Knowledge (XXG)

Carnosauria

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bases on their tails have anterior and posterior bone growth. The largest carnosaurs can reach up to 10 meters in length. The length of the body from the tail to the hip is between 54% and 62% of the total body length, and the length of the body from the head to the hip is between 38% and 46% of the
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Infante, P.; Canudo, José Ignacio; Ruiz Omeñaca, José Ignacio (2005). "Primera evidencia de dinosaurios terópodos en la Formación Mirambel (Barremiense inferior, Cretácico Inferior) en Castellote, Teruel" [First evidence of theropod dinosaurs from the Mirambel Formation (Lower Barremian, Lower
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Although the remains of other large theropods like tyrannosaurids bear evidence of fighting within their species and with other predators, the remains of allosauroids do not bear much evidence of injuries from theropod combat. Most notably, despite a good fossil record, allosauroid skulls lack the
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Multiple severe injuries have been found on allosauroid remains, which implies that allosauroids were frequently in dangerous situations and supports the hypothesis of an active, predatory lifestyle. Despite the multitude of injuries, only a few of those injuries show signs of infection. For those
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Allosauroids were present in both the northern and southern continents during the Jurassic and Early Cretaceous, but they were later displaced by the tyrannosauroids in North America and Asia during the Late Cretaceous. This is likely due to regional extinction events, which, along with increased
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was the only other taxon known to have that trait at the time. Neovenatorids were envisioned as the latest-surviving allosauroids, which were able to persist well into the Late Cretaceous due to their low profile and coelurosaur-like adaptations. Later studies supported this hypothesis, such as
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separated into the modern continents. By the Middle Jurassic period, tetanurans had spread to every continent and diverged into the allosauroids and the coelurosaurs. Allosauroids first appeared in the Middle Jurassic period and were the first giant taxa (weighing more than 2 tons) in theropod
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bone, and a short connection between the braincase and the palate. Allosauroid skulls are about 2.5 to 3 times longer as they are tall. Their narrow skull along with their serrated teeth allow carnosaurs to better slice flesh off of their prey. Carnosaur teeth are flat and have equally-sized
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across all sizes, which is found to be between 37% and 58% of the femoral length anterior to the hip. Other similarities across all carnosaurs include the structure of their hind limb and pelvis. The pelvis in particular is thought to be designed to reduce stress regardless of body size. In
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Fernandes de Azevedo, R. P.; Simbras, F. M.; Furtado, M. R.; Candeiro, C. R. A.; Bergqvist, L. P. (2013). "First Brazilian carcharodontosaurid and other new theropod dinosaur fossils from the Campanian–Maastrichtian Presidente Prudente Formation, São Paulo State, southeastern Brazil".
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was able to quickly stop any infection from spreading to the rest of the body. This type of immune response is similar to modern reptilian immune responses; reptiles secrete fibrin near infected areas and localize the infection before it can spread via the bloodstream.
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presented below illustrates the interrelationships between the four major groups (or families) of carnosaurs. It is a simplified version of the tree presented in the 2012 analysis by Carrano, Benson and Sampson after they excluded three "wildcard" taxa
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The following family tree illustrates the position of Carnosauria within Theropoda. It is a simplified version of the tree presented in a synthesis of the relationships of the major theropod groups based on various studies conducted in the 2010s.
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in 2021. This publication is also the origin of the hypothesis that several "compsognathids" from Europe may have been juvenile carnosaurs. The results of this analysis differ from those of Rauhut and Pol in that Cau finds Megalosauroidea to be
969:, did. During the Cretaceous, some carnosaurs grew to sizes similar to those of the largest tyrannosaurids. These large carnosaurs lived in the same time period as the other large theropods found in the upper Morrison and Tendaguru formations. 3524:, similar to non-avian reptiles, which means they require fewer nutrients in order to survive. A lower nutrient requirement means allosauroids do not need to undertake frequent hunts, which lowers their risk of sustaining traumatic injuries. 1145:, among others, have followed this node-based definition. Depending on the study, Carnosauria and Allosauroidea are sometimes considered synonymous. In such cases, several researchers have elected to use Allosauroidea over Carnosauria. 4165:
Paulina-Carabajal, Ariana & Currie, Philip. (2012). New information on the braincase and endocast of Sinraptor dongi (Theropoda: Allosauroidea): Ethmoidal region, endocranial anatomy and pneumaticity. Vertebrata PalAsiatica. 50.
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Méndez, Ariel H.; Novas, Fernando E.; Iori, Fabiano V. (May–June 2012). "Premier enregistrement de Megaraptora (Theropoda, Neovenatoridae) au Brésil" [First record of Megaraptora (Theropoda, Neovenatoridae) from Brazil].
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due to drought, the frequency of finding allosauroid remains in groups supports the social animal theory. As social animals, allosauroids would share the burden of hunting, allowing injured members of the pack to recover faster.
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have been interpreted by some authors as quill knobs theorized to have supported primitive quills; however this interpretation has been questioned, and they have been suggested to represent traces of ligaments instead.
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Hendrickx, Christophe; Bell, Phil R.; Pittman, Michael; Milner, Andrew R. C.; Cuesta, Elena; O'Connor, Jingmai; Loewen, Mark; Currie, Philip J.; Mateus, Octávio; Kaye, Thomas G.; Delcourt, Rafael (June 2022).
1748:. These genera were allied with the other neovenatorids on the basis of several features spread out throughout the skeleton, particularly the large amount of pneumatization present. The pneumatic ilium of 5336:
Benson, Roger B. J.; Carrano, Matthew T.; Brusatte, Stephen L. (January 2010). "A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic".
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distinctive face-biting wounds that are common in tyrannosaurid skulls, leaving open the question of if allosauroids engaged in interspecies and intraspecies fighting. Remains of the allosauroid
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on both edges. The flat side of the tooth face the sides of the skull, while the edges align on the same plane as the skull. From analyzing the skull of different carnosaurs, the volume of the
960:. They also have 3 fingers per hand, with the second and third digit being approximately equal in length. The femur is larger than the tibia. Another defining feature of carnosaurs is that the 3714:
Delcourt, R.; Grillo, O.N. (2017). "Carcharodontosaurids remained extinct in the Campanian-Maastrichtian: Reassessment of a fragmentary maxilla from Presidente Prudente Formation, Brazil".
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Bates, Karl T.; Benson, Roger B. J.; Falkingham, Peter L. (1 June 2012). "A computational analysis of locomotor anatomy and body mass evolution in Allosauroidea (Dinosauria: Theropoda)".
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Bybee, Paul J.; Lee, Andrew H.; Lamm, Ellen-Thérèse (2006). "Sizing the Jurassic theropod dinosaur Allosaurus: Assessing growth strategy and evolution of ontogenetic scaling of limbs".
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Mateus, Octávio. (1997). Lourinhanosaurus antunesi, A New Upper Jurassic Allosauroid (Dinosauria: Theropoda) from Lourinhã, Portugal. Mémorias da Academia Ciêncas de Lisboa. 37.
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Benson, R.B.J.; Carrano, M.T; Brusatte, S.L. (2010). "A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic".
3002:. This hypothesis has not been universally accepted, and it notably conflicts with Cau's 2021 publication by finding Megalosauroidea as monophyletic and the sister taxon of 5280: 5040: 984:
In addition to body similarities, most carnosaurs, especially most allosauroids are also united by certain skull features. Some of the defining ones include a smaller
3620:"Probable basal allosauroid from the early Middle Jurassic Cañadón Asfalto Formation of Argentina highlights phylogenetic uncertainty in tetanuran theropod dinosaurs" 1815:
The cladogram below follows a 2016 analysis by Sebastián Apesteguía, Nathan D. Smith, Rubén Juarez Valieri, and Peter J. Makovicky based on the dataset of Carrano
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In 2024, Andrea Cau published a paper which presented an analysis of theropod ontogeny which suggested that several theropods that were traditionally considered
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longus (CFL) muscle that allowed them to flex theirs. Larger carnosaurs are found to have a lower CFL muscle-to-body-mass proportion that smaller carnosaurs.
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Sereno, P. C.; Dutheil, D. B.; Iarochene, M.; Larsson, H. C. E.; Lyon, G. H.; Magwene, P. M.; Sidor, C. A.; Varricchio, D. J.; Wilson, J. A. (17 May 1996).
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are also often found in groups, which could imply the existence of social behavior. While there are alternative explanations for the groupings, like
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Carrano, Benson & Sampson large study of tetanuran relationships in 2012, and Zanno & Makovicky description of the newly discovered theropod
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in 1997. Sereno was the first to provide a stem-based definition for the Allosauroidea in 1998, defining the clade as "All neotetanurans closer to
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and a few other taxa as members of the Neovenatoridae. This would make neovenatorids the latest-surviving allosauroids; at least one megaraptoran,
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particular, the way the femur is inclined reduces the bending and torsion stress. Furthermore, like other animals with tails, carnosaurs possess a
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Canale, Juan Ignacio; Novas, Fernando Emilio; Salgado, Leonardo; Coria, Rodolfo Aníbal (December 1, 2015). "Cranial ontogenetic variation in
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species isolation through the severing of land connections between the continents, differentiated many dinosaurs in the Late Cretaceous.
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Foth, Christian; Evers, Serjoscha W.; Pabst, Ben; Mateus, Octávio; Flisch, Alexander; Patthey, Mike; Rauhut, Oliver W.M. (12 May 2015).
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and megaraptorans, which suggests that the latter were not carnosaurs or allosauroids. As a result of these findings, and the fact that
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with respect to Allosauroidea. Because the authors amended the definition of Allosauroidea to include all theropods that are closer to
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scales due to their similarity and the fact that non-theropod remains were discovered associated with the tail of this particular
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Carrano, Matthew T.; Benson, Roger B. J.; Sampson, Scott D. (1 June 2012). "The phylogeny of Tetanurae (Dinosauria: Theropoda)".
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Phylogenetic studies conducted by Benson, Carrano and Brusatte (2010) and Carrano, Benson and Sampson (2012) recovered the group
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Subsequent analyses have contradicted the above hypothesis. Novas and colleagues conducted an analysis in 2012 which found that
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total body length. Carnosaurs scaled their limbs relative to their body in a way similar to how other large theropods, like the
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preserves scutes on the underside of the tail, as well as scutes on the feet along with small scales. A series of knobs on the
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injuries that did become infected, the infections were usually local to the site of the injury, implying that the allosauroid
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Holtz, T. R., Jr. and Osmólska H. 2004. Saurischia; pp. 21–24 in D. B. Weishampel, P. Dodson, and H. Osmólska (eds.),
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Carrano, Matthew T.; Benson, Roger B. J.; Sampson, Scott D. (2012). "The phylogeny of Tetanurae (Dinosauria: Theropoda)".
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Novas, Fernando E. (2013). "Evolution of the carnivorous dinosaurs during the Cretaceous: The evidence from Patagonia".
3006:, a grouping which includes both carnosaurs (or allosauroids) and coelurosaurs. Notably, this analysis also treats the 1335: 38: 9025: 6780: 1130: 889:
Carnosauria has traditionally been used as a dumping ground for all large theropods. Even non-dinosaurs, such as the
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Hendrickx, C.; Hartman, S.A.; Mateus, O. (2015). "An overview of non-avian theropod discoveries and classification".
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subsequently found that megaraptorans lacked most of the key features in the hands of derived coelurosaurs including
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and the sister-taxon of Allosauroidea within Carnosauria. An abbreviated version of this phylogeny is shown below.
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The relationship between allosauroids and megalosauroids was also supported by a provisional analysis published by
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itself is the only uncontroversial neovenatorid, the family Neovenatoridae sees little use in recent publications.
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The injuries were also found to be mostly healed. This healing may indicate that allosauroids had an intermediate
3153: 915: 4490:(Theropoda, Dinosauria) from the Lower Cretaceous Antlers Formation (Lower Cretaceous, Aptian) of Oklahoma, USA" 2219:
was found to fall within Allosauroidea. A cladogram displaying the relationships they recovered is shown below.
5513: 3342: 997: 8259: 2861: 2348: 1674:. Different clades have been recovered by different authors, and a scientific consensus has yet to emerge. 9587: 5313: 5418: 4857: 3595: 1904: 1305: 5397: 4921: 4591:"A New Megaraptoran Dinosaur (Dinosauria, Theropoda, Megaraptoridae) from the Late Cretaceous of Patagonia" 3236: 2632: 2399: 1844: 1543: 9613: 3868: 3017: 847:
Carnosaurs first appeared in the Middle Jurassic, around 176 mya. The last definite known carnosaurs, the
4389:"A new carnosaur (Dinosauria, Theropoda) from the Upper Jurassic of Xinjiang, People's Republic of China" 855:
epoch of the Cretaceous, roughly 90 mya; reportedly later remains of carcharodontosaurids, from the late
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during the Early Cretaceous period; various dinosaur footprints found in Apulia support this theory.
3521: 3127: 2628: 2395: 2147:. Instead, their hands retain a number of primitive characteristics seen in basal tetanurans such as 1840: 1682: 1539: 491: 3873: 3845: 9688: 7997: 7784: 3539: 2441: 1695:, a group of theropods with controversial affinities. Other studies recover megaraptorans as basal 335: 8266: 3503: 2042: 9452: 7973: 7833: 7689: 7629: 7064: 6974: 6857: 5721: 5411: 5370: 5318: 5258: 5215:(Dinosauria: Theropoda) and the probable role of heterochrony in carcharodontosaurid evolution". 4858:"Megaraptorans as members of an unexpected evolutionary radiation of tyrant-reptiles in Gondwana" 4799:"An Unusual New Theropod with a Didactyl Manus from the Upper Cretaceous of Patagonia, Argentina" 4727: 4552: 4118: 4060: 3968: 3925: 3827: 3689: 3231: 2857: 2344: 2216: 2197: 750: 225: 80: 1670:
The composition of the clade Carnosauria has been controversial among scientists since at least
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Powell, a giant maniraptoran (Dinosauria, Theropoda) from the Late Cretaceous of Argentina."
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Rauhut, Oliver W. M.; Foth, Christian (2020), Foth, Christian; Rauhut, Oliver W. M. (eds.),
4245: 4190: 4146: 4102: 4052: 4000: 3952: 3917: 3819: 3759: 3723: 3647: 3639: 3336: 3148: 1515: 1331: 1138: 1098: 448: 205: 76: 9501: 9239: 9202: 9134: 8989: 8982: 8810: 8752: 8725: 8715: 8675: 8510: 8369: 8353: 8171: 8143: 8121: 8008: 7853: 7560: 7334: 7217: 7195: 7116: 6945: 6922: 6915: 6893: 6879: 6872: 6833: 6684: 6531: 6458: 6447: 6383: 6271: 6158: 5735: 5644: 5498: 5473: 3513: 3107: 3048: 2754: 2605: 2240: 2193: 2128: 2016: 1707: 1358: 1267: 1207: 1184: 1134: 1033: 966: 802: 635: 399: 366: 100: 96: 5281:"Phylogeny of Allosauroidea (Dinosauria: Theropoda): Comparative analysis and resolution" 817:. Other researchers have found Allosauroidea and Megalosauroidea to be unrelated groups. 5350: 5296: 5228: 5067: 4814: 4797:
Sebastián Apesteguía; Nathan D. Smith; Rubén Juárez Valieri; Peter J. Makovicky (2016).
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and suggests that the postcranial remains of this taxon belong to a carnosaur (possibly
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were consistently found to be close relatives of each other; this was also the case for
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unrelated to carcharodontosaurs. Other theropods with uncertain affinities such as
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Carnosaurs share certain distinctive features, one of which is a triangular-shaped
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used a node-based definition in his 2007 study which defined the Allosauroidea as
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region (the “heel” of the Italian peninsula), which was connected to Africa by a
881:, discovered in rocks dated to 75-70 mya, might potentially also be a carnosaur. 9645: 9534: 9515: 9445: 9395: 9381: 9323: 9264: 9209: 9086: 8880: 8772: 8741: 8658: 8644: 8376: 8324: 8192: 8114: 8077: 7959: 7931: 7884: 7804: 7682: 7390: 7363: 7263: 7210: 7146: 7125: 7098: 6849: 6662: 6655: 6641: 6467: 6314: 6305: 6207: 6178: 6108: 6073: 6066: 5866: 5844: 5837: 5796: 5788: 5764: 5749: 5728: 5636: 5628: 4330:
and Its Implications for the Phylogeny of Allosauroidea (Dinosauria: Theropoda)"
4292: 4080:"Predatory Dinosaurs from the Sahara and Late Cretaceous Faunal Differentiation" 3569: 2986: 2980: 2969: 2816: 2165: 2143: 2037: 1791: 1692: 1405: 1102: 1094: 907: 872: 860: 775: 735: 715: 675: 544: 113: 65: 4856:
F. E. Novas; F. L. Agnolín; M. D. Ezcurra; J. I. Canale; J. D. Porfiri (2012).
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Holtz, Thomas (December 1998). "A New Phylogeny of the Carnivorous Dinosaurs".
3643: 3026:). An abbreviated version of the cladogram from that analysis is shown below. 9473: 9402: 9347: 8968: 8783: 8601: 8587: 8572: 8524: 8458: 8384: 8206: 8199: 8185: 8091: 8024: 7952: 7916: 7869: 7811: 7797: 7764: 7756: 7749: 7730: 7553: 7518: 7306: 6907: 6698: 6648: 6585: 6556: 6511: 6489: 6482: 6474: 6398: 6294: 6253: 6246: 6238: 6222: 6142: 6013: 5889: 5742: 5605: 5464: 5358: 5304: 5236: 4907: 4540: 4280: 4225: 3530: 3394: 3366: 3022: 2685: 2656: 2534: 2149: 2119: 2113: 2022: 1987: 1809: 1796: 1687: 1112: 1107: 1025: 936: 927: 911: 890: 821: 787: 564: 554: 533: 377: 355: 288: 217: 158: 123: 9636: 5254: 5026: 4723: 4624: 4257: 4202: 9494: 9487: 9187: 9180: 8853: 8792: 8446: 8164: 7710: 7614: 7349: 6934: 6819: 6806: 6733: 6669: 6592: 6571: 6328: 6191: 5999: 5986: 5904: 5756: 5575: 5562: 5525: 5434: 4947:, a tyrannosauroid theropod from the Wealden Supergroup of southern England" 3585: 3253: 3034: 2992: 2550: 2201: 2189: 2162: 1497: 1254: 1224: 1125: 1045: 931: 899: 814: 705: 685: 438: 301: 242: 199: 163: 107: 5366: 5164: 4982: 4842: 4780: 4642: 4548: 4373: 4265: 4210: 4012: 3964: 3661: 1073: 948: 5392: 4114: 17: 9630: 9078: 8503: 8496: 8468: 8361: 7861: 7242: 6578: 6405: 6166: 5803: 5482: 5458: 5018: 4497:
Publications Scientifiques du Muséum national d'Histoire naturelle, Paris
3205: 3012: 2137: 1955: 1801: 1701: 1037: 988: 985: 852: 783: 779: 665: 275: 262: 175: 153: 148: 133: 128: 118: 5180:"Head-biting behavior in theropod dinosaurs: Paleopathological evidence" 5084: 4963: 4249: 8636: 8398: 8128: 8031: 7356: 6677: 5245: 5002: 4916: 4771: 4746: 4679: 3956: 3556: 3400: 1993: 1603: 809:
with respect to Allosauroidea), and thus including the majority of non-
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While Carnosauria was historically considered largely synonymous with
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Ortega, Francisco; Escaso, Fernando; Sanz, José L. (September 2010).
3565: 864: 833: 252: 9607: 4415: 3921: 875:, which may or may not be carnosaurs, became extinct around 66 mya. 8135: 7876: 7818: 4881:"Phylogenetic relationships of the Cretaceous Gondwanan theropods 3502: 3311: 2974: 2117:
was closely related to carcharodontosaurids, simultaneously found
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may be juvenile allosauroids or megalosauroids. These included
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carnosaurs uniting some primitive members of the group such as
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Within Carnosauria, there is a slightly more exclusive clade,
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measuring 1-3 mm are known from the side of the torso and the
5403: 3991:
Sereno, P. C. (25 June 1999). "The Evolution of Dinosaurs".
5003:"A Unified Framework for Predatory Dinosaur Macroevolution" 4285:
The Evolution of Feathers: From Their Origin to the Present
3688:
Hutchinson, John R. (7 November 2005) . Smith, Dave (ed.).
2171:
Several recent analyses do not find a relationship between
71:"Carnosaur" redirects here. For the 1984 horror novel, see 1720:
Neovenatoridae, as formulated by these authors, contained
5105:
Atti della Società dei Naturalisti e Matematici di Modena
3676:
A Unified Framework for Predatory Dinosaur Macroevolution
922:) that were more closely related to birds were placed in 5279:
Brusatte, Stephen L.; Sereno, Paul C. (1 January 2008).
5101:"Comments on the Mesozoic theropod dinosaurs from Italy" 820:
Distinctive characteristics of carnosaurs include large
3555:
history of allosauroids closely follows the order that
1077:
Cladogram of Allosauroidea after Eddy and Clarke (2011)
42: 4589:
Coria, Rodolfo A.; Currie, Philip J. (20 July 2016).
3618:
Rauhut, Oliver W. M.; Pol, Diego (11 December 2019).
2161:
have been alternatively recovered as coelurosaurs or
4879:
Novas, F.E.; Rolando, A.M.A.; Agnolin, F.L. (2016).
4474:(2nd ed.), University of California Press, Berkeley. 3800:"The phylogeny of Tetanurae (Dinosauria: Theropoda)" 731: 721: 711: 701: 691: 681: 671: 661: 651: 641: 631: 621: 611: 581: 570: 560: 550: 540: 529: 518: 508: 497: 487: 477: 466: 455: 444: 434: 424: 413: 395: 384: 373: 362: 351: 9620: 9533: 9428: 9371: 9333: 9250: 9223: 9133: 9103: 9062: 8960: 8928: 8870: 8836: 8809: 8782: 8751: 8724: 8713: 8673: 8538: 8486: 8457: 8419: 8408: 8308: 8220: 8154: 8076: 8007: 7983: 7895: 7843: 7783: 7739: 7728: 7672: 7640: 7587: 7537: 7502: 7420: 7373: 7320: 7282: 7228: 7179: 7115: 7074: 7050: 7019: 6973: 6943: 6932: 6817: 6732: 6631: 6542: 6521: 6457: 6360: 6304: 6270: 6177: 6122: 6091: 6031: 5997: 5923: 5903: 5827: 5774: 5655: 5573: 5524: 5497: 918:as more primitive theropods. Others (such as the 797:, some recent studies have revived Carnosauria as 5314:20.500.11820/5f3e6d44-fea6-468d-81d3-769f8c2830dd 4792: 4790: 4514: 4512: 4510: 4324:Eddy, Drew R.; Clarke, Julia A. (21 March 2011). 5007:Bollettino della Società Paleontologica Italiana 4889:: the evidence afforded by their manual anatomy" 3895: 3893: 2192:. Their phylogenetic analysis found traditional 3694:University of California Museum of Paleontology 3560:history. Along with members of the superfamily 1763:in 2013, which they placed within Megaraptora. 4484:Currie, Philip J.; Carpenter, Kenneth (2000). 5419: 4936: 4934: 4572:PalArch's Journal of Vertebrate Palaeontology 4438:Annual Review of Earth and Planetary Sciences 8: 5178:Tanke, Darren H.; Currie, Philip J. (1998). 5039:: CS1 maint: DOI inactive as of July 2024 ( 3613: 3611: 4326:"New Information on the Cranial Anatomy of 934:sharing a more recent common ancestor with 26: 9608: 9339: 9256: 9231: 9115: 9111: 9070: 9059: 9046: 8957: 8925: 8757: 8721: 8681: 8550: 8546: 8431: 8427: 8416: 8316: 8305: 8292: 8083: 8073: 7989: 7901: 7789: 7736: 7646: 7599: 7595: 7545: 7510: 7499: 7486: 7417: 7326: 7234: 7187: 7056: 7031: 7027: 6955: 6951: 6940: 6825: 6814: 6801: 6729: 6548: 6539: 6368: 6183: 6134: 6130: 6043: 6039: 6005: 5994: 5981: 5780: 5663: 5581: 5570: 5557: 5503: 5494: 5442: 5426: 5412: 5404: 4996: 4994: 4992: 1093:, but it was given a formal definition by 598: 188: 87: 5312: 5244: 5195: 5154: 5144: 5083: 4972: 4962: 4915: 4832: 4822: 4770: 4678: 4632: 4614: 4363: 4353: 4150: 4137:Cretaceous) from Castellote, Terue]. 3872: 3651: 1787:taxon difficult to place with certainty. 1024:preserve skin impressions showing their 32:This is an accepted version of this page 4745:Zanno, L. E.; Makovicky, P. J. (2013). 4431:"The origin and evolution of dinosaurs" 3607: 902:. Most former carnosaurs (such as the 28: 5032: 1089:Allosauroidea was originally named by 930:analysis defines Carnosauria as those 5274: 5272: 5122: 5120: 5118: 4319: 4317: 4030: 4028: 4026: 4024: 4022: 3986: 3984: 3982: 867:, were later interpreted as those of 7: 1812:period, about 70 million years ago. 75:. For the 1993 film adaptation, see 5285:Journal of Systematic Palaeontology 4696:Journal of Systematic Palaeontology 4037:Journal of Systematic Palaeontology 3804:Journal of Systematic Palaeontology 3387: 3358: 3304: 3223: 3195: 3140: 3133: 3123: 3099: 3069: 3041: 3029: 2877: 2850: 2791: 2767: 2760: 2677: 2648: 2621: 2597: 2587: 2557: 2545: 2415: 2388: 2364: 2337: 2313: 2283: 2256: 2232: 2222: 2009: 1980: 1930: 1923: 1893: 1863: 1833: 1822: 1592: 1562: 1532: 1522: 1378: 1351: 1324: 1290: 1260: 1230: 1200: 1171: 1157: 952:Size comparison of seven carnosaurs 871:. The phylogenetically problematic 61:Extinct group of theropod dinosaurs 4396:Canadian Journal of Earth Sciences 3591:Cenomanian-Turonian boundary event 2182:In 2019, Rauhut and Pol described 525:Benson, Carrano and Brusatte, 2010 197:Four carnosaurs (top to bottom): 57: 9709:Taxa named by Friedrich von Huene 4943:"The osteology and affinities of 3690:"Tremble Before the Carnosauria!" 1004:ranges between 95 milliliters in 805:(which is sometimes recovered as 801:including both Allosauroidea and 9592: 9586: 9579: 9031: 9024: 9017: 9011: 8277: 8270: 8265: 8258: 8252: 7471: 7464: 7457: 6786: 6779: 6772: 6765: 5967: 5961: 5391: 4941:Naish, D.; Cau, A. (July 2022). 4387:Currie, P. J.; Zhao, X. (1993). 3419: 3399: 3370: 3341: 3319: 3257: 3235: 3152: 3079: 3051: 2907: 2887: 2860: 2704: 2660: 2631: 2567: 2445: 2425: 2398: 2347: 2293: 2266: 2041: 2021: 1992: 1903: 1873: 1843: 1622: 1602: 1572: 1542: 1408: 1388: 1361: 1334: 1304: 1270: 1240: 1210: 1183: 972:Carnosaurs maintained a similar 229: 3782:Rev. Mus. Argentino Cienc. Nat. 1032:skin impressions showing small 4458:10.1146/annurev.earth.25.1.435 3507:Allosauroidea skull comparison 1133:, and all of its descendants. 1097:and Zhao, and later used as a 840:) being longer than the shin ( 1: 9704:Fossil taxa described in 1920 5076:10.1016/j.cretres.2013.04.001 4005:10.1126/science.284.5423.2137 3764:10.1016/j.cretres.2012.06.004 3728:10.1016/j.cretres.2017.09.008 1752:was particularly notable, as 1717:are also sometimes included. 603:Uncertain or possible members 5217:Paläontologische Zeitschrift 4824:10.1371/journal.pone.0157793 4716:10.1080/14772019.2011.630927 4616:10.1371/journal.pone.0157973 4355:10.1371/journal.pone.0017932 4107:10.1126/science.272.5264.986 4057:10.1080/14772019.2011.630927 3824:10.1080/14772019.2011.630927 3776:Novas and Agnolin, (2004). " 1800:, lived near the end of the 4293:10.1007/978-3-030-27223-4_3 1131:most recent common ancestor 79:. For the film series, see 9725: 9694:Aalenian first appearances 4896:Memoirs of Museum Victoria 4671:10.1016/j.crpv.2011.12.007 4488:Acrocanthosaurus atokensis 4328:Acrocanthosaurus atokensis 3644:10.1038/s41598-019-53672-7 813:members of theropod clade 70: 63: 9576: 9342: 9259: 9234: 9118: 9114: 9073: 9058: 9045: 9009: 8956: 8924: 8760: 8684: 8553: 8549: 8434: 8430: 8319: 8304: 8291: 8250: 8086: 8072: 7992: 7904: 7792: 7649: 7602: 7598: 7548: 7513: 7498: 7485: 7454: 7416: 7329: 7237: 7190: 7059: 7034: 7030: 6958: 6954: 6828: 6813: 6800: 6762: 6728: 6551: 6371: 6186: 6137: 6133: 6046: 6042: 6008: 5993: 5980: 5958: 5783: 5666: 5584: 5569: 5556: 5506: 5493: 5441: 5359:10.1007/s00114-009-0614-x 5305:10.1017/S1477201907002404 5237:10.1007/s12542-014-0251-3 4908:10.24199/j.mmv.2016.74.05 4541:10.1007/s00114-009-0614-x 3798:Matthew, Carrano (2012). 3499:Paleobiology and behavior 3414: 3392: 3385: 3363: 3356: 3334: 3309: 3302: 3250: 3228: 3221: 3200: 3193: 3167: 3145: 3138: 3131: 3121: 3104: 3097: 3074: 3067: 3046: 3039: 2902: 2882: 2875: 2855: 2848: 2813: 2796: 2789: 2772: 2765: 2758: 2699: 2682: 2675: 2653: 2646: 2626: 2619: 2602: 2595: 2585: 2562: 2555: 2440: 2420: 2413: 2393: 2386: 2369: 2362: 2342: 2335: 2318: 2311: 2288: 2281: 2261: 2254: 2237: 2230: 2123:and related genera to be 2036: 2014: 2007: 1985: 1978: 1952: 1935: 1928: 1921: 1898: 1891: 1868: 1861: 1838: 1831: 1804:era, dating to the early 1617: 1597: 1590: 1567: 1560: 1537: 1530: 1403: 1383: 1376: 1356: 1349: 1329: 1322: 1295: 1288: 1265: 1258: 1235: 1228: 1205: 1198: 1176: 1169: 828:and modifications of the 756: 749: 348: 343: 226:Scientific classification 224: 196: 187: 90: 2185:Asfaltovenator vialidadi 940:than with modern birds. 851:, became extinct in the 64:Not to be confused with 39:latest accepted revision 5021:(inactive 2024-07-19). 3778:Unquillosaurus ceibalii 3596:Theropod paleopathology 774:is an extinct group of 4659:Comptes Rendus Palevol 4429:Sereno, P. C. (1997). 3508: 1681:, a proposed clade of 1666:Alternative hypotheses 1149:Conventional phylogeny 1078: 1008:to 250 milliliters in 953: 782:that lived during the 9661:Paleobiology Database 7985:Carcharodontosaurinae 7897:Carcharodontosauridae 4751:Nature Communications 3945:Journal of Morphology 3506: 3416:Carcharodontosauridae 2701:Carcharodontosauridae 2196:to represent a basal 1901:Carcharodontosauridae 1619:Carcharodontosauridae 1091:Othniel Charles Marsh 1076: 951: 586:Carcharodontosauridae 9699:Turonian extinctions 8961:Dubious coelurosaurs 7674:Metriacanthosaurinae 7642:Metriacanthosauridae 5924:Dubious neotheropods 5400:at Wikimedia Commons 5099:Cau, Andrea (2021). 5019:10.4435/BSPI.2024.08 5001:Cau, Andrea (2024). 3553:paleobiogeographical 3128:Metriacanthosauridae 2629:Metriacanthosauridae 2396:Metriacanthosauridae 1841:Metriacanthosauridae 1808:stage of the latest 1683:carcharodontosaurian 1540:Metriacanthosauridae 1137:and colleagues and 492:Metriacanthosauridae 7998:Carcharodontosaurus 7785:Carcharodontosauria 5351:2010NW.....97...71B 5339:Naturwissenschaften 5297:2008JSPal...6..155B 5229:2015PalZ...89..983C 5068:2013CrRes..45..174N 5056:Cretaceous Research 4964:10.7717/peerj.12727 4815:2016PLoSO..1157793A 4763:2013NatCo...4.2827Z 4708:2012JSPal..10..211C 4607:2016PLoSO..1157973C 4533:2010NW.....97...71B 4521:Naturwissenschaften 4486:"A new specimen of 4450:1997AREPS..25..435S 4408:1993CaJES..30.2037C 4346:2011PLoSO...617932E 4250:10.1038/nature09181 4242:2010Natur.467..203O 4099:1996Sci...272..986S 4049:2012JSPal..10..211C 3999:(5423): 2137–2147. 3914:2012Pbio...38..486B 3816:2012JSPal..10..211C 3756:2013CrRes..40..131F 3744:Cretaceous Research 3716:Cretaceous Research 3636:2019NatSR...918826R 2442:Carcharodontosauria 2215:or Neornithes, the 2206:Allosaurus fragilis 2127:closely related to 1888:Carcharodontosauria 1587:Carcharodontosauria 836:such as the thigh ( 522:Carcharodontosauria 29:Page version status 9453:Archaeornithomimus 7974:Veterupristisaurus 7834:Veterupristisaurus 7690:Metriacanthosaurus 7630:Piatnitzkysauridae 7065:Eustreptospondylus 6975:Piatnitzkysauridae 6858:Chuandongocoelurus 5722:Notatesseraeraptor 4772:10.1038/ncomms3827 4183:Biological Reviews 3957:10.1002/jmor.10406 3624:Scientific Reports 3509: 3232:Metriacanthosaurus 2858:Piatnitzkysauridae 2345:Piatnitzkysauridae 2217:Piatnitzkysauridae 1677:One such clade is 1079: 954: 849:carcharodontosaurs 180:Late Maastrichtian 81:Carnosaur (series) 35: 9676: 9675: 9614:Taxon identifiers 9605: 9604: 9601: 9600: 9574: 9573: 9570: 9569: 9566: 9565: 9562: 9561: 9529: 9528: 9525: 9524: 9335:Macrocheiriformes 9252:Ornithomimosauria 9226:Maniraptoriformes 9219: 9218: 9065:Maniraptoromorpha 9052:Maniraptoromorpha 9041: 9040: 9007: 9006: 9003: 9002: 8999: 8998: 8952: 8951: 8948: 8947: 8930:Maniraptoromorpha 8920: 8919: 8916: 8915: 8912: 8911: 8908: 8907: 8904: 8903: 8669: 8668: 8534: 8533: 8488:Proceratosauridae 8440:Chingkankousaurus 8287: 8286: 8248: 8247: 8244: 8243: 8240: 8239: 8216: 8215: 8068: 8067: 8064: 8063: 8060: 8059: 8056: 8055: 8052: 8051: 8048: 8047: 7724: 7723: 7720: 7719: 7481: 7480: 7452: 7451: 7448: 7447: 7444: 7443: 7440: 7439: 7412: 7411: 7408: 7407: 7404: 7403: 7400: 7399: 7316: 7315: 7272:Vallibonavenatrix 7175: 7174: 7171: 7170: 7015: 7014: 6796: 6795: 6760: 6759: 6756: 6755: 6752: 6751: 6724: 6723: 6720: 6719: 6716: 6715: 6712: 6711: 6708: 6707: 6356: 6355: 6352: 6351: 6118: 6117: 5976: 5975: 5956: 5955: 5952: 5951: 5948: 5947: 5899: 5898: 5552: 5551: 5548: 5547: 5544: 5543: 5396:Media related to 5213:Mapusaurus roseae 5197:10.7939/r34t6fj1p 5146:10.7717/peerj.940 4402:(10): 2037–2081. 4302:978-3-030-27223-4 4236:(7312): 203–206. 4195:10.1111/brv.12829 4093:(5264): 986–991. 3547:Paleobiogeography 3540:habitat reduction 3495: 3494: 3486: 3485: 3477: 3476: 3468: 3467: 3459: 3458: 3450: 3449: 3441: 3440: 3432: 3431: 3317: 3288: 3287: 3279: 3278: 3270: 3269: 3182: 3181: 2965: 2964: 2956: 2955: 2947: 2946: 2938: 2937: 2929: 2928: 2920: 2919: 2837: 2836: 2828: 2827: 2744: 2743: 2735: 2734: 2726: 2725: 2717: 2716: 2530: 2529: 2521: 2520: 2512: 2511: 2503: 2502: 2494: 2493: 2485: 2484: 2476: 2475: 2467: 2466: 2458: 2457: 2131:. However, Novas 2108: 2107: 2099: 2098: 2090: 2089: 2081: 2080: 2072: 2071: 2063: 2062: 2054: 2053: 1967: 1966: 1662: 1661: 1653: 1652: 1644: 1643: 1635: 1634: 1493: 1492: 1484: 1483: 1475: 1474: 1466: 1465: 1457: 1456: 1448: 1447: 1439: 1438: 1430: 1429: 1421: 1420: 1385:Ornithomimosauria 769: 768: 764: 744: 743: 526: 505: 421: 410: 339: 73:Carnosaur (novel) 47:19 September 2024 16:(Redirected from 9716: 9669: 9668: 9656: 9655: 9654: 9641: 9640: 9639: 9609: 9596: 9590: 9583: 9439:Aepyornithomimus 9340: 9257: 9232: 9196:Sinocalliopteryx 9116: 9112: 9106:Neocoelurosauria 9071: 9060: 9047: 9035: 9028: 9021: 9015: 8958: 8926: 8895:Zhuchengtyrannus 8827:Thanatotheristes 8758: 8722: 8690:Appalachiosaurus 8682: 8551: 8547: 8541:Pantyrannosauria 8432: 8428: 8422:Tyrannosauroidea 8417: 8392:Xinjiangovenator 8317: 8306: 8293: 8281: 8274: 8269: 8262: 8256: 8084: 8074: 7990: 7910:Acrocanthosaurus 7902: 7790: 7737: 7663:Yangchuanosaurus 7647: 7600: 7596: 7546: 7527:Lourinhanosaurus 7511: 7500: 7487: 7475: 7468: 7461: 7418: 7384:Sigilmassasaurus 7327: 7284:Ceratosuchopsini 7235: 7188: 7133:Dubreuillosaurus 7057: 7040:Streptospondylus 7032: 7028: 6998:Piatnitzkysaurus 6956: 6952: 6941: 6826: 6815: 6802: 6790: 6783: 6776: 6769: 6730: 6565:Ekrixinatosaurus 6549: 6540: 6369: 6184: 6135: 6131: 6044: 6040: 6006: 5995: 5982: 5971: 5965: 5875:Procompsognathus 5781: 5664: 5582: 5571: 5558: 5504: 5495: 5488: 5487: 5443: 5428: 5421: 5414: 5405: 5395: 5379: 5378: 5333: 5327: 5326: 5316: 5276: 5267: 5266: 5248: 5208: 5202: 5201: 5199: 5175: 5169: 5168: 5158: 5148: 5124: 5113: 5112: 5096: 5090: 5089: 5087: 5051: 5045: 5044: 5038: 5030: 4998: 4987: 4986: 4976: 4966: 4945:Eotyrannus lengi 4938: 4929: 4928: 4926: 4920:. Archived from 4919: 4893: 4876: 4870: 4869: 4853: 4847: 4846: 4836: 4826: 4794: 4785: 4784: 4774: 4742: 4736: 4735: 4691: 4685: 4684: 4682: 4653: 4647: 4646: 4636: 4618: 4586: 4580: 4579: 4567: 4561: 4560: 4516: 4505: 4504: 4494: 4481: 4475: 4468: 4462: 4461: 4435: 4426: 4420: 4419: 4393: 4384: 4378: 4377: 4367: 4357: 4321: 4312: 4311: 4310: 4309: 4276: 4270: 4269: 4221: 4215: 4214: 4173: 4167: 4163: 4157: 4156: 4154: 4133: 4127: 4126: 4084: 4075: 4069: 4068: 4032: 4017: 4016: 3988: 3977: 3976: 3940: 3934: 3933: 3897: 3888: 3885: 3879: 3878: 3876: 3856: 3850: 3849: 3842: 3836: 3835: 3795: 3789: 3774: 3768: 3767: 3738: 3732: 3731: 3711: 3705: 3704: 3702: 3700: 3685: 3679: 3672: 3666: 3665: 3655: 3615: 3423: 3403: 3388: 3374: 3359: 3345: 3337:Lourinhanosaurus 3323: 3315: 3305: 3261: 3239: 3224: 3196: 3156: 3149:Yangchuanosaurus 3141: 3134: 3124: 3100: 3083: 3070: 3055: 3042: 3030: 2911: 2891: 2878: 2864: 2851: 2792: 2768: 2761: 2708: 2678: 2664: 2649: 2635: 2622: 2598: 2588: 2571: 2558: 2546: 2449: 2429: 2416: 2402: 2389: 2365: 2351: 2338: 2314: 2297: 2284: 2270: 2257: 2233: 2223: 2045: 2025: 2010: 1996: 1981: 1931: 1924: 1907: 1894: 1877: 1864: 1847: 1834: 1823: 1626: 1606: 1593: 1576: 1563: 1546: 1533: 1523: 1516:Streptospondylus 1412: 1392: 1379: 1365: 1352: 1338: 1332:Tyrannosauroidea 1325: 1308: 1291: 1274: 1261: 1244: 1231: 1214: 1201: 1187: 1172: 1158: 1099:stem-based taxon 885:History of study 824:, a long narrow 762: 733: 723: 713: 703: 693: 683: 673: 663: 653: 643: 633: 623: 613: 599: 583: 572: 562: 552: 542: 531: 524: 520: 510: 503: 499: 489: 479: 468: 457: 449:Lourinhanosaurus 446: 436: 426: 419: 415: 404: 397: 386: 375: 364: 353: 334: 329: 234: 233: 206:Acrocanthosaurus 192: 173: 110: 94:Temporal range: 88: 77:Carnosaur (film) 21: 9724: 9723: 9719: 9718: 9717: 9715: 9714: 9713: 9679: 9678: 9677: 9672: 9664: 9659: 9650: 9649: 9644: 9635: 9634: 9629: 9616: 9606: 9597: 9584: 9558: 9521: 9502:Sinornithomimus 9424: 9367: 9329: 9246: 9240:Compsognathidae 9228: 9215: 9203:Sinosauropteryx 9135:Compsognathidae 9129: 9108: 9099: 9067: 9054: 9037: 9036: 9022: 8995: 8990:Shanyangosaurus 8983:Phaedrolosaurus 8944: 8900: 8866: 8832: 8811:Daspletosaurini 8805: 8778: 8753:Tyrannosaurinae 8747: 8726:Albertosaurinae 8718: 8716:Tyrannosauridae 8709: 8678: 8676:Eutyrannosauria 8665: 8543: 8530: 8511:Proceratosaurus 8482: 8453: 8424: 8413: 8404: 8370:Richardoestesia 8354:Chilantaisaurus 8313: 8300: 8283: 8282: 8263: 8236: 8212: 8172:Australovenator 8150: 8144:Chilantaisaurus 8122:Phuwiangvenator 8044: 8009:Giganotosaurini 8003: 7979: 7891: 7854:Chilantaisaurus 7839: 7779: 7733: 7716: 7668: 7636: 7592: 7583: 7561:Monolophosaurus 7542: 7533: 7507: 7494: 7477: 7476: 7462: 7436: 7396: 7369: 7335:Camarillasaurus 7312: 7278: 7224: 7218:Ostafrikasaurus 7196:Camarillasaurus 7184: 7167: 7117:Afrovenatorinae 7111: 7070: 7046: 7024: 7011: 6969: 6948: 6946:Megalosauroidea 6937: 6928: 6923:Vectaerovenator 6916:Szechuanosaurus 6894:Monolophosaurus 6880:Kaijiangosaurus 6873:Cryolophosaurus 6834:Calamospondylus 6822: 6809: 6792: 6791: 6784: 6777: 6770: 6748: 6704: 6685:Pycnonemosaurus 6627: 6538: 6532:Majungasaurinae 6517: 6459:Majungasaurinae 6453: 6448:Xenotarsosaurus 6384:Dryptosauroides 6365: 6348: 6300: 6272:Elaphrosaurinae 6266: 6173: 6159:Ornithomimoides 6127: 6114: 6087: 6036: 6027: 6002: 5989: 5972: 5966: 5944: 5919: 5895: 5823: 5770: 5736:Shuangbaisaurus 5660: 5651: 5578: 5565: 5540: 5520: 5514:Avemetatarsalia 5499:Avemetatarsalia 5489: 5474:Avemetatarsalia 5448: 5447: 5437: 5432: 5388: 5383: 5382: 5335: 5334: 5330: 5278: 5277: 5270: 5210: 5209: 5205: 5177: 5176: 5172: 5126: 5125: 5116: 5098: 5097: 5093: 5053: 5052: 5048: 5031: 5000: 4999: 4990: 4940: 4939: 4932: 4924: 4891: 4887:Australovenator 4878: 4877: 4873: 4855: 4854: 4850: 4809:(7): e0157793. 4796: 4795: 4788: 4744: 4743: 4739: 4693: 4692: 4688: 4655: 4654: 4650: 4601:(7): e0157973. 4588: 4587: 4583: 4569: 4568: 4564: 4518: 4517: 4508: 4492: 4483: 4482: 4478: 4469: 4465: 4433: 4428: 4427: 4423: 4416:10.1139/e93-179 4391: 4386: 4385: 4381: 4323: 4322: 4315: 4307: 4305: 4303: 4278: 4277: 4273: 4223: 4222: 4218: 4189:(3): 960–1004. 4175: 4174: 4170: 4164: 4160: 4135: 4134: 4130: 4082: 4077: 4076: 4072: 4034: 4033: 4020: 3990: 3989: 3980: 3942: 3941: 3937: 3922:10.1666/10004.1 3899: 3898: 3891: 3886: 3882: 3874:10.1.1.211.3044 3858: 3857: 3853: 3844: 3843: 3839: 3797: 3796: 3792: 3775: 3771: 3740: 3739: 3735: 3713: 3712: 3708: 3698: 3696: 3687: 3686: 3682: 3674:Cau A. (2024). 3673: 3669: 3617: 3616: 3609: 3604: 3582: 3562:Megalosauroidea 3549: 3514:immune response 3501: 3496: 3487: 3478: 3469: 3460: 3451: 3442: 3433: 3424: 3404: 3346: 3324: 3289: 3280: 3271: 3183: 3108:Monolophosaurus 3049:Megalosauroidea 2966: 2957: 2948: 2939: 2930: 2921: 2838: 2829: 2755:Megalosauroidea 2745: 2736: 2727: 2718: 2709: 2606:Monolophosaurus 2531: 2522: 2513: 2504: 2495: 2486: 2477: 2468: 2459: 2450: 2241:Monolophosaurus 2208:than to either 2200:of carnosaurs, 2194:Megalosauroidea 2109: 2100: 2091: 2082: 2073: 2064: 2055: 2046: 2026: 2017:Chilantaisaurus 1997: 1968: 1769:Australovenator 1746:Australovenator 1726:Chilantaisaurus 1708:Chilantaisaurus 1668: 1663: 1654: 1645: 1636: 1627: 1607: 1494: 1485: 1476: 1467: 1458: 1449: 1440: 1431: 1422: 1393: 1359:Compsognathidae 1268:Megalosauroidea 1208:Dilophosauridae 1151: 1135:Thomas R. Holtz 1071: 946: 887: 803:Megalosauroidea 745: 636:Chilantaisaurus 604: 400:Megalosauroidea 367:Monolophosaurus 333: 327: 228: 183: 172: 171: 166: 161: 156: 151: 146: 141: 136: 131: 126: 121: 116: 105: 104: 101:Late Cretaceous 97:Middle Jurassic 95: 92: 84: 69: 62: 55: 54: 53: 52: 51: 50: 34: 22: 15: 12: 11: 5: 9722: 9720: 9712: 9711: 9706: 9701: 9696: 9691: 9681: 9680: 9674: 9673: 9671: 9670: 9657: 9642: 9626: 9624: 9618: 9617: 9612: 9603: 9602: 9599: 9598: 9578: 9577: 9575: 9572: 9571: 9568: 9567: 9564: 9563: 9560: 9559: 9557: 9556: 9555: 9554: 9548: 9539: 9537: 9531: 9530: 9527: 9526: 9523: 9522: 9520: 9519: 9512: 9505: 9498: 9491: 9484: 9477: 9470: 9467:Dromiceiomimus 9463: 9456: 9449: 9442: 9434: 9432: 9430:Ornithomimidae 9426: 9425: 9423: 9422: 9415: 9411:Paraxenisaurus 9407: 9399: 9392: 9385: 9377: 9375: 9373:Deinocheiridae 9369: 9368: 9366: 9365: 9362:Shenzhousaurus 9358: 9351: 9343: 9337: 9331: 9330: 9328: 9327: 9320: 9312: 9305: 9298: 9291: 9284: 9276: 9268: 9260: 9254: 9248: 9247: 9245: 9244: 9235: 9229: 9224: 9221: 9220: 9217: 9216: 9214: 9213: 9206: 9199: 9192: 9184: 9177: 9170: 9163: 9159:Beipiaognathus 9155: 9148: 9139: 9137: 9131: 9130: 9128: 9127: 9119: 9109: 9104: 9101: 9100: 9098: 9097: 9090: 9083: 9074: 9068: 9063: 9056: 9055: 9050: 9043: 9042: 9039: 9038: 9030: 9016: 9010: 9008: 9005: 9004: 9001: 9000: 8997: 8996: 8994: 8993: 8986: 8979: 8972: 8964: 8962: 8954: 8953: 8950: 8949: 8946: 8945: 8943: 8942: 8941: 8940: 8934: 8932: 8922: 8921: 8918: 8917: 8914: 8913: 8910: 8909: 8906: 8905: 8902: 8901: 8899: 8898: 8891: 8884: 8876: 8874: 8872:Tyrannosaurini 8868: 8867: 8865: 8864: 8857: 8850: 8842: 8840: 8838:Teratophoneini 8834: 8833: 8831: 8830: 8823: 8820:Daspletosaurus 8815: 8813: 8807: 8806: 8804: 8803: 8800:Qianzhousaurus 8796: 8788: 8786: 8780: 8779: 8777: 8776: 8769: 8761: 8755: 8749: 8748: 8746: 8745: 8738: 8730: 8728: 8719: 8714: 8711: 8710: 8708: 8707: 8700: 8697:Bistahieversor 8693: 8685: 8679: 8674: 8671: 8670: 8667: 8666: 8664: 8663: 8655: 8648: 8641: 8633: 8626: 8619: 8612: 8605: 8598: 8591: 8584: 8577: 8569: 8562: 8554: 8544: 8539: 8536: 8535: 8532: 8531: 8529: 8528: 8521: 8514: 8507: 8500: 8492: 8490: 8484: 8483: 8481: 8480: 8472: 8464: 8462: 8455: 8454: 8452: 8451: 8443: 8435: 8425: 8420: 8414: 8411:Tyrannoraptora 8409: 8406: 8405: 8403: 8402: 8395: 8388: 8381: 8373: 8366: 8358: 8350: 8343: 8335: 8328: 8320: 8314: 8309: 8302: 8301: 8296: 8289: 8288: 8285: 8284: 8276: 8257: 8251: 8249: 8246: 8245: 8242: 8241: 8238: 8237: 8235: 8234: 8233: 8232: 8226: 8224: 8218: 8217: 8214: 8213: 8211: 8210: 8203: 8196: 8189: 8182: 8175: 8168: 8160: 8158: 8156:Megaraptoridae 8152: 8151: 8149: 8148: 8140: 8132: 8125: 8118: 8111: 8103: 8095: 8087: 8081: 8070: 8069: 8066: 8065: 8062: 8061: 8058: 8057: 8054: 8053: 8050: 8049: 8046: 8045: 8043: 8042: 8035: 8028: 8021: 8018:Giganotosaurus 8013: 8011: 8005: 8004: 8002: 8001: 7993: 7987: 7981: 7980: 7978: 7977: 7970: 7963: 7956: 7949: 7942: 7935: 7928: 7921: 7913: 7905: 7899: 7893: 7892: 7890: 7889: 7881: 7873: 7866: 7858: 7849: 7847: 7845:Neovenatoridae 7841: 7840: 7838: 7837: 7830: 7827:Ulughbegsaurus 7823: 7815: 7808: 7801: 7793: 7787: 7781: 7780: 7778: 7777: 7769: 7761: 7753: 7745: 7743: 7734: 7729: 7726: 7725: 7722: 7721: 7718: 7717: 7715: 7714: 7707: 7700: 7693: 7686: 7678: 7676: 7670: 7669: 7667: 7666: 7659: 7655:Xuanhanosaurus 7650: 7644: 7638: 7637: 7635: 7634: 7626: 7622:Xuanhanosaurus 7618: 7611: 7608:Asfaltovenator 7603: 7593: 7588: 7585: 7584: 7582: 7581: 7573: 7569:Megalosauridae 7565: 7557: 7549: 7543: 7538: 7535: 7534: 7532: 7531: 7523: 7514: 7508: 7503: 7496: 7495: 7490: 7483: 7482: 7479: 7478: 7470: 7456: 7455: 7453: 7450: 7449: 7446: 7445: 7442: 7441: 7438: 7437: 7435: 7434: 7433: 7432: 7426: 7424: 7414: 7413: 7410: 7409: 7406: 7405: 7402: 7401: 7398: 7397: 7395: 7394: 7387: 7379: 7377: 7371: 7370: 7368: 7367: 7360: 7353: 7346: 7343:Ichthyovenator 7339: 7330: 7324: 7318: 7317: 7314: 7313: 7311: 7310: 7303: 7296: 7288: 7286: 7280: 7279: 7277: 7276: 7268: 7260: 7253: 7246: 7238: 7232: 7226: 7225: 7223: 7222: 7214: 7207: 7204:Cristatusaurus 7200: 7191: 7185: 7180: 7177: 7176: 7173: 7172: 7169: 7168: 7166: 7165: 7161:Poekilopleuron 7157: 7154:Piveteausaurus 7150: 7143: 7136: 7129: 7121: 7119: 7113: 7112: 7110: 7109: 7102: 7095: 7088: 7080: 7078: 7076:Megalosaurinae 7072: 7071: 7069: 7068: 7060: 7054: 7052:Megalosauridae 7048: 7047: 7045: 7044: 7035: 7025: 7020: 7017: 7016: 7013: 7012: 7010: 7009: 7005:Xuanhanosaurus 7001: 6994: 6987: 6979: 6977: 6971: 6970: 6968: 6967: 6964:Yunyangosaurus 6959: 6949: 6944: 6938: 6933: 6930: 6929: 6927: 6926: 6919: 6912: 6904: 6901:Pandoravenator 6897: 6890: 6887:Kayentavenator 6883: 6876: 6869: 6861: 6854: 6846: 6838: 6829: 6823: 6818: 6811: 6810: 6805: 6798: 6797: 6794: 6793: 6785: 6778: 6771: 6764: 6763: 6761: 6758: 6757: 6754: 6753: 6750: 6749: 6747: 6746: 6745: 6744: 6738: 6736: 6726: 6725: 6722: 6721: 6718: 6717: 6714: 6713: 6710: 6709: 6706: 6705: 6703: 6702: 6695: 6688: 6681: 6674: 6666: 6659: 6652: 6645: 6637: 6635: 6629: 6628: 6626: 6625: 6618: 6615:Skorpiovenator 6611: 6603: 6596: 6589: 6582: 6575: 6568: 6561: 6552: 6546: 6537: 6536: 6527: 6525: 6519: 6518: 6516: 6515: 6508: 6500: 6493: 6486: 6479: 6471: 6463: 6461: 6455: 6454: 6452: 6451: 6444: 6437: 6430: 6427:Spectrovenator 6423: 6416: 6409: 6402: 6395: 6388: 6380: 6372: 6366: 6361: 6358: 6357: 6354: 6353: 6350: 6349: 6347: 6346: 6339: 6332: 6325: 6318: 6310: 6308: 6302: 6301: 6299: 6298: 6291: 6284: 6276: 6274: 6268: 6267: 6265: 6264: 6261:Spinostropheus 6257: 6250: 6243: 6235: 6227: 6219: 6211: 6204: 6196: 6187: 6181: 6175: 6174: 6172: 6171: 6163: 6155: 6147: 6138: 6128: 6125:Abelisauroidea 6123: 6120: 6119: 6116: 6115: 6113: 6112: 6105: 6097: 6095: 6093:Ceratosauridae 6089: 6088: 6086: 6085: 6082:Saltriovenator 6078: 6070: 6063: 6059:Fosterovenator 6055: 6047: 6037: 6032: 6029: 6028: 6026: 6025: 6021:Lophostropheus 6017: 6009: 6003: 5998: 5991: 5990: 5985: 5978: 5977: 5974: 5973: 5960: 5959: 5957: 5954: 5953: 5950: 5949: 5946: 5945: 5943: 5942: 5935: 5927: 5925: 5921: 5920: 5918: 5917: 5916: 5915: 5909: 5907: 5901: 5900: 5897: 5896: 5894: 5893: 5886: 5882:Pterospondylus 5878: 5871: 5863: 5855: 5852:Lucianovenator 5848: 5841: 5833: 5831: 5825: 5824: 5822: 5821: 5814: 5807: 5800: 5793: 5784: 5778: 5776:Coelophysoidea 5772: 5771: 5769: 5768: 5761: 5753: 5746: 5739: 5732: 5725: 5718: 5715:Lophostropheus 5711: 5704: 5697: 5690: 5683: 5675: 5667: 5661: 5656: 5653: 5652: 5650: 5649: 5641: 5633: 5625: 5617: 5614:Erythrovenator 5610: 5602: 5594: 5585: 5579: 5574: 5567: 5566: 5561: 5554: 5553: 5550: 5549: 5546: 5545: 5542: 5541: 5539: 5538: 5537: 5536: 5530: 5528: 5522: 5521: 5519: 5518: 5517: 5516: 5507: 5501: 5491: 5490: 5486: 5485: 5476: 5467: 5461: 5455: 5446: 5439: 5438: 5433: 5431: 5430: 5423: 5416: 5408: 5402: 5401: 5387: 5386:External links 5384: 5381: 5380: 5328: 5291:(2): 155–182. 5268: 5223:(4): 983–993. 5203: 5170: 5114: 5091: 5046: 4988: 4930: 4927:on 2016-08-18. 4871: 4848: 4786: 4737: 4702:(2): 211–300. 4686: 4665:(4): 251–256. 4648: 4581: 4562: 4506: 4476: 4472:The Dinosauria 4463: 4421: 4379: 4313: 4301: 4271: 4216: 4168: 4158: 4141:(in Spanish). 4128: 4070: 4043:(2): 211–300. 4018: 3978: 3951:(3): 347–359. 3935: 3908:(3): 486–507. 3889: 3880: 3851: 3837: 3810:(2): 211–300. 3790: 3769: 3733: 3706: 3680: 3667: 3606: 3605: 3603: 3600: 3599: 3598: 3593: 3588: 3581: 3578: 3548: 3545: 3536:predator traps 3522:metabolic rate 3500: 3497: 3493: 3492: 3489: 3488: 3484: 3483: 3480: 3479: 3475: 3474: 3471: 3470: 3466: 3465: 3462: 3461: 3457: 3456: 3453: 3452: 3448: 3447: 3444: 3443: 3439: 3438: 3435: 3434: 3430: 3429: 3426: 3425: 3418: 3413: 3410: 3409: 3406: 3405: 3398: 3391: 3386: 3384: 3381: 3380: 3377: 3376: 3362: 3357: 3355: 3352: 3351: 3348: 3347: 3340: 3333: 3330: 3329: 3326: 3325: 3318: 3308: 3303: 3301: 3295: 3294: 3291: 3290: 3286: 3285: 3282: 3281: 3277: 3276: 3273: 3272: 3268: 3267: 3264: 3263: 3249: 3246: 3245: 3242: 3241: 3227: 3222: 3220: 3217: 3216: 3213: 3212: 3199: 3194: 3192: 3189: 3188: 3185: 3184: 3180: 3179: 3176: 3175: 3166: 3163: 3162: 3159: 3158: 3144: 3139: 3137: 3132: 3130: 3122: 3120: 3117: 3116: 3113: 3112: 3103: 3098: 3096: 3090: 3089: 3086: 3085: 3073: 3068: 3066: 3062: 3061: 3058: 3057: 3045: 3040: 3038: 3028: 2963: 2962: 2959: 2958: 2954: 2953: 2950: 2949: 2945: 2944: 2941: 2940: 2936: 2935: 2932: 2931: 2927: 2926: 2923: 2922: 2918: 2917: 2914: 2913: 2905:Megalosauridae 2901: 2898: 2897: 2894: 2893: 2881: 2876: 2874: 2871: 2870: 2867: 2866: 2854: 2849: 2847: 2844: 2843: 2840: 2839: 2835: 2834: 2831: 2830: 2826: 2825: 2822: 2821: 2812: 2809: 2808: 2805: 2804: 2800:Asfaltovenator 2795: 2790: 2788: 2785: 2784: 2781: 2780: 2771: 2766: 2764: 2759: 2757: 2751: 2750: 2747: 2746: 2742: 2741: 2738: 2737: 2733: 2732: 2729: 2728: 2724: 2723: 2720: 2719: 2715: 2714: 2711: 2710: 2703: 2698: 2695: 2694: 2691: 2690: 2681: 2676: 2674: 2671: 2670: 2667: 2666: 2652: 2647: 2645: 2642: 2641: 2638: 2637: 2625: 2620: 2618: 2615: 2614: 2611: 2610: 2601: 2596: 2594: 2586: 2584: 2578: 2577: 2574: 2573: 2561: 2556: 2554: 2544: 2528: 2527: 2524: 2523: 2519: 2518: 2515: 2514: 2510: 2509: 2506: 2505: 2501: 2500: 2497: 2496: 2492: 2491: 2488: 2487: 2483: 2482: 2479: 2478: 2474: 2473: 2470: 2469: 2465: 2464: 2461: 2460: 2456: 2455: 2452: 2451: 2444: 2439: 2436: 2435: 2432: 2431: 2419: 2414: 2412: 2409: 2408: 2405: 2404: 2392: 2387: 2385: 2382: 2381: 2378: 2377: 2373:Asfaltovenator 2368: 2363: 2361: 2358: 2357: 2354: 2353: 2341: 2336: 2334: 2331: 2330: 2327: 2326: 2322:Xuanhanosaurus 2317: 2312: 2310: 2304: 2303: 2300: 2299: 2291:Megalosauridae 2287: 2282: 2280: 2277: 2276: 2273: 2272: 2260: 2255: 2253: 2250: 2249: 2246: 2245: 2236: 2231: 2229: 2221: 2129:tyrannosaurids 2106: 2105: 2102: 2101: 2097: 2096: 2093: 2092: 2088: 2087: 2084: 2083: 2079: 2078: 2075: 2074: 2070: 2069: 2066: 2065: 2061: 2060: 2057: 2056: 2052: 2051: 2048: 2047: 2040: 2035: 2032: 2031: 2028: 2027: 2020: 2013: 2008: 2006: 2003: 2002: 1999: 1998: 1991: 1984: 1979: 1977: 1974: 1973: 1970: 1969: 1965: 1964: 1961: 1960: 1951: 1948: 1947: 1944: 1943: 1934: 1929: 1927: 1922: 1920: 1918:Neovenatoridae 1914: 1913: 1910: 1909: 1897: 1892: 1890: 1884: 1883: 1880: 1879: 1867: 1862: 1860: 1854: 1853: 1850: 1849: 1837: 1832: 1830: 1821: 1679:Neovenatoridae 1667: 1664: 1660: 1659: 1656: 1655: 1651: 1650: 1647: 1646: 1642: 1641: 1638: 1637: 1633: 1632: 1629: 1628: 1621: 1616: 1613: 1612: 1609: 1608: 1601: 1599:Neovenatoridae 1596: 1591: 1589: 1583: 1582: 1579: 1578: 1566: 1561: 1559: 1553: 1552: 1549: 1548: 1536: 1531: 1529: 1521: 1510:Xuanhanosaurus 1504:Poekilopleuron 1491: 1490: 1487: 1486: 1482: 1481: 1478: 1477: 1473: 1472: 1469: 1468: 1464: 1463: 1460: 1459: 1455: 1454: 1451: 1450: 1446: 1445: 1442: 1441: 1437: 1436: 1433: 1432: 1428: 1427: 1424: 1423: 1419: 1418: 1415: 1414: 1402: 1399: 1398: 1395: 1394: 1387: 1382: 1377: 1375: 1372: 1371: 1368: 1367: 1355: 1350: 1348: 1345: 1344: 1341: 1340: 1328: 1323: 1321: 1315: 1314: 1311: 1310: 1294: 1289: 1287: 1281: 1280: 1277: 1276: 1264: 1259: 1257: 1251: 1250: 1247: 1246: 1234: 1229: 1227: 1221: 1220: 1217: 1216: 1204: 1199: 1197: 1194: 1193: 1190: 1189: 1180:Coelophysoidea 1175: 1170: 1168: 1156: 1150: 1147: 1070: 1069:Classification 1067: 1010:Giganotosaurus 979:caudofemoralis 974:center of mass 967:tyrannosaurids 945: 942: 920:tyrannosaurids 886: 883: 878:Unquillosaurus 811:coleurosaurian 767: 766: 754: 753: 747: 746: 742: 741: 740: 739: 729: 719: 709: 699: 689: 679: 669: 659: 649: 639: 629: 619: 606: 605: 602: 597: 596: 595: 594: 593: 592: 591: 590: 589: 579: 575:Ulughbegsaurus 568: 558: 548: 538: 516: 495: 485: 475: 471:Xuanhanosaurus 464: 453: 442: 432: 429:Asfaltovenator 411: 393: 389:Poekilopleuron 382: 371: 360: 346: 345: 341: 340: 325: 318: 317: 312: 305: 304: 299: 292: 291: 286: 279: 278: 273: 266: 265: 260: 256: 255: 250: 246: 245: 240: 236: 235: 222: 221: 194: 193: 185: 184: 167: 162: 157: 152: 147: 142: 137: 132: 127: 122: 117: 112: 111: 93: 60: 56: 36: 30: 27: 25: 24: 23: 14: 13: 10: 9: 6: 4: 3: 2: 9721: 9710: 9707: 9705: 9702: 9700: 9697: 9695: 9692: 9690: 9687: 9686: 9684: 9667: 9662: 9658: 9653: 9647: 9643: 9638: 9632: 9628: 9627: 9625: 9623: 9619: 9615: 9610: 9595: 9591: 9589: 9582: 9553: 9549: 9547: 9543: 9542: 9541: 9540: 9538: 9536: 9532: 9518: 9517: 9513: 9511: 9510: 9509:Struthiomimus 9506: 9504: 9503: 9499: 9497: 9496: 9492: 9490: 9489: 9485: 9483: 9482: 9478: 9476: 9475: 9471: 9469: 9468: 9464: 9462: 9461: 9460:"Coelosaurus" 9457: 9455: 9454: 9450: 9448: 9447: 9443: 9441: 9440: 9436: 9435: 9433: 9431: 9427: 9421: 9420: 9416: 9413: 9412: 9408: 9405: 9404: 9400: 9398: 9397: 9393: 9391: 9390: 9386: 9384: 9383: 9379: 9378: 9376: 9374: 9370: 9364: 9363: 9359: 9357: 9356: 9355:Pelecanimimus 9352: 9350: 9349: 9345: 9344: 9341: 9338: 9336: 9332: 9326: 9325: 9321: 9318: 9317: 9316:Thecocoelurus 9313: 9311: 9310: 9306: 9304: 9303: 9299: 9297: 9296: 9295:Kinnareemimus 9292: 9290: 9289: 9285: 9282: 9281: 9277: 9274: 9273: 9269: 9267: 9266: 9262: 9261: 9258: 9255: 9253: 9249: 9242: 9241: 9237: 9236: 9233: 9230: 9227: 9222: 9212: 9211: 9207: 9205: 9204: 9200: 9198: 9197: 9193: 9190: 9189: 9185: 9183: 9182: 9178: 9176: 9175: 9174:Huaxiagnathus 9171: 9169: 9168: 9167:Compsognathus 9164: 9161: 9160: 9156: 9154: 9153: 9149: 9146: 9145: 9141: 9140: 9138: 9136: 9132: 9126: 9125: 9121: 9120: 9117: 9113: 9110: 9107: 9102: 9096: 9095: 9094:Ornitholestes 9091: 9089: 9088: 9084: 9081: 9080: 9076: 9075: 9072: 9069: 9066: 9061: 9057: 9053: 9048: 9044: 9034: 9029: 9027: 9020: 9014: 8992: 8991: 8987: 8985: 8984: 8980: 8978: 8977: 8973: 8971: 8970: 8966: 8965: 8963: 8959: 8955: 8938: 8937: 8936: 8935: 8933: 8931: 8927: 8923: 8897: 8896: 8892: 8890: 8889: 8888:Tyrannosaurus 8885: 8883: 8882: 8878: 8877: 8875: 8873: 8869: 8863: 8862: 8861:Teratophoneus 8858: 8856: 8855: 8851: 8849: 8848: 8844: 8843: 8841: 8839: 8835: 8829: 8828: 8824: 8822: 8821: 8817: 8816: 8814: 8812: 8808: 8802: 8801: 8797: 8795: 8794: 8790: 8789: 8787: 8785: 8781: 8775: 8774: 8770: 8768: 8767: 8763: 8762: 8759: 8756: 8754: 8750: 8744: 8743: 8739: 8737: 8736: 8735:Albertosaurus 8732: 8731: 8729: 8727: 8723: 8720: 8717: 8712: 8706: 8705: 8701: 8699: 8698: 8694: 8692: 8691: 8687: 8686: 8683: 8680: 8677: 8672: 8661: 8660: 8656: 8654: 8653: 8652:Xiongguanlong 8649: 8647: 8646: 8642: 8639: 8638: 8634: 8632: 8631: 8630:Suskityrannus 8627: 8625: 8624: 8623:Stokesosaurus 8620: 8618: 8617: 8616:Santanaraptor 8613: 8611: 8610: 8606: 8604: 8603: 8599: 8597: 8596: 8592: 8590: 8589: 8585: 8583: 8582: 8578: 8575: 8574: 8570: 8568: 8567: 8563: 8561: 8560: 8559:Alectrosaurus 8556: 8555: 8552: 8548: 8545: 8542: 8537: 8527: 8526: 8522: 8520: 8519: 8515: 8513: 8512: 8508: 8506: 8505: 8501: 8499: 8498: 8494: 8493: 8491: 8489: 8485: 8478: 8477: 8476:Tanycolagreus 8473: 8471: 8470: 8466: 8465: 8463: 8460: 8456: 8449: 8448: 8444: 8442: 8441: 8437: 8436: 8433: 8429: 8426: 8423: 8418: 8415: 8412: 8407: 8401: 8400: 8396: 8394: 8393: 8389: 8387: 8386: 8382: 8379: 8378: 8374: 8372: 8371: 8367: 8364: 8363: 8359: 8356: 8355: 8351: 8349: 8348: 8344: 8341: 8340: 8339:Bahariasaurus 8336: 8334: 8333: 8329: 8327: 8326: 8322: 8321: 8318: 8315: 8312: 8311:Coelurosauria 8307: 8303: 8299: 8298:Coelurosauria 8294: 8290: 8280: 8275: 8273: 8268: 8261: 8255: 8230: 8229: 8228: 8227: 8225: 8223: 8222:Coelurosauria 8219: 8209: 8208: 8204: 8202: 8201: 8197: 8195: 8194: 8190: 8188: 8187: 8183: 8181: 8180: 8176: 8174: 8173: 8169: 8167: 8166: 8162: 8161: 8159: 8157: 8153: 8147: 8145: 8141: 8139: 8137: 8133: 8131: 8130: 8126: 8124: 8123: 8119: 8117: 8116: 8112: 8109: 8108: 8104: 8101: 8100: 8099:Bahariasaurus 8096: 8094: 8093: 8089: 8088: 8085: 8082: 8079: 8075: 8071: 8041: 8040: 8036: 8034: 8033: 8029: 8027: 8026: 8022: 8020: 8019: 8015: 8014: 8012: 8010: 8006: 8000: 7999: 7995: 7994: 7991: 7988: 7986: 7982: 7976: 7975: 7971: 7969: 7968: 7964: 7962: 7961: 7957: 7955: 7954: 7950: 7948: 7947: 7943: 7941: 7940: 7939:Kelmayisaurus 7936: 7934: 7933: 7929: 7927: 7926: 7922: 7919: 7918: 7914: 7912: 7911: 7907: 7906: 7903: 7900: 7898: 7894: 7887: 7886: 7882: 7879: 7878: 7874: 7872: 7871: 7867: 7864: 7863: 7859: 7856: 7855: 7851: 7850: 7848: 7846: 7842: 7836: 7835: 7831: 7829: 7828: 7824: 7821: 7820: 7816: 7814: 7813: 7809: 7807: 7806: 7802: 7800: 7799: 7795: 7794: 7791: 7788: 7786: 7782: 7775: 7774: 7773:Saurophaganax 7770: 7767: 7766: 7762: 7759: 7758: 7754: 7752: 7751: 7747: 7746: 7744: 7742: 7738: 7735: 7732: 7727: 7713: 7712: 7708: 7706: 7705: 7704:Siamotyrannus 7701: 7699: 7698: 7694: 7692: 7691: 7687: 7685: 7684: 7680: 7679: 7677: 7675: 7671: 7665: 7664: 7660: 7657: 7656: 7652: 7651: 7648: 7645: 7643: 7639: 7632: 7631: 7627: 7624: 7623: 7619: 7617: 7616: 7612: 7610: 7609: 7605: 7604: 7601: 7597: 7594: 7591: 7590:Allosauroidea 7586: 7579: 7578: 7577:Spinosauridae 7574: 7571: 7570: 7566: 7563: 7562: 7558: 7556: 7555: 7551: 7550: 7547: 7544: 7541: 7536: 7529: 7528: 7524: 7521: 7520: 7516: 7515: 7512: 7509: 7506: 7501: 7497: 7493: 7488: 7484: 7474: 7469: 7467: 7460: 7430: 7429: 7428: 7427: 7425: 7423: 7419: 7415: 7393: 7392: 7388: 7386: 7385: 7381: 7380: 7378: 7376: 7372: 7366: 7365: 7361: 7359: 7358: 7354: 7352: 7351: 7347: 7345: 7344: 7340: 7337: 7336: 7332: 7331: 7328: 7325: 7323: 7322:Spinosaurinae 7319: 7309: 7308: 7304: 7302: 7301: 7300:Riparovenator 7297: 7295: 7294: 7293:Ceratosuchops 7290: 7289: 7287: 7285: 7281: 7274: 7273: 7269: 7266: 7265: 7261: 7259: 7258: 7257:Riojavenatrix 7254: 7252: 7251: 7247: 7245: 7244: 7240: 7239: 7236: 7233: 7231: 7230:Baryonychinae 7227: 7220: 7219: 7215: 7213: 7212: 7208: 7206: 7205: 7201: 7198: 7197: 7193: 7192: 7189: 7186: 7183: 7182:Spinosauridae 7178: 7163: 7162: 7158: 7156: 7155: 7151: 7149: 7148: 7144: 7142: 7141: 7137: 7135: 7134: 7130: 7128: 7127: 7123: 7122: 7120: 7118: 7114: 7108: 7107: 7106:Wiehenvenator 7103: 7101: 7100: 7096: 7094: 7093: 7089: 7087: 7086: 7082: 7081: 7079: 7077: 7073: 7067: 7066: 7062: 7061: 7058: 7055: 7053: 7049: 7042: 7041: 7037: 7036: 7033: 7029: 7026: 7023: 7018: 7007: 7006: 7002: 7000: 6999: 6995: 6993: 6992: 6988: 6986: 6985: 6981: 6980: 6978: 6976: 6972: 6966: 6965: 6961: 6960: 6957: 6953: 6950: 6947: 6942: 6939: 6936: 6931: 6925: 6924: 6920: 6918: 6917: 6913: 6910: 6909: 6905: 6903: 6902: 6898: 6896: 6895: 6891: 6889: 6888: 6884: 6882: 6881: 6877: 6875: 6874: 6870: 6867: 6866: 6862: 6860: 6859: 6855: 6852: 6851: 6847: 6844: 6843: 6842:Chienkosaurus 6839: 6836: 6835: 6831: 6830: 6827: 6824: 6821: 6816: 6812: 6808: 6803: 6799: 6789: 6782: 6775: 6768: 6742: 6741: 6740: 6739: 6737: 6735: 6731: 6727: 6701: 6700: 6696: 6694: 6693: 6689: 6687: 6686: 6682: 6680: 6679: 6675: 6672: 6671: 6667: 6665: 6664: 6660: 6658: 6657: 6653: 6651: 6650: 6646: 6644: 6643: 6639: 6638: 6636: 6634: 6633:Furileusauria 6630: 6624: 6623: 6619: 6617: 6616: 6612: 6609: 6608: 6607:Rahiolisaurus 6604: 6602: 6601: 6597: 6595: 6594: 6590: 6588: 6587: 6583: 6581: 6580: 6576: 6574: 6573: 6569: 6567: 6566: 6562: 6559: 6558: 6554: 6553: 6550: 6547: 6545: 6541: 6534: 6533: 6529: 6528: 6526: 6524: 6523:Carnotaurinae 6520: 6514: 6513: 6509: 6506: 6505: 6504:Rahiolisaurus 6501: 6499: 6498: 6497:Majungasaurus 6494: 6492: 6491: 6487: 6485: 6484: 6480: 6477: 6476: 6472: 6470: 6469: 6465: 6464: 6462: 6460: 6456: 6450: 6449: 6445: 6443: 6442: 6438: 6436: 6435: 6434:Tarascosaurus 6431: 6429: 6428: 6424: 6422: 6421: 6417: 6415: 6414: 6410: 6408: 6407: 6403: 6401: 6400: 6396: 6394: 6393: 6392:Eoabelisaurus 6389: 6386: 6385: 6381: 6379: 6378: 6377:Chenanisaurus 6374: 6373: 6370: 6367: 6364: 6363:Abelisauridae 6359: 6345: 6344: 6340: 6338: 6337: 6333: 6331: 6330: 6326: 6324: 6323: 6322:Masiakasaurus 6319: 6317: 6316: 6312: 6311: 6309: 6307: 6303: 6297: 6296: 6292: 6290: 6289: 6288:Huinculsaurus 6285: 6283: 6282: 6281:Elaphrosaurus 6278: 6277: 6275: 6273: 6269: 6263: 6262: 6258: 6256: 6255: 6251: 6249: 6248: 6244: 6241: 6240: 6236: 6233: 6232: 6228: 6225: 6224: 6220: 6217: 6216: 6212: 6210: 6209: 6205: 6202: 6201: 6200:Austrocheirus 6197: 6194: 6193: 6189: 6188: 6185: 6182: 6180: 6176: 6169: 6168: 6164: 6161: 6160: 6156: 6153: 6152: 6148: 6145: 6144: 6140: 6139: 6136: 6132: 6129: 6126: 6121: 6111: 6110: 6106: 6104: 6103: 6099: 6098: 6096: 6094: 6090: 6084: 6083: 6079: 6076: 6075: 6071: 6069: 6068: 6064: 6061: 6060: 6056: 6054: 6053: 6052:Berberosaurus 6049: 6048: 6045: 6041: 6038: 6035: 6030: 6023: 6022: 6018: 6016: 6015: 6011: 6010: 6007: 6004: 6001: 5996: 5992: 5988: 5983: 5979: 5970: 5964: 5941: 5940: 5939:Halticosaurus 5936: 5934: 5933: 5932:Teinurosaurus 5929: 5928: 5926: 5922: 5913: 5912: 5911: 5910: 5908: 5906: 5902: 5892: 5891: 5887: 5884: 5883: 5879: 5877: 5876: 5872: 5869: 5868: 5864: 5861: 5860: 5859:Megapnosaurus 5856: 5854: 5853: 5849: 5847: 5846: 5842: 5840: 5839: 5835: 5834: 5832: 5830: 5829:Coelophysidae 5826: 5820: 5819: 5818:Powellvenator 5815: 5813: 5812: 5808: 5806: 5805: 5801: 5799: 5798: 5794: 5791: 5790: 5786: 5785: 5782: 5779: 5777: 5773: 5767: 5766: 5762: 5759: 5758: 5754: 5752: 5751: 5747: 5745: 5744: 5740: 5738: 5737: 5733: 5731: 5730: 5726: 5724: 5723: 5719: 5717: 5716: 5712: 5710: 5709: 5708:Liliensternus 5705: 5703: 5702: 5698: 5696: 5695: 5691: 5689: 5688: 5684: 5681: 5680: 5679:Dolichosuchus 5676: 5674: 5673: 5672:Dilophosaurus 5669: 5668: 5665: 5662: 5659: 5654: 5647: 5646: 5645:Herrerasauria 5642: 5639: 5638: 5634: 5631: 5630: 5626: 5623: 5622: 5618: 5616: 5615: 5611: 5608: 5607: 5603: 5600: 5599: 5598:Daemonosaurus 5595: 5592: 5591: 5587: 5586: 5583: 5580: 5577: 5572: 5568: 5564: 5559: 5555: 5534: 5533: 5532: 5531: 5529: 5527: 5523: 5515: 5511: 5510: 5509: 5508: 5505: 5502: 5500: 5496: 5492: 5484: 5480: 5477: 5475: 5471: 5468: 5466: 5462: 5460: 5456: 5454: 5450: 5449: 5444: 5440: 5436: 5429: 5424: 5422: 5417: 5415: 5410: 5409: 5406: 5399: 5394: 5390: 5389: 5385: 5376: 5372: 5368: 5364: 5360: 5356: 5352: 5348: 5344: 5340: 5332: 5329: 5324: 5320: 5315: 5310: 5306: 5302: 5298: 5294: 5290: 5286: 5282: 5275: 5273: 5269: 5264: 5260: 5256: 5252: 5247: 5242: 5238: 5234: 5230: 5226: 5222: 5218: 5214: 5207: 5204: 5198: 5193: 5189: 5185: 5181: 5174: 5171: 5166: 5162: 5157: 5152: 5147: 5142: 5138: 5134: 5130: 5123: 5121: 5119: 5115: 5110: 5106: 5102: 5095: 5092: 5086: 5081: 5077: 5073: 5069: 5065: 5061: 5057: 5050: 5047: 5042: 5036: 5028: 5024: 5020: 5016: 5012: 5008: 5004: 4997: 4995: 4993: 4989: 4984: 4980: 4975: 4970: 4965: 4960: 4956: 4952: 4948: 4946: 4937: 4935: 4931: 4923: 4918: 4913: 4909: 4905: 4901: 4897: 4890: 4888: 4884: 4875: 4872: 4868:(Suppl): R33. 4867: 4863: 4859: 4852: 4849: 4844: 4840: 4835: 4830: 4825: 4820: 4816: 4812: 4808: 4804: 4800: 4793: 4791: 4787: 4782: 4778: 4773: 4768: 4764: 4760: 4756: 4752: 4748: 4741: 4738: 4733: 4729: 4725: 4721: 4717: 4713: 4709: 4705: 4701: 4697: 4690: 4687: 4681: 4676: 4672: 4668: 4664: 4660: 4652: 4649: 4644: 4640: 4635: 4630: 4626: 4622: 4617: 4612: 4608: 4604: 4600: 4596: 4592: 4585: 4582: 4577: 4573: 4566: 4563: 4558: 4554: 4550: 4546: 4542: 4538: 4534: 4530: 4526: 4522: 4515: 4513: 4511: 4507: 4503:(2): 207–246. 4502: 4498: 4491: 4489: 4480: 4477: 4473: 4467: 4464: 4459: 4455: 4451: 4447: 4443: 4439: 4432: 4425: 4422: 4417: 4413: 4409: 4405: 4401: 4397: 4390: 4383: 4380: 4375: 4371: 4366: 4361: 4356: 4351: 4347: 4343: 4340:(3): e17932. 4339: 4335: 4331: 4329: 4320: 4318: 4314: 4304: 4298: 4294: 4290: 4286: 4282: 4275: 4272: 4267: 4263: 4259: 4255: 4251: 4247: 4243: 4239: 4235: 4231: 4227: 4220: 4217: 4212: 4208: 4204: 4200: 4196: 4192: 4188: 4184: 4180: 4172: 4169: 4162: 4159: 4153: 4148: 4144: 4140: 4132: 4129: 4124: 4120: 4116: 4112: 4108: 4104: 4100: 4096: 4092: 4088: 4081: 4074: 4071: 4066: 4062: 4058: 4054: 4050: 4046: 4042: 4038: 4031: 4029: 4027: 4025: 4023: 4019: 4014: 4010: 4006: 4002: 3998: 3994: 3987: 3985: 3983: 3979: 3974: 3970: 3966: 3962: 3958: 3954: 3950: 3946: 3939: 3936: 3931: 3927: 3923: 3919: 3915: 3911: 3907: 3903: 3896: 3894: 3890: 3884: 3881: 3875: 3870: 3866: 3862: 3855: 3852: 3847: 3841: 3838: 3833: 3829: 3825: 3821: 3817: 3813: 3809: 3805: 3801: 3794: 3791: 3787: 3783: 3779: 3773: 3770: 3765: 3761: 3757: 3753: 3749: 3745: 3737: 3734: 3729: 3725: 3721: 3717: 3710: 3707: 3695: 3691: 3684: 3681: 3677: 3671: 3668: 3663: 3659: 3654: 3649: 3645: 3641: 3637: 3633: 3629: 3625: 3621: 3614: 3612: 3608: 3601: 3597: 3594: 3592: 3589: 3587: 3584: 3583: 3579: 3577: 3573: 3571: 3567: 3563: 3558: 3554: 3546: 3544: 3541: 3537: 3533: 3532: 3525: 3523: 3518: 3515: 3505: 3498: 3491: 3490: 3482: 3481: 3473: 3472: 3464: 3463: 3455: 3454: 3446: 3445: 3437: 3436: 3428: 3427: 3422: 3417: 3412: 3411: 3408: 3407: 3402: 3397: 3396: 3390: 3389: 3383: 3382: 3379: 3378: 3375: 3373: 3369: 3368: 3361: 3360: 3354: 3353: 3350: 3349: 3344: 3339: 3338: 3332: 3331: 3328: 3327: 3322: 3314: 3313: 3307: 3306: 3300: 3297: 3296: 3293: 3292: 3284: 3283: 3275: 3274: 3266: 3265: 3262: 3260: 3256: 3255: 3248: 3247: 3244: 3243: 3240: 3238: 3234: 3233: 3226: 3225: 3219: 3218: 3215: 3214: 3211: 3209: 3207: 3198: 3197: 3191: 3190: 3187: 3186: 3178: 3177: 3174: 3173: 3172: 3171:Saurophaganax 3165: 3164: 3161: 3160: 3157: 3155: 3151: 3150: 3143: 3142: 3136: 3135: 3129: 3126: 3125: 3119: 3118: 3115: 3114: 3111: 3110: 3109: 3102: 3101: 3095: 3094:Allosauroidea 3092: 3091: 3088: 3087: 3084: 3082: 3078: 3077:Coelurosauria 3072: 3071: 3065:Avetheropoda 3064: 3063: 3060: 3059: 3056: 3054: 3050: 3044: 3043: 3037: 3036: 3032: 3031: 3027: 3025: 3024: 3019: 3015: 3014: 3009: 3005: 3001: 3000: 2999:Compsognathus 2995: 2994: 2989: 2988: 2983: 2982: 2977: 2976: 2971: 2961: 2960: 2952: 2951: 2943: 2942: 2934: 2933: 2925: 2924: 2916: 2915: 2912: 2910: 2906: 2900: 2899: 2896: 2895: 2892: 2890: 2886: 2885:Spinosauridae 2880: 2879: 2873: 2872: 2869: 2868: 2865: 2863: 2859: 2853: 2852: 2846: 2845: 2842: 2841: 2833: 2832: 2824: 2823: 2820: 2819: 2818: 2811: 2810: 2807: 2806: 2803: 2802: 2801: 2794: 2793: 2787: 2786: 2783: 2782: 2779: 2778: 2777: 2770: 2769: 2763: 2762: 2756: 2753: 2752: 2749: 2748: 2740: 2739: 2731: 2730: 2722: 2721: 2713: 2712: 2707: 2702: 2697: 2696: 2693: 2692: 2689: 2688: 2687: 2680: 2679: 2673: 2672: 2669: 2668: 2665: 2663: 2659: 2658: 2651: 2650: 2644: 2643: 2640: 2639: 2636: 2634: 2630: 2624: 2623: 2617: 2616: 2613: 2612: 2609: 2608: 2607: 2600: 2599: 2593: 2592:Allosauroidea 2590: 2589: 2583: 2580: 2579: 2576: 2575: 2572: 2570: 2566: 2565:Coelurosauria 2560: 2559: 2553: 2552: 2548: 2547: 2543: 2541: 2536: 2526: 2525: 2517: 2516: 2508: 2507: 2499: 2498: 2490: 2489: 2481: 2480: 2472: 2471: 2463: 2462: 2454: 2453: 2448: 2443: 2438: 2437: 2434: 2433: 2430: 2428: 2424: 2418: 2417: 2411: 2410: 2407: 2406: 2403: 2401: 2397: 2391: 2390: 2384: 2383: 2380: 2379: 2376: 2375: 2374: 2367: 2366: 2360: 2359: 2356: 2355: 2352: 2350: 2346: 2340: 2339: 2333: 2332: 2329: 2328: 2325: 2324: 2323: 2316: 2315: 2309: 2308:Allosauroidea 2306: 2305: 2302: 2301: 2298: 2296: 2292: 2286: 2285: 2279: 2278: 2275: 2274: 2271: 2269: 2265: 2264:Spinosauridae 2259: 2258: 2252: 2251: 2248: 2247: 2244: 2243: 2242: 2235: 2234: 2228: 2225: 2224: 2220: 2218: 2214: 2212: 2207: 2203: 2199: 2195: 2191: 2187: 2186: 2180: 2178: 2174: 2169: 2167: 2164: 2160: 2156: 2152: 2151: 2146: 2145: 2140: 2139: 2134: 2130: 2126: 2122: 2121: 2116: 2115: 2104: 2103: 2095: 2094: 2086: 2085: 2077: 2076: 2068: 2067: 2059: 2058: 2050: 2049: 2044: 2039: 2034: 2033: 2030: 2029: 2024: 2019: 2018: 2012: 2011: 2005: 2004: 2001: 2000: 1995: 1990: 1989: 1983: 1982: 1976: 1975: 1972: 1971: 1963: 1962: 1959: 1958: 1957: 1950: 1949: 1946: 1945: 1942: 1941: 1940: 1933: 1932: 1926: 1925: 1919: 1916: 1915: 1912: 1911: 1908: 1906: 1902: 1896: 1895: 1889: 1886: 1885: 1882: 1881: 1878: 1876: 1872: 1866: 1865: 1859: 1856: 1855: 1852: 1851: 1848: 1846: 1842: 1836: 1835: 1828: 1827:Allosauroidea 1825: 1824: 1820: 1818: 1813: 1811: 1807: 1806:Maastrichtian 1803: 1799: 1798: 1793: 1788: 1786: 1782: 1778: 1774: 1770: 1766: 1762: 1761: 1755: 1751: 1747: 1743: 1739: 1735: 1731: 1727: 1723: 1718: 1716: 1715: 1710: 1709: 1704: 1703: 1698: 1694: 1690: 1689: 1684: 1680: 1675: 1673: 1665: 1658: 1657: 1649: 1648: 1640: 1639: 1631: 1630: 1625: 1620: 1615: 1614: 1611: 1610: 1605: 1600: 1595: 1594: 1588: 1585: 1584: 1581: 1580: 1577: 1575: 1571: 1565: 1564: 1558: 1555: 1554: 1551: 1550: 1547: 1545: 1541: 1535: 1534: 1528: 1527:Allosauroidea 1525: 1524: 1520: 1518: 1517: 1512: 1511: 1506: 1505: 1499: 1489: 1488: 1480: 1479: 1471: 1470: 1462: 1461: 1453: 1452: 1444: 1443: 1435: 1434: 1426: 1425: 1417: 1416: 1413: 1411: 1407: 1401: 1400: 1397: 1396: 1391: 1386: 1381: 1380: 1374: 1373: 1370: 1369: 1366: 1364: 1360: 1354: 1353: 1347: 1346: 1343: 1342: 1339: 1337: 1333: 1327: 1326: 1320: 1319:Coelurosauria 1317: 1316: 1313: 1312: 1309: 1307: 1303: 1302:Allosauroidea 1299: 1293: 1292: 1286: 1283: 1282: 1279: 1278: 1275: 1273: 1269: 1263: 1262: 1256: 1253: 1252: 1249: 1248: 1245: 1243: 1239: 1233: 1232: 1226: 1223: 1222: 1219: 1218: 1215: 1213: 1209: 1203: 1202: 1196: 1195: 1192: 1191: 1188: 1186: 1181: 1174: 1173: 1166: 1165: 1160: 1159: 1155: 1148: 1146: 1144: 1143:Ken Carpenter 1140: 1136: 1132: 1128: 1127: 1122: 1118: 1114: 1110: 1109: 1104: 1100: 1096: 1092: 1088: 1084: 1083:Allosauroidea 1075: 1068: 1066: 1063: 1059: 1055: 1051: 1047: 1043: 1039: 1035: 1031: 1027: 1023: 1022: 1017: 1013: 1011: 1007: 1003: 1002:cranial vault 999: 994: 990: 987: 982: 980: 975: 970: 968: 963: 959: 950: 943: 941: 939: 938: 933: 929: 925: 924:Coelurosauria 921: 917: 913: 909: 905: 904:megalosaurids 901: 897: 896: 892: 884: 882: 880: 879: 874: 873:megaraptorans 870: 866: 862: 858: 857:Maastrichtian 854: 850: 845: 843: 839: 835: 831: 827: 823: 818: 816: 812: 808: 804: 800: 796: 795:Allosauroidea 791: 789: 785: 781: 777: 773: 765: 760: 759:Allosauroidea 755: 752: 748: 738: 737: 730: 728: 727: 720: 718: 717: 710: 708: 707: 700: 698: 697: 690: 688: 687: 680: 678: 677: 670: 668: 667: 660: 658: 657: 650: 648: 647: 646:Compsognathus 640: 638: 637: 630: 628: 627: 626:Bahariasaurus 620: 618: 617: 610: 609: 608: 607: 601: 600: 588: 587: 580: 577: 576: 569: 567: 566: 559: 557: 556: 549: 547: 546: 539: 536: 535: 528: 527: 523: 517: 515: 514: 507: 506: 502: 496: 494: 493: 486: 484: 483: 482:Saurophaganax 476: 473: 472: 465: 462: 461: 460:Siamotyrannus 454: 451: 450: 443: 441: 440: 433: 431: 430: 423: 422: 418: 417:Allosauroidea 412: 408: 402: 401: 394: 391: 390: 383: 380: 379: 372: 369: 368: 361: 358: 357: 350: 349: 347: 342: 337: 332: 326: 323: 320: 319: 316: 313: 310: 307: 306: 303: 300: 297: 294: 293: 290: 287: 284: 281: 280: 277: 274: 271: 268: 267: 264: 261: 258: 257: 254: 251: 248: 247: 244: 241: 238: 237: 232: 227: 223: 220: 219: 214: 213: 208: 207: 202: 201: 195: 191: 186: 181: 177: 176:Late Toarcian 170: 165: 160: 155: 150: 145: 140: 135: 130: 125: 120: 115: 109: 102: 98: 89: 86: 82: 78: 74: 67: 59: 48: 44: 40: 33: 19: 9621: 9585: 9514: 9507: 9500: 9493: 9486: 9481:Ornithomimus 9479: 9472: 9465: 9458: 9451: 9444: 9437: 9419:Tyrannomimus 9417: 9409: 9401: 9394: 9389:Deinocheirus 9387: 9380: 9360: 9353: 9346: 9322: 9314: 9309:Nqwebasaurus 9307: 9302:Nedcolbertia 9300: 9293: 9286: 9280:Calamosaurus 9278: 9272:Aviatyrannis 9270: 9263: 9238: 9208: 9201: 9194: 9186: 9179: 9172: 9165: 9157: 9152:Aristosuchus 9150: 9144:Aniksosaurus 9142: 9124:Aniksosaurus 9122: 9092: 9085: 9077: 9023: 8988: 8981: 8974: 8967: 8893: 8886: 8879: 8859: 8852: 8847:Dynamoterror 8845: 8825: 8818: 8798: 8791: 8771: 8766:Asiatyrannus 8764: 8740: 8733: 8704:Dryptosaurus 8702: 8695: 8688: 8657: 8650: 8643: 8635: 8628: 8621: 8614: 8607: 8600: 8595:Jinbeisaurus 8593: 8586: 8579: 8571: 8566:Aviatyrannis 8564: 8557: 8523: 8518:Sinotyrannus 8516: 8509: 8502: 8495: 8474: 8467: 8445: 8438: 8397: 8390: 8383: 8375: 8368: 8360: 8352: 8347:Bicentenaria 8345: 8337: 8332:Asiamericana 8330: 8323: 8264: 8205: 8198: 8191: 8184: 8177: 8170: 8163: 8142: 8134: 8127: 8120: 8113: 8107:Deltadromeus 8105: 8097: 8090: 8039:Tyrannotitan 8037: 8030: 8023: 8016: 7996: 7972: 7967:Taurovenator 7965: 7958: 7951: 7946:Lajasvenator 7944: 7937: 7930: 7925:Concavenator 7923: 7915: 7908: 7883: 7875: 7868: 7860: 7852: 7832: 7825: 7817: 7810: 7803: 7796: 7771: 7763: 7755: 7748: 7741:Allosauridae 7709: 7702: 7697:Shidaisaurus 7695: 7688: 7681: 7661: 7653: 7628: 7620: 7613: 7606: 7575: 7567: 7559: 7552: 7539: 7525: 7517: 7505:Avetheropoda 7492:Avetheropoda 7463: 7422:Avetheropoda 7389: 7382: 7375:Spinosaurini 7362: 7355: 7348: 7341: 7333: 7305: 7298: 7291: 7270: 7262: 7255: 7250:Protathlitis 7248: 7241: 7216: 7209: 7202: 7194: 7159: 7152: 7145: 7140:Leshansaurus 7138: 7131: 7124: 7104: 7097: 7092:Megalosaurus 7090: 7085:Duriavenator 7083: 7063: 7038: 7022:Megalosauria 7003: 6996: 6991:Marshosaurus 6989: 6984:Condorraptor 6982: 6962: 6921: 6914: 6906: 6899: 6892: 6885: 6878: 6871: 6865:Cruxicheiros 6863: 6856: 6848: 6840: 6832: 6697: 6692:Quilmesaurus 6690: 6683: 6676: 6668: 6661: 6654: 6647: 6640: 6620: 6613: 6605: 6598: 6591: 6584: 6577: 6570: 6563: 6555: 6544:Brachyrostra 6530: 6510: 6502: 6495: 6488: 6481: 6473: 6466: 6446: 6441:Tralkasaurus 6439: 6432: 6425: 6418: 6411: 6404: 6397: 6390: 6382: 6375: 6343:Vespersaurus 6341: 6336:Velocisaurus 6334: 6327: 6320: 6313: 6293: 6286: 6279: 6259: 6252: 6245: 6237: 6231:Deltadromeus 6229: 6221: 6215:Compsosuchus 6213: 6206: 6198: 6190: 6165: 6157: 6151:Lametasaurus 6149: 6141: 6107: 6102:Ceratosaurus 6100: 6080: 6072: 6065: 6057: 6050: 6034:Ceratosauria 6019: 6012: 5937: 5930: 5888: 5880: 5873: 5865: 5857: 5850: 5843: 5836: 5816: 5811:Podokesaurus 5809: 5802: 5795: 5787: 5763: 5755: 5748: 5741: 5734: 5727: 5720: 5713: 5706: 5699: 5694:Gojirasaurus 5692: 5687:Dracovenator 5685: 5677: 5670: 5658:Neotheropoda 5643: 5635: 5627: 5621:Guaibasaurus 5619: 5612: 5604: 5596: 5590:Chindesaurus 5588: 5478: 5469: 5345:(1): 71–78. 5342: 5338: 5331: 5288: 5284: 5220: 5216: 5212: 5206: 5187: 5183: 5173: 5136: 5132: 5108: 5104: 5094: 5085:11336/102037 5059: 5055: 5049: 5035:cite journal 5010: 5006: 4954: 4950: 4944: 4922:the original 4899: 4895: 4886: 4882: 4874: 4865: 4861: 4851: 4806: 4802: 4754: 4750: 4740: 4699: 4695: 4689: 4662: 4658: 4651: 4598: 4594: 4584: 4575: 4571: 4565: 4527:(1): 71–78. 4524: 4520: 4500: 4496: 4487: 4479: 4471: 4466: 4441: 4437: 4424: 4399: 4395: 4382: 4337: 4333: 4327: 4306:, retrieved 4284: 4274: 4233: 4229: 4219: 4186: 4182: 4171: 4161: 4142: 4138: 4131: 4090: 4086: 4073: 4040: 4036: 3996: 3992: 3948: 3944: 3938: 3905: 3902:Paleobiology 3901: 3883: 3864: 3860: 3854: 3840: 3807: 3803: 3793: 3785: 3781: 3777: 3772: 3747: 3743: 3736: 3719: 3715: 3709: 3697:. Retrieved 3693: 3683: 3670: 3630:(1): 18826. 3627: 3623: 3574: 3561: 3550: 3529: 3526: 3519: 3510: 3393: 3365: 3364: 3335: 3310: 3298: 3252: 3251: 3230: 3229: 3203: 3202:MNN GAD1-2 ( 3201: 3169: 3168: 3147: 3146: 3106: 3105: 3093: 3075: 3047: 3033: 3021: 3011: 3008:abelisauroid 3004:Avetheropoda 2997: 2991: 2985: 2979: 2973: 2970:coelurosaurs 2967: 2903: 2883: 2856: 2815: 2814: 2798: 2797: 2776:Marshosaurus 2774: 2773: 2684: 2683: 2655: 2654: 2627: 2604: 2603: 2591: 2581: 2563: 2549: 2540:monophyletic 2532: 2423:Allosauridae 2421: 2394: 2371: 2370: 2343: 2320: 2319: 2307: 2289: 2262: 2239: 2238: 2226: 2211:Megalosaurus 2209: 2205: 2202:paraphyletic 2183: 2181: 2176: 2172: 2170: 2158: 2155:Deltadromeus 2154: 2148: 2142: 2136: 2132: 2125:coelurosaurs 2118: 2112: 2110: 2015: 1986: 1954: 1953: 1939:Deltadromeus 1937: 1936: 1917: 1899: 1887: 1871:Allosauridae 1869: 1857: 1839: 1826: 1816: 1814: 1795: 1789: 1780: 1776: 1772: 1768: 1764: 1758: 1753: 1749: 1745: 1741: 1737: 1733: 1729: 1725: 1721: 1719: 1714:Deltadromeus 1712: 1706: 1700: 1697:coelurosaurs 1686: 1678: 1676: 1669: 1586: 1570:Allosauridae 1568: 1556: 1538: 1526: 1514: 1508: 1502: 1495: 1404: 1357: 1330: 1301: 1297: 1296: 1285:Avetheropoda 1266: 1238:Ceratosauria 1236: 1206: 1177: 1164:Neotheropoda 1162: 1152: 1124: 1120: 1117:Kevin Padian 1106: 1082: 1080: 1062:Concavenator 1061: 1054:Concavenator 1053: 1049: 1029: 1021:Concavenator 1019: 1015: 1014: 1009: 1005: 983: 971: 955: 935: 916:reclassified 908:spinosaurids 900:polyphyletic 895:Teratosaurus 893: 888: 876: 869:abelisaurids 859:(70–66 mya) 846: 819: 807:paraphyletic 794: 792: 771: 770: 758: 757: 734: 724: 714: 704: 696:Nedcolbertia 694: 684: 674: 664: 656:Deltadromeus 654: 644: 634: 624: 614: 584: 573: 563: 553: 543: 532: 521: 513:Allosauridae 511: 500: 490: 480: 469: 458: 447: 437: 427: 416: 407:paraphyletic 398: 387: 376: 365: 354: 330: 321: 315:Avetheropoda 308: 295: 282: 269: 216: 212:Concavenator 210: 204: 198: 106:174–89  85: 58: 46: 37:This is the 31: 9652:Carnosauria 9646:Wikispecies 9622:Carnosauria 9546:Maniraptora 9535:Maniraptora 9516:Tototlmimus 9446:Anserimimus 9396:Garudimimus 9382:Beishanlong 9324:Valdoraptor 9265:Arkansaurus 9210:Xunmenglong 9087:Juravenator 8881:Tarbosaurus 8773:Nanuqsaurus 8742:Gorgosaurus 8659:Megaraptora 8645:Timurlengia 8377:Sciurumimus 8325:Aratasaurus 8193:Murusraptor 8115:Fukuiraptor 8078:Megaraptora 7960:Shaochilong 7932:Eocarcharia 7885:Megaraptora 7805:Lusovenator 7683:Alpkarakush 7540:Carnosauria 7391:Spinosaurus 7364:Siamosaurus 7264:Suchosaurus 7211:Iberospinus 7147:Magnosaurus 7126:Afrovenator 7099:Torvosaurus 6850:Chilesaurus 6663:Carnotaurus 6656:Caletodraco 6642:Abelisaurus 6468:Arcovenator 6315:Laevisuchus 6306:Noasaurinae 6208:Berthasaura 6179:Noasauridae 6109:Genyodectes 6074:Lukousaurus 6067:Jubbulpuria 5867:Panguraptor 5845:Coelophysis 5838:Camposaurus 5797:Panguraptor 5789:Dracoraptor 5765:Zupaysaurus 5750:Tachiraptor 5729:Sarcosaurus 5629:Nhandumirim 5398:Carnosauria 5246:11336/19258 5190:: 167–184. 5062:: 174–215. 5013:(1): 1–19. 4957:: e12727≠. 4917:11336/48895 4862:Ameghiniana 4680:11336/19445 4444:: 435–489. 3788:(1): 61-66. 3750:: 131–142. 3722:: 515–524. 3570:land bridge 2987:Sciurumimus 2981:Juravenator 2817:Sciurumimus 2582:Carnosauria 2227:Carnosauria 2166:ceratosaurs 2144:Deinonychus 2038:Megaraptora 1792:Megaraptora 1765:Fukuiraptor 1734:Fukuiraptor 1693:Megaraptora 1406:Maniraptora 1298:Carnosauria 1139:Phil Currie 1103:Paul Sereno 1095:Phil Currie 1030:Allosaurus, 912:ceratosaurs 891:rauisuchian 861:Bauru Group 822:eye sockets 776:carnivorous 772:Carnosauria 763:Marsh, 1878 736:Megaraptora 716:Sciurumimus 676:Juravenator 545:Lusovenator 420:Marsh, 1878 331:Carnosauria 66:Carnotaurus 9689:Carnosaurs 9683:Categories 9474:Gallimimus 9403:Harpymimus 9348:Harpymimus 8969:Iliosuchus 8939:see below↓ 8784:Alioramini 8602:Juratyrant 8588:Eotyrannus 8573:Bagaraatan 8525:Yutyrannus 8459:Coeluridae 8385:Vayuraptor 8231:see below↓ 8207:Tratayenia 8200:Orkoraptor 8186:Megaraptor 8092:Aoniraptor 8025:Mapusaurus 7953:Sauroniops 7917:Altispinax 7870:Neovenator 7812:Siamraptor 7798:Datanglong 7765:Epanterias 7757:Antrodemus 7750:Allosaurus 7731:Allosauria 7554:Altispinax 7519:Gasosaurus 7431:see below↓ 7307:Suchomimus 6908:Sinosaurus 6743:see below↓ 6699:Viavenator 6649:Aucasaurus 6586:Ilokelesia 6557:Dahalokely 6512:Rajasaurus 6490:Indosaurus 6483:Genusaurus 6475:Dahalokely 6399:Indosuchus 6295:Limusaurus 6254:Ligabueino 6247:Kiyacursor 6239:Genusaurus 6223:Dahalokely 6143:Betasuchus 6014:Dornraptor 5914:see below↓ 5890:Segisaurus 5743:Sinosaurus 5606:Eodromaeus 5535:see below↓ 5483:Dinosauria 5465:Sauropsida 4883:Megaraptor 4578:(1): 1–73. 4308:2024-03-01 4152:10272/8751 3602:References 3531:Mapusaurus 3395:Neovenator 3367:Allosaurus 3316:(juvenile) 3299:Allosauria 3023:Sauroniops 2686:Neovenator 2657:Allosaurus 2535:Andrea Cau 2213:bucklandii 2177:Neovenator 2173:Neovenator 2150:Allosaurus 2120:Megaraptor 2114:Neovenator 1988:Neovenator 1858:Allosauria 1810:Cretaceous 1797:Orkoraptor 1785:"wildcard" 1781:Orkoraptor 1777:Megaraptor 1754:Neovenator 1738:Orkoraptor 1730:Megaraptor 1722:Neovenator 1688:Neovenator 1557:Allosauria 1121:Allosaurus 1113:Neornithes 1108:Allosaurus 1052:specimen. 1050:Allosaurus 1026:integument 1016:Allosaurus 991:, a short 986:mandibular 958:pubic boot 937:Allosaurus 932:tetanurans 910:, and the 788:Cretaceous 565:Siamraptor 555:Neovenator 534:Datanglong 504:Paul, 1988 501:Allosauria 405:(possibly 378:Gasosaurus 356:Altispinax 344:Subgroups 289:Saurischia 276:Dinosauria 218:Allosaurus 91:Carnosaurs 18:Carnosaurs 9550:includes 9495:Rativates 9488:Qiupalong 9188:Scipionyx 9181:Mirischia 8854:Lythronax 8793:Alioramus 8447:Labocania 8165:Aerosteon 7711:Sinraptor 7615:Erectopus 7350:Irritator 6935:Orionides 6820:Tetanurae 6807:Tetanurae 6734:Tetanurae 6670:Elemgasem 6593:Llukalkan 6572:Elemgasem 6329:Noasaurus 6192:Afromimus 6000:Averostra 5987:Averostra 5905:Averostra 5757:Velocipes 5576:Theropoda 5563:Theropoda 5526:Theropoda 5451:Kingdom: 5435:Theropoda 5263:133485236 5255:0031-0220 5027:0375-7633 4902:: 49–61. 4724:1477-2019 4625:1932-6203 4258:1476-4687 4203:1464-7931 4139:Geogaceta 3869:CiteSeerX 3586:Bone Wars 3254:Sinraptor 3035:Tetanurae 2993:Scipionyx 2551:Tetanurae 2190:Tetanurae 2163:noasaurid 1773:Aerosteon 1750:Aerosteon 1742:Aerosteon 1691:with the 1498:cladogram 1255:Tetanurae 1225:Averostra 1126:Sinraptor 1006:Sinraptor 998:denticles 928:cladistic 926:. Modern 815:Tetanurae 790:periods. 780:dinosaurs 778:theropod 706:Scipionyx 686:Labocania 439:Erectopus 336:von Huene 302:Theropoda 249:Kingdom: 243:Eukaryota 200:Sinraptor 174:Possible 9631:Wikidata 9079:Coelurus 8504:Kileskus 8497:Guanlong 8469:Coelurus 8362:Gualicho 7862:Gualicho 7243:Baryonyx 6579:Guemesia 6406:Kryptops 6167:Ozraptor 5804:Pendraig 5459:Chordata 5457:Phylum: 5453:Animalia 5375:22646156 5367:19826771 5323:86314252 5165:26020001 5139:: e940. 5111:: 81–95. 4983:35821895 4843:27410683 4803:PLOS ONE 4781:24264527 4757:: 2827. 4732:85354215 4643:27439002 4595:PLOS ONE 4557:22646156 4549:19826771 4374:21445312 4334:PLOS ONE 4266:20829793 4211:34991180 4123:39658297 4065:85354215 4013:10381873 3973:35111050 3965:16380967 3930:86326069 3867:: 5–61. 3832:85354215 3662:31827108 3580:See also 3206:Kryptops 3013:Kryptops 2159:Gualicho 2138:Guanlong 1956:Gualicho 1819:(2012). 1802:Mesozoic 1702:Gualicho 1129:, their 1111:than to 1046:sauropod 1038:mandible 993:quadrate 989:fenestra 853:Turonian 784:Jurassic 751:Synonyms 666:Gualicho 263:Chordata 259:Phylum: 253:Animalia 239:Domain: 182:records. 43:reviewed 9637:Q131190 8637:Timimus 8399:Zuolong 8129:Rapator 8032:Meraxes 7357:Oxalaia 6678:Koleken 5701:Lepidus 5463:Class: 5347:Bibcode 5293:Bibcode 5225:Bibcode 5156:4435507 5064:Bibcode 4974:9271276 4834:4943716 4811:Bibcode 4759:Bibcode 4704:Bibcode 4634:4954680 4603:Bibcode 4529:Bibcode 4446:Bibcode 4404:Bibcode 4365:3061882 4342:Bibcode 4238:Bibcode 4166:85-101. 4115:8662584 4095:Bibcode 4087:Science 4045:Bibcode 3993:Science 3910:Bibcode 3812:Bibcode 3784:, n.s. 3752:Bibcode 3699:5 March 3653:6906444 3632:Bibcode 3557:Pangaea 3018:chimera 1829:  1182:  1167:  962:chevron 944:Anatomy 914:) were 9288:Hexing 8976:Kakuru 8581:Dilong 6622:Thanos 6600:Niebla 6420:Rugops 6413:Kurupi 5373:  5365:  5321:  5261:  5253:  5163:  5153:  5025:  4981:  4971:  4841:  4831:  4779:  4730:  4722:  4641:  4631:  4623:  4555:  4547:  4372:  4362:  4299:  4264:  4256:  4230:Nature 4209:  4201:  4121:  4113:  4063:  4011:  3971:  3963:  3928:  3871:  3830:  3660:  3650:  3566:Apulia 3010:genus 2996:, and 2133:et al. 1817:et al. 1783:was a 1744:, and 1513:, and 1178:  1161:  1085:. The 1042:scutes 1034:scales 906:, the 865:Brazil 834:pelvis 338:, 1920 9666:91967 9552:birds 8609:Moros 8136:Siats 7877:Siats 7819:Siats 5479:Clade 5470:Clade 5371:S2CID 5319:S2CID 5259:S2CID 5133:PeerJ 4951:PeerJ 4925:(PDF) 4892:(PDF) 4728:S2CID 4553:S2CID 4493:(PDF) 4434:(PDF) 4392:(PDF) 4119:S2CID 4083:(PDF) 4061:S2CID 3969:S2CID 3926:S2CID 3828:S2CID 3312:Aorun 3016:as a 2975:Aorun 2198:grade 1760:Siats 1087:clade 1028:. In 842:tibia 838:femur 826:skull 799:clade 726:Siats 616:Aorun 322:Clade 309:Clade 296:Clade 283:Clade 270:Clade 9544:see 8179:Maip 5637:Tawa 5512:see 5363:PMID 5251:ISSN 5184:Gaia 5161:PMID 5041:link 5023:ISSN 4979:PMID 4885:and 4839:PMID 4777:PMID 4720:ISSN 4639:PMID 4621:ISSN 4545:PMID 4370:PMID 4297:ISBN 4262:PMID 4254:ISSN 4207:PMID 4199:ISSN 4111:PMID 4009:PMID 3961:PMID 3861:Gaia 3701:2024 3658:PMID 3551:The 2157:and 2141:and 1775:and 1767:and 1711:and 1672:2010 1496:The 1141:and 1058:ulna 1018:and 832:and 830:legs 786:and 178:and 114:PreꞒ 5355:doi 5309:hdl 5301:doi 5241:hdl 5233:doi 5192:doi 5151:PMC 5141:doi 5109:152 5080:hdl 5072:doi 5015:doi 4969:PMC 4959:doi 4912:hdl 4904:doi 4829:PMC 4819:doi 4767:doi 4712:doi 4675:hdl 4667:doi 4629:PMC 4611:doi 4537:doi 4454:doi 4412:doi 4360:PMC 4350:doi 4289:doi 4246:doi 4234:467 4191:doi 4147:hdl 4103:doi 4091:272 4053:doi 4001:doi 3997:284 3953:doi 3949:267 3918:doi 3820:doi 3760:doi 3724:doi 3648:PMC 3640:doi 3538:or 1779:. 1115:." 1101:by 1060:of 863:in 844:). 45:on 9685:: 9663:: 9648:: 9633:: 5481:: 5472:: 5369:. 5361:. 5353:. 5343:97 5341:. 5317:. 5307:. 5299:. 5287:. 5283:. 5271:^ 5257:. 5249:. 5239:. 5231:. 5221:89 5219:. 5188:15 5186:. 5182:. 5159:. 5149:. 5135:. 5131:. 5117:^ 5107:. 5103:. 5078:. 5070:. 5060:45 5058:. 5037:}} 5033:{{ 5011:63 5009:. 5005:. 4991:^ 4977:. 4967:. 4955:10 4953:. 4949:. 4933:^ 4910:. 4900:74 4898:. 4894:. 4866:49 4864:. 4860:. 4837:. 4827:. 4817:. 4807:11 4805:. 4801:. 4789:^ 4775:. 4765:. 4753:. 4749:. 4726:. 4718:. 4710:. 4700:10 4698:. 4673:. 4663:11 4661:. 4637:. 4627:. 4619:. 4609:. 4599:11 4597:. 4593:. 4576:12 4574:. 4551:. 4543:. 4535:. 4525:97 4523:. 4509:^ 4501:22 4499:. 4495:. 4452:. 4442:25 4440:. 4436:. 4410:. 4400:30 4398:. 4394:. 4368:. 4358:. 4348:. 4336:. 4332:. 4316:^ 4295:, 4283:, 4260:. 4252:. 4244:. 4232:. 4228:. 4205:. 4197:. 4187:97 4185:. 4181:. 4145:. 4143:38 4117:. 4109:. 4101:. 4089:. 4085:. 4059:. 4051:. 4041:10 4039:. 4021:^ 4007:. 3995:. 3981:^ 3967:. 3959:. 3947:. 3924:. 3916:. 3906:38 3904:. 3892:^ 3865:15 3863:. 3826:. 3818:. 3808:10 3806:. 3802:. 3758:. 3748:40 3746:. 3720:84 3718:. 3692:. 3656:. 3646:. 3638:. 3626:. 3622:. 3610:^ 3210:) 2990:, 2984:, 2978:, 2168:. 1740:, 1736:, 1732:, 1724:, 1705:, 1519:. 1507:, 1300:/ 1123:, 1012:. 761:? 403:? 324:: 311:: 298:: 285:: 272:: 215:, 209:, 203:, 164:Pg 108:Ma 103:, 99:– 41:, 9414:? 9406:? 9319:? 9283:? 9275:? 9243:? 9191:? 9162:? 9147:? 9082:? 8662:? 8640:? 8576:? 8479:? 8461:? 8450:? 8380:? 8365:? 8357:? 8342:? 8146:? 8138:? 8110:? 8102:? 8080:? 7920:? 7888:? 7880:? 7865:? 7857:? 7822:? 7776:? 7768:? 7760:? 7658:? 7633:? 7625:? 7580:? 7572:? 7564:? 7530:? 7522:? 7338:? 7275:? 7267:? 7221:? 7199:? 7164:? 7043:? 7008:? 6911:? 6868:? 6853:? 6845:? 6837:? 6673:? 6610:? 6560:? 6535:? 6507:? 6478:? 6387:? 6242:? 6234:? 6226:? 6218:? 6203:? 6195:? 6170:? 6162:? 6154:? 6146:? 6077:? 6062:? 6024:? 5885:? 5870:? 5862:? 5792:? 5760:? 5682:? 5648:? 5640:? 5632:? 5624:? 5609:? 5601:? 5593:? 5427:e 5420:t 5413:v 5377:. 5357:: 5349:: 5325:. 5311:: 5303:: 5295:: 5289:6 5265:. 5243:: 5235:: 5227:: 5200:. 5194:: 5167:. 5143:: 5137:3 5088:. 5082:: 5074:: 5066:: 5043:) 5029:. 5017:: 4985:. 4961:: 4914:: 4906:: 4845:. 4821:: 4813:: 4783:. 4769:: 4761:: 4755:4 4734:. 4714:: 4706:: 4683:. 4677:: 4669:: 4645:. 4613:: 4605:: 4559:. 4539:: 4531:: 4460:. 4456:: 4448:: 4418:. 4414:: 4406:: 4376:. 4352:: 4344:: 4338:6 4291:: 4268:. 4248:: 4240:: 4213:. 4193:: 4155:. 4149:: 4125:. 4105:: 4097:: 4067:. 4055:: 4047:: 4015:. 4003:: 3975:. 3955:: 3932:. 3920:: 3912:: 3877:. 3848:. 3834:. 3822:: 3814:: 3786:6 3766:. 3762:: 3754:: 3730:. 3726:: 3703:. 3664:. 3642:: 3634:: 3628:9 3208:" 3204:" 732:† 722:† 712:† 702:† 692:† 682:† 672:† 662:† 652:† 642:† 632:† 622:† 612:† 582:† 578:? 571:† 561:† 551:† 541:† 537:? 530:† 519:† 509:† 498:† 488:† 478:† 474:? 467:† 463:? 456:† 452:? 445:† 435:† 425:† 414:† 409:) 396:† 392:? 385:† 381:? 374:† 370:? 363:† 359:? 352:† 328:† 169:N 159:K 154:J 149:T 144:P 139:C 134:D 129:S 124:O 119:Ꞓ 83:. 68:. 49:. 20:)

Index

Carnosaurs
latest accepted revision
reviewed
Carnotaurus
Carnosaur (novel)
Carnosaur (film)
Carnosaur (series)
Middle Jurassic
Late Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N
Late Toarcian
Late Maastrichtian

Sinraptor
Acrocanthosaurus
Concavenator
Allosaurus
Scientific classification

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