portal 3299: 3298: 3297: 3015:Titus Andronicus 2991:", exhibited at 2975:Titus (dinosaur) 2601:Science magazine 2289:Montana's T. rex 2125:"Sue" specimen, 2101:Jingmai O'Connor 2089:Stephen Brusatte 2049:Bozeman, Montana 1896:"Stan": BHI 3033 1864: 1862: 1861: 1856: 1854: 1853: 1850: 1845: 1844: 1843: 1736:Robert T. Bakker 1534:holotype CM 9380 1381:The holotype of 600:"C-rex/Celeste" 470: 468: 467: 462: 460: 459: 97:Specimen number 94: 66: 63: 45: 44: 38: 21: 7132: 7131: 7127: 7126: 7125: 7123: 7122: 7121: 7087: 7086: 7071:Wayback Machine 7050: 7045: 7044: 7012: 7003: 7002: 6998: 6958: 6953: 6952: 6948: 6938: 6936: 6923: 6922: 6918: 6908: 6906: 6902: 6901: 6897: 6887: 6885: 6877: 6876: 6872: 6862: 6860: 6851: 6850: 6846: 6836: 6834: 6825: 6824: 6820: 6810: 6808: 6801: 6792: 6791: 6784: 6774: 6772: 6759: 6758: 6754: 6744: 6742: 6738: 6734: 6733: 6729: 6691: 6690: 6686: 6676: 6674: 6667: 6662: 6661: 6657: 6647: 6645: 6636: 6635: 6631: 6621: 6619: 6610: 6609: 6605: 6595: 6593: 6584: 6583: 6579: 6569: 6567: 6555: 6554: 6550: 6540: 6538: 6527: 6526: 6522: 6512: 6510: 6501: 6500: 6496: 6486: 6484: 6475: 6474: 6470: 6460: 6458: 6457:on 8 March 2022 6454: 6447: 6438: 6437: 6428: 6418: 6416: 6414:Chicago Tribune 6407: 6406: 6402: 6395: 6391: 6381: 6379: 6370: 6369: 6365: 6356: 6355: 6351: 6337: 6336: 6332: 6322: 6320: 6312: 6311: 6307: 6294: 6293: 6289: 6276: 6275: 6271: 6261: 6259: 6250: 6249: 6242: 6232: 6230: 6222: 6221: 6217: 6207: 6205: 6198:"Dinosaur Hall" 6196: 6195: 6191: 6176: 6175: 6171: 6161: 6159: 6153: 6152: 6148: 6138: 6136: 6127: 6126: 6117: 6107: 6105: 6097: 6096: 6092: 6082: 6080: 6074: 6073: 6069: 6058: 6054: 6044: 6042: 6041:on 3 April 2019 6029: 6028: 6024: 6014: 6012: 6006: 6005: 5998: 5988: 5986: 5985:on 1 April 2019 5977: 5976: 5965: 5955: 5953: 5951:Financial Times 5944: 5943: 5934: 5924: 5922: 5913: 5912: 5908: 5901: 5897: 5886: 5882: 5868: 5867: 5858: 5848: 5846: 5844:Chicago Tribune 5837: 5836: 5832: 5822: 5820: 5803: 5802: 5793: 5783: 5781: 5772: 5771: 5767: 5757: 5755: 5754:. 25 March 2019 5746: 5745: 5738: 5728: 5726: 5713: 5712: 5705: 5657: 5656: 5645: 5635: 5633: 5625: 5624: 5620: 5610: 5608: 5606:www.bonhams.com 5600: 5599: 5595: 5587: 5583: 5571: 5564: 5502: 5501: 5497: 5487: 5485: 5484:on 22 July 2012 5475: 5474: 5470: 5420: 5419: 5415: 5394:PeerJ Preprints 5388:Yun, C (2015). 5387: 5386: 5382: 5376: 5372: 5357: 5353: 5343: 5341: 5332: 5331: 5327: 5318:Larson, Peter. 5317: 5316: 5312: 5302: 5300: 5299:on 15 June 2012 5287: 5286: 5282: 5272: 5271: 5267: 5257: 5255: 5247: 5246: 5242: 5232: 5231: 5227: 5217: 5215: 5213: 5198: 5197: 5186: 5176: 5175: 5168: 5160:Chicago Tribune 5153: 5152: 5148: 5139: 5138: 5134: 5124: 5122: 5121:. 13 April 2015 5113: 5112: 5108: 5099: 5098: 5094: 5084: 5082: 5069: 5068: 5064: 5054: 5052: 5047: 5046: 5042: 5032: 5030: 5021: 5020: 5016: 5010:Wayback Machine 5001: 4997: 4991:Wayback Machine 4982: 4978: 4973:Wayback Machine 4955: 4948: 4938: 4936: 4932: 4925: 4921: 4920: 4913: 4903: 4901: 4892: 4891: 4887: 4877: 4875: 4869: 4868: 4864: 4854: 4852: 4843: 4842: 4838: 4828: 4826: 4821: 4820: 4816: 4808: 4793: 4789: 4788: 4784: 4774: 4772: 4769: 4764: 4763: 4759: 4744:10.2307/3889334 4725: 4724: 4720: 4710: 4708: 4704: 4697: 4692: 4691: 4687: 4634: 4633: 4629: 4611: 4610: 4606: 4592: 4591: 4587: 4577: 4575: 4559: 4558: 4554: 4512: 4511: 4504: 4494: 4492: 4483: 4482: 4478: 4462: 4458: 4441: 4430: 4429: 4425: 4409: 4402: 4348: 4347: 4338: 4328: 4326: 4318: 4317: 4313: 4259: 4258: 4254: 4249:Wayback Machine 4236: 4232: 4215: 4214: 4210: 4196: 4195: 4191: 4187:, 21 July 2008. 4180: 4176: 4170:Wayback Machine 4155: 4151: 4139: 4135: 4125: 4123: 4114: 4107: 4105: 4096: 4090: 4086: 4032: 4031: 4027: 4017: 4015: 4002: 4001: 3997: 3987: 3985: 3976: 3975: 3971: 3961: 3959: 3945: 3944: 3940: 3930: 3929: 3920: 3911: 3904: 3864: 3863: 3859: 3825: 3824: 3820: 3778: 3777: 3773: 3755: 3754: 3750: 3736: 3735: 3731: 3705: 3704: 3700: 3693: 3669: 3668: 3664: 3610: 3609: 3605: 3598: 3577: 3576: 3572: 3559:(15): 322–323. 3546: 3545: 3541: 3517: 3516: 3512: 3478: 3477: 3470: 3441: 3440: 3436: 3426: 3424: 3406: 3405: 3401: 3346: 3345: 3338: 3329: 3325: 3320: 3308: 3307: 3300: 3295: 3293: 3288: 3236: 3185: 3179: 3152: 3112: 3076: 2977: 2971: 2908: 2906:Trix (dinosaur) 2902: 2890:Wichita, Kansas 2866: 2842: 2806: 2766: 2715: 2709: 2692:Wichita, Kansas 2688: 2644: 2521: 2425: 2408: 2321: 2283:III. Montana's 2275:and right hand 2206: 2119: 2113: 1997: 1904: 1902:Stan (dinosaur) 1898: 1835: 1830: 1829: 1789: 1769: 1695: 1585: 1524: 1457: 1400: 1303:AWMM-IL2022.21 1290:Lance Formation 1264:AWMM-IL 2022.9 1248: 1125:Jordan, Montana 989:Luis M. Chiappe 949: 786: 785:Robert Gebhardt 784: 742:Sue Hendrickson 619:Celeste Horner 451: 446: 445: 412:"Black Beauty" 392:Harley Garbani 254: 214: 206:Lance Formation 92: 67: 61: 58: 42: 34: 23: 22: 15: 12: 11: 5: 7130: 7128: 7120: 7119: 7114: 7109: 7104: 7099: 7089: 7088: 7085: 7084: 7078: 7073: 7061: 7049: 7048:External links 7046: 7043: 7042: 6996: 6946: 6916: 6895: 6870: 6844: 6818: 6782: 6752: 6727: 6700:(4): 164–173. 6684: 6655: 6629: 6618:. 28 June 2018 6603: 6577: 6548: 6520: 6494: 6468: 6426: 6400: 6389: 6363: 6349: 6330: 6305: 6287: 6269: 6240: 6215: 6189: 6169: 6146: 6115: 6099:"tristan-otto" 6090: 6067: 6052: 6022: 5996: 5963: 5932: 5906: 5895: 5880: 5856: 5830: 5791: 5765: 5736: 5703: 5674:(4): 656–672. 5643: 5618: 5593: 5581: 5562: 5495: 5468: 5413: 5380: 5370: 5358:News Release: 5351: 5325: 5310: 5280: 5265: 5240: 5225: 5211: 5184: 5166: 5146: 5132: 5106: 5092: 5062: 5051:. Science News 5040: 5014: 4995: 4976: 4946: 4911: 4885: 4862: 4836: 4814: 4782: 4757: 4718: 4685: 4656:(3): 327–341. 4627: 4604: 4585: 4552: 4525:(2): 156–179. 4502: 4476: 4456: 4423: 4400: 4361:(10): e26037. 4336: 4311: 4252: 4230: 4208: 4189: 4174: 4149: 4133: 4084: 4025: 3995: 3969: 3946:Edgar, Blake. 3938: 3918: 3902: 3875:Fossil Studies 3857: 3838:(3): 497–517. 3818: 3791:(3): 497–520. 3771: 3748: 3729: 3698: 3691: 3662: 3623:(10): e26037. 3603: 3596: 3570: 3539: 3510: 3491:(3): 259–265. 3468: 3434: 3399: 3336: 3322: 3321: 3319: 3316: 3315: 3314: 3306: 3305: 3290: 3289: 3287: 3284: 3235: 3234:: NMMNH P-3698 3229: 3181:Main article: 3178: 3175: 3151: 3148: 3135:exocannibalism 3111: 3108: 3075: 3072: 3063:photogrammetry 3047:paleontologist 3028:United Kingdom 2973:Main article: 2970: 2967: 2952:T. rex in Town 2926:) traveled to 2904:Main article: 2901: 2898: 2865: 2862: 2841: 2838: 2805: 2802: 2765: 2762: 2711:Main article: 2708: 2705: 2687: 2684: 2643: 2640: 2627:permineralized 2533:Fort Peck Lake 2520: 2517: 2489:is a juvenile 2424: 2421: 2407: 2404: 2392:Tyrannosaurus. 2320: 2317: 2255:with complete 2205: 2198: 2117:Sue (dinosaur) 2115:Main article: 2112: 2109: 2099:, David Hone, 2003:"The nation's 1996: 1993: 1900:Main article: 1897: 1894: 1842: 1838: 1795:"Black Beauty" 1788: 1785: 1768: 1765: 1694: 1688: 1584: 1581: 1523: 1520: 1456: 1450: 1399: 1393: 1390: 1389: 1379: 1374: 1369: 1365:Donald Staton 1363: 1358: 1353: 1350: 1348: 1343: 1339: 1338: 1336: 1331: 1326: 1323: 1318: 1313: 1310: 1307: 1304: 1300: 1299: 1297: 1292: 1287: 1284: 1279: 1274: 1271: 1268: 1265: 1261: 1260: 1258: 1255: 1250: 1245: 1240: 1235: 1232: 1229: 1226: 1222: 1221: 1214: 1211: 1206: 1205:Craig Pfister 1203: 1198: 1196: 1193: 1190: 1183: 1179: 1178: 1175: 1172: 1167: 1158: 1153: 1148: 1145: 1142: 1135: 1131: 1130: 1127: 1122: 1117: 1114: 1109: 1107: 1104: 1101: 1098: 1094: 1093: 1086: 1081: 1076: 1075:Craig Pfister 1073: 1068: 1063: 1060: 1057: 1054: 1050: 1049: 1040: 1037: 1032: 1029: 1024: 1019: 1016: 1013: 1006: 1002: 1001: 999: 996: 991: 986: 981: 976: 973: 970: 967: 963: 962: 959: 956: 951: 946: 941: 936: 933: 930: 927: 926:BMRP 2002.4.1 923: 922: 920: 915: 910: 907: 902: 897: 894: 891: 888: 884: 883: 881: 876: 871: 868: 863: 858: 855: 852: 849: 848:TCM 2001.90.1 845: 844: 840: 831: 826: 823: 818: 815: 812: 809: 807: 803: 802: 798: 793: 788: 781: 776: 771: 768: 765: 762: 758: 757: 754: 749: 744: 739: 734: 729: 726: 723: 716: 712: 711: 709: 704: 699: 698:Frank Stewart 696: 691: 686: 683: 680: 679:"F-rex/Frank" 677: 673: 672: 670: 665: 660: 657: 652: 647: 644: 641: 638: 634: 633: 630: 625: 620: 617: 612: 607: 604: 601: 598: 594: 593: 590:medullary bone 586: 583: 578: 575: 570: 565: 562: 559: 556: 552: 551: 548: 545: 540: 537: 532: 527: 524: 521: 518: 514: 513: 510: 505: 500: 499:Stan Sacrison 497: 492: 490: 487: 484: 477: 473: 472: 458: 454: 442: 437: 432: 429: 424: 419: 416: 413: 410: 409:RTMP 1981.6.1 406: 405: 403: 398: 393: 390: 385: 380: 377: 374: 372: 368: 367: 352: 347: 342: 337: 332: 327: 324: 322: 317: 313: 312: 305: 302: 297: 292: 287: 282: 279: 276: 274: 270: 269: 266: 263: 258: 249: 244: 239: 236: 233: 228: 224: 223: 211: 208: 203: 198: 193: 188: 185: 182: 177: 173: 172: 166: 161: 156: 151: 146: 141: 138: 135: 130: 126: 125: 122: 119: 116: 113: 110: 107: 104: 101: 98: 91: 88: 69: 68: 48: 46: 24: 14: 13: 10: 9: 6: 4: 3: 2: 7129: 7118: 7115: 7113: 7110: 7108: 7105: 7103: 7102:Tyrannosaurus 7100: 7098: 7095: 7094: 7092: 7082: 7079: 7077: 7074: 7072: 7068: 7065: 7062: 7059: 7057: 7052: 7051: 7047: 7038: 7034: 7030: 7026: 7022: 7018: 7011: 7009: 7000: 6997: 6992: 6988: 6984: 6980: 6976: 6972: 6968: 6964: 6957: 6950: 6947: 6934: 6930: 6926: 6920: 6917: 6905: 6899: 6896: 6884: 6880: 6874: 6871: 6859: 6855: 6848: 6845: 6833: 6829: 6822: 6819: 6807: 6800: 6798: 6789: 6787: 6783: 6771: 6767: 6765: 6756: 6753: 6737: 6731: 6728: 6723: 6719: 6715: 6711: 6707: 6703: 6699: 6695: 6688: 6685: 6673: 6666: 6659: 6656: 6644: 6640: 6633: 6630: 6617: 6613: 6607: 6604: 6591: 6587: 6581: 6578: 6566: 6562: 6560: 6552: 6549: 6537: 6536: 6531: 6524: 6521: 6509: 6505: 6498: 6495: 6482: 6481:Wollaton Hall 6478: 6472: 6469: 6453: 6446: 6444: 6435: 6433: 6431: 6427: 6415: 6411: 6404: 6401: 6398: 6393: 6390: 6377: 6373: 6367: 6364: 6359: 6353: 6350: 6345: 6341: 6334: 6331: 6319: 6315: 6309: 6306: 6301: 6297: 6291: 6288: 6283: 6279: 6273: 6270: 6257: 6253: 6247: 6245: 6241: 6229: 6225: 6219: 6216: 6203: 6199: 6193: 6190: 6185: 6184: 6179: 6173: 6170: 6158:. Spektrum.de 6157: 6150: 6147: 6135: 6131: 6124: 6122: 6120: 6116: 6104: 6100: 6094: 6091: 6078: 6071: 6068: 6064: 6063: 6059:Back in 2021 6056: 6053: 6040: 6036: 6032: 6026: 6023: 6010: 6003: 6001: 5997: 5984: 5980: 5974: 5972: 5970: 5968: 5964: 5952: 5948: 5941: 5939: 5937: 5933: 5921: 5917: 5910: 5907: 5904: 5899: 5896: 5893: 5891: 5884: 5881: 5876: 5872: 5865: 5863: 5861: 5857: 5845: 5841: 5834: 5831: 5819: 5815: 5811: 5807: 5800: 5798: 5796: 5792: 5780: 5776: 5769: 5766: 5753: 5749: 5743: 5741: 5737: 5724: 5720: 5716: 5710: 5708: 5704: 5699: 5695: 5691: 5687: 5682: 5677: 5673: 5669: 5665: 5663: 5654: 5652: 5650: 5648: 5644: 5632: 5628: 5622: 5619: 5607: 5603: 5597: 5594: 5590: 5585: 5582: 5578: 5574: 5569: 5567: 5563: 5558: 5554: 5549: 5544: 5540: 5536: 5531: 5526: 5522: 5518: 5514: 5510: 5506: 5499: 5496: 5483: 5479: 5472: 5469: 5464: 5460: 5456: 5452: 5448: 5444: 5440: 5436: 5432: 5428: 5424: 5417: 5414: 5408: 5403: 5399: 5395: 5391: 5384: 5381: 5374: 5371: 5368: 5367:archive.today 5364: 5361: 5355: 5352: 5339: 5335: 5329: 5326: 5321: 5314: 5311: 5298: 5294: 5290: 5284: 5281: 5276: 5269: 5266: 5254: 5250: 5244: 5241: 5236: 5229: 5226: 5214: 5208: 5204: 5203: 5195: 5193: 5191: 5189: 5185: 5180: 5173: 5171: 5167: 5162: 5161: 5156: 5150: 5147: 5142: 5136: 5133: 5120: 5116: 5110: 5107: 5102: 5096: 5093: 5080: 5076: 5074: 5066: 5063: 5050: 5044: 5041: 5028: 5026: 5018: 5015: 5011: 5007: 5004: 4999: 4996: 4992: 4988: 4985: 4980: 4977: 4974: 4970: 4967: 4965: 4959: 4953: 4951: 4947: 4931: 4924: 4918: 4916: 4912: 4900: 4896: 4889: 4886: 4873: 4866: 4863: 4850: 4846: 4840: 4837: 4824: 4818: 4815: 4807: 4803: 4799: 4792: 4786: 4783: 4768: 4761: 4758: 4753: 4749: 4745: 4741: 4737: 4733: 4729: 4722: 4719: 4703: 4696: 4689: 4686: 4681: 4677: 4672: 4667: 4663: 4659: 4655: 4651: 4647: 4645: 4644:Tyrannosaurus 4640: 4639:Tyrannosaurus 4631: 4628: 4623: 4619: 4615: 4608: 4605: 4600: 4596: 4589: 4586: 4573: 4569: 4564: 4556: 4553: 4548: 4544: 4540: 4536: 4532: 4528: 4524: 4520: 4516: 4509: 4507: 4503: 4490: 4486: 4480: 4477: 4473: 4472:27 June 2013. 4471: 4467: 4460: 4457: 4452: 4446: 4438: 4434: 4427: 4424: 4420: 4419:28 June 2013. 4418: 4414: 4407: 4405: 4401: 4396: 4392: 4387: 4382: 4377: 4372: 4368: 4364: 4360: 4356: 4352: 4345: 4343: 4341: 4337: 4325: 4322: 4315: 4312: 4307: 4303: 4298: 4293: 4288: 4283: 4279: 4275: 4271: 4267: 4263: 4256: 4253: 4250: 4246: 4243: 4239: 4234: 4231: 4226: 4222: 4218: 4212: 4209: 4204: 4200: 4193: 4190: 4186: 4183: 4178: 4175: 4171: 4167: 4164: 4161: 4160: 4153: 4150: 4147: 4144:'s homepage: 4143: 4137: 4134: 4121: 4117: 4103: 4099: 4094: 4088: 4085: 4080: 4076: 4071: 4066: 4061: 4056: 4052: 4048: 4044: 4040: 4036: 4029: 4026: 4013: 4009: 4007: 3999: 3996: 3983: 3981: 3973: 3970: 3957: 3953: 3951: 3942: 3939: 3934: 3927: 3925: 3923: 3919: 3915: 3909: 3907: 3903: 3898: 3894: 3889: 3884: 3880: 3876: 3872: 3870: 3861: 3858: 3853: 3849: 3845: 3841: 3837: 3833: 3829: 3822: 3819: 3814: 3810: 3806: 3802: 3798: 3794: 3790: 3786: 3782: 3775: 3772: 3767: 3763: 3759: 3752: 3749: 3744: 3740: 3733: 3730: 3725: 3721: 3717: 3713: 3709: 3708:Tyrannosaurus 3702: 3699: 3694: 3692:0-521-43810-1 3688: 3684: 3680: 3676: 3672: 3666: 3663: 3658: 3654: 3649: 3644: 3639: 3634: 3630: 3626: 3622: 3618: 3614: 3607: 3604: 3599: 3597:0-679-43386-4 3593: 3589: 3584: 3583: 3574: 3571: 3566: 3562: 3558: 3554: 3550: 3549:Tyrannosaurus 3543: 3540: 3534: 3529: 3525: 3521: 3514: 3511: 3506: 3502: 3498: 3494: 3490: 3486: 3482: 3481:Tyrannosaurus 3475: 3473: 3469: 3464: 3460: 3456: 3452: 3448: 3444: 3443:Osborn, H. F. 3438: 3435: 3422: 3418: 3414: 3412: 3403: 3400: 3395: 3391: 3386: 3381: 3377: 3373: 3368: 3363: 3359: 3355: 3351: 3343: 3341: 3337: 3333: 3327: 3324: 3317: 3313: 3310: 3309: 3303: 3292: 3285: 3283: 3281: 3277: 3276:T. mcraeensis 3273: 3272:Tyrannosaurus 3269: 3265: 3261: 3257: 3253: 3245: 3244:T. mcraeensis 3240: 3233: 3230: 3228: 3226: 3222: 3221: 3216: 3211: 3208: 3207:Tyrannosaurus 3204: 3200: 3199: 3194: 3190: 3184: 3176: 3174: 3171: 3169: 3165: 3156: 3149: 3147: 3145: 3141: 3136: 3131: 3128: 3125: 3116: 3109: 3107: 3105: 3100: 3096: 3092: 3085: 3080: 3073: 3071: 3068: 3064: 3060: 3056: 3055:Tyrannosaurus 3052: 3048: 3043: 3041: 3037: 3033: 3029: 3025: 3024:Tyrannosaurus 3021: 3017: 3016: 3011: 3007: 3006: 2998: 2994: 2993:Wollaton Hall 2990: 2986: 2981: 2976: 2968: 2966: 2964: 2963:Tyrannosaurus 2959: 2957: 2953: 2949: 2948:Queen Beatrix 2945: 2940: 2937: 2933: 2929: 2925: 2921: 2912: 2907: 2899: 2897: 2895: 2891: 2887: 2883: 2875: 2870: 2863: 2861: 2859: 2858:Tyrannosaurus 2851: 2846: 2839: 2837: 2835: 2831: 2826: 2824: 2819: 2810: 2803: 2801: 2799: 2794: 2793:Tyrannosaurus 2789: 2788:Tyrannosaurus 2784: 2780: 2770: 2763: 2761: 2759: 2754: 2749: 2746: 2743: 2740: 2736: 2732: 2724: 2723:Tyrannosaurus 2719: 2714: 2706: 2704: 2701: 2697: 2693: 2685: 2683: 2680: 2678: 2675: 2670: 2666: 2662: 2657: 2648: 2641: 2639: 2637: 2636:Tyrannosaurus 2633: 2628: 2623: 2619: 2615: 2614:blood vessels 2611: 2607: 2603: 2602: 2593: 2588: 2584: 2582: 2579: 2575: 2571: 2567: 2563: 2559: 2555: 2551: 2547: 2543: 2539: 2534: 2525: 2518: 2516: 2514: 2510: 2506: 2502: 2498: 2497: 2492: 2488: 2483: 2479: 2478: 2472: 2470: 2466: 2461: 2457: 2453: 2450: 2449:tyrannosaurid 2446: 2438: 2434: 2429: 2422: 2420: 2417: 2416:Tyrannosaurus 2413: 2405: 2403: 2401: 2397: 2393: 2389: 2384: 2382: 2378: 2377:Tyrannosaurus 2374: 2370: 2366: 2365: 2360: 2359: 2358:Edmontosaurus 2354: 2350: 2346: 2345:Tyrannosaurus 2342: 2338: 2334: 2325: 2318: 2316: 2314: 2310: 2309:apex predator 2306: 2301: 2296: 2294: 2290: 2286: 2282: 2279:, as well as 2278: 2274: 2270: 2266: 2262: 2258: 2254: 2250: 2246: 2241: 2239: 2235: 2231: 2227: 2226:Tyrannosaurus 2223: 2215: 2210: 2203: 2197: 2194: 2192: 2188: 2184: 2183:Tyrannosaurus 2180: 2179:Tyrannosaurus 2176: 2170: 2168: 2164: 2160: 2155: 2151: 2147: 2143: 2142:Tyrannosaurus 2139: 2135: 2128: 2123: 2118: 2110: 2108: 2106: 2105:Lindsay Zanno 2102: 2098: 2094: 2090: 2086: 2082: 2077: 2075: 2071: 2067: 2063: 2058: 2054: 2050: 2045: 2043: 2039: 2034: 2030: 2026: 2022: 2018: 2014: 2006: 2001: 1994: 1992: 1990: 1989: 1984: 1983:Tyrannosaurus 1980: 1976: 1975: 1974:Tyrannosaurus 1970: 1966: 1962: 1957: 1955: 1950: 1946: 1942: 1938: 1937: 1936:Tyrannosaurus 1932: 1928: 1924: 1920: 1913: 1908: 1903: 1895: 1893: 1890: 1889:individuals. 1888: 1887:Tyrannosaurus 1883: 1878: 1876: 1872: 1868: 1840: 1836: 1826: 1822: 1818: 1814: 1810: 1806: 1802: 1793: 1786: 1784: 1782: 1778: 1774: 1766: 1764: 1762: 1761:Tyrannosaurus 1758: 1753: 1749: 1745: 1741: 1737: 1733: 1729: 1725: 1724: 1719: 1715: 1711: 1703: 1699: 1692: 1689: 1687: 1683: 1681: 1680:tyrannosaurid 1677: 1676: 1671: 1670:Tyrannosaurus 1667: 1666: 1661: 1657: 1656:Tyrannosaurus 1649: 1644: 1640: 1638: 1634: 1633:Tyrannosaurus 1630: 1626: 1622: 1618: 1614: 1613:Tyrannosaurus 1610: 1606: 1599: 1595: 1594: 1593:Tyrannosaurus 1589: 1582: 1580: 1577: 1573: 1569: 1565: 1564:Tyrannosaurus 1561: 1558: 1554: 1550: 1549: 1544: 1540: 1533: 1528: 1521: 1519: 1517: 1513: 1509: 1505: 1501: 1497: 1493: 1489: 1485: 1481: 1477: 1469: 1465: 1461: 1454: 1451: 1449: 1447: 1446:Manospondylus 1443: 1439: 1435: 1431: 1427: 1423: 1419: 1412: 1408: 1404: 1397: 1396:Manospondylus 1394: 1388: 1384: 1383:T. mcraeensis 1380: 1378: 1375: 1373: 1370: 1368: 1364: 1362: 1359: 1357: 1354: 1351: 1349: 1347: 1344: 1342:NMMNH P-3698 1341: 1340: 1337: 1335: 1332: 1330: 1327: 1324: 1322: 1319: 1317: 1314: 1311: 1308: 1305: 1302: 1301: 1298: 1296: 1293: 1291: 1288: 1285: 1283: 1280: 1278: 1275: 1272: 1269: 1266: 1263: 1262: 1259: 1256: 1254: 1251: 1246: 1244: 1241: 1239: 1236: 1233: 1230: 1228:"Tufts-Love" 1227: 1224: 1223: 1219: 1215: 1212: 1210: 1207: 1204: 1202: 1199: 1197: 1194: 1191: 1188: 1184: 1181: 1180: 1176: 1173: 1171: 1168: 1166: 1162: 1159: 1157: 1154: 1152: 1149: 1146: 1143: 1140: 1136: 1133: 1132: 1128: 1126: 1123: 1121: 1118: 1115: 1113: 1110: 1108: 1105: 1102: 1099: 1096: 1095: 1091: 1087: 1085: 1082: 1080: 1077: 1074: 1072: 1069: 1067: 1064: 1061: 1058: 1055: 1052: 1051: 1047: 1046: 1041: 1038: 1036: 1033: 1030: 1028: 1025: 1023: 1020: 1017: 1014: 1011: 1007: 1004: 1003: 1000: 997: 995: 992: 990: 987: 985: 982: 980: 977: 974: 971: 968: 965: 964: 960: 957: 955: 952: 947: 945: 942: 940: 937: 934: 931: 928: 925: 924: 921: 919: 916: 914: 911: 908: 906: 903: 901: 898: 895: 892: 890:"E. D. Cope" 889: 886: 885: 882: 880: 877: 875: 872: 869: 867: 864: 862: 859: 856: 853: 850: 847: 846: 841: 839: 835: 834:Bowman County 832: 830: 827: 825:Dean Pearson 824: 822: 819: 816: 813: 810: 808: 805: 804: 799: 797: 794: 792: 789: 782: 780: 777: 775: 772: 769: 766: 763: 760: 759: 755: 753: 750: 748: 745: 743: 740: 738: 735: 733: 730: 727: 724: 721: 717: 714: 713: 710: 708: 705: 703: 700: 697: 695: 692: 690: 687: 684: 681: 678: 675: 674: 671: 669: 666: 664: 661: 658: 656: 653: 651: 648: 645: 642: 640:"G-rex/Greg" 639: 636: 635: 631: 629: 626: 624: 621: 618: 616: 613: 611: 608: 605: 602: 599: 596: 595: 591: 587: 584: 582: 579: 576: 574: 571: 569: 566: 563: 560: 557: 554: 553: 549: 546: 544: 541: 539:Kathy Wankel 538: 536: 533: 531: 528: 525: 522: 519: 516: 515: 511: 509: 506: 504: 501: 498: 496: 493: 491: 488: 485: 482: 478: 475: 474: 456: 452: 443: 441: 438: 436: 433: 430: 428: 425: 423: 420: 417: 414: 411: 408: 407: 404: 402: 399: 397: 394: 391: 389: 386: 384: 381: 378: 375: 373: 370: 369: 365: 361: 357: 354:Described as 353: 351: 348: 346: 343: 341: 338: 336: 333: 331: 328: 325: 323: 321: 318: 315: 314: 310: 306: 303: 301: 298: 296: 293: 291: 290:New York City 288: 286: 283: 280: 277: 275: 272: 271: 267: 264: 262: 259: 257: 253: 250: 248: 245: 243: 240: 237: 234: 232: 229: 226: 225: 222: 218: 215:Described as 212: 209: 207: 204: 202: 199: 197: 194: 192: 189: 186: 183: 181: 178: 175: 174: 171: 168:Described as 167: 165: 162: 160: 157: 155: 152: 150: 149:New York City 147: 145: 142: 139: 136: 134: 133:Manospondylus 131: 128: 127: 120: 117: 114: 111: 108: 105: 103:Completeness 102: 99: 96: 95: 90:Specimen data 89: 87: 85: 84: 83:Tyrannosaurus 75: 65: 56: 52: 49:This list is 47: 40: 39: 36: 32: 27: 19: 7055: 7020: 7016: 7007: 6999: 6969:(4): 72–77. 6966: 6962: 6949: 6937:. 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Specimens of Tyrannosaurus

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