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118:
384:
The dinosaur's tail is long and feathered, with end feathers up to 29.27 centimetres (11.52 in) long, equalling approximately 30% of the length of the animal's skeleton. This length exceeds the previous record length of 7 inches (18 cm) for feathers from non-avian dinosaurs. The elongated
526:
and other microraptorines add to the evidence that dinosaurs had evolved a host of morphological features and behaviours such as feathers, hollow bones, nesting behaviour, and possibly flight, before they evolved in birds.
380:
had feathers all over its body, including forelimbs and hindlimbs, which gives the appearance of having two pairs of wings. The presence of long feathers on all four limbs also suggests that these dinosaurs could fly.
635:
Nicholas R. Longrich; David M. Martill; Brian Andres (2018). "Late
Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary". PLOS Biology. 16 (3):
385:
tail feathers are thought to have helped provide softer and safer landings. They could have controlled pitch, a feature useful for a heavier animal, that would have attained a faster gliding speed.
510:
large feathered limbs and elongated bony tails with long tail feathers. These features caused experts to hypothesize that miniaturisation was an essential evolutionary step to enable flight.
394:, the morphology of microraptorines suggests a different aerodynamic model from that of modern birds, which have characteristically bald legs and exhibit stable flight using two wings only.
365:, South Africa reveals that the specimen was a fully grown adult, approximately 1.2 metres (3.9 ft) long and estimated to weigh 4 kilograms (8.8 lb), roughly the size of a
498:, a raven-sized dinosaur, approximately 1 kilogram (2.2 lb) in weight, was the first of the microraptorines to be discovered. It shared with species such as
721:
450:
1779:
839:
1794:
453:. The holotype consists of a rather complete skeleton with skull, compressed on a plate and counterplate. It shows extensive remains of the
353:
was the largest four-winged dinosaur known and among the largest
Mesozoic flying paravians, volant true birds seldom approaching its size.
1679:
773:
536:
483:
The Jehol Biota are found in the richest fossil beds for feathered dinosaurs in the world which have yielded notable species such as
1799:
569:
Gang Han; Luis M. Chiappe; Shu-An Ji; Michael Habib; Alan H. Turner; Anusuya
Chinsamy; Xueling Liu & Lizhuo Han (15 July 2014).
489:, a feathered tyrannosauroid with a length of 27 to 30 feet (8.2 to 9.1 m) and the largest feathered dinosaur known to date.
1656:
1645:
832:
1662:
1789:
138:
1651:
660:
571:"A new raptorial dinosaur with exceptionally long feathering provides insights into dromaeosaurid flight performance"
406:, in moist temperate forest, primarily vegetated by ginkgos and conifers, with hot, dry summers and frosty winters.
400:
is thought to have existed alongside a variety of predatory and herbivorous dinosaurs of the Jehol Biota, including
687:
446:
1784:
936:
825:
287:
258:
477:
373:
the largest four-winged microraptorine known, exceeding previously known specimens in size by at least 60%.
362:
1706:
1751:
1689:
1738:
990:
1407:
799:
1144:
963:
743:
458:
422:
133:
1756:
1604:
1495:
1431:
1327:
1120:
592:
441:
province of China. From 2012 it was studied by an international team of scientists led by Dr.
324:
1528:
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1312:
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30:
1588:
1442:
1345:
1058:
1036:
1002:
921:
848:
611:
222:
1246:
465:. The generic name of the dinosaur combines the Chinese words for "long feather", 長羽 (
1773:
1567:
1481:
1373:
1358:
1297:
1290:
1273:
1208:
1201:
1130:
390:
366:
343:
238:
68:
388:
While elongated feathers on the hindlimbs were present in many early birds, such as
1626:
1537:
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1416:
1395:
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1319:
1186:
1179:
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442:
270:
1729:
1581:
1559:
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1466:
1423:
1388:
1366:
1305:
1216:
1151:
975:
494:
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336:
43:
1574:
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1101:
1079:
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402:
196:
88:
53:
1612:
1544:
1239:
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1137:
948:
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209:
150:
93:
37:
596:
117:
1723:
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438:
415:
339:
306:
183:
170:
83:
78:
63:
58:
48:
1619:
905:
587:
570:
98:
73:
1194:
1086:
866:
310:
160:
1700:
461:. In 2014, the specimen was described as the new genus and species
817:
744:"New feathered dinosaur from China sheds light on dinosaur flight"
470:
328:
1743:
1704:
859:
821:
688:"New feathered predatory fossil sheds light on dinosaur flight"
518:
is also the second four-winged dinosaur to be discovered after
800:"'Shaggy' Tyrannosaur Now World's Biggest Feathered Beast"
722:"Four-winged dinosaur had record-breaking tail feathers"
514:
s larger size and weight contradicts that hypothesis.
661:"Bizarre Dinosaur Had 4 'Wings,' Long Tail Feathers"
277:
1713:
1527:
1441:
1406:
1344:
1272:
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1119:
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642:
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562:
560:
558:
556:
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552:
833:
8:
451:Natural History Museum of Los Angeles County
774:"New Four-Winged Dinosaur Built for Flight"
480:honours researcher Professor Yang Yandong.
1701:
1350:
1064:
1053:
980:
953:
926:
917:
856:
840:
826:
818:
116:
20:
715:
713:
711:
709:
682:
680:
678:
586:
767:
765:
612:"Four-winged dinosaur is 'biggest ever'"
548:
305:is a genus of "four-winged", predatory
7:
323:(125 million year old) deposits in
1780:Early Cretaceous dinosaurs of Asia
537:Timeline of dromaeosaurid research
313:specimen representing the species
14:
421:, was discovered by farmers near
1685:
1684:
1661:
1655:
1650:
1644:
798:Welsh, Jennifer (4 April 2012).
137:
772:Iacurci, Jenna (15 July 2014).
724:. Australian Geographic Society
720:Pickrell, John (16 July 2014).
659:Choi, Charles (15 July 2014).
610:Morgan, James (16 July 2014).
361:Analysis of the fossil at the
349:At the time of its discovery,
1:
1795:Fossil taxa described in 2014
447:Dinosaur Institute Department
376:Like other microraptorines,
319:, which was discovered from
309:. It is known from a single
1816:
476:("robber", "seizer"). The
16:Extinct genus of dinosaurs
1674:
1642:
1353:
1067:
1052:
1033:
983:
956:
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855:
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134:Scientific classification
132:
124:
115:
23:
1800:Paleontology in Liaoning
369:. These dimensions make
335:belongs to the group of
363:University of Cape Town
1752:Paleobiology Database
575:Nature Communications
429:fossil deposits, the
342:dinosaurs called the
125:Life restoration of
1790:Feathered dinosaurs
1408:Saurornitholestinae
463:Changyuraptor yangi
459:pennaceous feathers
316:Changyuraptor yangi
282:Changyuraptor yangi
588:10.1038/ncomms5382
445:, Director of the
1767:
1766:
1707:Taxon identifiers
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1697:
1670:
1669:
1640:
1639:
1636:
1635:
1605:Saurornitholestes
1496:Saurornitholestes
1432:Saurornitholestes
1121:Halszkaraptorinae
1048:
1047:
1044:
1043:
1029:
1028:
1025:
1024:
1021:
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802:. LiveScience.com
778:Nature World News
663:. LiveScience.com
437:, located in the
325:Liaoning Province
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1529:Velociraptorinae
1475:Dromaeosauroides
1351:
1335:Zhongjianosaurus
1313:Sinornithosaurus
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486:Yutyrannus huali
431:Yixian Formation
427:Early Cretaceous
321:Early Cretaceous
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31:Early Cretaceous
29:Temporal range:
21:
1815:
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1785:Microraptorians
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1589:Luanchuanraptor
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1443:Dromaeosaurinae
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1402:
1346:Eudromaeosauria
1340:
1268:
1157:
1115:
1061:
1059:Dromaeosauridae
1040:
1037:Dromaeosauridae
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1003:Dromaeosauridae
997:
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937:Avemetatarsalia
922:Avemetatarsalia
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457:in the form of
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223:Dromaeosauridae
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1302:
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1259:
1251:
1247:Unquillosaurus
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1228:
1220:
1213:
1205:
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1191:
1183:
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750:. 15 July 2014
735:
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694:. 15 July 2014
674:
638:
628:
602:
547:
546:
544:
541:
540:
539:
532:
529:
512:Changyuraptor'
411:
408:
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296:
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28:
15:
13:
10:
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6:
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3:
2:
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1801:
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1786:
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1781:
1778:
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1775:
1758:
1753:
1749:
1745:
1740:
1736:
1731:
1725:
1721:
1720:
1718:
1716:
1715:Changyuraptor
1712:
1708:
1703:
1691:
1683:
1681:
1677:
1676:
1673:
1664:
1660:
1658:
1653:
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1591:
1590:
1586:
1584:
1583:
1579:
1577:
1576:
1572:
1570:
1569:
1568:Kansaignathus
1565:
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1557:
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1508:
1506:
1505:
1501:
1498:
1497:
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1491:
1490:
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1482:Dromaeosaurus
1479:
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1468:
1464:
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1460:
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1405:
1398:
1397:
1393:
1391:
1390:
1386:
1383:
1382:
1378:
1376:
1375:
1374:Dineobellator
1371:
1369:
1368:
1364:
1361:
1360:
1356:
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1322:
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1317:
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1310:
1308:
1307:
1303:
1300:
1299:
1298:Hesperonychus
1295:
1293:
1292:
1291:Graciliraptor
1288:
1286:
1285:
1284:Changyuraptor
1281:
1280:
1278:
1275:
1274:Microraptoria
1271:
1265:
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1260:
1257:
1256:
1252:
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1248:
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1241:
1237:
1234:
1233:
1229:
1226:
1225:
1221:
1219:
1218:
1214:
1211:
1210:
1209:Ornithodesmus
1206:
1204:
1203:
1202:Neuquenraptor
1199:
1197:
1196:
1192:
1189:
1188:
1184:
1182:
1181:
1177:
1175:
1174:
1170:
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1164:
1160:
1154:
1153:
1149:
1147:
1146:
1142:
1140:
1139:
1135:
1133:
1132:
1131:Halszkaraptor
1128:
1127:
1125:
1122:
1118:
1112:
1111:
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1104:
1103:
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1096:
1092:
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1038:
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824:
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820:
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745:
739:
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723:
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689:
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563:
561:
559:
557:
555:
553:
549:
542:
538:
535:
534:
530:
528:
525:
524:Changyuraptor
521:
517:
516:Changyuraptor
513:
509:
508:
503:
502:
497:
496:
490:
488:
487:
481:
479:
478:specific name
475:
472:
468:
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460:
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448:
444:
440:
436:
432:
428:
424:
420:
417:
409:
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404:
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398:Changyuraptor
395:
393:
392:
391:Archaeopteryx
386:
382:
379:
378:Changyuraptor
374:
372:
371:Changyuraptor
368:
364:
356:
354:
352:
347:
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344:Microraptoria
341:
338:
337:dromaeosaurid
334:
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322:
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317:
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308:
304:
303:
302:Changyuraptor
292:
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275:
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268:
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253:Changyuraptor
247:
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239:Microraptoria
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128:
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119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
25:Changyuraptor
22:
19:
1714:
1649:
1627:Velociraptor
1625:
1618:
1611:
1603:
1595:
1587:
1580:
1573:
1566:
1558:
1550:
1543:
1538:Acheroraptor
1536:
1516:
1509:
1502:
1494:
1487:
1480:
1473:
1465:
1459:Dakotaraptor
1457:
1452:Achillobator
1450:
1430:
1422:
1417:Atrociraptor
1415:
1396:Zhenyuanlong
1394:
1387:
1381:Tianyuraptor
1379:
1372:
1365:
1357:
1333:
1326:
1320:Tianyuraptor
1318:
1311:
1304:
1296:
1289:
1283:
1282:
1261:
1253:
1245:
1238:
1230:
1222:
1215:
1207:
1200:
1193:
1187:Dakotaraptor
1185:
1180:Buitreraptor
1178:
1173:Austroraptor
1171:
1163:Unenlagiinae
1150:
1143:
1136:
1129:
1110:Zhenyuanlong
1108:
1100:
1095:Tianyuraptor
1093:
1085:
1078:
1071:
1035:
901:
892:
883:
804:. Retrieved
793:
781:. Retrieved
777:
752:. Retrieved
748:ScienceDaily
747:
738:
726:. Retrieved
696:. Retrieved
691:
665:. Retrieved
631:
619:. Retrieved
615:
605:
578:
574:
523:
519:
515:
511:
505:
499:
493:
491:
484:
482:
473:
466:
462:
443:Luis Chiappe
418:
414:The fossil,
413:
401:
397:
396:
389:
387:
383:
377:
375:
370:
360:
350:
348:
332:
315:
314:
301:
300:
299:
290:
281:
280:
271:Type species
261:
252:
251:
229:
203:
190:
177:
126:
24:
18:
1582:Linheraptor
1560:Deinonychus
1511:Yurgovuchia
1467:Deinonychus
1424:Bambiraptor
1389:Vectiraptor
1367:Bambiraptor
1306:Microraptor
1217:Pamparaptor
1152:Natovenator
991:Maniraptora
976:Maniraptora
520:Microraptor
495:Microraptor
435:Jehol Biota
423:Xijianchang
357:Description
1774:Categories
1678:See also:
1552:Boreonykus
1504:Utahraptor
1255:Variraptor
1224:Pyroraptor
1102:Variraptor
1080:Pyroraptor
1012:see below↓
888:Dinosauria
879:Sauropsida
543:References
507:Xiaotingia
501:Anchiornis
455:integument
403:Yutyrannus
197:Saurischia
184:Dinosauria
1730:Q17363721
1545:Adasaurus
1240:Unenlagia
1232:Rahonavis
1138:Hulsanpes
964:Theropoda
949:Theropoda
897:Theropoda
865:Kingdom:
636:e2001663.
433:, of the
410:Discovery
307:dinosaurs
210:Theropoda
157:Kingdom:
151:Eukaryota
36:125
1744:47043812
1724:Wikidata
1690:Category
1680:Timeline
1597:Nuthetes
1518:Zapsalis
1489:Itemirus
1263:Ypupiara
1145:Mahakala
1073:Daurlong
873:Chordata
871:Phylum:
867:Animalia
692:Phys.org
616:BBC News
597:25025742
581:: 4382.
531:See also
492:In 2003
469:), with
467:cháng yǔ
439:Liaoning
416:holotype
351:C. yangi
340:theropod
333:C. yangi
216:Family:
171:Chordata
167:Phylum:
161:Animalia
147:Domain:
127:C. yangi
1620:Tsaagan
906:Paraves
877:Class:
806:16 July
783:16 July
754:16 July
728:16 July
698:16 July
667:16 July
621:16 July
449:at the
425:in the
419:HG B016
245:Genus:
105:↓
1757:299732
1359:Balaur
1328:Wulong
1195:Diuqin
1087:Shanag
595:
474:raptor
367:turkey
311:fossil
293:, 2014
291:et al.
264:, 2014
262:et al.
902:Clade
893:Clade
884:Clade
471:Latin
329:China
230:Clade
204:Clade
191:Clade
178:Clade
1613:Shri
1575:Kuru
989:see
962:see
935:see
808:2014
785:2014
756:2014
730:2014
700:2014
669:2014
623:2014
593:PMID
504:and
44:PreꞒ
1739:EoL
583:doi
288:Han
259:Han
1776::
1754::
1741::
1726::
904::
895::
886::
776:.
764:^
746:.
708:^
690:.
677:^
641:^
614:.
591:.
577:.
573:.
551:^
522:.
346:.
331:.
327:,
232::
206::
193::
180::
94:Pg
38:Ma
33:,
1608:?
1600:?
1592:?
1563:?
1555:?
1499:?
1470:?
1462:?
1427:?
1399:?
1384:?
1362:?
1323:?
1301:?
1276:?
1258:?
1250:?
1235:?
1227:?
1212:?
1190:?
1165:?
1123:?
1105:?
1090:?
841:e
834:t
827:v
810:.
787:.
758:.
732:.
702:.
671:.
625:.
599:.
585::
579:5
278:†
249:†
236:†
220:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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