46:
33:
535:
342:, and features eleven-segment antennae. Little sexual dimorphism exists between the male and female specimen, beyond a longer fifth abdominal
334:
are divided into five segments, of increasing length, with the fifth segment as long as the others combined. It has long, narrow, light brown
361:
narrows to a slightly pointed tip. It has a U-shaped depression between the V-shaped notches on the tip of the genitalia. The genitals'
581:
242:
462:
45:
460:
Matsumoto, Keita (2021). "Six new species of the genus
Caccothryptus from the Himalayas (Coleoptera: Limnichidae)".
227:
266:
identified a number of distinct species from these specimens. A male specimen from
Kathmandu was labeled as the
586:
246:
189:
253:
493:
323:
259:
230:
in 1902. In 2014, the genus was divided into five species groups by Carles
Hernando and Ignacio Ribera.
140:
553:
386:
576:
263:
181:
294:
204:
188:, Nepal in 1983. Six of Brendell's specimens were distinguished from its initial classification of
358:
40:
475:
467:
362:
331:
349:
The beetle's genitals display a number of differences from the original classification of
241:, Nepal by Martin J. D. Brendell, curator of Coleoptera Department of Entomology at the
289:
262:
in 1923. These were stored at the
Natural History Museum. In 2021, museum entomologist
195:
32:
387:"Sharp, David. 1902. Descriptions of Oriental Limnichini (Coleoptera, Fam. Byrrhidae)"
570:
222:
117:
516:
335:
330:, which are distributed across both the upper and lower sides of the beetle. Its
217:
173:
169:
107:
471:
366:
479:
315:
238:
185:
77:
57:
527:
338:. The beetle's head, smooth and punctured, is slightly retracted into its
510:
343:
339:
275:
267:
540:
391:
274:. Five specimens (comprising three males and two females) were labeled
314:
of 3.39–3.68 mm, and elytral widths of 2.32–2.52 mm. Their
311:
97:
87:
67:
487:
548:
199:
327:
491:
322:
has an elongated oval body, brown in colour except for reddish
438:
436:
194:
due to distinct genital structures. The species was named for
184:
in 2021, using specimens gathered by Martin J. D. Brendell at
318:
are 0.90–1 mm in length and 2.03–2.11 mm in width.
411:
409:
365:
are wavy, forming two concave curves. Females have a long
353:, which enabled its classification as a separate species.
500:
176:. The species was described alongside five other
180:species by Natural History Museum entomologist
8:
488:
31:
20:
442:
427:
415:
378:
310:specimens identified by Matsumoto have
7:
554:FB863BA1-2C04-435A-869C-5C6A16C82964
288:and named for Japanese manga artist
237:specimens were collected in 1983 in
220:(minute marsh-loving beetle) genus
14:
284:was placed in the species group
44:
258:, first described by forester
243:Natural History Museum, London
1:
463:European Journal of Taxonomy
603:
170:minute marsh-loving beetle
582:Beetles described in 2021
472:10.5852/ejt.2021.739.1275
146:
139:
41:Scientific classification
39:
30:
23:
245:. These were labeled as
226:was first described by
502:Caccothryptus arakawae
308:Caccothryptus arakawae
272:Caccothryptus arakawae
165:Caccothryptus arakawae
150:Caccothryptus arakawae
25:Caccothryptus arakawae
260:Harry George Champion
191:Caccothryptus testudo
430:, pp. 168, 171.
445:, pp. 171–173.
295:Fullmetal Alchemist
205:Fullmetal Alchemist
564:
563:
494:Taxon identifiers
292:, the creator of
198:, the creator of
172:in the subfamily
161:
160:
16:Species of beetle
594:
557:
556:
544:
543:
531:
530:
521:
520:
519:
489:
483:
466:(739): 168–184.
446:
440:
431:
425:
419:
413:
404:
403:
401:
399:
383:
326:and long, white
168:is a species of
152:
132:C. arakawae
49:
48:
35:
21:
602:
601:
597:
596:
595:
593:
592:
591:
587:Beetles of Asia
567:
566:
565:
560:
552:
547:
539:
534:
526:
524:
515:
514:
509:
496:
486:
459:
455:
450:
449:
441:
434:
426:
422:
414:
407:
397:
395:
385:
384:
380:
375:
346:on the female.
312:elytral lengths
304:
264:Keita Matsumoto
214:
182:Keita Matsumoto
157:
156:Matsumoto, 2021
154:
148:
135:
43:
17:
12:
11:
5:
600:
598:
590:
589:
584:
579:
569:
568:
562:
561:
559:
558:
545:
532:
522:
506:
504:
498:
497:
492:
485:
484:
456:
454:
451:
448:
447:
443:Matsumoto 2021
432:
428:Matsumoto 2021
420:
418:, p. 168.
416:Matsumoto 2021
405:
377:
376:
374:
371:
303:
300:
290:Hiromu Arakawa
213:
210:
196:Hiromu Arakawa
159:
158:
155:
144:
143:
137:
136:
129:
127:
123:
122:
115:
111:
110:
105:
101:
100:
95:
91:
90:
85:
81:
80:
75:
71:
70:
65:
61:
60:
55:
51:
50:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
599:
588:
585:
583:
580:
578:
575:
574:
572:
555:
550:
546:
542:
537:
533:
529:
523:
518:
512:
508:
507:
505:
503:
499:
495:
490:
481:
477:
473:
469:
465:
464:
458:
457:
452:
444:
439:
437:
433:
429:
424:
421:
417:
412:
410:
406:
394:
393:
388:
382:
379:
372:
370:
368:
364:
360:
356:
352:
347:
345:
341:
337:
333:
329:
325:
321:
317:
313:
309:
301:
299:
297:
296:
291:
287:
283:
280:
277:
273:
269:
265:
261:
257:
256:
251:
250:
244:
240:
236:
235:Caccothryptus
231:
229:
225:
224:
223:Caccothryptus
219:
211:
209:
208:
206:
201:
197:
193:
192:
187:
183:
179:
178:Caccothryptus
175:
171:
167:
166:
153:
151:
145:
142:
141:Binomial name
138:
134:
133:
128:
125:
124:
121:
120:
119:Caccothryptus
116:
113:
112:
109:
106:
103:
102:
99:
96:
93:
92:
89:
86:
83:
82:
79:
76:
73:
72:
69:
66:
63:
62:
59:
56:
53:
52:
47:
42:
38:
34:
29:
26:
22:
19:
501:
461:
453:Bibliography
423:
396:. Retrieved
390:
381:
354:
350:
348:
336:tarsal claws
319:
307:
305:
293:
285:
281:
278:
271:
254:
248:
234:
232:
221:
215:
203:
190:
177:
164:
163:
162:
149:
147:
131:
130:
118:
24:
18:
577:Limnichidae
359:median lobe
355:C. arakawae
320:C. arakawae
316:prothoraces
302:Description
282:C. arakawae
233:A group of
228:David Sharp
218:Limnichidae
174:Limnichinae
108:Limnichidae
571:Categories
517:Q127204631
373:References
367:ovipositor
351:C. testudo
286:C. testudo
255:C. testudo
98:Coleoptera
78:Arthropoda
480:2118-9773
276:paratypes
239:Kathmandu
186:Kathmandu
126:Species:
64:Kingdom:
58:Eukaryota
541:11160488
525:BioLib:
511:Wikidata
344:ventrite
340:pronotum
268:holotype
249:ripicola
212:Taxonomy
104:Family:
74:Phylum:
68:Animalia
54:Domain:
528:1833955
392:ZooBank
202:series
114:Genus:
94:Order:
88:Insecta
84:Class:
478:
398:5 July
363:dorsum
549:Plazi
332:tarsi
328:setae
324:tibia
200:manga
536:GBIF
476:ISSN
400:2024
306:The
270:for
252:and
216:The
468:doi
357:'s
247:C.
573::
551::
538::
513::
474:.
435:^
408:^
389:.
369:.
298:.
482:.
470::
402:.
279:.
207:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
