127:
153:
996:
2922:
2945:
2450:
2442:
2429:
2422:
2435:
551:
pholidosaur features and has a robust tubular and slightly flattened rostrum widening abruptly around the jugals. The nasals are separated from the nares by the premaxilla, which tapers at its posterior end and extends far between the maxilla, and the supratemporal fenestrae are moderately sized, making up 25% of the skull table, differing from
483:. Shortly afterwards more material of the same specimen was found and described. In the early 2000s another locality of the Phu Kradung Formation was discovered near the village of Kham Phok, Mukdahan Province, yielding amongst other remains the anterior tip of a crocodylomorph mandible which was assigned to
538:
The holotype mandible is very robust, and its tip is spoon shaped and wider than the portion of the jaw immediately behind it. The mandible of PRC102-143 is similar in size, though less complete, and preserves the same spatulate appearance. The 3rd and 4th dentary alveoli are raised and contiguous
550:
The referred cranial material suggests a possible skull length of 1.1 m (3 ft 7 in). The complete animal could reach over 10 m (33 ft) in body length, although other estimates suggest 7–8 m (23–26 ft). The specimen shows a combination of goniopholidid and
563:
whose fenestrae take up more of the skull table's surface. The premaxillary teeth are located on the anterior half of the premaxilla and when seen in lateral view the anterior margin extends further down than the rest of the bone, creating a characteristic "beak" shape also seen in
487:
and described the pholidosaurid affinity of the specimen. This interpretation remained tentative until the discovery of previously unnoticed skull material associated with the Kham Phok mandible (specimen PRC102-143) which led to the reassignment of
576:. This along with the hook-like shape, whereas traditional goniopholidids possess a notch in this area. Both the presence of a diastema and the hook-like anterior margin of the premaxilla are considered to be pholidosaurid characteristics.
583:
is uncertain, as the Phu
Kradung Formation may either date to the Late Jurassic or Early Cretaceous. While vertebrate fossil discoveries point at a Late Jurassic origin,
3043:
2997:
595:
The phylogenetic tree below depicts
Pholidosauridae as recovered by Jouve & Jalil (2020) obtained from a matrix including 224 characters. In this tree
1206:
3073:
3048:
3078:
3068:
499:
Based on the new cranial material, Jeremy E. Martin, Komsorn
Lauprasert, Eric Buffetaut, Romain Liard and Varavudh Suteethorn erected the new genus
1244:
3063:
1457:
1299:
Martin, Jeremy E.; Antoine, Pierre-Olivier; Perrier, Vincent; Welcomme, Jean-Loup; Metais, Gregoire; Marivaux, Laurent (2019-07-04).
1340:
Racey and
Goodall (2009). "Late Palaeozoic and Mesozoic Ecosystems in SE Asia". Geological Society. London. Special Publication
1406:"A re-evaluation of goniopholidid crocodylomorph material from Central Asia: Biogeographic and phylogenetic implications"
3058:
3053:
1300:
2900:
1355:
1044:, comprising the rostrum, the preorbital region of the skull roof, the quadrates and parts of the quadratojugal, the
513:
2921:
152:
1218:
415:
1450:
2449:
2428:
3038:
2965:
2421:
3010:
1545:
1263:
455:
419:
411:
2441:
1356:"Paleocene resurrection of a crocodylomorph taxon: Biotic crises, climatic and sea level fluctuations"
2948:
1370:
467:
310:
284:
2896:
2158:
1057:
463:
2434:
1053:
458:. This mandible was at first known from the posterior portion of the bone, described and named by
2288:
2193:
1443:
1386:
339:
147:
3015:
1281:
126:
547:, Chalawan does not possess a diastema between the enlarged 4th and small 5th dentary alveoli.
3002:
2938:
2757:
2669:
2318:
1323:
1174:
1092:, suggesting that it might belong to a different species. Using an updated version of Andrade
1045:
2705:
2677:
2638:
2623:
2528:
2302:
2047:
1872:
1420:
1378:
1315:
1186:
408:
400:
45:
2830:
2772:
2612:
2559:
2508:
2272:
2142:
2082:
1956:
1893:
1858:
1752:
1089:
505:
493:
451:
259:
246:
233:
1374:
995:
2851:
2750:
2736:
2721:
2662:
2569:
2518:
2221:
2179:
2112:
2019:
1914:
1865:
1838:
1728:
1695:
1674:
1664:
1405:
1282:"Evolution and palaeoecology of crocodiles in the Mesozoic of Khorat Plateau, Thailand"
1081:
771:
459:
3032:
2837:
2781:
2743:
2655:
2601:
2590:
2498:
2311:
2165:
2098:
2010:
1992:
1963:
1935:
1900:
1886:
1879:
1810:
1796:
1775:
1654:
1644:
1390:
1173:
Martin, J. E.; Lauprasert, K.; Buffetaut, E.; Liard, R. & Suteethorn, V. (2013).
722:
705:
636:
471:
404:
397:
385:
377:
84:
41:
17:
572:. The transverse area of the premaxilla is separated from the rest of the bone by a
2860:
2538:
2374:
2367:
2332:
2325:
2281:
2257:
2235:
2228:
2151:
2128:
2091:
2068:
2061:
2026:
1985:
1977:
1928:
1849:
1824:
1803:
1782:
1530:
1505:
1138:. As the paper synonymizing the two crocodylomorphs appeared shortly before Martin
1110:
1105:
845:
805:
788:
559:
553:
292:
220:
1404:
Halliday, T.; Brandalise de
Andrade, M.; Benton, M.J. & Efimov, M.B. (2015) .
1319:
539:
and hosted fang like teeth, after which the dentary constricts, much like seen in
2988:
2874:
2867:
2809:
2728:
2714:
2684:
2381:
2360:
2353:
2346:
2243:
2214:
2207:
2200:
2186:
2172:
2121:
2054:
2040:
1970:
1949:
1907:
1831:
1817:
1761:
1743:
1717:
1624:
919:
902:
587:
data instead suggests that the
Formation represents Early Cretaceous sediments.
59:
2885:
2816:
2788:
2579:
2548:
2339:
2250:
2033:
1942:
1921:
1706:
1685:
1634:
1514:
1496:
1175:"A large pholidosaurid in the Phu Kradung Formation of north-eastern Thailand"
1115:
1005:
584:
518:
475:
348:
104:
69:
1382:
1327:
2691:
1789:
1607:
1594:
1557:
1466:
1049:
523:
447:
207:
164:
109:
53:
1425:
2982:
2823:
2647:
2481:
2468:
2392:
1490:
1135:
1061:
1037:
573:
184:
99:
94:
79:
74:
64:
1122:(based on the holotype) to the analysis did not confirm the referral of
1191:
527:
390:
194:
114:
89:
1048:
and nearly complete mandibles. It was collected from "Layer 2" of the
516:
is derived from the name of
Chalawan, a giant in the Thai folktale of
1484:
1301:"A large crocodyloid from the Oligocene of the Bugti Hills, Pakistan"
174:
2959:
1041:
994:
393:
2963:
2466:
1592:
1477:
1439:
1142:
s description of the cranial material, the taxonomic status of
138:
specimen PRC102-143 (left), and reconstruction of the skull of
450:
collected in the early 1980s from a road-cut near the town of
1435:
1249:, a mesosuchian crocodilian from the Jurassic of Thailand"
1076:, and can not be differentiated from the holotype of
2848:
2799:
2769:
2702:
2635:
2619:
2608:
2597:
2586:
2575:
2565:
2555:
2544:
2534:
2524:
2514:
2504:
2494:
2299:
2269:
2139:
2109:
2079:
2007:
1740:
1724:
1713:
1702:
1691:
1681:
1670:
1660:
1650:
1640:
1630:
1620:
299:
2972:
2884:
2847:
2798:
2768:
2701:
2634:
2479:
2391:
2298:
2268:
2138:
2108:
2078:
2006:
1848:
1751:
1739:
1605:
1556:
1529:
1354:Jouve, Stéphane & Jalil, Nour-Eddine (2020).
1451:
599:was recovered as a sister taxon to the genus
8:
1072:shares some features with other species of
2960:
2487:
2476:
2463:
1748:
1613:
1602:
1589:
1535:
1526:
1474:
1458:
1444:
1436:
1168:
1166:
1164:
1162:
1160:
1158:
125:
31:
1424:
1190:
3044:Early Cretaceous crocodylomorphs of Asia
1243:Buffetaut, E. & Ingavat, R. (1984).
1056:beds, in the Shar Teeg locality, of the
1154:
1096:(2011) phylogenetic analysis, Halliday
1238:
1236:
384:
7:
1286:PHD Thesis, Chulalongkorn University
1080:. Nevertheless, it lacks definitive
894:
870:
837:
830:
763:
756:
697:
677:
670:
628:
621:
614:
606:
1308:Journal of Vertebrate Paleontology
1064:, embedded in grey clay. Halliday
25:
3074:Taxa named by Varavudh Suteethorn
3049:Early Cretaceous reptiles of Asia
3079:Prehistoric pseudosuchian genera
2944:
2943:
2920:
2448:
2440:
2433:
2427:
2420:
422:. It contains a single species,
151:
1020:(2013) tentatively synonymized
522:that could take the form of a
434:as a possible second species.
1:
3064:Fossil taxa described in 2013
1413:Acta Palaeontologica Polonica
1320:10.1080/02724634.2019.1671427
1217:. Fossilworks. Archived from
1088:, and possibly even these of
706:Pholidosaurus schaumbergensis
3069:Taxa named by Éric Buffetaut
1280:Lauprasert, Komsorn (2006).
1262:(1): 199–206. Archived from
1134:, as it resulted in a large
772:Woodbinesuchus byersmauricei
637:Meridiosaurus vallisparadisi
454:, in the upper part of the
354:Buffetaut and Ingavat, 1980
3095:
723:Pholidosaurus purbeckensis
2933:
2918:
2490:
2475:
2462:
2417:
1616:
1601:
1588:
1538:
1525:
1473:
1040:PIN 4174‒1, a fragmented
1036:is known solely from the
1022:"Sunosuchus" shartegensis
990:"Sunosuchus" shartegensis
916:
899:
892:
875:
868:
842:
835:
828:
802:
789:Sabinosuchus coahuilensis
785:
768:
761:
754:
719:
702:
695:
682:
675:
668:
633:
626:
619:
612:
381:
345:
338:
323:
318:
298:
291:
148:Scientific classification
146:
133:
124:
34:
27:Extinct genus of reptiles
1383:10.1016/j.gr.2020.03.010
470:as a new species of the
416:Nong Bua Lamphu Province
309:(Buffetaut and Ingavat,
1247:Sunosuchus thailandicus
485:Sunosuchus thailandicus
481:Sunosuchus thailandicus
360:Sunosuchus shartegensis
1013:
806:Oceanosuchus boecensis
610:Pholidosauridae
446:was a nearly complete
3011:Paleobiology Database
2899:for extant taxa, and
1426:10.4202/app.2013.0018
1215:Paleobiology Database
998:
903:Sarcosuchus imperator
879:Chalawan thailandicus
510:Chalawan thailandicus
456:Phu Kradung Formation
431:Chalawan shartegensis
425:Chalawan thailandicus
420:northeastern Thailand
412:Phu Kradung Formation
386:[t͡ɕʰāːlāwān]
327:Chalawan shartegensis
304:Chalawan thailandicus
18:Chalawan thailandicus
1221:on 22 September 2022
570:Terminonaris robusta
442:The first fossil of
438:Discovery and naming
3059:Fossils of Thailand
3054:Cretaceous Thailand
2159:Anteophthalmosuchus
1375:2020GondR..85....1J
1211:Martin et al. 2013"
1068:(2013) stated that
1058:Govi-Altai Province
684:French Pholidosaur
2901:Extinct Crocodilia
2289:Koumpiodontosuchus
2194:Eutretauranosuchus
1245:"The lower jaw of
1192:10.1111/pala.12086
1118:. The addition of
1024:(Efimov, 1988) as
1014:
920:Sarcosuchus hartii
3026:
3025:
2966:Taxon identifiers
2957:
2956:
2939:Coelognathosuchia
2929:
2928:
2916:
2915:
2912:
2911:
2758:Musturzabalsuchus
2670:Pachycheilosuchus
2458:
2457:
2415:
2414:
2411:
2410:
2319:Brillanceausuchus
2002:
2001:
1584:
1583:
1580:
1579:
1576:
1575:
1363:Gondwana Research
1146:remains unknown.
1144:"S." shartegensis
1124:"S." shartegensis
1102:"S." shartegensis
1070:"S." shartegensis
1046:occipital condyle
1034:"S." shartegensis
985:
984:
976:
975:
967:
966:
958:
957:
949:
948:
940:
939:
931:
930:
857:
856:
846:Terminonaris spp.
817:
816:
743:
742:
734:
733:
657:
656:
648:
647:
579:The exact age of
543:. However unlike
401:mesoeucrocodylian
369:
368:
364:
355:
332:
287:
134:Known remains of
16:(Redirected from
3086:
3019:
3018:
3006:
3005:
2993:
2992:
2991:
2961:
2947:
2946:
2924:
2903:for extinct ones
2850:
2801:
2771:
2706:Allodaposuchidae
2704:
2678:Pietraroiasuchus
2639:Hylaeochampsidae
2637:
2624:Paralligatoridae
2621:
2610:
2599:
2588:
2577:
2567:
2557:
2546:
2536:
2529:Gilchristosuchus
2526:
2516:
2506:
2496:
2488:
2477:
2464:
2453:
2452:
2445:
2444:
2438:
2437:
2431:
2425:
2424:
2303:Paralligatoridae
2301:
2271:
2141:
2111:
2081:
2048:Knoetschkesuchus
2009:
1873:Arambourgisuchus
1749:
1742:
1726:
1715:
1704:
1693:
1683:
1672:
1662:
1652:
1642:
1632:
1622:
1614:
1603:
1590:
1536:
1527:
1520:
1519:
1475:
1460:
1453:
1446:
1437:
1431:
1430:
1428:
1410:
1401:
1395:
1394:
1360:
1351:
1345:
1338:
1332:
1331:
1305:
1296:
1290:
1289:
1277:
1271:
1270:
1268:
1253:
1240:
1231:
1230:
1228:
1226:
1203:
1197:
1196:
1194:
1170:
1052:(Late Jurassic)
895:
871:
838:
831:
764:
757:
698:
678:
671:
629:
622:
615:
607:
409:Early Cretaceous
388:
383:
362:
353:
330:
301:
278:
271:
258:
156:
155:
129:
119:
56:
46:Early Cretaceous
40:Temporal range:
32:
21:
3094:
3093:
3089:
3088:
3087:
3085:
3084:
3083:
3029:
3028:
3027:
3022:
3014:
3009:
3001:
2996:
2987:
2986:
2981:
2968:
2958:
2953:
2925:
2908:
2880:
2843:
2831:Eothoracosaurus
2794:
2773:Aegyptosuchidae
2764:
2697:
2630:
2613:Bernissartiidae
2560:Portugalosuchus
2509:Confractosuchus
2484:
2471:
2454:
2447:
2439:
2432:
2419:
2407:
2387:
2294:
2273:Bernissartiidae
2264:
2143:Goniopholididae
2134:
2104:
2083:Stomatosuchidae
2074:
1998:
1957:Phosphatosaurus
1894:Cerrejonisuchus
1859:Acherontisuchus
1844:
1753:Pholidosauridae
1735:
1610:
1597:
1572:
1552:
1521:
1480:
1479:
1469:
1464:
1434:
1408:
1403:
1402:
1398:
1358:
1353:
1352:
1348:
1339:
1335:
1314:(4): e1671427.
1303:
1298:
1297:
1293:
1279:
1278:
1274:
1266:
1251:
1242:
1241:
1234:
1224:
1222:
1205:
1204:
1200:
1172:
1171:
1156:
1152:
1120:S. thailandicus
1090:Goniopholididae
1086:S. thailandicus
1078:S. thailandicus
993:
986:
977:
968:
959:
950:
941:
932:
858:
818:
744:
735:
658:
649:
593:
536:
506:combinatio nova
494:Pholidosauridae
452:Nong Bua Lamphu
440:
403:known from the
314:
307:
277:
269:
260:Pholidosauridae
256:
247:Crocodyliformes
234:Crocodylomorpha
150:
140:C. thailandicus
136:C. thailandicus
120:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
51:
50:
48:
38:
28:
23:
22:
15:
12:
11:
5:
3092:
3090:
3082:
3081:
3076:
3071:
3066:
3061:
3056:
3051:
3046:
3041:
3031:
3030:
3024:
3023:
3021:
3020:
3007:
2994:
2978:
2976:
2970:
2969:
2964:
2955:
2954:
2952:
2951:
2941:
2934:
2931:
2930:
2927:
2926:
2919:
2917:
2914:
2913:
2910:
2909:
2907:
2906:
2905:
2904:
2890:
2888:
2882:
2881:
2879:
2878:
2871:
2864:
2856:
2854:
2852:Planocraniidae
2845:
2844:
2842:
2841:
2834:
2827:
2820:
2813:
2805:
2803:
2802:"Thoracosaurs"
2796:
2795:
2793:
2792:
2785:
2777:
2775:
2766:
2765:
2763:
2762:
2754:
2751:Massaliasuchus
2747:
2740:
2737:Ischyrochampsa
2733:
2725:
2722:Allodaposuchus
2718:
2710:
2708:
2699:
2698:
2696:
2695:
2688:
2681:
2674:
2666:
2663:Iharkutosuchus
2659:
2652:
2643:
2641:
2632:
2631:
2629:
2628:
2617:
2606:
2595:
2584:
2573:
2570:Pristichampsus
2563:
2553:
2542:
2532:
2522:
2519:Dolichochampsa
2512:
2502:
2491:
2485:
2480:
2473:
2472:
2467:
2460:
2459:
2456:
2455:
2418:
2416:
2413:
2412:
2409:
2408:
2406:
2405:
2404:
2403:
2397:
2395:
2389:
2388:
2386:
2385:
2378:
2371:
2364:
2357:
2350:
2343:
2336:
2329:
2322:
2315:
2307:
2305:
2296:
2295:
2293:
2292:
2285:
2277:
2275:
2266:
2265:
2263:
2262:
2254:
2247:
2240:
2232:
2225:
2222:Ophiussasuchus
2218:
2211:
2204:
2197:
2190:
2183:
2180:Denazinosuchus
2176:
2169:
2162:
2155:
2147:
2145:
2136:
2135:
2133:
2132:
2125:
2117:
2115:
2113:Paluxysuchidae
2106:
2105:
2103:
2102:
2095:
2087:
2085:
2076:
2075:
2073:
2072:
2065:
2058:
2051:
2044:
2037:
2030:
2023:
2020:Alligatorellus
2015:
2013:
2004:
2003:
2000:
1999:
1997:
1996:
1989:
1982:
1974:
1967:
1960:
1953:
1946:
1939:
1932:
1925:
1918:
1915:Dorbignysuchus
1911:
1904:
1897:
1890:
1883:
1876:
1869:
1866:Anthracosuchus
1862:
1854:
1852:
1846:
1845:
1843:
1842:
1839:Woodbinesuchus
1835:
1828:
1821:
1814:
1807:
1800:
1793:
1786:
1779:
1772:
1765:
1757:
1755:
1746:
1737:
1736:
1734:
1733:
1729:Thalattosuchia
1722:
1711:
1700:
1696:Tadzhikosuchus
1689:
1679:
1675:Stolokrosuchus
1668:
1665:Montsecosuchus
1658:
1648:
1638:
1628:
1617:
1611:
1606:
1599:
1598:
1593:
1586:
1585:
1582:
1581:
1578:
1577:
1574:
1573:
1571:
1570:
1569:
1568:
1562:
1560:
1554:
1553:
1551:
1550:
1549:
1548:
1539:
1533:
1523:
1522:
1518:
1517:
1508:
1499:
1493:
1487:
1478:
1471:
1470:
1465:
1463:
1462:
1455:
1448:
1440:
1433:
1432:
1419:(2): 291–312.
1396:
1346:
1333:
1291:
1272:
1269:on 2012-03-09.
1232:
1198:
1185:(4): 757–769.
1153:
1151:
1148:
1082:synapomorphies
992:
987:
983:
982:
979:
978:
974:
973:
970:
969:
965:
964:
961:
960:
956:
955:
952:
951:
947:
946:
943:
942:
938:
937:
934:
933:
929:
928:
925:
924:
915:
912:
911:
908:
907:
898:
893:
891:
888:
887:
884:
883:
874:
869:
867:
864:
863:
860:
859:
855:
854:
851:
850:
841:
836:
834:
829:
827:
824:
823:
820:
819:
815:
814:
811:
810:
801:
798:
797:
794:
793:
784:
781:
780:
777:
776:
767:
762:
760:
755:
753:
750:
749:
746:
745:
741:
740:
737:
736:
732:
731:
728:
727:
718:
715:
714:
711:
710:
701:
696:
694:
691:
690:
687:
686:
681:
676:
674:
669:
667:
664:
663:
660:
659:
655:
654:
651:
650:
646:
645:
642:
641:
632:
627:
625:
620:
618:
613:
611:
605:
592:
589:
535:
532:
460:Eric Buffetaut
439:
436:
367:
366:
343:
342:
336:
335:
334:
333:
321:
320:
319:Other species
316:
315:
308:
296:
295:
289:
288:
267:
263:
262:
254:
250:
249:
244:
237:
236:
231:
224:
223:
218:
211:
210:
205:
198:
197:
192:
188:
187:
182:
178:
177:
172:
168:
167:
162:
158:
157:
144:
143:
131:
130:
122:
121:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
57:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3091:
3080:
3077:
3075:
3072:
3070:
3067:
3065:
3062:
3060:
3057:
3055:
3052:
3050:
3047:
3045:
3042:
3040:
3039:Tethysuchians
3037:
3036:
3034:
3017:
3012:
3008:
3004:
2999:
2995:
2990:
2984:
2980:
2979:
2977:
2975:
2971:
2967:
2962:
2950:
2942:
2940:
2936:
2935:
2932:
2923:
2902:
2898:
2894:
2893:
2892:
2891:
2889:
2887:
2883:
2877:
2876:
2872:
2870:
2869:
2865:
2863:
2862:
2858:
2857:
2855:
2853:
2846:
2840:
2839:
2838:Thoracosaurus
2835:
2833:
2832:
2828:
2826:
2825:
2821:
2819:
2818:
2814:
2812:
2811:
2807:
2806:
2804:
2797:
2791:
2790:
2786:
2784:
2783:
2782:Aegyptosuchus
2779:
2778:
2776:
2774:
2767:
2760:
2759:
2755:
2753:
2752:
2748:
2746:
2745:
2744:Lohuecosuchus
2741:
2739:
2738:
2734:
2731:
2730:
2726:
2724:
2723:
2719:
2717:
2716:
2712:
2711:
2709:
2707:
2700:
2694:
2693:
2689:
2687:
2686:
2682:
2680:
2679:
2675:
2672:
2671:
2667:
2665:
2664:
2660:
2658:
2657:
2656:Hylaeochampsa
2653:
2650:
2649:
2645:
2644:
2642:
2640:
2633:
2626:
2625:
2618:
2615:
2614:
2607:
2604:
2603:
2602:Atoposauridae
2596:
2593:
2592:
2591:Titanochampsa
2585:
2582:
2581:
2574:
2572:
2571:
2564:
2562:
2561:
2554:
2551:
2550:
2543:
2541:
2540:
2533:
2531:
2530:
2523:
2521:
2520:
2513:
2511:
2510:
2503:
2501:
2500:
2499:Borealosuchus
2493:
2492:
2489:
2486:
2483:
2478:
2474:
2470:
2465:
2461:
2451:
2446:
2443:
2436:
2430:
2423:
2401:
2400:
2399:
2398:
2396:
2394:
2390:
2384:
2383:
2379:
2377:
2376:
2372:
2370:
2369:
2365:
2363:
2362:
2358:
2356:
2355:
2351:
2349:
2348:
2344:
2342:
2341:
2337:
2335:
2334:
2330:
2328:
2327:
2323:
2321:
2320:
2316:
2314:
2313:
2312:Batrachomimus
2309:
2308:
2306:
2304:
2297:
2291:
2290:
2286:
2284:
2283:
2279:
2278:
2276:
2274:
2267:
2260:
2259:
2255:
2253:
2252:
2248:
2246:
2245:
2241:
2238:
2237:
2233:
2231:
2230:
2226:
2224:
2223:
2219:
2217:
2216:
2212:
2210:
2209:
2205:
2203:
2202:
2198:
2196:
2195:
2191:
2189:
2188:
2184:
2182:
2181:
2177:
2175:
2174:
2170:
2168:
2167:
2166:Calsoyasuchus
2163:
2161:
2160:
2156:
2154:
2153:
2149:
2148:
2146:
2144:
2137:
2131:
2130:
2126:
2124:
2123:
2119:
2118:
2116:
2114:
2107:
2101:
2100:
2099:Stomatosuchus
2096:
2094:
2093:
2089:
2088:
2086:
2084:
2077:
2071:
2070:
2066:
2064:
2063:
2059:
2057:
2056:
2052:
2050:
2049:
2045:
2043:
2042:
2038:
2036:
2035:
2031:
2029:
2028:
2024:
2022:
2021:
2017:
2016:
2014:
2012:
2011:Atoposauridae
2005:
1995:
1994:
1993:Tilemsisuchus
1990:
1988:
1987:
1983:
1980:
1979:
1975:
1973:
1972:
1968:
1966:
1965:
1964:Rhabdognathus
1961:
1959:
1958:
1954:
1952:
1951:
1947:
1945:
1944:
1940:
1938:
1937:
1936:Guarinisuchus
1933:
1931:
1930:
1926:
1924:
1923:
1919:
1917:
1916:
1912:
1910:
1909:
1905:
1903:
1902:
1901:Chenanisuchus
1898:
1896:
1895:
1891:
1889:
1888:
1887:Brachiosuchus
1884:
1882:
1881:
1880:Atlantosuchus
1877:
1875:
1874:
1870:
1868:
1867:
1863:
1861:
1860:
1856:
1855:
1853:
1851:
1847:
1841:
1840:
1836:
1834:
1833:
1829:
1827:
1826:
1822:
1820:
1819:
1815:
1813:
1812:
1811:Pholidosaurus
1808:
1806:
1805:
1801:
1799:
1798:
1797:Meridiosaurus
1794:
1792:
1791:
1787:
1785:
1784:
1780:
1778:
1777:
1776:Crocodilaemus
1773:
1771:
1770:
1766:
1764:
1763:
1759:
1758:
1756:
1754:
1750:
1747:
1745:
1738:
1731:
1730:
1723:
1720:
1719:
1712:
1709:
1708:
1701:
1698:
1697:
1690:
1688:
1687:
1680:
1677:
1676:
1669:
1667:
1666:
1659:
1657:
1656:
1655:Khoratosuchus
1649:
1647:
1646:
1645:Karatausuchus
1639:
1637:
1636:
1629:
1627:
1626:
1619:
1618:
1615:
1612:
1609:
1604:
1600:
1596:
1591:
1587:
1566:
1565:
1564:
1563:
1561:
1559:
1555:
1547:
1543:
1542:
1541:
1540:
1537:
1534:
1532:
1528:
1524:
1516:
1512:
1509:
1507:
1503:
1500:
1498:
1494:
1492:
1488:
1486:
1482:
1481:
1476:
1472:
1468:
1461:
1456:
1454:
1449:
1447:
1442:
1441:
1438:
1427:
1422:
1418:
1414:
1407:
1400:
1397:
1392:
1388:
1384:
1380:
1376:
1372:
1368:
1364:
1357:
1350:
1347:
1343:
1337:
1334:
1329:
1325:
1321:
1317:
1313:
1309:
1302:
1295:
1292:
1287:
1283:
1276:
1273:
1265:
1261:
1257:
1256:Palaeontology
1250:
1248:
1239:
1237:
1233:
1220:
1216:
1212:
1210:
1202:
1199:
1193:
1188:
1184:
1180:
1179:Palaeontology
1176:
1169:
1167:
1165:
1163:
1161:
1159:
1155:
1149:
1147:
1145:
1141:
1137:
1133:
1129:
1125:
1121:
1117:
1113:
1112:
1107:
1103:
1100:(2013) found
1099:
1095:
1091:
1087:
1083:
1079:
1075:
1071:
1067:
1063:
1059:
1055:
1051:
1047:
1043:
1039:
1035:
1031:
1027:
1023:
1019:
1012:
1008:
1007:
1002:
997:
991:
988:
981:
980:
972:
971:
963:
962:
954:
953:
945:
944:
936:
935:
927:
926:
923:
922:
921:
914:
913:
910:
909:
906:
905:
904:
897:
896:
890:
889:
886:
885:
882:
881:
880:
873:
872:
866:
865:
862:
861:
853:
852:
849:
848:
847:
840:
839:
833:
832:
826:
825:
822:
821:
813:
812:
809:
808:
807:
800:
799:
796:
795:
792:
791:
790:
783:
782:
779:
778:
775:
774:
773:
766:
765:
759:
758:
752:
751:
748:
747:
739:
738:
730:
729:
726:
725:
724:
717:
716:
713:
712:
709:
708:
707:
700:
699:
693:
692:
689:
688:
685:
680:
679:
673:
672:
666:
665:
662:
661:
653:
652:
644:
643:
640:
639:
638:
631:
630:
624:
623:
617:
616:
609:
608:
604:
602:
598:
590:
588:
586:
582:
577:
575:
571:
567:
562:
561:
556:
555:
548:
546:
542:
533:
531:
529:
525:
521:
520:
515:
511:
508:
507:
502:
497:
495:
491:
486:
482:
478:
477:
473:
472:goniopholidid
469:
465:
464:Rucha Ingavat
461:
457:
453:
449:
445:
437:
435:
433:
432:
427:
426:
421:
417:
413:
410:
406:
405:Late Jurassic
402:
399:
398:pholidosaurid
395:
392:
387:
379:
375:
374:
365:
361:
356:
352:
350:
344:
341:
337:
328:
325:
324:
322:
317:
312:
306:
305:
297:
294:
290:
286:
282:
276:
275:
268:
265:
264:
261:
255:
252:
251:
248:
245:
242:
239:
238:
235:
232:
229:
226:
225:
222:
219:
216:
213:
212:
209:
206:
203:
200:
199:
196:
193:
190:
189:
186:
183:
180:
179:
176:
173:
170:
169:
166:
163:
160:
159:
154:
149:
145:
141:
137:
132:
128:
123:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
55:
52:150–140
47:
43:
42:Late Jurassic
37:
33:
30:
19:
2973:
2873:
2866:
2861:Boverisuchus
2859:
2836:
2829:
2822:
2815:
2808:
2787:
2780:
2756:
2749:
2742:
2735:
2727:
2720:
2713:
2690:
2683:
2676:
2668:
2661:
2654:
2646:
2622:
2611:
2600:
2589:
2578:
2568:
2558:
2547:
2539:Heterosuchus
2537:
2527:
2517:
2507:
2497:
2426:
2380:
2375:Wannchampsus
2373:
2368:Turanosuchus
2366:
2359:
2352:
2345:
2338:
2333:Paralligator
2331:
2326:Kansajsuchus
2324:
2317:
2310:
2287:
2282:Bernissartia
2280:
2258:Symptosuchus
2256:
2249:
2242:
2236:Pinacosuchus
2234:
2229:Oweniasuchus
2227:
2220:
2213:
2206:
2199:
2192:
2185:
2178:
2171:
2164:
2157:
2152:Amphicotylus
2150:
2129:Paluxysuchus
2127:
2120:
2097:
2092:Laganosuchus
2090:
2069:Varanosuchus
2067:
2062:Theriosuchus
2060:
2053:
2046:
2039:
2032:
2027:Alligatorium
2025:
2018:
1991:
1986:Sokotosuchus
1984:
1978:Sabinosuchus
1976:
1969:
1962:
1955:
1948:
1941:
1934:
1929:Fortignathus
1927:
1920:
1913:
1906:
1899:
1892:
1885:
1878:
1871:
1864:
1857:
1850:Dyrosauridae
1837:
1830:
1825:Terminonaris
1823:
1816:
1809:
1804:Oceanosuchus
1802:
1795:
1788:
1783:Dakotasuchus
1781:
1774:
1768:
1767:
1760:
1727:
1716:
1705:
1694:
1684:
1673:
1663:
1653:
1643:
1633:
1623:
1546:Pseudosuchia
1531:Pseudosuchia
1510:
1506:Pseudosuchia
1501:
1416:
1412:
1399:
1366:
1362:
1349:
1341:
1336:
1311:
1307:
1294:
1285:
1275:
1264:the original
1259:
1255:
1246:
1223:. Retrieved
1219:the original
1214:
1208:
1201:
1182:
1178:
1143:
1139:
1132:thailandicus
1131:
1127:
1123:
1119:
1111:Kansajsuchus
1109:
1106:sister taxon
1101:
1097:
1093:
1085:
1077:
1073:
1069:
1065:
1054:Ulan Malgait
1033:
1030:thailandicus
1029:
1026:"Sunosuchus"
1025:
1021:
1017:
1015:
1011:shartegensis
1010:
1004:
1000:
999:Holotype of
989:
918:
917:
901:
900:
878:
877:
876:
844:
843:
804:
803:
787:
786:
770:
769:
721:
720:
704:
703:
683:
635:
634:
600:
596:
594:
580:
578:
569:
565:
560:Oceanosuchus
558:
554:Terminonaris
552:
549:
544:
540:
537:
517:
514:generic name
509:
504:
500:
498:
489:
484:
480:
474:
443:
441:
430:
429:
424:
423:
372:
371:
370:
363:Efimov, 1988
359:
357:
351:thailandicus
347:
346:
331:Efimov, 1988
326:
303:
302:
293:Type species
280:
273:
272:
240:
227:
221:Pseudosuchia
214:
201:
139:
135:
35:
29:
2875:Planocrania
2868:Duerosuchus
2810:Argochampsa
2729:Arenysuchus
2715:Agaresuchus
2685:Turcosuchus
2382:Yanjisuchus
2361:Tarsomordeo
2354:Shamosuchus
2347:Scolomastax
2244:Siamosuchus
2215:Nannosuchus
2208:Hulkepholis
2201:Goniopholis
2187:Diplosaurus
2173:Coelosuchus
2122:Deltasuchus
2055:Sabresuchus
2041:Atoposaurus
1971:Rodeosuchus
1950:Luciasuchus
1908:Congosaurus
1832:Vectisuchus
1818:Sarcosuchus
1762:Anglosuchus
1744:Tethysuchia
1718:Zhyrasuchus
1625:Burkesuchus
1225:17 December
601:Sarcosuchus
566:Sarcosuchus
545:Sarcosuchus
541:Sarcosuchus
534:Description
208:Archosauria
3033:Categories
2937:See also:
2897:Crocodilia
2886:Crocodilia
2817:Eogavialis
2789:Aegisuchus
2580:Susisuchus
2549:Isisfordia
2402:see below↓
2340:Rugosuchus
2251:Sunosuchus
2034:Aprosuchus
1943:Hyposaurus
1922:Dyrosaurus
1707:Wahasuchus
1686:Susisuchus
1635:Isisfordia
1567:see below↓
1515:Metasuchia
1497:Sauropsida
1150:References
1116:Tajikistan
1104:to be the
1074:Sunosuchus
1006:Sunosuchus
585:palynology
519:Krai Thong
476:Sunosuchus
349:Sunosuchus
2989:Q15221406
2692:Unasuchus
1790:Elosuchus
1608:Neosuchia
1595:Neosuchia
1558:Neosuchia
1483:Kingdom:
1467:Neosuchia
1391:219451890
1344:Pp 69-84.
1328:0272-4634
1060:of Outer
1050:Tithonian
1016:Halliday
591:Phylogeny
524:crocodile
448:lower jaw
171:Kingdom:
165:Eukaryota
2983:Wikidata
2974:Chalawan
2949:Category
2824:Eosuchus
2648:Acynodon
2482:Eusuchia
2469:Eusuchia
2393:Eusuchia
1769:Chalawan
1491:Chordata
1489:Phylum:
1485:Animalia
1369:: 1–18.
1209:Chalawan
1136:polytomy
1062:Mongolia
1038:holotype
1001:Chalawan
597:Chalawan
581:Chalawan
574:diastema
503:and the
501:Chalawan
490:Chalawan
444:Chalawan
389:) is an
373:Chalawan
340:Synonyms
274:Chalawan
253:Family:
195:Reptilia
185:Chordata
181:Phylum:
175:Animalia
161:Domain:
142:(right)
36:Chalawan
3003:1480518
1495:Class:
1371:Bibcode
1140:et al.'
530:teeth.
528:diamond
492:to the
428:, with
391:extinct
382:ชาละวัน
279:Martin
266:Genus:
191:Class:
3016:283703
1389:
1326:
1288:: 239.
1098:et al.
1094:et al.
1066:et al.
1018:et al.
512:. The
376:(from
281:et al.
2998:IRMNG
1511:Clade
1502:Clade
1409:(PDF)
1387:S2CID
1359:(PDF)
1304:(PDF)
1267:(PDF)
1252:(PDF)
1114:from
1042:skull
526:with
394:genus
241:Clade
228:Clade
215:Clade
202:Clade
2895:see
1544:see
1324:ISSN
1227:2021
1130:cf.
1128:"S."
1028:cf.
568:and
557:and
468:1980
462:and
378:Thai
311:1980
285:2013
60:PreꞒ
1421:doi
1379:doi
1342:315
1316:doi
1187:doi
1126:to
1108:of
1084:of
466:in
414:of
407:or
396:of
3035::
3013::
3000::
2985::
1513::
1504::
1417:60
1415:.
1411:.
1385:.
1377:.
1367:85
1365:.
1361:.
1322:.
1312:39
1310:.
1306:.
1284:.
1260:27
1258:.
1254:.
1235:^
1213:.
1207:"†
1183:57
1181:.
1177:.
1157:^
1032:.
1009:)
603:.
496:.
479:,
418:,
380::
358:?
329:?
283:,
243::
230::
217::
204::
110:Pg
54:Ma
2849:†
2800:†
2770:†
2761:?
2732:?
2703:†
2673:?
2651:?
2636:†
2627:?
2620:†
2616:?
2609:†
2605:?
2598:†
2594:?
2587:†
2583:?
2576:†
2566:†
2556:†
2552:?
2545:†
2535:†
2525:†
2515:†
2505:†
2495:†
2300:†
2270:†
2261:?
2239:?
2140:†
2110:†
2080:†
2008:†
1981:?
1741:†
1732:?
1725:†
1721:?
1714:†
1710:?
1703:†
1699:?
1692:†
1682:†
1678:?
1671:†
1661:†
1651:†
1641:†
1631:†
1621:†
1459:e
1452:t
1445:v
1429:.
1423::
1393:.
1381::
1373::
1330:.
1318::
1229:.
1195:.
1189::
1003:(
313:)
300:†
270:†
257:†
115:N
105:K
100:J
95:T
90:P
85:C
80:D
75:S
70:O
65:Ꞓ
49:~
44:-
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
