316:
102:
645:
124:
3284:
879:
3158:
3197:
3325:
1028:
1011:
994:
2943:
3132:
3171:
977:
1056:
304:. The head of most auger beetles cannot be seen from above, as it is downwardly directed and hidden by the thorax. Exceptions are the
960:
716:"The earliest occurrence and remarkable stasis of the family Bostrichidae (Coleoptera: Polyphaga) in Cretaceous Charentes amber"
3318:
444:
3176:
3184:
688:
3354:
123:
3041:
3018:
696:
328:
has large, distinctive thoracic horns, and is found in parts of Africa and Arabia; it is often imported to Europe as
3311:
3111:
630:
3202:
335:
The fossil record of the family extends to the
Cretaceous, with the oldest records being from the Cenomanian aged
3124:
637:
220:
952:
The
Powderpost Beetles of the World (Coleoptera: Bostrichidae): Keys for the Identification of Species, Volume 1
3349:
1049:
589:
315:
1017:
715:
1000:
663:
471:
451:
555:
2958:
582:
545:
392:
382:
101:
3046:
983:
822:"The first record of subfamily Polycaoninae (Coleoptera; Bostrichidae) from mid-Cretaceous Burmese amber"
525:
461:
3344:
3233:
1024:
1007:
990:
515:
535:
3251:
565:
439:
3212:
3080:
3005:
2195:
910:
833:
778:
623:
491:
2414:
1042:
644:
402:
362:
926:
857:
802:
616:
499:
481:
118:
3238:
2996:
3220:
3067:
2683:
2100:
956:
849:
794:
747:
599:
419:
305:
3295:
3261:
3163:
3225:
2780:
2559:
2554:
2477:
2235:
1251:
918:
841:
786:
737:
727:
336:
110:
2469:
2422:
2267:
2251:
1927:
1687:
1656:
1571:
1489:
1380:
1348:
606:
505:
429:
372:
195:
914:
837:
782:
3119:
3106:
2872:
2823:
2678:
2485:
2153:
2137:
2076:
1971:
1942:
1870:
1834:
1818:
1734:
1679:
1584:
1481:
1330:
1292:
1272:
1168:
345:
292:
are a family of beetles with more than 700 described species. They are commonly called
3338:
3010:
2854:
2658:
2649:
2390:
2361:
2353:
2345:
2319:
2294:
2190:
2177:
2116:
2034:
1937:
1932:
1813:
1747:
1718:
1702:
1666:
1636:
1589:
1543:
1510:
1426:
1418:
1389:
1227:
861:
806:
340:
205:
59:
3023:
930:
875:
3072:
2831:
2755:
2501:
2493:
2448:
2432:
2382:
2303:
2259:
2211:
2129:
2042:
1911:
1898:
1878:
1839:
1739:
1726:
1697:
1631:
1619:
1515:
1472:
1455:
1447:
1235:
1205:
1136:
1125:
1109:
821:
766:
275:
261:
236:
3283:
845:
790:
950:
922:
3189:
3145:
2990:
2917:
2888:
2862:
2849:
2789:
2747:
2731:
2723:
2715:
2699:
2691:
2627:
2585:
2577:
2546:
2461:
2440:
2406:
2398:
2311:
2243:
2227:
2219:
2182:
2169:
2145:
2063:
2055:
2005:
1997:
1955:
1919:
1890:
1862:
1857:
1773:
1768:
1760:
1710:
1646:
1641:
1555:
1520:
1505:
1497:
1413:
1405:
1222:
241:
34:
767:"New auger beetle (Coleoptera; Bostrichidae) from mid-Cretaceous Burmese amber"
2844:
2805:
2797:
2739:
2602:
2564:
2517:
2509:
2329:
2277:
2161:
2092:
1963:
1947:
1903:
1844:
1808:
1788:
1752:
1661:
1651:
1525:
1259:
1243:
700:
251:
79:
44:
2981:
1034:
853:
798:
751:
3093:
2909:
2901:
2893:
2880:
2836:
2635:
2610:
2538:
2366:
2337:
2021:
1852:
1826:
1778:
1692:
1594:
1563:
1535:
1368:
1343:
1338:
1319:
1284:
1173:
1156:
1148:
1097:
1085:
246:
185:
155:
135:
84:
3054:
2975:
2813:
2763:
2707:
2666:
2615:
2272:
2203:
2084:
1796:
1602:
1576:
1434:
1267:
1214:
1193:
1103:
256:
74:
69:
54:
49:
39:
3033:
3059:
2530:
2522:
2374:
2108:
2029:
2013:
742:
309:
266:
89:
64:
3150:
2121:
1981:
1614:
1091:
1079:
1066:
175:
165:
145:
3137:
2952:
3098:
714:
Delclòs, Xavier; Soriano, Carmen; Perrichot, Vincent (2014-03-31).
3246:
643:
329:
314:
732:
2956:
1365:
1316:
1190:
1122:
1038:
901:
n. gen., n. sp. (Coleoptera: Ecumenidae) in
Dominican amber".
3085:
903:
Historical
Biology: An International Journal of Paleobiology
349:
and the extant subfamilies
Dinoderinae and Polycaoninae.
1437:(furniture beetles, death watch beetles, spider beetles)
3299:
332:
in
African wooden bowls ("ethnic souvenirs") .
2965:
2871:
2822:
2788:
2779:
2657:
2648:
2601:
2576:
2460:
2431:
2328:
2302:
2293:
2075:
2054:
1980:
1787:
1678:
1534:
1480:
1471:
1446:
1388:
1379:
1204:
693:(Olivier) (Coleoptera: Bostrichidae: Bostrichinae)"
308:(subfamily Lyctinae), and members of the subfamily
2314:(jewel beetles, or metallic wood-boring beetles)
897:n. gen., n. sp., (Coleoptera: Bostrichidae) and
820:Legalov, Andrei A.; Háva, Jiří (December 2020).
2037:(true weevils, bark beetles, ambrosia beetles)
3319:
1050:
949:Borowski, Jerzy; Węgrzynowicz, Piotr (2012).
8:
2280:(ironclad beetles, cylindrical bark beetles)
3326:
3312:
2953:
2785:
2654:
2299:
1477:
1385:
1376:
1362:
1313:
1201:
1187:
1119:
1057:
1043:
1035:
100:
20:
2944:List of subgroups of the order Coleoptera
2024:(straight snout weevils, New York weevil)
741:
731:
2451:(cicada beetle, cicada parasite beetles)
679:
2541:(false fireflies, long-lipped beetles)
343:ambers, belonging to the extant genus
7:
3280:
3278:
3252:396787A7-5C54-1B5D-FF1D-F97FFE5CF66C
3213:137206b0-7cdb-4776-a05f-83aee4d5196f
3125:5fbfc7d4-7fc1-459b-848c-860892c59681
2567:(false metallic wood-boring beetles)
765:Legalov, Andrei A. (December 2018).
576:Species found in the United Kingdom
3298:. You can help Knowledge (XXG) by
14:
3282:
2178:Perimylopidae, or Promecheilidae
1763:(minute brown scavenger beetles)
122:
2393:(variegated mud-loving beetles)
697:Pest and Diseases Image Library
1:
2222:(narrow-waisted bark beetles)
1914:(parasitic flat bark beetles)
1865:(short-winged flower beetles)
1721:(lady beetles, or God's cows)
1458:(tooth-necked fungus beetles)
846:10.1016/j.cretres.2020.104620
791:10.1016/j.cretres.2018.08.014
2734:(sand-loving scarab beetles)
2124:(minute tree-fungus beetles)
1950:(silvanid flat bark beetles)
923:10.1080/08912963.2012.712518
2883:(primitive carrion beetles)
2839:(minute mud-loving beetles)
2702:(bumble bee scarab beetles)
2472:(soft-bodied plant beetles)
2443:(soft bodied plant beetles)
1246:(predaceous diving beetles)
1031:Featured Creatures Web site
1021:, a false powderpost beetle
1014:Featured Creatures Web site
997:Featured Creatures Web site
893:Poinar Jr, George (2013). "
638:Trogoxylon parallelopipedum
3371:
3277:
2718:(scavenger scarab beetles)
2710:(enigmatic scarab beetles)
1883:Lamingtonium binnaberrense
1597:(soft-wing flower beetles)
720:Palaeontologia Electronica
687:Walker, Ken (2006-11-23).
631:Stephanopachys substriatus
413:Species found in Australia
2941:
2857:(water scavenger beetles)
2590:Rhinorhipus tamborinensis
2369:(long-toed water beetles)
2262:(polypore fungus beetles)
2164:(palm and flower beetles)
2148:(tumbling flower beetles)
2095:(ant-like flower beetles)
1873:(lined flat bark beetles)
1847:(pleasing fungus beetles)
1742:(handsome fungus beetles)
1375:
1361:
1326:
1312:
1287:(burrowing water beetles)
1200:
1186:
1132:
1118:
1074:
985:Heterobostrychus aequalis
649:Heterobostrychus aequalis
357:This list is incomplete:
302:horned powderpost beetles
281:
274:
233:
228:
119:Scientific classification
117:
108:
99:
23:
2904:(feather-winged beetles)
2238:(false longhorn beetles)
2185:(jugular-horned beetles)
2132:(false darkling beetles)
2103:(cryptic fungus beetles)
1922:(shining flower beetles)
1262:(crawling water beetles)
1161:Sikhotealinia zhiltzovae
1004:, southern lyctus beetle
987:, an oriental wood borer
691:Bostrychoplites cornutus
590:Bostrychoplites cornutus
326:Bostrychoplites cornutus
298:false powderpost beetles
2385:(forest stream beetles)
2246:(synchroa bark beetles)
2206:(dead log bark beetles)
2198:(fire-coloured beetles)
2087:(ant-like leaf beetles)
1821:(silken fungus beetles)
1755:(well polished beetles)
1729:(minute fungus beetles)
1548:Acanthocnemus nigricans
895:Discoclavata dominicana
664:Discoclavata dominicana
3294:-related article is a
2896:(round fungus beetles)
2230:(false flower beetles)
2214:(wedge-shaped beetles)
2172:(false blister beetle)
2156:(hairy fungus beetles)
2111:(conifer bark beetles)
2008:(leaf-rolling weevils)
1669:(bark-gnawing beetles)
1429:(wounded-tree beetles)
1295:(false ground beetles)
1217:(trout-stream beetles)
652:
546:Xylopsocus gibbicollis
472:Mesoxylion cylindricus
452:Lyctus parallelocollis
393:Prostephanus truncatus
383:Lichenophanes bicornis
322:
3234:Paleobiology Database
2808:(false clown beetles)
2630:(plate-thigh beetles)
2512:(false click beetles)
2417:(water-penny beetles)
2066:(ship-timber beetles)
2045:(pine flower weevils)
1893:(root-eating beetles)
1713:(minute bark beetles)
1624:Phloiophilus edwardsi
1151:(reticulated beetles)
647:
556:Xylothrips religiosus
318:
3120:Fauna Europaea (new)
2686:(false stag beetles)
2671:Belohina inexpectata
2533:(net-winged beetles)
2496:(rare click beetles)
2409:(travertine beetles)
2401:(minute mud beetles)
1966:(dry-fungus beetles)
1801:Agapytho foveicollis
1558:(false skin beetles)
1421:(Jacobson's beetles)
1141:Crowsoniella relicta
1019:Xylopsocus capucinus
899:Lissantauga epicrana
583:Bostrichus capucinus
492:Rhyzopertha dominica
3355:Bostrichoidea stubs
2684:Diphyllostomatidae
2016:(primitive weevils)
1829:(flat bark beetles)
1579:(checkered beetles)
1566:(fruitworm beetles)
1254:(whirligig beetles)
915:2013HBio...25..107P
838:2020CrRes.11604620L
826:Cretaceous Research
783:2018CrRes..92..210L
771:Cretaceous Research
526:Xylobosca bispinosa
462:Mesoxylion collaris
403:Xylobiops basilaris
363:Amphicerus cornutus
2549:(glowworm beetles)
2254:(darkling beetles)
2101:Archeocrypticidae
1958:(palmetto beetles)
1705:(dry bark beetles)
1492:(longhorn beetles)
1002:Lyctus planicollis
653:
617:Lyctus planicollis
516:Xylion cylindricus
500:lesser grain borer
482:Minthea rugicollis
323:
306:powderpost beetles
282:About 700 species
3307:
3306:
3272:
3271:
3221:Open Tree of Life
2959:Taxon identifiers
2950:
2949:
2937:
2936:
2933:
2932:
2929:
2928:
2921:
2913:
2912:(carrion beetles)
2905:
2897:
2884:
2858:
2840:
2809:
2801:
2775:
2774:
2767:
2759:
2751:
2743:
2735:
2727:
2719:
2711:
2703:
2695:
2687:
2674:
2644:
2643:
2631:
2623:
2593:
2568:
2550:
2542:
2534:
2526:
2513:
2505:
2497:
2489:
2488:(soldier beetles)
2481:
2478:Brachypsectridae
2473:
2452:
2444:
2418:
2410:
2402:
2394:
2386:
2378:
2370:
2357:
2349:
2341:
2315:
2289:
2288:
2281:
2263:
2255:
2247:
2239:
2236:Stenotrachelidae
2231:
2223:
2215:
2207:
2199:
2186:
2173:
2165:
2157:
2149:
2141:
2140:(blister beetles)
2133:
2125:
2112:
2104:
2096:
2088:
2067:
2046:
2038:
2025:
2017:
2009:
2001:
1991:
1967:
1959:
1951:
1923:
1915:
1907:
1894:
1886:
1874:
1866:
1848:
1830:
1822:
1804:
1764:
1756:
1743:
1730:
1722:
1714:
1706:
1688:Akalyptoischiidae
1670:
1627:
1610:
1598:
1580:
1567:
1559:
1551:
1501:
1493:
1467:
1466:
1459:
1438:
1430:
1422:
1409:
1401:
1357:
1356:
1334:
1308:
1307:
1304:
1303:
1296:
1288:
1280:
1263:
1255:
1247:
1239:
1231:
1218:
1182:
1181:
1164:
1152:
1144:
669:
612:
600:Lyctus cavicollis
595:
571:
561:
551:
541:
536:Xylodeleis obsipa
531:
521:
511:
497:
487:
477:
467:
457:
447:
435:
425:
420:Dinoderus minutus
408:
398:
388:
378:
368:
286:
285:
224:
30:Cenomanian–Recent
16:Family of beetles
3362:
3328:
3321:
3314:
3286:
3279:
3265:
3264:
3255:
3254:
3242:
3241:
3229:
3228:
3216:
3215:
3206:
3205:
3193:
3192:
3190:NHMSYS0001717355
3180:
3179:
3167:
3166:
3154:
3153:
3141:
3140:
3128:
3127:
3115:
3114:
3102:
3101:
3089:
3088:
3076:
3075:
3063:
3062:
3050:
3049:
3037:
3036:
3027:
3026:
3014:
3013:
3001:
3000:
2999:
2986:
2985:
2984:
2954:
2919:
2911:
2903:
2895:
2882:
2856:
2838:
2807:
2799:
2786:
2781:Staphyliniformia
2765:
2757:
2749:
2741:
2733:
2725:
2717:
2709:
2701:
2693:
2685:
2668:
2655:
2629:
2620:Declinia relicta
2617:
2587:
2566:
2560:Sinopyrophoridae
2555:Rhagophthalmidae
2548:
2540:
2532:
2524:
2511:
2503:
2495:
2487:
2479:
2471:
2470:Artematopodidae
2450:
2442:
2416:
2408:
2400:
2392:
2384:
2377:(riffle beetles)
2376:
2368:
2356:(turtle beetles)
2355:
2347:
2339:
2313:
2300:
2279:
2261:
2253:
2245:
2237:
2229:
2221:
2213:
2205:
2197:
2184:
2171:
2163:
2155:
2147:
2139:
2131:
2123:
2110:
2102:
2094:
2086:
2065:
2044:
2036:
2023:
2015:
2007:
2000:(fungus weevils)
1999:
1990:
1989:
1985:
1972:Tasmosalpingidae
1965:
1957:
1949:
1921:
1913:
1905:
1892:
1880:
1872:
1864:
1846:
1828:
1820:
1798:
1762:
1754:
1741:
1728:
1720:
1712:
1704:
1668:
1621:
1604:
1596:
1578:
1565:
1557:
1545:
1544:Acanthocnemidae
1499:
1491:
1478:
1457:
1436:
1428:
1420:
1407:
1399:
1386:
1377:
1363:
1332:
1314:
1294:
1286:
1274:
1261:
1253:
1245:
1237:
1230:(ground beetles)
1229:
1216:
1202:
1188:
1158:
1150:
1138:
1137:Crowsoniellidae
1120:
1059:
1052:
1045:
1036:
966:
935:
934:
890:
884:
883:
872:
866:
865:
817:
811:
810:
762:
756:
755:
745:
735:
711:
705:
704:
699:. Archived from
684:
668:Poinar Jr., 2013
667:
610:
593:
569:
566:Xylotillus lindi
559:
549:
539:
529:
519:
509:
495:
485:
475:
465:
455:
443:
440:Lyctus discedens
433:
423:
406:
396:
386:
376:
366:
353:Selected species
320:Dinoderus brevis
219:
127:
126:
111:Euderia squamosa
104:
94:
31:
27:Temporal range:
21:
3370:
3369:
3365:
3364:
3363:
3361:
3360:
3359:
3350:Beetle families
3335:
3334:
3333:
3332:
3275:
3273:
3268:
3260:
3258:
3250:
3245:
3237:
3232:
3224:
3219:
3211:
3209:
3201:
3196:
3188:
3183:
3175:
3170:
3162:
3157:
3149:
3144:
3136:
3131:
3123:
3118:
3110:
3105:
3097:
3092:
3084:
3079:
3071:
3066:
3058:
3053:
3045:
3040:
3032:
3030:
3022:
3017:
3009:
3004:
2995:
2994:
2989:
2980:
2979:
2974:
2961:
2951:
2946:
2925:
2867:
2818:
2800:(clown beetles)
2771:
2742:(betsy beetles)
2650:Scarabaeiformia
2640:
2597:
2572:
2504:(click beetles)
2480:(Texas beetles)
2456:
2427:
2423:Ptilodactylidae
2348:(cedar beetles)
2324:
2285:
2268:Trictenotomidae
2154:Mycetophagidae
2071:
2050:
1987:
1986:
1984:
1976:
1928:Phloeostichidae
1879:Lamingtoniidae
1871:Laemophloeidae
1819:Cryptophagidae
1783:
1674:
1657:Thanerocleridae
1620:Phloiophilidae
1607:Metaxina ornata
1572:Chaetosomatidae
1530:
1463:
1442:
1400:(auger beetles)
1381:Bostrichiformia
1371:
1353:
1349:Torridincolidae
1333:(skiff beetles)
1331:Hydroscaphidae
1322:
1300:
1293:Trachypachidae
1238:(tiger beetles)
1196:
1178:
1128:
1114:
1070:
1063:
974:
969:
963:
948:
944:
939:
938:
892:
891:
887:
874:
873:
869:
819:
818:
814:
764:
763:
759:
713:
712:
708:
686:
685:
681:
676:
624:Lyctus sinensis
607:Lyctus linearis
594:(Olivier, 1790)
506:Sinoxylon anale
430:Lyctus brunneus
373:Apate terebrans
355:
218:
196:Bostrichiformia
121:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
3368:
3366:
3358:
3357:
3352:
3347:
3337:
3336:
3331:
3330:
3323:
3316:
3308:
3305:
3304:
3287:
3270:
3269:
3267:
3266:
3256:
3243:
3230:
3217:
3207:
3194:
3181:
3168:
3155:
3142:
3129:
3116:
3107:Fauna Europaea
3103:
3090:
3077:
3064:
3051:
3038:
3028:
3015:
3002:
2987:
2971:
2969:
2963:
2962:
2957:
2948:
2947:
2942:
2939:
2938:
2935:
2934:
2931:
2930:
2927:
2926:
2924:
2923:
2920:(rove beetles)
2918:Staphylinidae
2915:
2907:
2899:
2891:
2886:
2877:
2875:
2873:Staphylinoidea
2869:
2868:
2866:
2865:
2860:
2855:Hydrophilidae
2852:
2847:
2842:
2834:
2828:
2826:
2824:Hydrophiloidea
2820:
2819:
2817:
2816:
2811:
2803:
2794:
2792:
2783:
2777:
2776:
2773:
2772:
2770:
2769:
2766:(hide beetles)
2761:
2753:
2750:(rain beetles)
2745:
2737:
2729:
2726:(stag beetles)
2721:
2713:
2705:
2697:
2689:
2681:
2679:Bolboceratidae
2676:
2663:
2661:
2652:
2646:
2645:
2642:
2641:
2639:
2638:
2633:
2625:
2613:
2607:
2605:
2599:
2598:
2596:
2595:
2586:Rhinorhipidae
2582:
2580:
2578:Rhinorhipoidea
2574:
2573:
2571:
2570:
2562:
2557:
2552:
2544:
2536:
2528:
2520:
2515:
2507:
2499:
2491:
2483:
2475:
2466:
2464:
2458:
2457:
2455:
2454:
2446:
2437:
2435:
2429:
2428:
2426:
2425:
2420:
2412:
2404:
2396:
2391:Heteroceridae
2388:
2380:
2372:
2364:
2359:
2354:Chelonariidae
2351:
2346:Callirhipidae
2343:
2340:(pill beetles)
2334:
2332:
2326:
2325:
2323:
2322:
2317:
2308:
2306:
2297:
2291:
2290:
2287:
2286:
2284:
2283:
2275:
2270:
2265:
2257:
2252:Tenebrionidae
2249:
2241:
2233:
2225:
2217:
2209:
2201:
2193:
2188:
2180:
2175:
2167:
2159:
2151:
2143:
2135:
2127:
2119:
2114:
2106:
2098:
2090:
2081:
2079:
2077:Tenebrionoidea
2073:
2072:
2070:
2069:
2060:
2058:
2052:
2051:
2049:
2048:
2040:
2035:Curculionidae
2032:
2027:
2019:
2011:
2003:
1994:
1992:
1982:Curculionoidea
1978:
1977:
1975:
1974:
1969:
1961:
1953:
1945:
1943:Protocucujidae
1940:
1935:
1930:
1925:
1917:
1909:
1901:
1896:
1888:
1876:
1868:
1860:
1855:
1850:
1842:
1837:
1835:Cybocephalidae
1832:
1824:
1816:
1811:
1806:
1793:
1791:
1785:
1784:
1782:
1781:
1776:
1771:
1766:
1758:
1750:
1745:
1737:
1735:Discolomatidae
1732:
1724:
1719:Coccinellidae
1716:
1708:
1703:Bothrideridae
1700:
1695:
1690:
1684:
1682:
1680:Coccinelloidea
1676:
1675:
1673:
1672:
1667:Trogossitidae
1664:
1659:
1654:
1649:
1644:
1639:
1634:
1629:
1617:
1612:
1600:
1592:
1587:
1585:Lophocateridae
1582:
1574:
1569:
1561:
1553:
1540:
1538:
1532:
1531:
1529:
1528:
1523:
1518:
1513:
1508:
1503:
1500:(leaf beetles)
1498:Chrysomelidae
1495:
1486:
1484:
1482:Chrysomeloidea
1475:
1469:
1468:
1465:
1464:
1462:
1461:
1452:
1450:
1444:
1443:
1441:
1440:
1432:
1427:Nosodendridae
1424:
1419:Jacobsoniidae
1416:
1411:
1408:(skin beetles)
1403:
1394:
1392:
1383:
1373:
1372:
1366:
1359:
1358:
1355:
1354:
1352:
1351:
1346:
1341:
1336:
1327:
1324:
1323:
1317:
1310:
1309:
1306:
1305:
1302:
1301:
1299:
1298:
1290:
1282:
1277:Meru phyllisae
1270:
1265:
1257:
1249:
1241:
1233:
1225:
1220:
1211:
1209:
1198:
1197:
1191:
1184:
1183:
1180:
1179:
1177:
1176:
1171:
1169:Micromalthidae
1166:
1154:
1146:
1133:
1130:
1129:
1123:
1116:
1115:
1113:
1112:
1106:
1100:
1094:
1088:
1082:
1075:
1072:
1071:
1064:
1062:
1061:
1054:
1047:
1039:
1033:
1032:
1015:
998:
981:
973:
972:External links
970:
968:
967:
961:
945:
943:
940:
937:
936:
909:(1): 107–113.
885:
876:"Bostrichidae"
867:
812:
757:
706:
703:on 2007-09-09.
689:"Auger beetle
678:
677:
675:
672:
671:
670:
658:
657:
656:Fossil species
642:
641:
634:
627:
620:
613:
603:
596:
586:
578:
577:
573:
572:
562:
552:
542:
532:
522:
512:
502:
488:
478:
468:
458:
448:
436:
426:
415:
414:
410:
409:
399:
389:
379:
377:(Pallas, 1772)
369:
367:(Pallas, 1772)
354:
351:
346:Stephanopachys
284:
283:
279:
278:
272:
271:
270:
269:
264:
259:
254:
249:
244:
239:
231:
230:
226:
225:
213:
209:
208:
203:
199:
198:
193:
189:
188:
183:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
143:
139:
138:
133:
129:
128:
115:
114:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
32:
26:
15:
13:
10:
9:
6:
4:
3:
2:
3367:
3356:
3353:
3351:
3348:
3346:
3343:
3342:
3340:
3329:
3324:
3322:
3317:
3315:
3310:
3309:
3303:
3301:
3297:
3293:
3288:
3285:
3281:
3276:
3263:
3257:
3253:
3248:
3244:
3240:
3235:
3231:
3227:
3222:
3218:
3214:
3208:
3204:
3199:
3195:
3191:
3186:
3182:
3178:
3173:
3169:
3165:
3160:
3156:
3152:
3147:
3143:
3139:
3134:
3130:
3126:
3121:
3117:
3113:
3108:
3104:
3100:
3095:
3091:
3087:
3082:
3078:
3074:
3069:
3065:
3061:
3056:
3052:
3048:
3043:
3039:
3035:
3029:
3025:
3020:
3016:
3012:
3007:
3003:
2998:
2992:
2988:
2983:
2977:
2973:
2972:
2970:
2968:
2964:
2960:
2955:
2945:
2940:
2922:
2916:
2914:
2908:
2906:
2900:
2898:
2892:
2890:
2887:
2885:
2879:
2878:
2876:
2874:
2870:
2864:
2861:
2859:
2853:
2851:
2848:
2846:
2843:
2841:
2835:
2833:
2830:
2829:
2827:
2825:
2821:
2815:
2812:
2810:
2806:Sphaeritidae
2804:
2802:
2796:
2795:
2793:
2791:
2787:
2784:
2782:
2778:
2768:
2762:
2760:
2756:Scarabaeidae
2754:
2752:
2746:
2744:
2738:
2736:
2730:
2728:
2722:
2720:
2714:
2712:
2706:
2704:
2698:
2696:
2694:(dor beetles)
2690:
2688:
2682:
2680:
2677:
2675:
2672:
2665:
2664:
2662:
2660:
2659:Scarabaeoidea
2656:
2653:
2651:
2647:
2637:
2634:
2632:
2626:
2624:
2621:
2614:
2612:
2609:
2608:
2606:
2604:
2600:
2594:
2591:
2584:
2583:
2581:
2579:
2575:
2569:
2563:
2561:
2558:
2556:
2553:
2551:
2545:
2543:
2537:
2535:
2529:
2527:
2521:
2519:
2516:
2514:
2508:
2506:
2500:
2498:
2494:Cerophytidae
2492:
2490:
2484:
2482:
2476:
2474:
2468:
2467:
2465:
2463:
2459:
2453:
2449:Rhipiceridae
2447:
2445:
2439:
2438:
2436:
2434:
2430:
2424:
2421:
2419:
2413:
2411:
2405:
2403:
2397:
2395:
2389:
2387:
2383:Eulichadidae
2381:
2379:
2373:
2371:
2365:
2363:
2362:Cneoglossidae
2360:
2358:
2352:
2350:
2344:
2342:
2336:
2335:
2333:
2331:
2327:
2321:
2320:Schizopodidae
2318:
2316:
2310:
2309:
2307:
2305:
2301:
2298:
2296:
2295:Elateriformia
2292:
2282:
2276:
2274:
2271:
2269:
2266:
2264:
2260:Tetratomidae
2258:
2256:
2250:
2248:
2242:
2240:
2234:
2232:
2226:
2224:
2218:
2216:
2212:Ripiphoridae
2210:
2208:
2202:
2200:
2196:Pyrochroidae
2194:
2192:
2191:Pterogeniidae
2189:
2187:
2181:
2179:
2176:
2174:
2168:
2166:
2160:
2158:
2152:
2150:
2144:
2142:
2136:
2134:
2130:Melandryidae
2128:
2126:
2120:
2118:
2117:Chalcodryidae
2115:
2113:
2107:
2105:
2099:
2097:
2091:
2089:
2083:
2082:
2080:
2078:
2074:
2068:
2062:
2061:
2059:
2057:
2053:
2047:
2043:Nemonychidae
2041:
2039:
2033:
2031:
2028:
2026:
2020:
2018:
2012:
2010:
2004:
2002:
1996:
1995:
1993:
1983:
1979:
1973:
1970:
1968:
1962:
1960:
1954:
1952:
1946:
1944:
1941:
1939:
1938:Propalticidae
1936:
1934:
1933:Priasilphidae
1931:
1929:
1926:
1924:
1918:
1916:
1912:Passandridae
1910:
1908:
1906:(sap beetles)
1902:
1900:
1897:
1895:
1889:
1887:
1884:
1877:
1875:
1869:
1867:
1861:
1859:
1856:
1854:
1851:
1849:
1843:
1841:
1838:
1836:
1833:
1831:
1825:
1823:
1817:
1815:
1814:Cavognathidae
1812:
1810:
1807:
1805:
1802:
1795:
1794:
1792:
1790:
1786:
1780:
1777:
1775:
1772:
1770:
1767:
1765:
1759:
1757:
1751:
1749:
1748:Eupsilobiidae
1746:
1744:
1740:Endomychidae
1738:
1736:
1733:
1731:
1727:Corylophidae
1725:
1723:
1717:
1715:
1709:
1707:
1701:
1699:
1696:
1694:
1691:
1689:
1686:
1685:
1683:
1681:
1677:
1671:
1665:
1663:
1660:
1658:
1655:
1653:
1650:
1648:
1645:
1643:
1642:Protopeltidae
1640:
1638:
1637:Prionoceridae
1635:
1633:
1630:
1628:
1625:
1618:
1616:
1613:
1611:
1608:
1601:
1599:
1593:
1591:
1590:Mauroniscidae
1588:
1586:
1583:
1581:
1575:
1573:
1570:
1568:
1562:
1560:
1554:
1552:
1549:
1542:
1541:
1539:
1537:
1533:
1527:
1524:
1522:
1519:
1517:
1514:
1512:
1511:Megalopodidae
1509:
1507:
1504:
1502:
1496:
1494:
1490:Cerambycidae
1488:
1487:
1485:
1483:
1479:
1476:
1474:
1470:
1460:
1456:Derodontidae
1454:
1453:
1451:
1449:
1448:Derodontoidea
1445:
1439:
1433:
1431:
1425:
1423:
1417:
1415:
1414:Endecatomidae
1412:
1410:
1404:
1402:
1398:Bostrichidae
1396:
1395:
1393:
1391:
1390:Bostrichoidea
1387:
1384:
1382:
1378:
1374:
1370:
1364:
1360:
1350:
1347:
1345:
1344:Sphaeriusidae
1342:
1340:
1337:
1335:
1329:
1328:
1325:
1321:
1315:
1311:
1297:
1291:
1289:
1283:
1281:
1278:
1271:
1269:
1266:
1264:
1258:
1256:
1250:
1248:
1242:
1240:
1236:Cicindelidae
1234:
1232:
1226:
1224:
1221:
1219:
1213:
1212:
1210:
1207:
1203:
1199:
1195:
1189:
1185:
1175:
1172:
1170:
1167:
1165:
1162:
1155:
1153:
1147:
1145:
1142:
1135:
1134:
1131:
1127:
1121:
1117:
1111:
1107:
1105:
1101:
1099:
1095:
1093:
1089:
1087:
1083:
1081:
1077:
1076:
1073:
1068:
1060:
1055:
1053:
1048:
1046:
1041:
1040:
1037:
1030:
1026:
1022:
1020:
1016:
1013:
1009:
1005:
1003:
999:
996:
992:
988:
986:
982:
979:
976:
975:
971:
964:
962:9788362860036
958:
954:
953:
947:
946:
941:
932:
928:
924:
920:
916:
912:
908:
904:
900:
896:
889:
886:
882:. 2004-11-11.
881:
877:
871:
868:
863:
859:
855:
851:
847:
843:
839:
835:
831:
827:
823:
816:
813:
808:
804:
800:
796:
792:
788:
784:
780:
776:
772:
768:
761:
758:
753:
749:
744:
739:
734:
729:
725:
721:
717:
710:
707:
702:
698:
694:
692:
683:
680:
673:
666:
665:
660:
659:
655:
654:
650:
646:
640:
639:
635:
633:
632:
628:
626:
625:
621:
619:
618:
614:
611:(Goeze, 1877)
609:
608:
604:
602:
601:
597:
592:
591:
587:
585:
584:
580:
579:
575:
574:
568:
567:
563:
558:
557:
553:
548:
547:
543:
538:
537:
533:
528:
527:
523:
518:
517:
513:
508:
507:
503:
501:
494:
493:
489:
484:
483:
479:
474:
473:
469:
464:
463:
459:
454:
453:
449:
446:
442:
441:
437:
432:
431:
427:
422:
421:
417:
416:
412:
411:
405:
404:
400:
395:
394:
390:
387:(Weber, 1801)
385:
384:
380:
375:
374:
370:
365:
364:
360:
359:
358:
352:
350:
348:
347:
342:
338:
333:
331:
327:
321:
317:
313:
311:
307:
303:
299:
295:
294:auger beetles
291:
280:
277:
273:
268:
265:
263:
260:
258:
255:
253:
250:
248:
245:
243:
240:
238:
235:
234:
232:
227:
222:
217:
214:
211:
210:
207:
206:Bostrichoidea
204:
202:Superfamily:
201:
200:
197:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
147:
144:
141:
140:
137:
134:
131:
130:
125:
120:
116:
113:
112:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
3345:Bostrichidae
3300:expanding it
3292:Bostrichidae
3291:
3289:
3274:
3024:Bostrichidae
3011:Bostrichidae
2997:Bostrichidae
2967:Bostrichidae
2966:
2845:Helophoridae
2837:Georissidae
2832:Epimetopidae
2748:Pleocomidae
2732:Ochodaeidae
2716:Hybosoridae
2700:Glaphyridae
2692:Geotrupidae
2670:
2667:Belohinidae
2628:Eucinetidae
2619:
2616:Decliniidae
2589:
2547:Phengodidae
2486:Cantharidae
2441:Dascillidae
2433:Dascilloidea
2415:Psephenidae
2407:Lutrochidae
2399:Limnichidae
2312:Buprestidae
2304:Buprestoidea
2244:Synchroidae
2228:Scraptiidae
2220:Salpingidae
2183:Prostomidae
2170:Oedemeridae
2146:Mordellidae
2064:Lymexylidae
2056:Lymexyloidea
2006:Attelabidae
1998:Anthribidae
1956:Smicripidae
1920:Phalacridae
1904:Nitidulidae
1899:Myraboliidae
1891:Monotomidae
1882:
1863:Kateretidae
1840:Cyclaxyridae
1800:
1797:Agapythidae
1761:Latridiidae
1711:Cerylonidae
1698:Anamorphidae
1632:Phycosecidae
1623:
1606:
1603:Metaxinidae
1556:Biphyllidae
1547:
1516:Orsodacnidae
1473:Cucujiformia
1406:Dermestidae
1397:
1276:
1215:Amphizoidae
1160:
1140:
1126:Archostemata
1110:Holometabola
1108:Superorder:
1102:Infraclass:
1018:
1001:
984:
951:
942:Bibliography
906:
902:
898:
894:
888:
870:
829:
825:
815:
774:
770:
760:
733:10.26879/408
723:
719:
709:
701:the original
690:
682:
662:
648:
636:
629:
622:
615:
605:
598:
588:
581:
564:
554:
544:
534:
524:
514:
504:
490:
480:
470:
460:
450:
438:
428:
418:
401:
397:(Horn, 1878)
391:
381:
371:
361:
356:
344:
334:
325:
324:
319:
301:
297:
293:
290:Bostrichidae
289:
287:
262:Polycaoninae
237:Bostrichinae
229:Subfamilies
216:Bostrichidae
215:
192:Infraorder:
109:
24:Bostrichidae
18:
3146:iNaturalist
2991:Wikispecies
2889:Hydraenidae
2863:Spercheidae
2850:Hydrochidae
2814:Synteliidae
2798:Histeridae
2790:Histeroidea
2740:Passalidae
2708:Glaresidae
2565:Throscidae
2525:(fireflies)
2523:Lampyridae
2510:Eucnemidae
2502:Elateridae
2462:Elateroidea
2278:Zopheridae
2162:Mycteridae
2093:Anthicidae
1964:Sphindidae
1948:Silvanidae
1858:Hobartiidae
1845:Erotylidae
1774:Mycetaeidae
1769:Murmidiidae
1753:Euxestidae
1647:Rentoniidae
1521:Oxypeltidae
1506:Disteniidae
1339:Lepiceridae
1268:Hygrobiidae
1260:Haliplidae
1244:Dytiscidae
1223:Aspidytidae
777:: 210–213.
743:2445/120503
570:(Blackburn)
560:(Boisduval)
496:(Fabricius)
424:(Fabricius)
407:(Say, 1824)
242:Dinoderinae
3339:Categories
2910:Silphidae
2902:Ptiliidae
2894:Leiodidae
2881:Agyrtidae
2724:Lucanidae
2603:Scirtoidea
2539:Omethidae
2518:Jurasaidae
2367:Dryopidae
2338:Byrrhidae
2330:Byrrhoidea
2022:Brentidae
1827:Cucujidae
1809:Boganiidae
1789:Cucujoidea
1662:Thymalidae
1652:Rhadalidae
1595:Melyridae
1564:Byturidae
1526:Vesperidae
1435:Ptiniidae
1285:Noteridae
1252:Gyrinidae
1228:Carabidae
1157:Jurodidae
1149:Cupedidae
1096:Subclass:
1086:Arthropoda
1067:Coleoptera
980:Fact Sheet
955:. Mantis.
832:: 104620.
726:(1): 1–8.
674:References
466:(Erichson)
434:(Stephens)
337:Charentese
252:Euderiinae
182:Suborder:
176:Coleoptera
156:Arthropoda
2764:Trogidae
2758:(scarabs)
2636:Scirtidae
2611:Clambidae
2204:Pythidae
2138:Meloidae
2085:Aderidae
1988:(weevils)
1853:Helotidae
1779:Teredidae
1693:Alexiidae
1577:Cleridae
1536:Cleroidea
1369:Polyphaga
1367:Suborder
1320:Myxophaga
1318:Suborder
1273:Meruidae
1192:Suborder
1174:Ommatidae
1124:Suborder
1098:Pterygota
1078:Kingdom:
862:224965515
854:0195-6671
807:133930742
799:0195-6671
752:1094-8074
550:(Macleay)
530:(Macleay)
476:(Macleay)
456:Blackburn
445:Blackburn
276:Diversity
247:Dysidinae
221:Latreille
186:Polyphaga
142:Kingdom:
136:Eukaryota
3055:BugGuide
3031:BioLib:
2976:Wikidata
2531:Lycidae
2375:Elmidae
2273:Ulodidae
2109:Boridae
2014:Belidae
1615:Peltidae
1208:families
1194:Adephaga
1104:Neoptera
1084:Phylum:
1080:Animalia
1069:families
931:86040146
486:(Walker)
257:Lyctinae
212:Family:
152:Phylum:
146:Animalia
132:Domain:
3262:4779743
2982:Q874723
2122:Ciidae
2030:Caridae
1092:Insecta
1090:Class:
1065:Extant
1023:on the
1006:on the
989:on the
911:Bibcode
834:Bibcode
779:Bibcode
520:Macleay
510:(Lesne)
341:Burmese
310:Psoinae
267:Psoinae
172:Order:
166:Insecta
162:Class:
3259:uBio:
3226:633649
3210:NZOR:
3177:114476
3164:108553
3099:1BOSTF
3047:111683
1206:Extant
959:
929:
860:
852:
805:
797:
750:
540:Germar
330:larvae
223:, 1802
3290:This
3247:Plazi
3239:69386
3203:92690
3159:IRMNG
3151:47625
3112:11022
3073:622G8
3060:18100
978:Lucid
927:S2CID
880:CSIRO
858:S2CID
803:S2CID
651:larva
300:, or
3296:stub
3198:NCBI
3172:ITIS
3138:7773
3133:GBIF
3094:EPPO
3086:7477
3042:BOLD
3034:9767
1029:IFAS
1012:IFAS
995:IFAS
957:ISBN
850:ISSN
795:ISSN
748:ISSN
339:and
288:The
35:PreꞒ
3185:NBN
3081:EoL
3068:CoL
3019:AFD
3006:ADW
919:doi
842:doi
830:116
787:doi
738:hdl
728:doi
3341::
3249::
3236::
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3008::
2993::
2978::
1027:/
1025:UF
1010:/
1008:UF
993:/
991:UF
925:.
917:.
907:25
905:.
878:.
856:.
848:.
840:.
828:.
824:.
801:.
793:.
785:.
775:92
773:.
769:.
746:.
736:.
724:17
722:.
718:.
695:.
498:-
312:.
296:,
85:Pg
3327:e
3320:t
3313:v
3302:.
2673:)
2669:(
2622:)
2618:(
2592:)
2588:(
1885:)
1881:(
1803:)
1799:(
1626:)
1622:(
1609:)
1605:(
1550:)
1546:(
1279:)
1275:(
1163:)
1159:(
1143:)
1139:(
1058:e
1051:t
1044:v
965:.
933:.
921::
913::
864:.
844::
836::
809:.
789::
781::
754:.
740::
730::
661:†
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:Ꞓ
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