64:
1280:
1296:
1264:
277:
1508:
Köhler F., Holford, Do V. T. & Ho T. H. (2009). "Exploring a largely unknown fauna: On the diversity of pachychilid freshwater gastropods in
Vietnam (Caenogastropoda: Cerithioidea)".
1572:
Köhler F. & Deein G. (2010). "Hybridisation as potential source of incongruence in the morphological and mitochondrial diversity of a Thai freshwater gastropod (Pachychilidae,
1714:
1325:
1727:
1688:
344:
1231:
1203:
1217:
336:
1189:
1245:
340:
316:
41:
1766:
1349:
195:
1578:
1545:
1479:
1649:
1804:
1732:
1794:
1539:
Köhler F. & Glaubrecht M. (2001). "Toward a systematic revision of the
Southeast Asian freshwater gastropod
63:
1295:
1279:
371:
Species are found predominantly in fast flowing, well oxygenated rivers, occasionally also in lakes. They are
1150:
1102:
1085:
831:
1136:
1054:
556:
505:
152:
1001:
821:
1602:
771:
1444:
Köhler F. & Glaubrecht M. (2006). "A systematic revision of the
Southeast Asian freshwater gastropod
1263:
1119:
811:
791:
781:
739:
698:
676:
666:
606:
525:
1753:
1543:
H. – Adams, 1866 (Cerithioidea: Pachychilidae): an account of species from around the South China Sea".
1068:
1019:
933:
883:
626:
1654:
984:
861:
801:
709:
687:
586:
493:
439:
418:
1333:
851:
841:
646:
545:
1771:
1675:
1458:
1037:
903:
719:
656:
636:
472:
408:
923:
872:
576:
483:
943:
749:
729:
616:
398:
1510:
1345:
1235:
1207:
893:
566:
270:
250:
191:
179:
58:
1758:
1740:
1719:
1662:
1745:
1586:
1553:
1487:
450:
253:
239:
1799:
1302:
1286:
1249:
1230:
1221:
1202:
967:
913:
759:
596:
461:
115:
50:
1518:
1216:
1193:
1167:
535:
515:
308:
304:
236:
1244:
1788:
1270:
429:
372:
256:
135:
1188:
1667:
1640:
276:
267:
161:
125:
1701:
1634:
1450:
1557:
376:
324:
297:
105:
40:
1625:
1323:
Adams H. (1866). "Description of a new genus and a new species of mollusks".
352:
243:
75:
1590:
1491:
1619:
348:
328:
95:
1693:
1352:(1892). "Mission scientifique au Mexique et dans l'Amérique Centrale".
356:
320:
312:
300:
246:
30:
1706:
360:
85:
1596:
1243:
1229:
1215:
1201:
1187:
275:
1680:
332:
1600:
1473:
Köhler F., Du L.-N. & Yang J.-X. (2010). "A new species of
379:, retaining developing eggs and youngs in special brood pouch.
1477:
from Yunnan, China (Caenogastropoda, Pachychilidae)".
1609:
1181:(images in the first row are in the same ratio):
1326:Proceedings of the Zoological Society of London
1440:
1438:
1436:
1434:
1432:
1430:
1428:
1426:
1424:
1422:
1420:
1418:
1416:
1414:
1412:
1410:
1408:
1406:
1404:
1402:
1400:
1398:
1396:
1394:
1392:
1390:
1388:
1386:
1535:
1533:
1531:
1529:
1527:
1504:
1502:
1500:
1469:
1467:
1384:
1382:
1380:
1378:
1376:
1374:
1372:
1370:
1368:
1366:
8:
1597:
39:
20:
1316:
1259:
776:Gurung, Takayasu & Matsuoka, 1997
413:Köhler, Holford M., Do & Ho, 2009
7:
1107:Köhler, Holford, Do & Ho, 2009
611:Köhler, Holford, Do & Ho, 2009
266:is apparently in honor of a Swiss
14:
315:in the east. This range includes
1448:(Cerithioidea: Pachychilidae)".
1294:
1278:
1262:
62:
1098:(Bavay & Dautzenberg, 1908)
591:(Griffith & Pidgeon, 1834)
1:
816:Köhler & Glaubrecht, 2002
1579:Zoosystematics and Evolution
1546:Journal of Molluscan Studies
1480:Zoosystematics and Evolution
938:Köhler, Du & Yang, 2010
571:(H. Lea & I. Lea, 1851)
1821:
1177:Variability of shells of
1012:Brotia escheri aquitanica
387:Species within the genus
185:
178:
167:
160:
59:Scientific classification
57:
47:
38:
23:
1558:10.1093/mollus/67.3.281
1151:Sulcospira tourannensis
1137:Sulcospira testudinaria
1103:Sulcospira dakrongensis
1086:Sulcospira infracostata
1030:Brotia escheri auingeri
832:Brotia pseudosulcospira
311:in the west through to
1591:10.1002/zoos.201000013
1492:10.1002/zoos.201000012
1354:Recherches zoologiques
1253:
1239:
1225:
1211:
1197:
1055:Sulcospira hainanensis
557:Brotia dautzenbergiana
506:Brotia castellaunensis
284:
1754:Paleobiology Database
1247:
1233:
1219:
1205:
1191:
1141:(von dem Busch, 1842)
1132:(von dem Busch, 1842)
1069:Sulcospira tonkiniana
1002:Sulcospira delavayana
908:(von dem Busch, 1842)
877:(Godwin-Austen, 1872)
822:Brotia pseudoasperata
403:(F. Sandberger, 1871)
279:
1066: : synonym of
999: : synonym of
772:Brotia palaeocostula
1576:H. Adarns, 1866)".
1146:Brotia tourannensis
1129:Brotia testudinaria
1120:Sulcospira swinhoei
1078:Brotia infracostata
812:Brotia praetermissa
792:Brotia peninsularis
782:Brotia paludiformis
740:Brotia microsculpta
699:Brotia lombersensis
677:Brotia laodelectata
667:Brotia kelantanesis
631:(von Martens, 1900)
607:Brotia hoabinhensis
526:Brotia clavaeformis
16:Genus of gastropods
1511:Molluscan Research
1254:
1240:
1236:Brotia sumatrensis
1226:
1212:
1208:Brotia episcopalis
1198:
1172:(Rafinesque, 1833)
1064:Brotia hamonvillei
1047:Brotia hainanensis
1020:Tinnyea aquitanica
934:Brotia yunnanensis
894:Brotia sumatrensis
884:Brotia subgloriosa
627:Brotia indragirica
567:Brotia episcopalis
540:(Rafinesque, 1833)
285:
271:Auguste Louis Brot
1805:Freshwater snails
1782:
1781:
1741:Open Tree of Life
1603:Taxon identifiers
1258:
1257:
1173:
1164:
1160:Brotia variabilis
1156:
1142:
1133:
1125:
1116:
1108:
1099:
1091:
1082:
1074:
1060:
1051:
1043:
1034:
1025:
1016:
1007:
998:
994:Brotia delavayana
990:
985:Brotia henriettae
981:
973:
964:
949:
939:
929:
919:
909:
899:
889:
878:
867:
862:Brotia sooloensis
857:
847:
837:
827:
817:
807:
802:Brotia persculpta
797:
787:
777:
765:
755:
745:
735:
725:
715:
714:(Oppenheim, 1901)
710:Brotia majevitzae
704:
693:
688:Brotia lhazeensis
682:
672:
662:
652:
642:
632:
622:
612:
602:
592:
587:Brotia henriettae
582:
572:
562:
551:
541:
531:
521:
511:
499:
494:Brotia borneensis
489:
479:
467:
456:
445:
440:Brotia assamensis
435:
424:
419:Brotia angulifera
414:
404:
262:The generic name
240:freshwater snails
228:
227:
222:
214:
206:
156:
1812:
1795:Gastropod genera
1775:
1774:
1762:
1761:
1749:
1748:
1736:
1735:
1723:
1722:
1710:
1709:
1697:
1696:
1684:
1683:
1671:
1670:
1658:
1657:
1645:
1644:
1643:
1630:
1629:
1628:
1598:
1560:
1537:
1522:
1506:
1495:
1471:
1462:
1442:
1361:
1343:
1337:
1321:
1298:
1282:
1266:
1184:
1183:
1171:
1162:
1155:(Souleyet, 1852)
1154:
1140:
1131:
1124:(H. Adams, 1870)
1123:
1115:(H. Adams, 1870)
1114:
1106:
1097:
1089:
1080:
1072:
1058:
1049:
1042:(Matheron, 1842)
1041:
1033:(Handmann, 1882)
1032:
1023:
1017:: synonym of †
1014:
1005:
996:
988:
979:
971:
962:
950:- incertae sedis
947:
937:
927:
917:
907:
897:
887:
879:- incertae sedis
876:
868:- incertae sedis
865:
855:
852:Brotia solemiana
845:
842:Brotia siamensis
835:
825:
815:
805:
795:
785:
775:
763:
753:
743:
733:
723:
713:
702:
691:
680:
670:
661:(Deshayes, 1874)
660:
651:(Blanford, 1869)
650:
647:Brotia iravadica
640:
630:
620:
610:
600:
590:
580:
570:
560:
552:- incertae sedis
549:
546:Brotia cylindrus
539:
529:
519:
509:
500:- incertae sedis
498:(Schepman, 1896)
497:
487:
478:Oppenheim, 1895)
477:
466:(Blanford, 1903)
465:
457:- incertae sedis
454:
451:Brotia beaumetzi
446:- incertae sedis
443:
433:
425:- incertae sedis
422:
412:
402:
280:Distribution of
220:
212:
210:Melania (Brotia)
204:
169:Melania pagodula
151:
67:
66:
43:
29:Temporal range:
21:
1820:
1819:
1815:
1814:
1813:
1811:
1810:
1809:
1785:
1784:
1783:
1778:
1770:
1765:
1757:
1752:
1744:
1739:
1731:
1726:
1718:
1713:
1705:
1700:
1692:
1687:
1679:
1674:
1666:
1661:
1653:
1648:
1639:
1638:
1633:
1624:
1623:
1618:
1605:
1569:
1564:
1563:
1538:
1525:
1507:
1498:
1472:
1465:
1443:
1364:
1344:
1340:
1322:
1318:
1313:
1306:
1303:Brotia herculea
1299:
1290:
1287:Brotia binodosa
1283:
1274:
1267:
1250:Brotia verbecki
1222:Brotia pagodula
1112:Brotia swinhoei
1090:(Mousson, 1848)
1081:(Mousson, 1848)
1073:(Morelet, 1886)
1038:Tinnyea lauraea
1035:: synonym of †
972:(Lamarck, 1822)
968:Jagora asperata
963:(Lamarck, 1822)
960:Brotia asperata
953:
914:Brotia verbecki
904:Brotia torquata
806:(Ehrmann, 1922)
760:Brotia pagodula
720:Brotia manningi
671:(Preston, 1907)
657:Brotia jullieni
637:Brotia insolita
597:Brotia herculea
510:(Boussac, 1911)
473:Brotia bittneri
462:Brotia binodosa
409:Brotia annamita
385:
369:
307:, ranging from
290:
237:Southeast Asian
202:Brotia (Brotia)
174:
171:
150:
116:Caenogastropoda
61:
51:Brotia pagodula
34:
27:
17:
12:
11:
5:
1818:
1816:
1808:
1807:
1802:
1797:
1787:
1786:
1780:
1779:
1777:
1776:
1763:
1750:
1737:
1724:
1711:
1698:
1685:
1672:
1659:
1646:
1631:
1615:
1613:
1607:
1606:
1601:
1595:
1594:
1585:(2): 301-314.
1568:
1567:External links
1565:
1562:
1561:
1552:(3): 281-318.
1523:
1517:(3): 121-146.
1496:
1486:(2): 295-300.
1463:
1362:
1360:(13): 117-392.
1338:
1315:
1314:
1312:
1309:
1308:
1307:
1300:
1293:
1291:
1284:
1277:
1275:
1268:
1261:
1256:
1255:
1241:
1227:
1213:
1199:
1194:Brotia citrina
1175:
1174:
1168:Brotia costula
1163:(Benson, 1836)
1157:
1143:
1126:
1117:: synonym of
1109:
1100:: synonym of
1095:Brotia proteus
1092:
1075:
1061:
1052:: synonym of
1044:
1026:
1024:(Noulet, 1846)
1015:(Noulet, 1846)
1008:
991:
977:Brotia baccata
974:
952:
951:
948:(Benson, 1836)
940:
930:
928:(Brandt, 1974)
924:Brotia wykoffi
920:
910:
900:
890:
888:(Brandt, 1968)
880:
873:Brotia spinata
869:
858:
856:(Brandt, 1968)
848:
838:
836:(Brandt, 1968)
828:
818:
808:
798:
796:(Brandt, 1974)
788:
778:
767:
766:- type species
756:
754:(Thiele, 1908)
746:
736:
726:
716:
705:
703:(Noulet, 1867)
694:
683:
673:
663:
653:
643:
633:
623:
613:
603:
593:
583:
577:Brotia godwini
573:
563:
561:(Morlet, 1884)
553:
542:
536:Brotia costula
532:
522:
516:Brotia citrina
512:
501:
490:
484:Brotia boenana
480:
468:
458:
447:
444:(Nevill, 1885)
436:
434:(Brandt, 1874)
426:
415:
405:
393:
384:
381:
368:
365:
309:Northern India
305:Southeast Asia
289:
286:
235:is a genus of
226:
225:
224:
223:
215:
207:
205:H. Adams, 1866
199:
183:
182:
176:
175:
172:
165:
164:
158:
157:
143:
139:
138:
133:
129:
128:
123:
119:
118:
113:
109:
108:
103:
99:
98:
93:
89:
88:
83:
79:
78:
73:
69:
68:
55:
54:
45:
44:
36:
35:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1817:
1806:
1803:
1801:
1798:
1796:
1793:
1792:
1790:
1773:
1768:
1764:
1760:
1755:
1751:
1747:
1742:
1738:
1734:
1729:
1725:
1721:
1716:
1712:
1708:
1703:
1699:
1695:
1690:
1686:
1682:
1677:
1673:
1669:
1664:
1660:
1656:
1651:
1647:
1642:
1636:
1632:
1627:
1621:
1617:
1616:
1614:
1612:
1608:
1604:
1599:
1592:
1588:
1584:
1581:
1580:
1575:
1571:
1570:
1566:
1559:
1555:
1551:
1548:
1547:
1542:
1536:
1534:
1532:
1530:
1528:
1524:
1520:
1516:
1513:
1512:
1505:
1503:
1501:
1497:
1493:
1489:
1485:
1482:
1481:
1476:
1470:
1468:
1464:
1460:
1456:
1453:
1452:
1447:
1441:
1439:
1437:
1435:
1433:
1431:
1429:
1427:
1425:
1423:
1421:
1419:
1417:
1415:
1413:
1411:
1409:
1407:
1405:
1403:
1401:
1399:
1397:
1395:
1393:
1391:
1389:
1387:
1385:
1383:
1381:
1379:
1377:
1375:
1373:
1371:
1369:
1367:
1363:
1359:
1355:
1351:
1347:
1342:
1339:
1335:
1331:
1328:
1327:
1320:
1317:
1310:
1305:
1304:
1297:
1292:
1289:
1288:
1281:
1276:
1273:
1272:
1271:Brotia armata
1265:
1260:
1252:
1251:
1246:
1242:
1238:
1237:
1232:
1228:
1224:
1223:
1218:
1214:
1210:
1209:
1204:
1200:
1196:
1195:
1190:
1186:
1185:
1182:
1180:
1170:
1169:
1165:: synonym of
1161:
1158:
1153:
1152:
1148:: synonym of
1147:
1144:
1139:
1138:
1134:: synonym of
1130:
1127:
1122:
1121:
1113:
1110:
1105:
1104:
1096:
1093:
1088:
1087:
1083:: synonym of
1079:
1076:
1071:
1070:
1065:
1062:
1057:
1056:
1048:
1045:
1040:
1039:
1031:
1027:
1022:
1021:
1013:
1009:
1006:(Heude, 1888)
1004:
1003:
995:
992:
987:
986:
982:: synonym of
980:(Gould, 1847)
978:
975:
970:
969:
965:: synonym of
961:
958:
957:
956:
946:
945:
944:Brotia zonata
941:
936:
935:
931:
926:
925:
921:
916:
915:
911:
906:
905:
901:
896:
895:
891:
886:
885:
881:
875:
874:
870:
866:(Reeve, 1859)
864:
863:
859:
854:
853:
849:
844:
843:
839:
834:
833:
829:
824:
823:
819:
814:
813:
809:
804:
803:
799:
794:
793:
789:
786:(Solem, 1966)
784:
783:
779:
774:
773:
768:
764:(Gould, 1847)
762:
761:
757:
752:
751:
750:Brotia pageli
747:
742:
741:
737:
732:
731:
730:Brotia mariae
727:
722:
721:
717:
712:
711:
706:
701:
700:
695:
690:
689:
684:
679:
678:
674:
669:
668:
664:
659:
658:
654:
649:
648:
644:
639:
638:
634:
629:
628:
624:
619:
618:
617:Brotia huberi
614:
609:
608:
604:
601:(Gould, 1846)
599:
598:
594:
589:
588:
584:
579:
578:
574:
569:
568:
564:
559:
558:
554:
548:
547:
543:
538:
537:
533:
528:
527:
523:
518:
517:
513:
508:
507:
502:
496:
495:
491:
486:
485:
481:
475:
474:
469:
464:
463:
459:
453:
452:
448:
442:
441:
437:
432:
431:
430:Brotia armata
427:
421:
420:
416:
411:
410:
406:
401:
400:
399:Brotia alpina
395:
394:
392:
390:
382:
380:
378:
374:
373:gonochoristic
366:
364:
362:
358:
354:
350:
346:
343:(1 species),
342:
338:
334:
330:
326:
322:
318:
314:
310:
306:
302:
299:
295:
287:
283:
278:
274:
273:(1821-1896).
272:
269:
265:
260:
258:
257:Pachychilidae
255:
252:
248:
245:
241:
238:
234:
233:
219:
216:
211:
208:
203:
200:
197:
193:
190:
187:
186:
184:
181:
177:
170:
166:
163:
159:
154:
149:
148:
144:
141:
140:
137:
136:Pachychilidae
134:
131:
130:
127:
124:
122:Superfamily:
121:
120:
117:
114:
111:
110:
107:
104:
101:
100:
97:
94:
91:
90:
87:
84:
81:
80:
77:
74:
71:
70:
65:
60:
56:
53:
52:
46:
42:
37:
32:
26:
22:
19:
1610:
1582:
1577:
1573:
1549:
1544:
1540:
1514:
1509:
1483:
1478:
1474:
1454:
1449:
1445:
1357:
1356:, Partie 7,
1353:
1341:
1329:
1324:
1319:
1301:
1285:
1269:
1248:
1234:
1220:
1206:
1192:
1178:
1176:
1166:
1159:
1149:
1145:
1135:
1128:
1118:
1111:
1101:
1094:
1084:
1077:
1067:
1063:
1059:(Brot, 1874)
1053:
1050:(Brot, 1872)
1046:
1036:
1029:
1018:
1011:
1000:
993:
989:(Gray, 1834)
983:
976:
966:
959:
954:
942:
932:
922:
918:(Brot, 1886)
912:
902:
898:(Brot, 1875)
892:
882:
871:
860:
850:
846:(Brot, 1886)
840:
830:
826:Brandt, 1968
820:
810:
800:
790:
780:
770:
758:
748:
744:Brandt, 1968
738:
734:Köhler, 2008
728:
724:Brandt, 1968
718:
708:
697:
686:
681:Köhler, 2008
675:
665:
655:
645:
641:(Brot, 1868)
635:
625:
615:
605:
595:
585:
581:(Brot, 1875)
575:
565:
555:
550:(Brot, 1886)
544:
534:
530:(Brot, 1874)
524:
520:(Brot, 1868)
514:
504:
492:
488:(Brot, 1881)
482:
471:
460:
455:(Brot, 1887)
449:
438:
428:
417:
407:
397:
388:
386:
370:
293:
291:
288:Distribution
281:
268:malacologist
263:
261:
231:
230:
229:
217:
209:
201:
188:
168:
162:Type species
146:
145:
126:Cerithioidea
49:
24:
18:
1702:iNaturalist
1635:Wikispecies
1451:Malacologia
997:Heude, 1888
621:Thach, 2021
292:Species of
213:Adams, 1866
189:Antimelania
173:Gould, 1847
1789:Categories
1346:Fischer P.
1311:References
955:Synonyms:
423:Brot, 1872
377:viviparous
325:Bangladesh
298:freshwater
221:Chen, 1943
192:P. Fischer
112:Subclass:
106:Gastropoda
1350:Crosse H.
391:include:
353:Indonesia
296:occur in
251:taxonomic
244:gastropod
82:Kingdom:
76:Eukaryota
48:Shell of
1620:Wikidata
1519:abstract
692:Yü, 1982
349:Malaysia
345:Cambodia
329:Thailand
301:habitats
249:in the
247:molluscs
180:Synonyms
153:H. Adams
132:Family:
96:Mollusca
92:Phylum:
86:Animalia
72:Domain:
33:- recent
1720:1272976
1694:4617180
1681:4873612
1626:Q990852
383:Species
367:Biology
363:only).
357:Sumatra
337:Vietnam
321:Myanmar
313:Sumatra
142:Genus:
102:Class:
31:Miocene
1800:Brotia
1772:719138
1759:104654
1746:929584
1733:218535
1641:Brotia
1611:Brotia
1574:Brotia
1541:Brotia
1475:Brotia
1446:Brotia
1348:&
1179:Brotia
389:Brotia
375:, and
361:Borneo
294:Brotia
282:Brotia
264:Brotia
254:family
232:Brotia
198:, 1892
196:Crosse
194:&
155:, 1866
147:Brotia
25:Brotia
1767:WoRMS
1715:IRMNG
1707:87102
1668:7NV63
1461:-251.
1336:-151.
341:China
317:India
218:Wanga
1728:NCBI
1689:GBIF
1655:6169
1650:BOLD
1330:1866
359:and
351:and
333:Laos
1676:EoL
1663:CoL
1587:doi
1554:doi
1488:doi
1459:159
1334:150
303:of
1791::
1769::
1756::
1743::
1730::
1717::
1704::
1691::
1678::
1665::
1652::
1637::
1622::
1583:86
1550:67
1526:^
1515:29
1499:^
1484:86
1466:^
1457::
1455:48
1365:^
1332::
1028:†
1010:†
769:†
707:†
696:†
685:†
503:†
470:†
396:†
347:,
339:,
335:,
331:,
327:,
323:,
319:,
259:.
242:,
1593:.
1589::
1556::
1521:.
1494:.
1490::
1358:2
476:(
355:(
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