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species living in open habitats and shrublands from
Bolivia and Paraguay to Argentina, southern Brazil and Uruguay. Although it was suggested that
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293:(Coleoptera: Scarabaeidae: Phanaeini): its phylogenetic significance, geographical polychromatism and the subspecies problem"
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might actually represent two distinct subspecies based on dorsal colouration, a 2022 study concluded that the
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Revista del Museo
Argentino de Ciencias Naturales 'Bernardino Rivadavia', Ciencias ZoolĂłgicas
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Cupello, Mario; Ribeiro-Costa, Cibele S.; Vaz-De-Mello, Fernando Z. (2022).
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MartĂnez, A. (1959). "Catálogo de los
Scarabaeidae Argentinos".
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should continue to be considered a single monotypic species.
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269:: CS1 maint: archived copy as title (
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167:or scarab beetles in the superfamily
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503:-related article is a
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107:Infraorder:
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376:Wikispecies
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256:2014-04-21
217:2011-02-04
193:References
95:Suborder:
88:Coleoptera
64:Arthropoda
100:Polyphaga
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390:BioLib:
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367:Q4938995
361:Wikidata
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330:: 1–126.
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52:Animalia
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143:Genus:
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