127:
151:
2182:
1745:
1751:
1757:
108:
2802:
360:. However, the only confidently identified Late Miocene borhyaenid specimen, Stylocynus paranensis, comes from a site which is known to have Early Miocene fossils mixed in with Late Miocene ones, and so it may be that this group did not survive the end of the Early Miocene.
280:, Sparassodonta. Like most metatherians, borhyaenids and other sparassodonts are thought to have had a pouch to carry their offspring around. Borhyaenids had strong and powerful
2735:
310:. However, in recent years, with the elevation of most sparassodont subfamilies to family rank and the discovery that borhyaenids are more closely related to
2843:
2722:
306:
Originally, the borhyaenids were considered one of the most diverse groups of sparassodonts, including all species not originally included in the
2877:
611:
494:
352:
borhyaenoids were once considered to be borhyaenids, all other unambiguous members of the group are now considered to be restricted to the
558:
2882:
2867:
356:. The fossil record of this group after the Early Miocene is poor, and only fragmentary remains attest to their presence in the
2836:
2872:
150:
2829:
2181:
584:
604:
563:
Ameghino and its implications for the knowledge of the Mio-Pliocene large carnivorous mammals of South
America"
1744:
1756:
126:
2862:
2662:
490:(Mammalia, Metatheria, Sparassodonta) and phylogeny of Cenozoic metatherian carnivores from South America"
2621:
2337:
2774:
2761:
2283:
1750:
349:
1535:
1005:
2527:
2346:
2027:
885:
2709:
2269:
2241:
2234:
1246:
1076:
404:
2612:
2496:
2448:
2410:
2312:
2290:
2020:
1964:
1926:
1854:
1836:
1549:
1419:
1412:
1357:
1343:
1135:
1066:
2403:
2382:
2373:
2248:
1919:
1884:
1877:
1521:
1336:
1308:
1301:
1259:
1145:
864:
844:
716:
597:
244:
223:
2587:
2464:
2437:
2421:
2255:
1870:
1808:
1614:
1592:
1556:
1327:
1175:
1056:
2809:
2628:
2580:
2545:
2512:
2471:
2326:
2319:
2054:
2006:
1635:
1577:
1542:
1528:
1433:
1371:
854:
689:
374:
145:
2766:
2601:
2594:
2573:
2566:
2478:
2457:
2276:
2262:
2075:
1971:
1940:
1843:
1822:
1801:
1651:
1491:
1484:
1350:
1286:
1232:
458:
2779:
2700:
2485:
2297:
2226:
1933:
1905:
1584:
1315:
1106:
520:(Mammalia, Metatheria, Sparassodonta), a large predator from the Late Oligocene of Patagonia".
2748:
2559:
2212:
2199:
2047:
1912:
1898:
1665:
1606:
1364:
1126:
1117:
834:
743:
396:
2813:
2753:
2740:
2305:
1829:
1708:
1514:
1505:
1039:
945:
537:
529:
261:
134:
1863:
2128:
2087:
1717:
1644:
1447:
1096:
315:
311:
307:
2552:
2355:
2096:
2061:
1563:
1470:
1426:
1378:
969:
962:
954:
420:
318:
than other sparassodonts, the family has been reduced to seven species in four genera.
2856:
2536:
2158:
2137:
2110:
2013:
1951:
1787:
1774:
1461:
1270:
1216:
1193:
1086:
997:
920:
874:
322:
295:
269:
205:
65:
2151:
2117:
1999:
1815:
1694:
1403:
976:
933:
895:
357:
277:
34:
107:
533:
2694:
2389:
1985:
1891:
1599:
1570:
1279:
1225:
1165:
1019:
983:
905:
567:
285:
40:
2505:
2396:
2144:
1621:
1440:
1385:
1155:
813:
800:
755:
620:
258:
232:
85:
50:
2685:
303:. Borhyaenids grew up to an average of 5 to 6 feet (1.5 to 1.8 m) long.
2430:
2362:
2103:
2068:
1978:
1685:
1294:
990:
673:
436:
341:
333:
289:
273:
239:
162:
90:
2679:
1992:
1477:
1012:
701:
644:
345:
182:
80:
75:
60:
55:
45:
2727:
1701:
557:
Forasiepi, Analía M.; Agustin G. Martinelli; Francisco J. Goin (2007).
542:
353:
95:
70:
728:
656:
650:
638:
329:
265:
192:
172:
30:
2801:
2656:
485:
118:
516:
Analía M. Forasiepi, M. Judith Babot and
Natalia Zimicz (2014). "
2714:
325:
300:
2660:
2197:
1772:
798:
631:
593:
281:
589:
2817:
495:
Monografías del Museo
Argentino de Ciencias Naturales
1662:
1632:
1502:
1458:
1400:
1243:
1213:
1190:
1171:
1161:
1151:
1141:
1131:
1123:
1113:
1102:
1092:
1082:
1072:
1062:
1052:
917:
901:
891:
881:
870:
860:
850:
840:
830:
2669:
2611:
2535:
2526:
2495:
2447:
2420:
2372:
2345:
2336:
2210:
2127:
2086:
2037:
1950:
1853:
1785:
1684:
1661:
1631:
1501:
1457:
1399:
1326:
1269:
1242:
1212:
1189:
1037:
944:
916:
811:
754:
727:
700:
671:
511:
509:
2837:
605:
8:
479:
477:
2844:
2830:
2657:
2532:
2342:
2218:
2207:
2194:
1956:
1793:
1782:
1769:
1251:
1186:
1045:
925:
823:
819:
808:
795:
733:
706:
679:
668:
628:
612:
598:
590:
125:
106:
20:
541:
473:
585:Borhyaenidae - Encyclopædia Britannica
264:of low-slung, heavily built predatory
321:The most studied borhyaenids are the
7:
2798:
2796:
522:Journal of Systematic Palaeontology
2816:. You can help Knowledge (XXG) by
332:sites in the southernmost part of
14:
2800:
2180:
1755:
1749:
1743:
149:
284:, like those of the unrelated
1:
484:Forasiepi, Analía M. (2009).
534:10.1080/14772019.2014.926403
2878:Prehistoric mammal families
348:); although some Oligocene
272:. Borhyaenids are not true
2899:
2795:
571:(in Spanish and English).
2221:
2206:
2193:
2178:
1959:
1796:
1781:
1768:
1741:
1254:
1185:
1048:
928:
826:
822:
807:
794:
736:
709:
682:
667:
627:
238:
231:
146:Scientific classification
144:
133:
124:
114:
105:
23:
16:Extinct family of mammals
2883:Prehistoric mammal stubs
2868:Eocene first appearances
561:Parahyaenodon argentinus
559:"Taxonomic revision of
2812:-related article is a
518:Australohyaena antiqua
428:Australohyaena antiqua
338:Australohyaena antiqua
2775:Paleobiology Database
2762:Paleobiology Database
488:Arctodictis sinclairi
388:Arctodictis sinclairi
369:†Family Borhyaenidae
450:Borhyaena macrodonta
340:, is known from the
328:, particularly from
2873:Miocene extinctions
2622:Roberthoffstetteria
2338:Bonapartheriiformes
276:, but members of a
2810:prehistoric mammal
2284:Procaroloameghinia
1636:Anatoliadelphyidae
1578:Maastrichtidelphys
444:Borhyaena tuberata
412:Acrocyon sectorius
382:Arctodictis munizi
139:Borhyaena tuberata
116:Borhyaena tuberata
2825:
2824:
2790:
2789:
2749:Open Tree of Life
2663:Taxon identifiers
2654:
2653:
2650:
2649:
2646:
2645:
2642:
2641:
2638:
2637:
2522:
2521:
2213:Polydolopimorphia
2200:Polydolopimorphia
2189:
2188:
2176:
2175:
2172:
2171:
2168:
2167:
1764:
1763:
1739:
1738:
1735:
1734:
1731:
1730:
1727:
1726:
1666:Polydolopimorphia
1536:Ectocentrocristus
1395:
1394:
1127:Gurlin Tsav skull
1033:
1032:
1029:
1028:
1006:Prodeltitheridium
790:
789:
786:
785:
782:
781:
778:
777:
774:
773:
252:
251:
227:
2890:
2846:
2839:
2832:
2804:
2797:
2783:
2782:
2770:
2769:
2757:
2756:
2744:
2743:
2731:
2730:
2718:
2717:
2705:
2704:
2703:
2690:
2689:
2688:
2658:
2533:
2528:Polydolopiformes
2347:Bonapartheriidae
2343:
2219:
2208:
2195:
2184:
2028:Pseudothylacynus
1957:
1794:
1783:
1770:
1759:
1753:
1747:
1709:Paucituberculata
1664:
1634:
1515:Amphiperatherium
1506:Herpetotheriidae
1504:
1460:
1402:
1252:
1245:
1215:
1192:
1187:
1173:
1163:
1153:
1143:
1133:
1125:
1115:
1104:
1094:
1084:
1074:
1064:
1054:
1046:
1040:Marsupialiformes
946:Deltatheridiidae
926:
919:
903:
893:
886:Kasserinotherium
883:
872:
862:
852:
842:
832:
824:
820:
809:
796:
734:
707:
680:
669:
662:
661:
629:
614:
607:
600:
591:
577:
576:
554:
548:
547:
545:
513:
504:
503:
481:
222:
217:
204:
154:
153:
135:Life restoration
129:
110:
100:
37:
27:Temporal range:
21:
2898:
2897:
2893:
2892:
2891:
2889:
2888:
2887:
2853:
2852:
2851:
2850:
2793:
2791:
2786:
2778:
2773:
2765:
2760:
2752:
2747:
2739:
2734:
2726:
2721:
2713:
2708:
2699:
2698:
2693:
2684:
2683:
2678:
2665:
2655:
2634:
2607:
2518:
2491:
2443:
2416:
2368:
2332:
2270:Hatcheritherium
2242:Caroloameghinia
2235:Bobbschaefferia
2215:
2202:
2185:
2164:
2129:Thylacosmilidae
2123:
2088:Proborhyaenidae
2082:
2033:
1946:
1849:
1790:
1777:
1760:
1723:
1718:Australidelphia
1702:Didelphimorphia
1680:
1657:
1645:Anatoliadelphys
1627:
1497:
1453:
1448:Siamoperadectes
1391:
1322:
1265:
1247:Archimetatheria
1238:
1208:
1181:
1097:Arcantiodelphys
1077:Anchistodelphys
1042:
1025:
940:
912:
816:
803:
770:
750:
723:
696:
676:
663:
634:
633:
623:
618:
581:
580:
556:
555:
551:
528:(#6): 503–525.
515:
514:
507:
483:
482:
475:
470:
405:Acrocyon riggsi
366:
336:. One species,
308:Thylacosmilidae
299:, for crushing
248:
242:
221:
215:
202:
148:
101:
99:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
29:
28:
25:
17:
12:
11:
5:
2896:
2894:
2886:
2885:
2880:
2875:
2870:
2865:
2855:
2854:
2849:
2848:
2841:
2834:
2826:
2823:
2822:
2805:
2788:
2787:
2785:
2784:
2771:
2758:
2745:
2732:
2719:
2706:
2691:
2675:
2673:
2667:
2666:
2661:
2652:
2651:
2648:
2647:
2644:
2643:
2640:
2639:
2636:
2635:
2633:
2632:
2625:
2617:
2615:
2613:Sillustaniidae
2609:
2608:
2606:
2605:
2598:
2591:
2584:
2577:
2570:
2563:
2556:
2553:Antarctodolops
2549:
2541:
2539:
2530:
2524:
2523:
2520:
2519:
2517:
2516:
2509:
2501:
2499:
2497:Rosendolopidae
2493:
2492:
2490:
2489:
2482:
2475:
2468:
2461:
2453:
2451:
2449:Prepidolopidae
2445:
2444:
2442:
2441:
2434:
2426:
2424:
2418:
2417:
2415:
2414:
2411:Proargyrolagus
2407:
2400:
2393:
2386:
2378:
2376:
2370:
2369:
2367:
2366:
2359:
2356:Bonapartherium
2351:
2349:
2340:
2334:
2333:
2331:
2330:
2323:
2316:
2313:Robertbutleria
2309:
2302:
2294:
2291:Protodidelphis
2287:
2280:
2273:
2266:
2259:
2252:
2245:
2238:
2231:
2222:
2216:
2211:
2204:
2203:
2198:
2191:
2190:
2187:
2186:
2179:
2177:
2174:
2173:
2170:
2169:
2166:
2165:
2163:
2162:
2155:
2148:
2141:
2133:
2131:
2125:
2124:
2122:
2121:
2114:
2107:
2100:
2097:Arminiheringia
2092:
2090:
2084:
2083:
2081:
2080:
2072:
2065:
2062:Australohyaena
2058:
2051:
2043:
2041:
2035:
2034:
2032:
2031:
2024:
2021:Pseudolycopsis
2017:
2010:
2003:
1996:
1989:
1982:
1975:
1968:
1965:Angelocabrerus
1960:
1954:
1948:
1947:
1945:
1944:
1937:
1930:
1927:Pseudonotictis
1923:
1916:
1909:
1902:
1895:
1888:
1881:
1874:
1867:
1859:
1857:
1855:Hathliacynidae
1851:
1850:
1848:
1847:
1840:
1837:Procladosictis
1833:
1826:
1819:
1812:
1805:
1797:
1791:
1786:
1779:
1778:
1773:
1766:
1765:
1762:
1761:
1742:
1740:
1737:
1736:
1733:
1732:
1729:
1728:
1725:
1724:
1722:
1721:
1714:
1713:
1712:
1705:
1690:
1688:
1682:
1681:
1679:
1678:
1677:
1676:
1670:
1668:
1659:
1658:
1656:
1655:
1648:
1640:
1638:
1629:
1628:
1626:
1625:
1618:
1611:
1603:
1596:
1589:
1581:
1574:
1567:
1564:Herpetotherium
1560:
1553:
1550:Galatiadelphys
1546:
1539:
1532:
1525:
1518:
1510:
1508:
1499:
1498:
1496:
1495:
1488:
1481:
1474:
1471:Albertatherium
1466:
1464:
1455:
1454:
1452:
1451:
1444:
1437:
1430:
1427:Mimoperadectes
1423:
1420:Didelphidectes
1416:
1413:Armintodelphys
1408:
1406:
1397:
1396:
1393:
1392:
1390:
1389:
1382:
1379:Scalaridelphys
1375:
1368:
1361:
1358:Monodelphopsis
1354:
1347:
1344:Austropediomys
1340:
1332:
1330:
1324:
1323:
1321:
1320:
1312:
1305:
1298:
1291:
1283:
1275:
1273:
1267:
1266:
1264:
1263:
1255:
1249:
1240:
1239:
1237:
1236:
1229:
1221:
1219:
1210:
1209:
1207:
1206:
1205:
1204:
1198:
1196:
1183:
1182:
1180:
1179:
1169:
1159:
1149:
1139:
1136:Iugomortiferum
1129:
1121:
1111:
1100:
1090:
1080:
1070:
1067:Aenigmadelphys
1060:
1049:
1043:
1038:
1035:
1034:
1031:
1030:
1027:
1026:
1024:
1023:
1016:
1009:
1002:
994:
987:
980:
973:
970:Deltatheroides
966:
963:Deltatheridium
959:
955:Atokatheridium
950:
948:
942:
941:
939:
938:
929:
923:
914:
913:
911:
910:
899:
889:
879:
868:
858:
848:
838:
827:
817:
812:
805:
804:
799:
792:
791:
788:
787:
784:
783:
780:
779:
776:
775:
772:
771:
769:
768:
767:
766:
760:
758:
752:
751:
749:
748:
747:
746:
737:
731:
725:
724:
722:
721:
720:
719:
710:
704:
698:
697:
695:
694:
693:
692:
683:
677:
672:
665:
664:
660:
659:
653:
647:
641:
632:
625:
624:
619:
617:
616:
609:
602:
594:
588:
587:
579:
578:
549:
505:
486:"Osteology of
472:
471:
469:
466:
465:
464:
463:
462:
454:
453:
452:
446:
432:
431:
430:
421:Australohyaena
416:
415:
414:
408:
392:
391:
390:
384:
365:
364:Classification
362:
316:thylacosmilids
312:proborhyaenids
257:is an extinct
250:
249:
243:
236:
235:
229:
228:
213:
209:
208:
200:
196:
195:
190:
186:
185:
180:
176:
175:
170:
166:
165:
160:
156:
155:
142:
141:
131:
130:
122:
121:
112:
111:
103:
102:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
38:
26:
15:
13:
10:
9:
6:
4:
3:
2:
2895:
2884:
2881:
2879:
2876:
2874:
2871:
2869:
2866:
2864:
2863:Sparassodonts
2861:
2860:
2858:
2847:
2842:
2840:
2835:
2833:
2828:
2827:
2821:
2819:
2815:
2811:
2806:
2803:
2799:
2794:
2781:
2776:
2772:
2768:
2763:
2759:
2755:
2750:
2746:
2742:
2737:
2733:
2729:
2724:
2720:
2716:
2711:
2707:
2702:
2696:
2692:
2687:
2681:
2677:
2676:
2674:
2672:
2668:
2664:
2659:
2631:
2630:
2626:
2624:
2623:
2619:
2618:
2616:
2614:
2610:
2604:
2603:
2599:
2597:
2596:
2592:
2590:
2589:
2585:
2583:
2582:
2578:
2576:
2575:
2571:
2569:
2568:
2564:
2562:
2561:
2557:
2555:
2554:
2550:
2548:
2547:
2543:
2542:
2540:
2538:
2537:Polydolopidae
2534:
2531:
2529:
2525:
2515:
2514:
2510:
2508:
2507:
2503:
2502:
2500:
2498:
2494:
2488:
2487:
2483:
2481:
2480:
2476:
2474:
2473:
2469:
2467:
2466:
2462:
2460:
2459:
2455:
2454:
2452:
2450:
2446:
2440:
2439:
2435:
2433:
2432:
2428:
2427:
2425:
2423:
2419:
2413:
2412:
2408:
2406:
2405:
2404:Microtragulus
2401:
2399:
2398:
2394:
2392:
2391:
2387:
2385:
2384:
2383:Anargyrolagus
2380:
2379:
2377:
2375:
2374:Argyrolagidae
2371:
2365:
2364:
2360:
2358:
2357:
2353:
2352:
2350:
2348:
2344:
2341:
2339:
2335:
2329:
2328:
2324:
2322:
2321:
2317:
2315:
2314:
2310:
2308:
2307:
2303:
2300:
2299:
2295:
2293:
2292:
2288:
2286:
2285:
2281:
2279:
2278:
2274:
2272:
2271:
2267:
2265:
2264:
2260:
2258:
2257:
2253:
2251:
2250:
2249:Carolocoutoia
2246:
2244:
2243:
2239:
2237:
2236:
2232:
2229:
2228:
2224:
2223:
2220:
2217:
2214:
2209:
2205:
2201:
2196:
2192:
2183:
2161:
2160:
2159:Thylacosmilus
2156:
2154:
2153:
2149:
2147:
2146:
2142:
2140:
2139:
2138:Anachlysictis
2135:
2134:
2132:
2130:
2126:
2120:
2119:
2115:
2113:
2112:
2111:Paraborhyaena
2108:
2106:
2105:
2101:
2099:
2098:
2094:
2093:
2091:
2089:
2085:
2078:
2077:
2073:
2071:
2070:
2066:
2064:
2063:
2059:
2057:
2056:
2052:
2050:
2049:
2045:
2044:
2042:
2040:
2036:
2030:
2029:
2025:
2023:
2022:
2018:
2016:
2015:
2014:Prothylacinus
2011:
2009:
2008:
2004:
2002:
2001:
1997:
1995:
1994:
1990:
1988:
1987:
1983:
1981:
1980:
1976:
1974:
1973:
1969:
1967:
1966:
1962:
1961:
1958:
1955:
1953:
1952:Borhyaenoidea
1949:
1943:
1942:
1938:
1936:
1935:
1931:
1929:
1928:
1924:
1922:
1921:
1920:Perathereutes
1917:
1915:
1914:
1910:
1908:
1907:
1903:
1901:
1900:
1896:
1894:
1893:
1889:
1887:
1886:
1885:Chasicostylus
1882:
1880:
1879:
1878:Borhyaenidium
1875:
1873:
1872:
1868:
1866:
1865:
1861:
1860:
1858:
1856:
1852:
1846:
1845:
1841:
1839:
1838:
1834:
1832:
1831:
1827:
1825:
1824:
1820:
1818:
1817:
1813:
1811:
1810:
1806:
1804:
1803:
1799:
1798:
1795:
1792:
1789:
1788:Sparassodonta
1784:
1780:
1776:
1775:Sparassodonta
1771:
1767:
1758:
1754:
1752:
1746:
1720:
1719:
1715:
1711:
1710:
1706:
1704:
1703:
1699:
1698:
1697:
1696:
1692:
1691:
1689:
1687:
1683:
1674:
1673:
1672:
1671:
1669:
1667:
1660:
1654:
1653:
1649:
1647:
1646:
1642:
1641:
1639:
1637:
1630:
1624:
1623:
1619:
1617:
1616:
1612:
1609:
1608:
1604:
1602:
1601:
1597:
1595:
1594:
1590:
1587:
1586:
1582:
1580:
1579:
1575:
1573:
1572:
1568:
1566:
1565:
1561:
1559:
1558:
1554:
1552:
1551:
1547:
1545:
1544:
1540:
1538:
1537:
1533:
1531:
1530:
1526:
1524:
1523:
1522:Asiadidelphis
1519:
1517:
1516:
1512:
1511:
1509:
1507:
1500:
1494:
1493:
1489:
1487:
1486:
1482:
1480:
1479:
1475:
1473:
1472:
1468:
1467:
1465:
1463:
1462:Alphadontidae
1456:
1450:
1449:
1445:
1443:
1442:
1438:
1436:
1435:
1431:
1429:
1428:
1424:
1422:
1421:
1417:
1415:
1414:
1410:
1409:
1407:
1405:
1398:
1388:
1387:
1383:
1381:
1380:
1376:
1374:
1373:
1369:
1367:
1366:
1362:
1360:
1359:
1355:
1353:
1352:
1348:
1346:
1345:
1341:
1339:
1338:
1337:Aquiladelphis
1334:
1333:
1331:
1329:
1325:
1318:
1317:
1313:
1311:
1310:
1309:Hoodootherium
1306:
1304:
1303:
1302:Fumodelphodon
1299:
1297:
1296:
1292:
1289:
1288:
1284:
1282:
1281:
1277:
1276:
1274:
1272:
1271:Stagodontidae
1268:
1262:
1261:
1260:Iqualadelphis
1257:
1256:
1253:
1250:
1248:
1241:
1235:
1234:
1230:
1228:
1227:
1223:
1222:
1220:
1218:
1217:Asiatheriidae
1211:
1202:
1201:
1200:
1199:
1197:
1195:
1194:Sparassodonta
1188:
1184:
1178:
1177:
1170:
1168:
1167:
1160:
1158:
1157:
1150:
1148:
1147:
1146:Jaskhadelphys
1140:
1138:
1137:
1130:
1128:
1122:
1120:
1119:
1112:
1109:
1108:
1101:
1099:
1098:
1091:
1089:
1088:
1087:Andinodelphys
1081:
1079:
1078:
1071:
1069:
1068:
1061:
1059:
1058:
1051:
1050:
1047:
1044:
1041:
1036:
1022:
1021:
1017:
1015:
1014:
1010:
1008:
1007:
1003:
1000:
999:
998:Oklatheridium
995:
993:
992:
988:
986:
985:
981:
979:
978:
974:
972:
971:
967:
965:
964:
960:
957:
956:
952:
951:
949:
947:
943:
936:
935:
931:
930:
927:
924:
922:
921:Deltatheroida
915:
908:
907:
900:
898:
897:
890:
888:
887:
880:
877:
876:
875:Holoclemensia
869:
867:
866:
865:Ghamidtherium
859:
857:
856:
849:
847:
846:
845:Archaeonothos
839:
837:
836:
829:
828:
825:
821:
818:
815:
810:
806:
802:
797:
793:
764:
763:
762:
761:
759:
757:
753:
745:
741:
740:
739:
738:
735:
732:
730:
726:
718:
714:
713:
712:
711:
708:
705:
703:
699:
691:
687:
686:
685:
684:
681:
678:
675:
670:
666:
658:
654:
652:
648:
646:
642:
640:
636:
635:
630:
626:
622:
615:
610:
608:
603:
601:
596:
595:
592:
586:
583:
582:
574:
570:
569:
564:
562:
553:
550:
544:
539:
535:
531:
527:
523:
519:
512:
510:
506:
501:
497:
496:
491:
489:
480:
478:
474:
467:
461:
460:
456:†Genus ?
455:
451:
447:
445:
441:
440:
439:
438:
433:
429:
425:
424:
423:
422:
417:
413:
409:
407:
406:
401:
400:
399:
398:
393:
389:
385:
383:
379:
378:
377:
376:
371:
370:
368:
367:
363:
361:
359:
355:
351:
347:
343:
339:
335:
331:
327:
324:
323:Early Miocene
319:
317:
313:
309:
304:
302:
298:
297:
296:Andrewsarchus
292:
291:
287:
286:placentalians
283:
279:
275:
271:
270:Sparassodonta
268:in the order
267:
263:
260:
256:
246:
241:
237:
234:
230:
225:
220:
214:
211:
210:
207:
206:Sparassodonta
201:
198:
197:
194:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
164:
161:
158:
157:
152:
147:
143:
140:
136:
132:
128:
123:
120:
117:
113:
109:
104:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
36:
32:
22:
19:
2818:expanding it
2807:
2792:
2701:Borhyaenidae
2671:Borhyaenidae
2670:
2627:
2620:
2600:
2593:
2588:Parabderites
2586:
2579:
2572:
2565:
2558:
2551:
2544:
2511:
2504:
2484:
2477:
2470:
2465:Perrodelphys
2463:
2456:
2438:Thylacotinga
2436:
2429:
2422:Chulpasiinae
2409:
2402:
2395:
2388:
2381:
2361:
2354:
2325:
2318:
2311:
2304:
2296:
2289:
2282:
2275:
2268:
2261:
2256:Guggenheimia
2254:
2247:
2240:
2233:
2225:
2157:
2152:Patagosmilus
2150:
2143:
2136:
2118:Proborhyaena
2116:
2109:
2102:
2095:
2074:
2067:
2060:
2053:
2046:
2039:Borhyaenidae
2038:
2026:
2019:
2012:
2005:
2000:Pharsophorus
1998:
1991:
1984:
1977:
1970:
1963:
1939:
1932:
1925:
1918:
1911:
1904:
1897:
1890:
1883:
1876:
1871:Australogale
1869:
1862:
1842:
1835:
1828:
1821:
1816:Hondadelphys
1814:
1809:Argyrolestes
1807:
1800:
1748:
1716:
1707:
1700:
1695:Ameridelphia
1693:
1650:
1643:
1620:
1615:Swaindelphys
1613:
1605:
1598:
1593:Nortedelphys
1591:
1583:
1576:
1569:
1562:
1557:Golerdelphys
1555:
1548:
1541:
1534:
1527:
1520:
1513:
1490:
1483:
1476:
1469:
1446:
1439:
1432:
1425:
1418:
1411:
1404:Peradectidae
1384:
1377:
1370:
1363:
1356:
1349:
1342:
1335:
1328:Pediomyoidea
1314:
1307:
1300:
1293:
1285:
1278:
1258:
1231:
1224:
1176:Sinbadelphys
1174:
1164:
1154:
1144:
1134:
1116:
1105:
1095:
1085:
1075:
1065:
1057:Adelodelphys
1055:
1018:
1011:
1004:
996:
989:
982:
977:Gurbanodelta
975:
968:
961:
953:
934:Pappotherium
932:
904:
896:Protalphadon
894:
884:
873:
863:
853:
843:
833:
572:
566:
560:
552:
525:
521:
517:
499:
493:
487:
457:
449:
443:
435:
427:
419:
411:
403:
395:
387:
381:
373:
358:Late Miocene
337:
320:
305:
294:
288:
278:sister taxon
255:Borhyaenidae
254:
253:
219:Borhyaenidae
218:
138:
115:
35:Late Miocene
24:Borhyaenidae
18:
2695:Wikispecies
2629:Sillustania
2581:Kramadolops
2546:Amphidolops
2513:Rosendolops
2472:Prepidolops
2390:Argyrolagus
2327:Zeusdelphys
2320:Wamradolops
2055:Arctodictis
2007:Plesiofelis
1986:Fredszalaya
1892:Cladosictis
1686:Marsupialia
1600:Peratherium
1571:Indodelphis
1543:Entomacodon
1529:Copedelphys
1434:Nanodelphys
1372:Protolambda
1280:Didelphodon
1226:Asiatherium
1166:Pucadelphys
1020:Tsagandelta
984:Lotheridium
906:Sinodelphys
855:Esteslestes
568:Ameghiniana
543:11336/59430
375:Arctodictis
259:metatherian
2857:Categories
2602:Pseudolops
2595:Polydolops
2574:Hypodolops
2567:Eurydolops
2506:Hondonadia
2479:Punadolops
2458:Incadolops
2397:Hondalagus
2277:Palangania
2263:Gashternia
2145:Eomakhaira
2076:Eutemnodus
1972:Chlorocyon
1941:Sipalocyon
1844:Stylocynus
1823:Nemolestes
1802:Allqokirus
1675:see below↓
1652:Orhaniyeia
1622:Thylacodon
1492:Turgidodon
1485:Eoalphadon
1441:Peradectes
1386:Unnuakomys
1351:Dakotadens
1287:Eobrasilia
1233:Kokopellia
1203:see below↓
1156:Mayulestes
814:Metatheria
801:Metatheria
765:see below↓
756:Metatheria
717:Cynodontia
702:Cynodontia
655:Subclass:
621:Metatheria
575:: 143–159.
468:References
459:Eutemnodus
274:marsupials
233:Type genus
2486:Seumandia
2431:Chulpasia
2363:Epidolops
2298:Pujatodon
2227:Apeirodon
2104:Callistoe
2069:Borhyaena
1979:Dukecynus
1934:Sallacyon
1906:Notocynus
1585:Morotodon
1316:Pariadens
1295:Eodelphis
1107:Camptomus
991:Nanocuris
690:Synapsida
674:Synapsida
637:Kingdom:
437:Borhyaena
342:Oligocene
334:Patagonia
290:Hyaenodon
240:Borhyaena
169:Kingdom:
163:Eukaryota
2680:Wikidata
2560:Eudolops
2048:Acrocyon
1993:Lycopsis
1913:Notogale
1899:Notictis
1607:Rumiodon
1478:Alphadon
1365:Pediomys
1118:Glasbius
1013:Sulestes
835:Adinodon
744:Mammalia
729:Mammalia
651:Mammalia
645:Chordata
643:Phylum:
639:Animalia
502:: 1–174.
397:Acrocyon
346:Deseadan
245:Ameghino
224:Ameghino
212:Family:
193:Mammalia
183:Chordata
179:Phylum:
173:Animalia
159:Domain:
2754:4943605
2728:3239401
2715:4471526
2686:Q139151
649:Class:
434:†Genus
418:†Genus
394:†Genus
372:†Genus
354:Miocene
266:mammals
199:Order:
189:Class:
2767:423457
2741:119011
2306:Reigia
1830:Patene
657:Theria
330:fossil
262:family
247:, 1887
226:, 1894
31:Danian
2808:This
2780:40010
2736:IRMNG
1864:Acyon
350:basal
301:bones
119:skull
2814:stub
2723:GBIF
742:see
715:see
688:see
326:taxa
314:and
293:and
282:jaws
41:PreꞒ
2710:EoL
538:hdl
530:doi
137:of
2859::
2777::
2764::
2751::
2738::
2725::
2712::
2697::
2682::
573:44
565:.
536:.
526:13
524:.
508:^
498:.
492:.
476:^
91:Pg
2845:e
2838:t
2831:v
2820:.
2301:?
2230:?
2079:?
1663:†
1633:†
1610:?
1588:?
1503:†
1459:†
1401:†
1319:?
1290:?
1244:†
1214:†
1191:†
1172:†
1162:†
1152:†
1142:†
1132:†
1124:†
1114:†
1110:?
1103:†
1093:†
1083:†
1073:†
1063:†
1053:†
1001:?
958:?
937:?
918:†
909:?
902:†
892:†
882:†
878:?
871:†
861:†
851:†
841:†
831:†
613:e
606:t
599:v
546:.
540::
532::
500:6
448:†
442:†
426:†
410:†
402:†
386:†
380:†
344:(
216:†
203:†
96:N
86:K
81:J
76:T
71:P
66:C
61:D
56:S
51:O
46:Ꞓ
33:–
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