1691:
1696:
133:
115:
1719:
467:
435:
would presumably have resembled other oviraptorosaurs, which were feathered, bird-like dinosaurs with beaked skulls, long necks, and long limbs. Recent studies suggest it was one of the more primitive members of the
Caenagnathidae.
426:
is the smallest known oviraptorosaur and one of the smallest non-avian dinosaurs. The jaw fragments are only a few centimetres long and total skull length has been estimated at 3 inches (7.6 cm). A 2010 estimate by
403:, making it the oldest known caenagnathoid. A second specimen was referred to the species, N 402/12457, a right dentary of a slightly smaller individual. Both individuals were adult. In 2015, new material of
779:
Currie, P.J.; Godfrey, S.J.; Nessov, L. (1994). "New caenagnathid (Dinosauria, Theropoda) specimens from the Upper
Cretaceous of North America and Asia".
1843:
980:
1784:
1863:
1858:
1712:
759:
853:
Sues, Hans-Dieter (1997). "On
Chirostenotes, a Late Cretaceous oviraptorosaur (Dinosauria: Theropoda) from western North America".
1848:
1690:
973:
132:
1853:
1695:
353:
369:
813:(Dinosauria: Theropoda: Oviraptorosauria) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan".
966:
535:
447:
was originally assigned to the
Caenagnathidae. It was even included in the definition of that clade published by
1722:
1077:
596:
889:"A New Large-Bodied Oviraptorosaurian Theropod Dinosaur from the Latest Cretaceous of Western North America"
365:
875:
1838:
1739:
1810:
1362:
660:
276:
1771:
1131:
900:
1253:
1104:
511:
388:
384:
127:
1815:
357:
1797:
1281:
1246:
928:
407:
was described. From the same site as the holotype, the material includes various vertebrae, a
1802:
1466:
1199:
1177:
1143:
989:
918:
908:
822:
788:
452:
448:
349:
319:
249:
114:
1391:
1304:
1288:
1062:
487:
476:
456:
428:
326:
904:
1659:
1645:
1429:
1314:
1239:
1224:
949:
923:
888:
497:
361:
316:
216:
873:
Dinosaurs: The Most
Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,
1832:
1631:
1609:
1550:
1534:
1481:
1355:
1341:
1265:
730:
72:
34:
360:. The generic name is a combination of a reference to the species' placement in the
1602:
1558:
1516:
1459:
1402:
1376:
1046:
342:
229:
826:
913:
451:. It has been suggested however, that it might have a more basal position in the
431:
gave it a length of 2 feet (0.61 m) and a weight of 3 pounds (1.4 kg).
1652:
1585:
1523:
1445:
1438:
1274:
1116:
643:
47:
1666:
1617:
1494:
1418:
1348:
1334:
1327:
1028:
1019:
400:
330:
190:
92:
57:
1762:
1624:
1573:
1566:
1501:
1369:
1231:
1089:
1037:
466:
396:
203:
144:
97:
41:
932:
1756:
1543:
1508:
1383:
1217:
1013:
392:
373:
322:
177:
164:
87:
82:
67:
62:
52:
30:
1789:
1411:
408:
380:
102:
77:
1007:
154:
1733:
887:
Lamanna, M. C.; Sues, H. D.; Schachner, E. R.; Lyson, T. R. (2014).
792:
474:
The below cladogram is based on that resolved in the description of
958:
459:. In 2015 after the description of new material, it was found that
1638:
1594:
465:
412:
1776:
312:
1737:
1000:
962:
804:
802:
809:
Sues, H.-D.; Averianov, A. (2015). "New material of
1746:
1584:
1533:
1480:
1428:
1401:
1313:
1302:
1264:
1197:
1142:
1115:
1088:
1061:
364:, with Asia, the continent of its provenance. The
470:Life restoration with purely hypothetical ecology
463:could confidently be referred to Caenagnathidae.
383:of the lower jaws. The specimen was found near
974:
8:
1734:
1486:
1319:
1310:
1209:
1205:
1194:
1121:
1094:
1067:
1058:
997:
981:
967:
959:
113:
20:
922:
912:
771:
841:The Princeton Field Guide to Dinosaurs
7:
843:, Princeton University Press p. 152
760:Timeline of oviraptorosaur research
635:
612:
575:
551:
527:
503:
493:
483:
348:was named and described in 1994 by
855:Journal of Vertebrate Paleontology
781:Canadian Journal of Earth Sciences
14:
1844:Late Cretaceous dinosaurs of Asia
1718:
1717:
1694:
1689:
131:
1864:Taxa named by Philip J. Currie
370:Gerbert Genrikhovich Martinson
122:Illustration of the lower jaw
1:
1859:Fossil taxa described in 1994
827:10.1016/j.cretres.2014.12.001
914:10.1371/journal.pone.0092022
871:Holtz, Thomas R. Jr. (2010)
1880:
16:Extinct species of reptile
1707:
1687:
1489:
1322:
1212:
1208:
1193:
1174:
1124:
1097:
1070:
1057:
996:
811:Caenagnathasia martinsoni
728:
657:
640:
633:
617:
610:
593:
580:
573:
560:Caenagnathasia martinsoni
556:
549:
536:Gigantoraptor erlianensis
532:
525:
508:
501:
491:
346:Caenagnathasia martinsoni
289:Caenagnathasia martinsoni
282:
275:
128:Scientific classification
126:
121:
112:
23:
582:Alberta dentary morph 3
372:. The genus is based on
471:
1849:Fossils of Uzbekistan
1811:Paleobiology Database
876:Winter 2010 Appendix.
661:Caenagnathus collinsi
597:Leptorhynchos gaddisi
469:
787:(10β11): 2255β2272.
1254:Scansoriopterygidae
905:2014PLoSO...992022L
815:Cretaceous Research
1854:Bissekty Formation
839:Paul, G.S., 2010,
512:Microvenator celer
472:
389:Bissekty Formation
379:, a pair of fused
311:('recent jaw from
268:C. martinsoni
1826:
1825:
1798:Open Tree of Life
1740:Taxon identifiers
1731:
1730:
1703:
1702:
1685:
1684:
1681:
1680:
1677:
1676:
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1297:
1282:Similicaudipteryx
1247:Protarchaeopteryx
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1170:
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954:at The Dinosauria
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455:, outside of the
387:in layers of the
320:oviraptorosaurian
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1720:
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1487:
1467:Epichirostenotes
1320:
1311:
1210:
1206:
1200:Oviraptorosauria
1195:
1178:Oviraptorosauria
1144:Oviraptorosauria
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1095:
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1051:
998:
990:Oviraptorosauria
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453:Oviraptorosauria
449:Hans-Dieter Sues
391:, dating to the
377:N 401/12457
350:Philip J. Currie
291:
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136:
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44:
29:Temporal range:
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1727:
1699:
1673:
1580:
1529:
1472:
1424:
1397:
1392:Ojoraptorsaurus
1307:
1305:Caenagnathoidea
1294:
1289:Xingtianosaurus
1260:
1202:
1181:
1158:
1138:
1111:
1084:
1078:Avemetatarsalia
1063:Avemetatarsalia
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793:10.1139/e93-196
778:
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756:
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718:
709:
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682:
673:
619:"Caenagnathus"
488:Caenagnathoidea
457:Caenagnathoidea
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429:Gregory S. Paul
421:
354:Stephen Godfrey
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327:Late Cretaceous
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1824:
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1821:
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1807:
1794:
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1768:
1752:
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1748:Caenagnathasia
1744:
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1705:
1704:
1701:
1700:
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1683:
1682:
1679:
1678:
1675:
1674:
1672:
1671:
1663:
1656:
1649:
1646:Machairasaurus
1642:
1635:
1628:
1621:
1614:
1606:
1599:
1590:
1588:
1582:
1581:
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1578:
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1505:
1498:
1490:
1484:
1478:
1477:
1474:
1473:
1471:
1470:
1463:
1456:
1453:Caenagnathasia
1449:
1442:
1434:
1432:
1430:Caenagnathinae
1426:
1425:
1423:
1422:
1415:
1407:
1405:
1399:
1398:
1396:
1395:
1388:
1380:
1373:
1366:
1359:
1352:
1345:
1338:
1331:
1323:
1317:
1315:Caenagnathidae
1308:
1303:
1300:
1299:
1296:
1295:
1293:
1292:
1285:
1278:
1270:
1268:
1262:
1261:
1259:
1258:
1250:
1243:
1240:Ningyuansaurus
1236:
1228:
1225:Incisivosaurus
1221:
1213:
1203:
1198:
1191:
1190:
1187:
1186:
1183:
1182:
1175:
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1016:
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952:C. martinsonii
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440:Classification
438:
433:Caenagnathasia
424:Caenagnathasia
420:
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405:Caenagnathasia
362:Caenagnathidae
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315:') is a small
308:Caenagnathasia
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1551:Corythoraptor
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1535:Oviraptorinae
1532:
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1503:
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1482:Oviraptoridae
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1360:
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1342:Chirostenotes
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139:
134:
129:
125:
120:
116:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
43:
40:93β70.6
36:
35:Maastrichtian
32:
26:
22:
19:
1747:
1665:
1658:
1651:
1644:
1637:
1630:
1623:
1616:
1608:
1603:Conchoraptor
1601:
1593:
1572:
1565:
1559:Huanansaurus
1557:
1549:
1542:
1522:
1517:Tongtianlong
1515:
1507:
1500:
1493:
1465:
1460:Caenagnathus
1458:
1452:
1451:
1444:
1437:
1417:
1410:
1403:Elmisaurinae
1390:
1382:
1377:Microvenator
1375:
1368:
1361:
1354:
1347:
1340:
1333:
1326:
1287:
1280:
1273:
1252:
1245:
1238:
1230:
1223:
1216:
1176:
1047:Pennaraptora
1042:
1033:
1024:
951:
896:
892:
882:
872:
867:
858:
854:
848:
840:
835:
818:
814:
810:
784:
780:
774:
729:
659:
658:
642:
641:
620:
618:
595:
594:
581:
559:
558:
557:
534:
533:
510:
509:
475:
473:
460:
444:
443:
432:
423:
422:
404:
399:, around 90
376:
345:
343:type species
340:
317:caenagnathid
307:
306:
305:
296:
288:
283:
267:
266:
252:
243:
242:
230:Elmisaurinae
197:
184:
171:
24:
18:
1653:Nemegtomaia
1586:Heyuanninae
1446:Apatoraptor
1275:Caudipteryx
1132:Maniraptora
1117:Maniraptora
644:Anzu wyliei
419:Description
385:Dzharakuduk
223:Subfamily:
1833:Categories
1711:See also:
1667:Shixinggia
1618:Gobiraptor
1495:Luoyanggia
1419:Elmisaurus
1349:Eoneophron
1335:Beibeilong
1328:Anomalipes
1153:see belowβ
1029:Dinosauria
1020:Sauropsida
766:References
621:sternbergi
331:Uzbekistan
191:Saurischia
178:Dinosauria
1625:Heyuannia
1574:Rinchenia
1567:Oviraptor
1502:Nankangia
1370:Hagryphus
1105:Theropoda
1090:Theropoda
1038:Theropoda
1006:Kingdom:
950:Image of
821:: 50β59.
397:Coniacian
381:dentaries
358:Lev Nesov
337:Discovery
325:from the
261:Species:
204:Theropoda
151:Kingdom:
145:Eukaryota
1777:47458439
1757:Wikidata
1723:Category
1713:Timeline
1544:Citipati
1509:Nomingia
1384:Nomingia
1218:Avimimus
1014:Chordata
1012:Phylum:
1008:Animalia
933:24647078
893:PLOS ONE
754:See also
411:, and a
393:Turonian
374:holotype
368:honours
323:theropod
210:Family:
165:Chordata
161:Phylum:
155:Animalia
141:Domain:
31:Turonian
1803:4126525
1790:4822720
1763:Q134763
1412:Citipes
1018:Class:
924:3960162
901:Bibcode
409:dentary
295:Currie
286:†
265:†
236:Genus:
1660:Oksoko
1524:Yulong
931:
921:
299:, 1994
297:et al.
255:, 1994
253:et al.
250:Currie
1816:54542
1639:Khaan
1595:Banji
1043:Clade
1034:Clade
1025:Clade
413:femur
198:Clade
185:Clade
172:Clade
1785:GBIF
1439:Anzu
1130:see
1103:see
1076:see
929:PMID
477:Anzu
356:and
341:The
313:Asia
48:Preκ
1772:EoL
1232:Kol
919:PMC
909:doi
823:doi
789:doi
401:mya
329:of
1835::
1813::
1800::
1787::
1774::
1759::
1045::
1036::
1027::
927:.
917:.
907:.
895:.
891:.
859:17
857:.
819:54
817:.
801:^
785:30
783:.
480:.
415:.
352:,
333:.
200::
187::
174::
98:Pg
42:Ma
37:,
1670:?
1613:?
1598:?
1577:?
1562:?
1554:?
1512:?
1387:?
1257:?
1235:?
982:e
975:t
968:v
935:.
911::
903::
897:9
829:.
825::
795:.
791::
395:-
240:β
227:β
214:β
103:N
93:K
88:J
83:T
78:P
73:C
68:D
63:S
58:O
53:κ
33:-
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