1555:
1512:
1322:
1370:
1301:
1483:
1425:
731:
149:
1906:
1454:
722:. A midsized group like the suckers, they are rather similar to catfish in appearance and behaviour, feeding primarily off the substrate and equipped with barbels to help them locate food at night or in murky conditions. Fishes in the families Cobitidae, Balitoridae, Botiidae, and Gyrinocheilidae are called loaches, although the last do not seem to belong to the lineage of "true" loaches, but are related to the suckers.
967:
744:
758:
601:
115:
1701:. If this were the case, they would have spread to Asia through Africa or North America before the continents split up, for these are purely freshwater fishes. As the Characiformes began to diversify and spread, they may have outcompeted South American basal cypriniforms in Africa, where more advanced cypriniforms survive and coexist with characiforms.
1952:
had to be rescued into captivity from imminent extinction; new populations have apparently been established again successfully from captive stock. The
Balitoridae and Cobitidae, meanwhile, contain a very large number of species about which essentially nothing is known except how they look and where
1838:
in biological and biochemical sciences, being used in many kinds of experiments. During early development, the zebrafish has a nearly transparent body, so it is ideal for studying developmental biology. It is also used for the elucidation of biochemical signaling pathways. They are also good pets,
898:
of the
Cobitoidei, but it could be split off the Catostomidae and Gyrinocheilidae in a distinct superfamily; the Catostomoidea might be closer relatives of the carps and minnows than of the "true" loaches. While the Cyprinioidea seem more "primitive" than the loach-like forms, they were apparently
1848:
2485:
Slechtová, V.; Bohlen, J.; Tan, H. H. (2007). "Families of
Cobitoidea (Teleostei; Cypriniformes) as revealed from nuclear genetic data and the position of the mysterious genera Barbucca, Psilorhynchus, Serpenticobitis and Vaillantella".
960:
are rather close relatives and stand apart from
Cyprininae – no good consensus exists yet on how the main lineages are interrelated. A systematic list, from the most ancient to the most modern lineages, can thus be given as:
1939:
fame possibly only survives in captivity. Ironically, while pollution and other forms of overuse by humans have driven it from its native home, it is bred for the aquarium fish trade by the thousands. The
1650:). This is evidenced not only by physiological details, but also by their great distribution, which indicates they had the longest time to spread. The earliest that Cypriniformes might have diverged from
3003:
1776:
The
Cyprinidae in particular are important in a variety of ways. Many species are important food fish, particularly in Europe and Asia. Some are also important as aquarium fish, of which the
2194:"A new family of Cypriniformes (Teleostei, Ostariophysi) based on a redescription of † Jianghanichthys hubeiensis (Lei, 1977) from the Eocene Yangxi Formation of China"
3885:
2954:
3924:
4064:
2529:
Chen, W.-J.; Lheknim, V.; Mayden, R. L. (2009). "Molecular phylogeny of the
Cobitoidea (Teleostei: Cypriniformes) revisited: Position of enigmatic loach
2876:
1823:
have a negative impact on the environment. Carp in particular stir up the riverbed, reducing the clarity of the water, making plant growth difficult.
597:, also possess pharyngeal teeth, the cypriniformes' teeth grind against a chewing pad on the base of the skull, rather than an upper pharyngeal jaw.
3859:
3102:
3898:
2893:
2605:
2914:
2966:
2864:
2852:
2840:
2828:
2804:
498:
fishes. The order contains 11-12 families (with some authorities having listed as many as 23), over 400 genera, and more than 4,250 named
2989:
2947:
2313:
627:), which make up two-thirds of the order's diversity. This is one of the largest families of fish, and is widely distributed across
2587:
4059:
3996:
2630:
1890:
have been severely affected; a considerable number went entirely extinct after settlement by
Europeans. For example, in 1900 the
2899:
1784:
are perhaps the most celebrated. The other families are of less commercial importance. The
Catostomidae have some importance in
4029:
3075:
253:
4054:
3016:
2348:"Bigmouth Buffalo Ictiobus cyprinellus sets freshwater teleost record as improved age analysis reveals centenarian longevity"
3903:
1756:, causing a decline of the suckers. Cyprinids reached North America and Europe about the same time, and Africa in the early
3911:
3095:
1935:
148:
3807:
3794:
3046:(2018): Phylogenetic classification of extant genera of fishes of the order Cypriniformes (Teleostei: Ostariophysi).
2346:
Lackmann, Alec R.; Andrews, Allen H.; Butler, Malcolm G.; Bielak-Lackmann, Ewelina S.; Clark, Mark E. (23 May 2019).
3929:
1941:
1720:(55-35 Mya), catostomids and cyprinids spread throughout Asia; the earliest members of the cyprinid subfamilies
1554:
1511:
735:
502:; new species are regularly described, and new genera are recognized frequently. Cyprinids are most diverse in
4044:
4039:
3088:
2778:
2691:"Early Cenozoic Cyprinoids (Ostariophysi: Cypriniformes: Cyprinidae and Danionidae) from Sumatra, Indonesia"
1321:
749:
4024:
3734:
1993:
1922:
1910:
1729:
1693:, where the most diversity of this group is found today. The alternative hypothesis is that they began in
915:
4049:
4034:
3960:
2246:
1961:
904:
3640:
2267:
3978:
3812:
3665:
3620:
2967:
Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei).
1482:
1369:
4001:
3939:
3833:
3781:
2702:
2690:
2542:
2495:
2205:
2087:
2042:
2000:
1646:
Cypriniformes include the most primitive of the
Ostariophysi in the narrow sense (i.e. excluding the
930:
923:
586:
486:
3690:
3014:(2003): Mitochondrial Genomics of Ostariophysan Fishes: Perspectives on Phylogeny and Biogeography.
2193:
1764:, which formed and disappeared again several times during the many millions of years of cypriniform
1737:
1424:
710:
and eastern Asia. These large fishes are similar to carps in appearance and ecology. Members of the
3650:
3635:
3373:
2979:
1867:
891:
653:
3655:
1651:
1300:
825:
3772:
3590:
3515:
3469:
3320:
2295:
1983:
1977:
1745:
980:
895:
570:
319:
143:
2816:
730:
582:
3965:
3947:
3820:
3685:
3292:
3284:
3032:
2985:
2943:
2718:
2558:
2511:
2385:
2367:
2317:
2287:
2221:
2105:
1234:
881:
786:
343:
3952:
3988:
3890:
3695:
3670:
3555:
3520:
3325:
3269:
3225:
3196:
3057:
3025:
2923:
2906:
He, Shunping; Gub, Xun; Mayden, Richard L.; Chen, Wei-Jen; Conway, Kevin W. & Chen, Yiyu
2787:
2710:
2550:
2503:
2375:
2359:
2279:
2213:
1820:
1705:
1647:
1535:
1214:
1204:
1003:
911:
900:
869:
865:
837:
817:
762:
590:
519:
450:
394:
304:
300:
268:
3080:
2583:
Ichthyological
Exploration of Freshwaters. Bd. 20, Nr. 2, 2009, S. 157-162 (
585:, a small median bone in the snout, and the lack of teeth in the mouth. Instead, they have
3705:
3645:
3605:
3545:
3502:
3381:
2591:
2115:
1690:
1675:
1448:
1393:
1316:
1244:
1147:
1087:
1079:
1075:
1066:
972:
885:
873:
833:
821:
673:
442:
359:
347:
339:
335:
312:
242:
3012:
Saitoh, Kenji; Miya, Masaki; Inoue, Jun G.; Ishiguro, Naoya B. & Nishida, Mutsuminame
2776:(2005): The biogeography of otophysan fishes (Ostariophysi: Otophysi): a new appraisal.
2706:
2546:
2499:
2209:
1905:
1453:
3585:
3560:
3530:
3477:
3449:
3439:
3430:
3363:
3355:
3297:
3112:
2584:
2380:
2347:
2096:
2069:
2060:
2051:
2033:
2024:
1967:
1891:
1852:
1835:
1752:, around 30 Mya, advanced cyprinids began to outcompete catostomids wherever they were
1659:
1114:
507:
446:
363:
225:
190:
3786:
844:
and under his revision it now consists of the following families: hillstream loaches (
4018:
3799:
3610:
3600:
3570:
3550:
3525:
3340:
3335:
3143:
2886:
2791:
2554:
2242:
2192:
Liu, Juan; Chang, Mee-Mann; Wilson, Mark V. H.; Murray, Alison M. (2 November 2015).
2078:
1887:
1694:
1655:
1550:
1224:
1096:
1048:
949:
877:
802:
798:
790:
707:
662:
636:
546:
538:
526:
515:
351:
331:
323:
249:
210:
72:
2299:
966:
3675:
3630:
3615:
3595:
3580:
3510:
3454:
3444:
3419:
3414:
3315:
3149:
2268:"Phylogenetic Classification of Extant Genera of Fishes of the Order Cypriniformes"
2015:
2006:
1949:
1365:
1130:
907:
857:
806:
782:
778:
774:
743:
719:
700:
681:
644:
605:
566:
542:
490:
376:
284:
234:
2714:
2217:
3825:
739:
is a stone loach. Closely related to true loaches, like these, they have barbels.
3916:
3872:
3766:
3700:
3660:
3625:
3540:
3535:
3487:
3396:
3259:
3251:
3207:
3188:
3178:
2927:
2507:
1879:
1796:
1506:
1174:
1164:
1057:
957:
849:
845:
689:
669:
578:
466:
327:
272:
123:
47:
757:
600:
3680:
3492:
3391:
3386:
3307:
3264:
3230:
3137:
3061:
3029:
2363:
2283:
1899:
1816:
1804:
1741:
1663:
1405:
1295:
1039:
1030:
1012:
941:
937:
927:
841:
829:
771:
695:. While the former are a speciose group, the latter contain only a handful of
658:
648:
616:
574:
503:
485:(among others). Cypriniformes is an “order-within-an-order”, placed under the
308:
296:
292:
135:
131:
92:
57:
3757:
2722:
2371:
2225:
1666:). However, their divergence probably occurred only with the splitting-up of
3846:
3482:
3459:
3159:
1930:
1875:
1847:
1827:
1812:
1765:
1749:
1733:
1709:
1477:
1194:
987:
953:
861:
810:
711:
704:
511:
288:
160:
97:
3036:
2562:
2515:
2389:
2291:
17:
639:. Most species are strictly freshwater inhabitants, but some are found in
114:
3751:
3575:
3565:
3406:
3217:
3131:
2999:
2871:
2859:
2847:
2835:
2823:
2811:
2799:
2251:
1882:
for food or the pet trade have driven some Cypriniformes to the brink of
1789:
1777:
1761:
1753:
1725:
1721:
1698:
1683:
1679:
1671:
1419:
1184:
1105:
1023:
945:
853:
794:
685:
661:, swimming upstream into rivers to spawn. Sometimes separated as family
640:
530:
518:. At 112 years old, the longest-lived cypriniform fish documented is the
470:
355:
280:
276:
200:
180:
87:
82:
67:
62:
52:
952:
structures of this massively diverse family are known – e.g. that
918:
branched off. These diversified from the lowlands into torrential river
753:) is one of the sucking loaches, which are distant from other "loaches".
3345:
3241:
3048:
2912:(Ostariophysi: Cypriniformes): Evidence from the mitochondrial genome.
1883:
1871:
1860:
1826:
In science, one of the most famous members of the Cypriniformes is the
1785:
1760:(some 23-20 Mya). The cypriniforms spread to North America through the
1757:
1713:
1667:
919:
715:
696:
632:
594:
581:
behind the dorsal fin. Other differences are the Cypriniformes' unique
577:
on their backs; most other fishes of Ostariophysi have a small, fleshy
554:
550:
499:
102:
77:
1704:
The earliest cypriniform fossils are already assignable to the living
3877:
3167:
3125:
1717:
991:
628:
624:
482:
170:
3864:
3728:
3851:
3973:
1904:
1846:
965:
756:
742:
729:
651:. At least one species is found in saltwater, the Pacific redfin,
599:
534:
478:
474:
454:
692:
677:
620:
462:
3732:
3084:
3274:
1781:
1019:
665:, they seem to be specially adapted fishes of the Cyprinidae.
430:
3838:
770:
Historically, these included all the forms now placed in the
573:. They differ from most of their relatives in having only a
424:
1902:; 70 years later, not a single living individual existed.
1886:
or even beyond. In particular, Cyprinidae of southwestern
2883:
Florida Fish and Wildlife Conservation Commission (FFWCC)
1834:). The zebrafish is one of the most important vertebrate
418:
412:
400:
2581:(Teleostei: Cypriniformes) using molecular genetic data.
2320:, California Academy of Sciences (retrieved 2 July 2015)
914:
conservative main lineage apparently at least two major
1795:
Accidentally or deliberately introduced populations of
2970:
The Raffles Bulletin of Zoology, Suppl. No. 26: 1-199.
2745:
2908:(2008): Phylogenetic position of the enigmatic genus
1674:, maybe 160 million years ago (Mya). By 110 Mya, the
657:. Brackish water and marine cyprinids are invariably
433:
421:
415:
397:
2151:
1788:, and some "loaches" are bred for the international
1716:, they are roughly 60 million years old. During the
1689:
The Cypriniformes are thought to have originated in
820:
of Cypriniformes are traditionally divided into two
409:
406:
316:
3741:
3501:
3468:
3428:
3405:
3372:
3354:
3306:
3283:
3250:
3239:
3216:
3205:
3187:
3176:
3158:
2149:
2147:
2145:
2143:
2141:
2139:
2137:
2135:
2133:
2131:
1261:Phylogeny based on the work of the following works
899:successful enough never to shift from the original
777:except the catfish, which were placed in the order
593:in the throat. While other groups of fish, such as
427:
403:
1839:but can be shy in bright light and crowded tanks.
676:are families of mountain-stream fishes feeding on
1682:Cypriniformes must have been distinct from their
2879:. Version of 2005-FEB-15. Retrieved 2007-03-05.
2867:. Version of 2004-NOV-22. Retrieved 2007-03-05.
2855:. Version of 2004-NOV-22. Retrieved 2007-03-05.
2843:. Version of 2004-NOV-22. Retrieved 2007-03-05.
2831:. Version of 2004-NOV-22. Retrieved 2007-03-05.
2819:. Version of 2004-NOV-22. Retrieved 2007-03-05.
2807:. Version of 2004-NOV-22. Retrieved 2007-03-05.
2266:Milton Tan & Jonathan W. Ambruster (2018).
1898:) was the most common freshwater fish found in
2955:International Union for Conservation of Nature
2902:. Version of 2005-08-03. Retrieved 2007-05-03.
2173:
2171:
2169:
2167:
2165:
2163:
2161:
3096:
2454:
2452:
781:. By this definition, the Cypriniformes were
8:
1956:Globally extinct Cypriniformes species are:
832:) and also the mountain carps as the family
809:) have been separated out to form their own
2757:
2755:
615:The most notable family placed here is the
3729:
3247:
3213:
3184:
3103:
3089:
3081:
1748:is known from the Eocene of China. In the
1658:and relatives) is thought to be about the
113:
33:
3044:Tan, Milton & Armbruster, Jonathan W.
2935:Helfman, G.; Collette, B. & Facey, D.
2675:
2673:
2379:
2631:"ITIS Standard Report Page: Cyprinoidea"
2577:Phylogenetic position of the fish genus
2332:
2330:
2328:
2326:
2894:Gulf States Marine Fisheries Commission
2865:Family Psilorhynchidae - Mountain carps
2127:
525:Their closest living relatives are the
2606:"Cobitoidei – loach-like cypriniforms"
1210:(square-head/sturgeon-mouthed loaches)
2853:Family Gyrinocheilidae - Algae eaters
2488:Molecular Phylogenetics and Evolution
2237:
2235:
984:Liu, Chang, Wilson & Murray, 2015
7:
3979:497DDA6B-FF94-F111-43AB-B2E14EAEC3ED
3940:314d31c8-0cc4-4b9c-98ac-c447e885270f
2996:Orlov, Alexei & Sa-a, Pascualita
2841:Family Cyprinidae - Minnows or carps
2689:Murray, Alison M. (2 January 2020).
2187:
2185:
2183:
569:, fishes of Cypriniformes possess a
494:—which is also made up of cyprinid,
2312:Eschmeyer, W.N., Fong, J.D. (2015)
1528:
1499:
1470:
1441:
1412:
1385:
1357:
1288:
1280:
1272:
1264:
4065:Extant Thanetian first appearances
2887:Florida's Exotic Freshwater Fishes
2805:Family Balitoridae - River loaches
2695:Journal of Vertebrate Paleontology
2198:Journal of Vertebrate Paleontology
936:The mountain carps are the highly
25:
2900:Cyprinus carpio (Linnaeus, 1758)
2792:10.1111/j.1365-2699.2004.01170.x
2746:"Biochemical Signaling Pathways"
2575:Jörg Bohlen, Vendula Šlechtová:
2555:10.1111/j.1095-8649.2009.02398.x
1553:
1510:
1481:
1452:
1423:
1368:
1320:
1299:
976:, the earliest known cypriniform
828:contains the carps and minnows (
393:
368:
147:
2877:Order Summary for Cypriniformes
1662:, about 250 million years ago (
1153:(algae eaters; sucking loaches)
948:. While some details about the
510:, but are entirely absent from
2983:. John Wiley & Sons, Inc.
944:(Cyprininae), or maybe to the
1:
2817:Family Catostomidae - Suckers
2715:10.1080/02724634.2020.1762627
2218:10.1080/02724634.2015.1004073
1361:Catostomoidea (suckers)
940:Cyprinidae, perhaps close to
465:and their kin) fish, such as
2667:Briggs (2005), Nelson (2006)
1936:The Bridge on the River Kwai
1678:evidence indicates that the
1248:Nalbant & Bănărescu 1977
2928:10.1016/j.ympev.2007.10.012
2508:10.1016/j.ympev.2007.02.019
2314:Species by family/subfamily
2155:Tan & Armbruster (2018)
1917:) remain in the wild today.
1208:Bohlen & Šlechtová 2009
1180:(scooter/fire-eyed loaches)
884:, and long-finned loaches (
4081:
3076:Cypriniformes Tree of Life
2829:Family Cobitidae - Loaches
1736:, in addition to possibly
1728:are known from the Eocene
785:, so recently, the orders
3120:
3062:10.11646/zootaxa.4476.1.4
3030:10.1007/s00239-002-2417-y
2610:Mikko's Phylogeny Archive
2590:24 September 2015 at the
2533:with six nuclear genes".
2364:10.1038/s42003-019-0452-0
2284:10.11646/zootaxa.4476.1.4
1946:Acanthobrama telavivensis
1548:
1533:
1526:
1504:
1497:
1475:
1468:
1446:
1439:
1417:
1410:
1391:
1383:
1363:
1355:
1314:
1293:
1286:
1278:
1270:
1142:Suborder Gyrinocheiloidei
840:reviewed the superfamily
747:The Chinese algae eater (
684:. They are found only in
565:Like other orders of the
382:
375:
266:
261:
240:
233:
144:Scientific classification
142:
121:
112:
36:
2941:. Blackwell Publishing.
2255:. December 2012 version.
2245:; Pauly, Daniel (eds.).
1772:Relationship with humans
894:is usually treated as a
736:Nemacheilus chrysolaimos
4060:Fish of Central America
3008:. Retrieved 2007-03-05.
2939:The Diversity of Fishes
2889:. Retrieved 2007-03-05.
2535:Journal of Fish Biology
2424:FishBase (2004d,f), He
2415:Orlov & Sa-a {2007]
1953:they were first found.
1927:Epalzeorhynchos bicolor
1915:Epalzeorhynchos bicolor
1911:red-tailed black sharks
1819:. In some cases, these
1809:Ctenopharyngodon idella
1708:Catostomidae; from the
1697:, similar to the other
1240:(snake/serpent loaches)
1100:Mayden & Chen, 2010
1091:Mayden & Chen, 2010
1070:Mayden & Chen, 2010
750:Gyrinocheilus aymonieri
4030:Ray-finned fish orders
2915:Mol. Phylogenet. Evol.
2352:Communications Biology
2092:Pogonichthys ciscoides
1994:Chondrostoma scodrense
1923:red-tailed black shark
1918:
1864:
1843:Threats and extinction
1730:Sangkarewang Formation
1298:(carps & minnows)
1125:Suborder Catostomoidei
977:
767:
754:
740:
612:
449:, which includes many
4055:Fish of North America
3961:Paleobiology Database
2965:Kottelat, M. (2012):
2406:(2003), Briggs (2005)
2247:"Order Cypriniformes"
1962:Acanthobrama hulensis
1908:
1850:
1744:. The extinct family
1230:(brook/stone loaches)
969:
760:
746:
733:
603:
383:Around 4,205 species
2735:GSMFC (2005), FFWCC
2088:Clear Lake splittail
2047:Lepidomeda altivelis
2043:Pahranagat spinedace
2001:Cyprinus yilongensis
1878:, and in some cases
1170:(hillstream loaches)
998:Suborder Cyprinoidei
922:, acquiring similar
547:American knifefishes
537:and their kin), the
3374:Protacanthopterygii
3364:Lepidogalaxiiformes
3005:Tribolodon brandtii
2980:Fishes of the World
2707:2020JVPal..40E2627M
2547:2009JFBio..75.2197C
2531:Ellopostomaresolved
2500:2007MolPE..44.1358S
2210:2015JVPal..35E4073L
2177:Kottelat, M. (2012)
2101:Rhinichthys deaconi
2029:Evarra tlahuacensis
1868:Habitat destruction
1811:) are found on all
1190:(pointface loaches)
1159:Suborder Cobitoidei
872:, sucking loaches (
654:Tribolodon brandtii
604:A true loach - the
3591:Cyprinodontiformes
3516:Trachichthyiformes
3470:Paracanthopterygii
3321:Alepocephaliformes
3115:orders by subclass
2475:(1997): pp.228-229
2446:FishBase (2004b,c)
2437:FishBase (2004a,e)
2083:Notropis saladonis
2011:Evarra bustamantei
1984:Snake River sucker
1978:Barbus microbarbis
1919:
1874:of upland rivers,
1865:
1762:Bering land bridge
1746:Jianghanichthyidae
981:Jianghanichthyidae
978:
912:ecomorphologically
876:), stone loaches (
768:
755:
741:
714:are common across
613:
589:structures called
571:Weberian apparatus
320:Jianghanichthyidae
4012:
4011:
3948:Open Tree of Life
3735:Taxon identifiers
3726:
3725:
3722:
3721:
3718:
3717:
3714:
3713:
3686:Tetraodontiformes
3641:Chaetodontiformes
3576:Pleuronectiformes
3531:Batrachoidiformes
3293:Osteoglossiformes
3285:Osteoglossomorpha
3064:
3039:
2992:
2975:Nelson, Joseph S.
2950:
2930:
2794:
2318:Catalog of Fishes
2110:Stypodon signifer
2106:Stumptooth minnow
2065:Notropis aulidion
2056:Moxostoma lacerum
2020:Evarra eigenmanni
1988:Chasmistes muriei
1863:since about 1960.
1638:
1637:
1629:
1628:
1620:
1619:
1611:
1610:
1602:
1601:
1593:
1592:
1584:
1583:
1575:
1574:
1566:
1565:
1389:Gyrinocheiloidea
1343:
1342:
1334:
1333:
1250:(longfin loaches)
1249:
1239:
1235:Serpenticobitidae
1229:
1219:
1209:
1199:
1189:
1179:
1169:
1152:
1135:
1119:
1110:
1101:
1092:
1083:
1071:
1062:
1053:
1044:
1035:
1017:
1008:
985:
882:Serpenticobitidae
860:), true loaches (
797:and allies), and
787:Gonorhynchiformes
387:
386:
344:Serpenticobitidae
229:
16:(Redirected from
4072:
4005:
4004:
3992:
3991:
3982:
3981:
3969:
3968:
3956:
3955:
3943:
3942:
3933:
3932:
3920:
3919:
3917:NBNSYS0000172142
3907:
3906:
3894:
3893:
3881:
3880:
3868:
3867:
3855:
3854:
3842:
3841:
3829:
3828:
3816:
3815:
3803:
3802:
3790:
3789:
3777:
3776:
3775:
3762:
3761:
3760:
3730:
3696:Centrarchiformes
3671:Priacanthiformes
3666:Scatophagiformes
3621:Uranoscopiformes
3566:Istiophoriformes
3556:Synbranchiformes
3521:Holocentriformes
3488:Stylephoriformes
3440:Ateleopodiformes
3326:Gonorynchiformes
3270:Notacanthiformes
3248:
3244:
3226:Lepisosteiformes
3214:
3210:
3197:Acipenseriformes
3185:
3181:
3105:
3098:
3091:
3082:
3056:
3045:
3024:
3013:
2997:
2984:
2976:
2958:
2942:
2936:
2922:
2907:
2897:
2884:
2874:
2862:
2850:
2838:
2826:
2814:
2802:
2786:
2775:
2762:
2759:
2750:
2749:
2742:
2736:
2733:
2727:
2726:
2686:
2680:
2677:
2668:
2665:
2659:
2652:
2646:
2645:
2643:
2641:
2627:
2621:
2620:
2618:
2616:
2601:
2595:
2573:
2567:
2566:
2541:(9): 2197–2208.
2526:
2520:
2519:
2482:
2476:
2469:
2463:
2456:
2447:
2444:
2438:
2435:
2429:
2422:
2416:
2413:
2407:
2400:
2394:
2393:
2383:
2343:
2337:
2334:
2321:
2310:
2304:
2303:
2263:
2257:
2256:
2239:
2230:
2229:
2189:
2178:
2175:
2156:
2153:
2038:Gila crassicauda
1896:Gila crassicauda
1857:Gila crassicauda
1648:Gonorynchiformes
1557:
1536:Ellopostomatidae
1529:
1515:
1514:
1500:
1486:
1485:
1471:
1457:
1456:
1442:
1428:
1427:
1413:
1408: (loaches)
1386:
1372:
1358:
1325:
1324:
1303:
1289:
1281:
1273:
1265:
1247:
1237:
1227:
1220:(sucker loaches)
1217:
1215:Gastromyzontidae
1207:
1205:Ellopostomatidae
1197:
1187:
1177:
1167:
1150:
1133:
1117:
1108:
1099:
1090:
1078:
1069:
1060:
1051:
1042:
1033:
1016:Rafinesque, 1815
1015:
1006:
1004:Acheilognathidae
983:
910:. Yet, from the
901:ecological niche
870:Gastromyzontidae
866:Ellopostomatidae
838:Maurice Kottelat
766:, a small sucker
763:Erimyzon sucetta
591:pharyngeal teeth
520:bigmouth buffalo
440:
439:
436:
435:
432:
429:
426:
423:
420:
417:
414:
411:
408:
405:
402:
399:
318:
305:Gastromyzontidae
301:Ellopostomatidae
269:Acheilognathidae
224:
152:
151:
117:
107:
44:
43:Paleocene–recent
40:Temporal range:
34:
21:
4080:
4079:
4075:
4074:
4073:
4071:
4070:
4069:
4015:
4014:
4013:
4008:
4000:
3995:
3987:
3985:
3977:
3972:
3964:
3959:
3951:
3946:
3938:
3936:
3928:
3923:
3915:
3910:
3902:
3897:
3889:
3884:
3876:
3871:
3863:
3858:
3850:
3845:
3837:
3832:
3824:
3819:
3811:
3806:
3798:
3793:
3785:
3780:
3771:
3770:
3765:
3756:
3755:
3750:
3737:
3727:
3710:
3706:Scorpaeniformes
3646:Acanthuriformes
3606:Gobiesociformes
3546:Syngnathiformes
3503:Acanthopterygii
3497:
3464:
3460:Polymixiiformes
3424:
3401:
3382:Argentiniformes
3368:
3350:
3302:
3279:
3240:
3235:
3206:
3201:
3177:
3172:
3168:Polypteriformes
3154:
3116:
3109:
3072:
3065:(HTML abstract)
3043:
3040:(HTML abstract)
3011:
2995:
2974:
2953:
2934:
2931:(HTML abstract)
2905:
2892:
2882:
2870:
2858:
2846:
2834:
2822:
2810:
2798:
2795:(HTML abstract)
2774:Briggs, John C.
2773:
2770:
2765:
2760:
2753:
2744:
2743:
2739:
2734:
2730:
2701:(1): e1762627.
2688:
2687:
2683:
2678:
2671:
2666:
2662:
2653:
2649:
2639:
2637:
2629:
2628:
2624:
2614:
2612:
2604:Mikko Haaramo.
2603:
2602:
2598:
2592:Wayback Machine
2574:
2570:
2528:
2527:
2523:
2484:
2483:
2479:
2470:
2466:
2457:
2450:
2445:
2441:
2436:
2432:
2423:
2419:
2414:
2410:
2401:
2397:
2345:
2344:
2340:
2335:
2324:
2311:
2307:
2265:
2264:
2260:
2241:
2240:
2233:
2204:(6): e1004073.
2191:
2190:
2181:
2176:
2159:
2154:
2129:
2125:
2116:Telestes ukliva
1921:The well-known
1845:
1836:model organisms
1801:Cyprinus carpio
1774:
1691:South-east Asia
1676:plate tectonics
1644:
1639:
1630:
1621:
1612:
1603:
1594:
1585:
1576:
1567:
1509:
1480:
1451:
1449:Vaillantellidae
1422:
1394:Gyrinocheilidae
1344:
1335:
1319:
1317:Psilorhynchidae
1259:
1245:Vaillantellidae
1148:Gyrinocheilidae
1088:Sundadanionidae
1076:Psilorhynchidae
1067:Paedocyprididae
1061:Bonaparte, 1853
973:Jianghanichthys
886:Vaillantellidae
874:Gyrinocheilidae
834:Psilorhynchidae
728:
703:) are found in
699:. The suckers (
674:Gyrinocheilidae
663:Psilorhynchidae
643:water, such as
563:
447:ray-finned fish
396:
392:
366:
362:
360:Vaillantellidae
358:
354:
350:
348:Sundadanionidae
346:
342:
340:Psilorhynchidae
338:
336:Paedocyprididae
334:
330:
326:
322:
315:
313:Gyrinocheilidae
311:
307:
303:
299:
295:
291:
287:
283:
279:
275:
271:
257:
246:
243:Cyprinus carpio
223:
146:
128:Cyprinus carpio
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
42:
41:
38:
30:
29:Order of fishes
23:
22:
15:
12:
11:
5:
4078:
4076:
4068:
4067:
4062:
4057:
4052:
4047:
4045:Fish of Europe
4042:
4040:Fish of Africa
4037:
4032:
4027:
4017:
4016:
4010:
4009:
4007:
4006:
3993:
3983:
3970:
3957:
3944:
3934:
3921:
3908:
3895:
3882:
3869:
3856:
3843:
3830:
3817:
3804:
3791:
3778:
3763:
3747:
3745:
3739:
3738:
3733:
3724:
3723:
3720:
3719:
3716:
3715:
3712:
3711:
3709:
3708:
3703:
3698:
3693:
3691:Pempheriformes
3688:
3683:
3678:
3673:
3668:
3663:
3658:
3653:
3648:
3643:
3638:
3633:
3628:
3623:
3618:
3613:
3608:
3603:
3598:
3593:
3588:
3586:Atheriniformes
3583:
3578:
3573:
3568:
3563:
3561:Anabantiformes
3558:
3553:
3548:
3543:
3538:
3533:
3528:
3523:
3518:
3513:
3507:
3505:
3499:
3498:
3496:
3495:
3490:
3485:
3480:
3478:Percopsiformes
3474:
3472:
3466:
3465:
3463:
3462:
3457:
3452:
3450:Myctophiformes
3447:
3442:
3436:
3434:
3431:incertae sedis
3426:
3425:
3423:
3422:
3417:
3411:
3409:
3403:
3402:
3400:
3399:
3394:
3389:
3384:
3378:
3376:
3370:
3369:
3367:
3366:
3360:
3358:
3352:
3351:
3349:
3348:
3343:
3338:
3333:
3328:
3323:
3318:
3312:
3310:
3304:
3303:
3301:
3300:
3298:Hiodontiformes
3295:
3289:
3287:
3281:
3280:
3278:
3277:
3275:Anguilliformes
3272:
3267:
3262:
3256:
3254:
3245:
3237:
3236:
3234:
3233:
3228:
3222:
3220:
3211:
3203:
3202:
3200:
3199:
3193:
3191:
3182:
3174:
3173:
3171:
3170:
3164:
3162:
3156:
3155:
3153:
3152:
3146:
3140:
3134:
3128:
3121:
3118:
3117:
3113:Actinopterygii
3110:
3108:
3107:
3100:
3093:
3085:
3079:
3078:
3071:
3070:External links
3068:
3067:
3066:
3055:(1): 006–039.
3041:
3023:(4): 464–472.
3009:
2993:
2972:
2963:
2951:
2932:
2903:
2890:
2880:
2868:
2856:
2844:
2832:
2820:
2808:
2796:
2785:(2): 287–294.
2769:
2766:
2764:
2763:
2751:
2737:
2728:
2681:
2669:
2660:
2647:
2622:
2596:
2568:
2521:
2494:(3): 1358–65.
2477:
2464:
2448:
2439:
2430:
2417:
2408:
2395:
2338:
2322:
2305:
2278:(1): 006–039.
2258:
2243:Froese, Rainer
2231:
2179:
2157:
2126:
2124:
2121:
2120:
2119:
2112:
2103:
2097:Las Vegas dace
2094:
2085:
2076:
2070:Phantom shiner
2067:
2061:Durango shiner
2058:
2052:Harelip sucker
2049:
2040:
2034:Thicktail chub
2031:
2025:Endorheic chub
2022:
2013:
2004:
1997:
1990:
1981:
1974:
1972:Alburnus akili
1965:
1892:thicktail chub
1859:) is globally
1853:thicktail chub
1844:
1841:
1821:exotic species
1773:
1770:
1738:Smilogastrinae
1660:Early Triassic
1643:
1640:
1636:
1635:
1632:
1631:
1627:
1626:
1623:
1622:
1618:
1617:
1614:
1613:
1609:
1608:
1605:
1604:
1600:
1599:
1596:
1595:
1591:
1590:
1587:
1586:
1582:
1581:
1578:
1577:
1573:
1572:
1569:
1568:
1564:
1563:
1560:
1559:
1547:
1544:
1543:
1540:
1539:
1532:
1527:
1525:
1522:
1521:
1518:
1517:
1503:
1498:
1496:
1493:
1492:
1489:
1488:
1474:
1469:
1467:
1464:
1463:
1460:
1459:
1445:
1440:
1438:
1435:
1434:
1431:
1430:
1416:
1411:
1409:
1402:
1401:
1398:
1397:
1390:
1384:
1382:
1379:
1378:
1375:
1374:
1362:
1356:
1354:
1350:
1349:
1346:
1345:
1341:
1340:
1337:
1336:
1332:
1331:
1328:
1327:
1313:
1310:
1309:
1306:
1305:
1292:
1287:
1285:
1279:
1277:
1271:
1269:
1268:Cypriniformes
1263:
1258:
1255:
1254:
1253:
1252:
1251:
1241:
1231:
1221:
1211:
1201:
1200:(true loaches)
1191:
1181:
1171:
1156:
1155:
1154:
1139:
1138:
1137:
1122:
1121:
1120:
1115:Xenocyprididae
1111:
1102:
1093:
1084:
1072:
1063:
1054:
1045:
1036:
1027:
1009:
995:
931:in the process
896:junior synonym
824:. Superfamily
727:
724:
610:Cobitis taenia
562:
559:
508:Southeast Asia
385:
384:
380:
379:
373:
372:
364:Xenocyprididae
264:
263:
259:
258:
247:
238:
237:
231:
230:
218:
214:
213:
208:
204:
203:
198:
194:
193:
191:Actinopterygii
188:
184:
183:
178:
174:
173:
168:
164:
163:
158:
154:
153:
140:
139:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
39:
28:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4077:
4066:
4063:
4061:
4058:
4056:
4053:
4051:
4048:
4046:
4043:
4041:
4038:
4036:
4033:
4031:
4028:
4026:
4025:Cypriniformes
4023:
4022:
4020:
4003:
3998:
3994:
3990:
3984:
3980:
3975:
3971:
3967:
3962:
3958:
3954:
3949:
3945:
3941:
3935:
3931:
3926:
3922:
3918:
3913:
3909:
3905:
3900:
3896:
3892:
3887:
3883:
3879:
3874:
3870:
3866:
3861:
3857:
3853:
3848:
3844:
3840:
3835:
3831:
3827:
3822:
3818:
3814:
3809:
3805:
3801:
3800:Cypriniformes
3796:
3792:
3788:
3787:Cypriniformes
3783:
3779:
3774:
3773:Cypriniformes
3768:
3764:
3759:
3753:
3749:
3748:
3746:
3744:
3743:Cypriniformes
3740:
3736:
3731:
3707:
3704:
3702:
3699:
3697:
3694:
3692:
3689:
3687:
3684:
3682:
3679:
3677:
3674:
3672:
3669:
3667:
3664:
3662:
3659:
3657:
3654:
3652:
3651:Lutjaniformes
3649:
3647:
3644:
3642:
3639:
3637:
3636:Ephippiformes
3634:
3632:
3629:
3627:
3624:
3622:
3619:
3617:
3614:
3612:
3611:Blenniiformes
3609:
3607:
3604:
3602:
3599:
3597:
3594:
3592:
3589:
3587:
3584:
3582:
3579:
3577:
3574:
3572:
3571:Carangiformes
3569:
3567:
3564:
3562:
3559:
3557:
3554:
3552:
3551:Scombriformes
3549:
3547:
3544:
3542:
3539:
3537:
3534:
3532:
3529:
3527:
3526:Ophidiiformes
3524:
3522:
3519:
3517:
3514:
3512:
3509:
3508:
3506:
3504:
3500:
3494:
3491:
3489:
3486:
3484:
3481:
3479:
3476:
3475:
3473:
3471:
3467:
3461:
3458:
3456:
3453:
3451:
3448:
3446:
3443:
3441:
3438:
3437:
3435:
3433:
3432:
3427:
3421:
3418:
3416:
3413:
3412:
3410:
3408:
3404:
3398:
3395:
3393:
3392:Salmoniformes
3390:
3388:
3387:Galaxiiformes
3385:
3383:
3380:
3379:
3377:
3375:
3371:
3365:
3362:
3361:
3359:
3357:
3356:Lepidogalaxii
3353:
3347:
3344:
3342:
3341:Gymnotiformes
3339:
3337:
3336:Characiformes
3334:
3332:
3331:Cypriniformes
3329:
3327:
3324:
3322:
3319:
3317:
3314:
3313:
3311:
3309:
3305:
3299:
3296:
3294:
3291:
3290:
3288:
3286:
3282:
3276:
3273:
3271:
3268:
3266:
3263:
3261:
3258:
3257:
3255:
3253:
3249:
3246:
3243:
3238:
3232:
3229:
3227:
3224:
3223:
3221:
3219:
3215:
3212:
3209:
3204:
3198:
3195:
3194:
3192:
3190:
3186:
3183:
3180:
3175:
3169:
3166:
3165:
3163:
3161:
3157:
3151:
3147:
3145:
3144:Gnathostomata
3141:
3139:
3135:
3133:
3129:
3127:
3123:
3122:
3119:
3114:
3106:
3101:
3099:
3094:
3092:
3087:
3086:
3083:
3077:
3074:
3073:
3069:
3063:
3059:
3054:
3051:
3050:
3042:
3038:
3034:
3031:
3027:
3022:
3019:
3018:
3017:J. Mol. Evol.
3010:
3007:
3006:
3001:
2994:
2991:
2990:0-471-25031-7
2987:
2982:
2981:
2973:
2971:
2968:
2964:
2961:
2956:
2952:
2949:
2948:0-86542-256-7
2945:
2940:
2933:
2929:
2925:
2920:
2917:
2916:
2911:
2910:Psilorhynchus
2904:
2901:
2895:
2891:
2888:
2881:
2878:
2873:
2869:
2866:
2861:
2857:
2854:
2849:
2845:
2842:
2837:
2833:
2830:
2825:
2821:
2818:
2813:
2809:
2806:
2801:
2797:
2793:
2789:
2784:
2781:
2780:
2772:
2771:
2767:
2758:
2756:
2752:
2747:
2741:
2738:
2732:
2729:
2724:
2720:
2716:
2712:
2708:
2704:
2700:
2696:
2692:
2685:
2682:
2679:Briggs (2005)
2676:
2674:
2670:
2664:
2661:
2657:
2651:
2648:
2636:
2632:
2626:
2623:
2611:
2607:
2600:
2597:
2593:
2589:
2586:
2582:
2578:
2572:
2569:
2564:
2560:
2556:
2552:
2548:
2544:
2540:
2536:
2532:
2525:
2522:
2517:
2513:
2509:
2505:
2501:
2497:
2493:
2489:
2481:
2478:
2474:
2468:
2465:
2461:
2455:
2453:
2449:
2443:
2440:
2434:
2431:
2427:
2421:
2418:
2412:
2409:
2405:
2399:
2396:
2391:
2387:
2382:
2377:
2373:
2369:
2365:
2361:
2357:
2353:
2349:
2342:
2339:
2336:Nelson (2006)
2333:
2331:
2329:
2327:
2323:
2319:
2315:
2309:
2306:
2301:
2297:
2293:
2289:
2285:
2281:
2277:
2273:
2269:
2262:
2259:
2254:
2253:
2248:
2244:
2238:
2236:
2232:
2227:
2223:
2219:
2215:
2211:
2207:
2203:
2199:
2195:
2188:
2186:
2184:
2180:
2174:
2172:
2170:
2168:
2166:
2164:
2162:
2158:
2152:
2150:
2148:
2146:
2144:
2142:
2140:
2138:
2136:
2134:
2132:
2128:
2122:
2118:
2117:
2113:
2111:
2107:
2104:
2102:
2098:
2095:
2093:
2089:
2086:
2084:
2080:
2079:Salado shiner
2077:
2075:
2074:Notropis orca
2071:
2068:
2066:
2062:
2059:
2057:
2053:
2050:
2048:
2044:
2041:
2039:
2035:
2032:
2030:
2026:
2023:
2021:
2017:
2014:
2012:
2008:
2005:
2003:
2002:
1998:
1996:
1995:
1991:
1989:
1985:
1982:
1980:
1979:
1975:
1973:
1969:
1966:
1964:
1963:
1959:
1958:
1957:
1954:
1951:
1947:
1943:
1938:
1937:
1932:
1928:
1924:
1916:
1912:
1907:
1903:
1901:
1897:
1893:
1889:
1888:North America
1885:
1881:
1877:
1873:
1869:
1862:
1858:
1854:
1849:
1842:
1840:
1837:
1833:
1829:
1824:
1822:
1818:
1814:
1810:
1806:
1802:
1798:
1793:
1791:
1787:
1783:
1779:
1771:
1769:
1767:
1763:
1759:
1755:
1751:
1747:
1743:
1739:
1735:
1731:
1727:
1723:
1719:
1715:
1711:
1707:
1702:
1700:
1696:
1695:South America
1692:
1687:
1685:
1681:
1677:
1673:
1669:
1665:
1661:
1657:
1656:Characiformes
1653:
1649:
1641:
1634:
1633:
1625:
1624:
1616:
1615:
1607:
1606:
1598:
1597:
1589:
1588:
1580:
1579:
1571:
1570:
1562:
1561:
1558:
1556:
1552:
1551:Nemacheilidae
1546:
1545:
1542:
1541:
1538:
1537:
1531:
1530:
1524:
1523:
1520:
1519:
1516:
1513:
1508:
1502:
1501:
1495:
1494:
1491:
1490:
1487:
1484:
1479:
1473:
1472:
1466:
1465:
1462:
1461:
1458:
1455:
1450:
1444:
1443:
1437:
1436:
1433:
1432:
1429:
1426:
1421:
1415:
1414:
1407:
1404:
1403:
1400:
1399:
1396:
1395:
1388:
1387:
1381:
1380:
1377:
1376:
1373:
1371:
1367:
1360:
1359:
1352:
1351:
1348:
1347:
1339:
1338:
1330:
1329:
1326:
1323:
1318:
1312:
1311:
1308:
1307:
1304:
1302:
1297:
1291:
1290:
1283:
1282:
1275:
1274:
1267:
1266:
1262:
1256:
1246:
1242:
1238:Kottelat 2012
1236:
1232:
1226:
1225:Nemacheilidae
1222:
1216:
1212:
1206:
1202:
1198:Swainson 1838
1196:
1192:
1186:
1182:
1178:Kottelat 2012
1176:
1172:
1168:Swainson 1839
1166:
1162:
1161:
1160:
1157:
1149:
1145:
1144:
1143:
1140:
1132:
1128:
1127:
1126:
1123:
1118:Günther, 1868
1116:
1112:
1107:
1103:
1098:
1097:Tanichthyidae
1094:
1089:
1085:
1081:
1077:
1073:
1068:
1064:
1059:
1055:
1052:Bleeker, 1864
1050:
1049:Leptobarbidae
1046:
1043:Bleeker, 1863
1041:
1037:
1034:Bleeker, 1863
1032:
1028:
1026:) and minnows
1025:
1021:
1014:
1010:
1007:Bleeker, 1863
1005:
1001:
1000:
999:
996:
993:
989:
982:
975:
974:
968:
964:
963:
962:
959:
955:
951:
947:
943:
939:
934:
932:
929:
925:
921:
917:
913:
909:
906:
902:
897:
893:
892:Catostomoidea
889:
887:
883:
879:
878:Nemacheilidae
875:
871:
867:
863:
859:
855:
851:
847:
843:
839:
835:
831:
827:
823:
819:
814:
812:
808:
807:electric eels
804:
800:
799:Gymnotiformes
796:
792:
791:Characiformes
788:
784:
780:
776:
773:
765:
764:
759:
752:
751:
745:
738:
737:
732:
725:
723:
721:
718:and parts of
717:
713:
709:
708:North America
706:
702:
698:
694:
691:
687:
683:
682:invertebrates
679:
675:
671:
666:
664:
660:
656:
655:
650:
646:
642:
638:
637:North America
634:
630:
626:
622:
618:
611:
607:
602:
598:
596:
592:
588:
584:
580:
576:
572:
568:
560:
558:
556:
552:
548:
544:
540:
539:Gymnotiformes
536:
532:
528:
527:Characiformes
523:
521:
517:
516:South America
513:
509:
505:
501:
497:
496:ostariophysin
493:
492:
488:
484:
480:
476:
472:
468:
464:
460:
456:
452:
448:
444:
438:
391:
390:Cypriniformes
381:
378:
374:
371:
370:
365:
361:
357:
353:
352:Tanichthyidae
349:
345:
341:
337:
333:
332:Nemacheilidae
329:
325:
324:Leptobarbidae
321:
314:
310:
306:
302:
298:
294:
290:
286:
282:
278:
274:
270:
265:
260:
255:
251:
245:
244:
239:
236:
232:
227:
222:
221:Cypriniformes
219:
216:
215:
212:
211:Cypriniphysae
209:
206:
205:
202:
199:
196:
195:
192:
189:
186:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
150:
145:
141:
137:
133:
129:
125:
120:
116:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
37:Cypriniformes
35:
32:
27:
19:
4050:Fish of Asia
4035:Ostariophysi
3742:
3681:Lophiiformes
3676:Caproiformes
3656:Lobotiformes
3631:Moroniformes
3616:Gerreiformes
3601:Mugiliformes
3596:Beloniformes
3581:Cichliformes
3511:Beryciformes
3455:Lampriformes
3445:Aulopiformes
3429:
3420:Stomiiformes
3415:Osmeriformes
3346:Siluriformes
3330:
3316:Clupeiformes
3265:Albuliformes
3150:Osteichthyes
3142:Infraphylum
3052:
3047:
3020:
3015:
3004:
2978:
2969:
2960:
2938:
2918:
2913:
2909:
2782:
2779:J. Biogeogr.
2777:
2740:
2731:
2698:
2694:
2684:
2663:
2655:
2650:
2638:. Retrieved
2635:www.itis.gov
2634:
2625:
2613:. Retrieved
2609:
2599:
2580:
2576:
2571:
2538:
2534:
2530:
2524:
2491:
2487:
2480:
2472:
2467:
2459:
2442:
2433:
2425:
2420:
2411:
2403:
2398:
2355:
2351:
2341:
2308:
2275:
2271:
2261:
2250:
2201:
2197:
2114:
2109:
2100:
2091:
2082:
2073:
2064:
2055:
2046:
2037:
2028:
2019:
2016:Plateau chub
2010:
2007:Mexican dace
1999:
1992:
1987:
1976:
1971:
1968:Gökçe balığı
1960:
1955:
1950:Yarqon River
1945:
1942:Yarqon bleak
1934:
1926:
1920:
1914:
1895:
1866:
1856:
1831:
1825:
1808:
1800:
1794:
1792:fish trade.
1775:
1703:
1688:
1652:Characiphysi
1645:
1549:
1534:
1505:
1476:
1447:
1418:
1392:
1366:Catostomidae
1364:
1315:
1294:
1284:Cyprinoidea
1276:Cyprinoidei
1260:
1158:
1141:
1134:Agassiz 1850
1131:Catostomidae
1124:
1109:Jordan, 1878
997:
971:
950:phylogenetic
935:
908:Ostariophysi
890:
858:Catostomidae
826:Cyprinioidea
815:
811:monophyletic
783:paraphyletic
779:Siluriformes
775:Ostariophysi
769:
761:
748:
734:
720:North Africa
701:Catostomidae
667:
652:
614:
609:
606:spined loach
567:Ostariophysi
564:
551:Siluriformes
543:electric eel
524:
495:
491:Ostariophysi
489:
458:
389:
388:
285:Catostomidae
267:
241:
235:Type species
220:
207:(unranked):
197:(unranked):
127:
122:A wild-type
31:
26:
3873:iNaturalist
3767:Wikispecies
3701:Perciformes
3661:Spariformes
3626:Labriformes
3541:Gobiiformes
3536:Kurtiformes
3397:Esociformes
3260:Elopiformes
3252:Elopomorpha
3208:Neopterygii
3189:Chondrostei
3179:Actinopteri
3148:Superclass
2921:: 419–425.
2761:IUCN (2007)
2579:Ellopostoma
1948:) from the
1929:) from the
1909:Few if any
1880:overfishing
1832:Danio rerio
1797:common carp
1686:relatives.
1507:Balitoridae
1353:Cobitoidei
1218:Fowler 1905
1175:Barbuccidae
1165:Balitoridae
1058:Leuciscidae
958:Leuciscinae
928:adaptations
856:, suckers (
850:Barbuccidae
846:Balitoridae
836:. In 2012,
803:knifefishes
726:Systematics
690:subtropical
670:Balitoridae
579:adipose fin
561:Description
549:), and the
328:Leuciscidae
277:Barbuccidae
273:Balitoridae
124:common carp
18:Cyprinoidea
4019:Categories
3493:Gadiformes
3308:Otocephala
3231:Amiiformes
3138:Vertebrata
3136:Subphylum
2768:References
2615:26 October
2358:(1): 197.
1900:California
1884:extinction
1817:Antarctica
1813:continents
1805:grass carp
1742:Labeoninae
1726:Danioninae
1699:otophysans
1406:Cobitoidea
1296:Cyprinidae
1228:Regan 1911
1040:Gobionidae
1031:Danionidae
1013:Cyprinidae
970:Fossil of
942:true carps
938:apomorphic
916:radiations
842:Cobitoidei
830:Cyprinidae
772:superorder
680:and small
659:anadromous
617:Cyprinidae
587:convergent
583:kinethmoid
575:dorsal fin
487:superorder
309:Gobionidae
297:Danionidae
293:Cyprinidae
136:Cyprininae
132:Cyprinidae
3483:Zeiformes
3242:Teleostei
3160:Cladistia
2863:(2004f):
2851:(2004e):
2839:(2004d):
2827:(2004c):
2815:(2004b):
2803:(2004a):
2723:0272-4634
2594:; 1,8 MB)
2372:2399-3642
2226:0272-4634
1933:River of
1931:Mae Klong
1876:pollution
1828:zebrafish
1766:evolution
1754:sympatric
1750:Oligocene
1734:Indonesia
1710:Paleocene
1684:Gondwanan
1680:Laurasian
1642:Evolution
1478:Cobitidae
1257:Phylogeny
1195:Cobitidae
1188:Berg 1940
1151:Gill 1905
1136:(suckers)
994:of China)
988:Paleocene
954:Cultrinae
946:danionins
862:Cobitidae
822:suborders
795:characins
712:Cobitidae
705:temperate
555:catfishes
531:characins
512:Australia
377:Diversity
289:Cobitidae
262:Families
167:Kingdom:
161:Eukaryota
3752:Wikidata
3407:Stomiati
3218:Holostei
3132:Chordata
3126:Animalia
3124:Kingdom
3037:12664166
3000:FishBase
2977:(2006):
2959:(2007):
2937:(1997):
2898:(2005):
2885:(2006):
2875:(2005):
2872:FishBase
2860:FishBase
2848:FishBase
2836:FishBase
2824:FishBase
2812:FishBase
2800:FishBase
2588:Archived
2563:20738682
2516:17433724
2471:Helfman
2390:31149641
2300:52976511
2292:30313339
2252:FishBase
1790:aquarium
1778:goldfish
1722:Barbinae
1672:Jurassic
1420:Botiidae
1185:Botiidae
1106:Tincidae
1024:goldfish
1018:(carps,
979:Family †
920:habitats
854:Botiidae
818:families
813:orders.
686:tropical
641:brackish
595:cichlids
459:cyprinid
451:families
367:and see
356:Tincidae
281:Botiidae
250:Linnaeus
201:Otophysi
181:Chordata
177:Phylum:
171:Animalia
157:Domain:
3953:1005931
3758:Q177879
3130:Phylum
3111:Extant
3049:Zootaxa
2896:(GSMFC)
2748:. ZFIN.
2703:Bibcode
2654:Saitoh
2640:26 July
2543:Bibcode
2496:Bibcode
2402:Saitoh
2381:6533251
2316:in the
2272:Zootaxa
2206:Bibcode
1872:damming
1861:extinct
1815:except
1786:angling
1758:Miocene
1714:Alberta
1670:in the
1668:Pangaea
1243:Family
1233:Family
1223:Family
1213:Family
1203:Family
1193:Family
1183:Family
1173:Family
1163:Family
1146:Family
1129:Family
1113:Family
1104:Family
1095:Family
1086:Family
1074:Family
1065:Family
1056:Family
1047:Family
1038:Family
1029:Family
1011:Family
1002:Family
924:habitus
903:of the
716:Eurasia
697:species
633:Eurasia
625:minnows
500:species
483:minnows
475:loaches
226:Bleeker
217:Order:
187:Class:
4002:154156
3989:113893
3986:uBio:
3937:NZOR:
3904:162846
3852:1CYPNO
3035:
2988:
2957:(IUCN)
2946:
2721:
2658:(2003)
2656:et al.
2561:
2514:
2473:et al.
2462:(2008)
2460:et al.
2428:(2008)
2426:et al.
2404:et al.
2388:
2378:
2370:
2298:
2290:
2224:
1803:) and
1718:Eocene
1706:family
1082:, 1926
992:Eocene
635:, and
629:Africa
535:tetras
481:, and
479:botias
471:gobies
455:genera
441:is an
228:, 1859
3997:WoRMS
3974:Plazi
3966:64193
3891:12345
3886:IRMNG
3878:49216
2296:S2CID
2123:Notes
905:basal
678:algae
649:bream
645:roach
621:carps
504:South
467:barbs
463:carps
443:order
3930:7952
3925:NCBI
3899:ITIS
3865:1153
3860:GBIF
3847:EPPO
3839:3194
3808:BOLD
3053:4476
3033:PMID
2986:ISBN
2944:ISBN
2719:ISSN
2642:2019
2617:2013
2559:PMID
2512:PMID
2386:PMID
2368:ISSN
2288:PMID
2276:4476
2222:ISSN
1851:The
1780:and
1740:and
1724:and
1080:Hora
956:and
926:and
816:The
805:and
693:Asia
688:and
672:and
668:The
647:and
623:and
545:and
514:and
506:and
453:and
369:text
254:1758
48:PreꞒ
3912:NBN
3834:EoL
3826:334
3821:CoL
3813:243
3795:AFD
3782:ADW
3058:doi
3026:doi
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