586:
1507:
3758:
3764:
689:
3047:
1433:
ten centimetres. They were hexagonal or pentagonal, thus with five or six sides. Each of these sides touched somewhat smaller scales, forming a rosette. Small, non-overlapping convex scales of about one centimetre in diameter surrounded the whole. The larger scales were wrinkled due to straight grooves orientated perpendicular to their edges. From top to bottom, the large scale rows gradually declined in size. Unfortunately, nothing can as yet be learned about the coloration of
4314:
2060:
147:
1491:
2128:
4344:
1096:
915:
1273:
1446:
1366:
4320:
1303:
863:
122:
3035:
2158:
from the matrix that it was buried in indicated that the juvenile ceratopsian drowned during a possible river crossing. Further study of the specimen revealed that juvenile chasmosaurs had a frill that was narrower in the back than that of adults, as well as being proportionately shorter in relation to the skull.
2103:
The function of the frill and horns is problematic. The horns are rather short and the frill had such large fenestrae that it could not have offered much functional defense. Paul suggested that the beak was the main defensive weapon. It is possible that the frill was simply used to appear imposing or
1432:
impressions. The area conserved, from the right hip region, measured about one by 0.5 metres. The skin appears to have had large scales in evenly spaced horizontal rows among smaller scales. The larger scales had a diameter of up to fifty-five millimetres and were distanced from each other by five to
2157:
was not fast moving, and that juveniles did not need to be fast moving either to keep pace with adults. The fossil was complete save for its missing front limbs, which had fallen into a sinkhole before the specimen was uncovered. Skin impressions were also uncovered beneath the skeleton and evidence
1361:
the inner one. The remainder of the rear edge lacked osteoderms. The parietal bones of the frill were pierced by very large openings, after which the genus was named: the parietal fenestrae. These were not oval in shape, as with most relatives, but triangular, with one point orientated towards the
585:
1207:
are known. These include AMNH 5422, ROM 843 (earlier ROM 5499) and NHMUK R4948, all (partial) skeletons with skull. The skull YPM 2016 and the skull and skeleton AMNH 5402 were noted by
Campbell et al. (2016) as differing from other
2308:
Lambe, L.M., 1914, "On the forelimb of a carnivorous dinosaur from the Belly River
Formation of Alberta, and a new genus of Ceratopsia from the same horizon, with remarks on the integument of some Cretaceous herbivorous dinosaurs",
1212:
referred specimens in having more epiparietals, although the authors interpreted them as individual variation, but this was reconsidered when
Campbell et al. (2019) interpreted these specimens as an indeterminate
1118:, several teams of researchers published work questioning the validity of this new genus. In 2011, Maidment & Barrett failed to confirm the presence of any supposedly unique features, and argued that
2665:"Temporal range extension and evolution of the chasmosaurine ceratopsid 'Vagaceratops' irvinensis (Dinosauria: Ornithischia) in the Upper Cretaceous (Campanian) Dinosaur Park Formation of Alberta"
1130:
specimens, agreed with the conclusions of
Maidment & Barrett, adding that some supposedly unique features, such as grooves on the parietal bone, were actually also present in the holotype of
1229:
species by Fowler and
Freedman Fowler (2020), who noted that "given the similarity between these two specimens (YPM 2016 and AMNH 5402) and CMN 2245, it is not clear why CMN 2245 was left in
620:
that was part of a neck frill. Although recognizing that his find represented a new species, Lambe thought this could be placed in a previously known short-frilled ceratopsian genus:
4496:
2774:"Revised geochronology, correlation, and dinosaur stratigraphic ranges of the Santonian-Maastrichtian (Late Cretaceous) formations of the Western Interior of North America"
1034:
and therefore was considered a possible synonym. However, the parietal (back margin of the frill) is not preserved, and was restored with plaster based on specimens of
4541:
2555:
Lucas SG, Sullivan RM, Hunt AP (2006). "Re-evaluation of
Pentaceratops and Chasmosaurus (Ornithischia: Ceratopsidae) in the Upper Cretaceous of the Western Interior".
1038:, which caused confusion among scientists in previous decades, because the parietal bone is critical for determining differences between species in ceratopsids like
4437:
955: 5656, NMC 34832 and TMP 1979.11.147, and (tentatively) AMNH 5401 and NMC 1254 were also referred to the genus. All specimens assigned to
1341:
the rear edge is shaped as a shallow U, and the sides are more convex. The sides were adorned by six to nine smaller skin ossifications (called episquamosals) or
2149:, is hypothesized to have done. The juvenile measured five feet long and was estimated to be three years of age and had similar limb proportions to the adult
1385:. There are five other cervicals preserved in this specimen, for a total of eight, which likely represents a complete neck. Cervicals four to eight are
2507:
Paul, G.S., 1987, "The science and art of reconstructing the life appearance of dinosaurs and their relatives: a rigorous how-to guide", pp 4–49 in:
2108:. The frill may also have been brightly colored, to draw attention to its size or as part of a mating display. However, it is difficult to prove any
4424:
1068:. He considered it too poorly preserved for a reliable determination, especially as it belonged to a juvenile individual, and regarded it too as a
4531:
4501:
2898:(Ornithischia: Ceratopsidae), a revision of the genus, and the utility of postcrania in the taxonomy and systematics of ceratopsid dinosaurs".
4526:
4506:
3757:
1499:
951:
TMP 1999.55.292, an isolated lateral ramus of a right parietal, and NMC 8803, central bar and lateral rami of parietals. Specimens
2368:
T.M. Lehman, 1990, "The ceratopsian subfamily
Chasmosaurinae: sexual dimorphism and systematics", In: K. Carpenter and P. J. Currie (eds.),
3079:
940:
2124:, the smaller one, NMC 2245, was the male and the larger, NMC 2280, the female. However, today the two are referred to different species.
710:
have been named within the genus. Today some of these are considered to only reflect a morphological variation among the known sample of
3763:
1027:
952:
3004:"Charting the growth of one of the world's oldest babies: Late Cretaceous Chasmosaurus fills in missing pieces of dinosaur evolution"
2224:
1506:
467:
4536:
4337:
2601:
Scott D. Sampson; Mark A. Loewen; Andrew A. Farke; Eric M. Roberts; Catherine A. Forster; Joshua A. Smith; Alan L. Titus (2010).
2167:
2329:, a new genus and species of trachodont dinosaur from the Belly River Formation of Alberta, with a description of the skull of
1166:
is actually from the upper
Dinosaur Park Formation, according to recent fieldwork. This situation is further complicated since
1004:
685:, specimen NMC 2245 found by the Sternbergs in 1913 and consisting of a largely complete skeleton, including skin impressions.
2978:
2120:
of which the females would have short ones. In 1927 C.M. Sternberg concluded that of the two skeletons he had mounted in the
4521:
2663:
Campbell, James
Alexander; Ryan, Michael J.; Schroder-Adams, Claudia J.; Holmes, Robert B.; Evans, David C. (2019-08-08).
3025:
4511:
4313:
1415:
1403:
1387:
1313:
The known differences between the two species mainly pertain to the horn and frill shape, as the referred postcrania of
146:
681:, "opening" or "divide" and refers to the very large parietal fenestrae in the skull frill. Lambe now also assigned a
605:
1333:
is very elongated and broader at the rear than at the front. It is hardly elevated from the plane of the snout. With
638:
2845:- new information from the Peabody Museum skull, and the use of phylogenetic analysis for dinosaur alpha taxonomy"
2715:"Transitional evolutionary forms in chasmosaurine ceratopsid dinosaurs: evidence from the Campanian of New Mexico"
4516:
2121:
1454:
871:
801:
330:
3170:
3072:
562:
2664:
2187:
Ryan, M.J. & Evans, D.C. (2005). "Ornithischian dinosaurs". In Currie, P.J. & Koppelhus, E.B. (eds.).
4491:
3288:
2064:
1518:
1008:
960:
882:
809:
642:
631:
558:
2446:(Dinosauria: Ornithischia) from the Upper Cretaceous (Campanian) Dinosaur Park Formation of Western Canada.
2441:
4364:
813:
786:
in 1933 named an unusual, short-muzzled skull, specimen ROM 839 (earlier ROM 5436) collected in 1926, as
4463:
4347:
3197:
543:
4319:
688:
536:
was of a slightly smaller to ‘average’ size, especially when compared to larger ceratopsians (such as
4411:
2785:
2614:
2400:
2275:
992:
696:
392:
381:
357:
334:
314:
289:
133:
1199:
are all from the lower
Dinosaur Park Formation, stratigraphically and morphologically separate from
1026:, known from specimen AMNH 5401, a nearly complete (but partially restored) skull on display at the
553:, but this name had been previously published for another animal. All of the excavated specimens of
3453:
3065:
589:
727:
4173:
4123:
3555:
3003:
2416:
2291:
2093:
1378:
783:
502:
367:
141:
2059:
4468:
4402:
1011:
means "conspicuous pride" in Greek, both referring to the skull frill. The species is based on
4450:
4442:
4256:
4194:
4157:
3631:
3518:
2821:
2803:
2754:
2736:
2686:
2642:
2574:
2220:
2109:
1490:
972:
911:, as its placement would probably remain unstable until chasmosaurines are better understood.
4455:
1329:
they are somewhat longer, especially the brow horns, and more curved backwards. The frill of
1325:- one on the nose and two on the brow. In both species these horns are quite short, but with
4286:
4084:
3909:
3611:
3306:
3039:
2907:
2856:
2811:
2793:
2744:
2726:
2676:
2632:
2622:
2456:
2408:
2283:
2105:
1720:
1410:
1394:
702:
Since that date, more remains, including skulls, have been found that have been referred to
494:
426:
4180:
4132:
4091:
4027:
3930:
3881:
3583:
3507:
3462:
3336:
3322:
3281:
3155:
3139:
2259:
2239:
Lambe, L.M., 1902, "New genera and species from the Belly River Series (mid-Cretaceous)",
1825:
817:
566:
30:
2789:
2618:
2603:"New Horned Dinosaurs from Utah Provide Evidence for Intracontinental Dinosaur Endemism"
2404:
2279:
2100:
had a longer snout and jaws and might have been more selective about the plants it ate.
4187:
4146:
4077:
4064:
4035:
4020:
3990:
3951:
3937:
3854:
3815:
3694:
3604:
3569:
3562:
3442:
3392:
3315:
3295:
2816:
2773:
2749:
2714:
2637:
2602:
1576:
1563:
1476:
1468:
1322:
1260:
791:
601:
285:
265:
223:
2127:
1244:
are synonymous, while splitting some remains assigned to the latter to a new species,
4485:
4265:
4244:
4214:
4205:
4105:
3999:
3944:
3923:
3895:
3888:
3874:
3867:
3673:
3662:
3655:
3597:
3590:
3576:
3534:
3469:
3435:
3406:
3359:
3051:
2420:
2262:; Burns, M. E.; Sissons, R. L. (2009). "A redescription of the ankylosaurid dinosaur
1866:
1801:
1751:
1687:
1639:
1537:
1514:
1510:
1416:
1382:
1350:
855:
822:
665:
617:
506:
498:
72:
2295:
1195:
belong to a distinct species, if not genus, of chasmosaurine. Specimens referred to
1095:
914:
4279:
4230:
4223:
4139:
4112:
4098:
4006:
3979:
3972:
3965:
3916:
3794:
3781:
3712:
3701:
3686:
3648:
3640:
3622:
3527:
3493:
3476:
3428:
3421:
3377:
3182:
3130:
2088:
1914:
1768:
1670:
1615:
1548:
1277:
1176:
1052:
996:
980:
924:
887:
841:
719:
484:
399:
345:
252:
207:
2861:
1445:
1272:
2961:
Sternberg, C.M., 1927, "Horned dinosaur group in the National Museum of Canada",
2798:
2627:
2460:
1388:
738:('from Canada') named by Thomas M. Lehman in 1990. The latter species, originally
4396:
4272:
4166:
4049:
4013:
3836:
3548:
3413:
2581:(Dinosauria: Ceratopsia) from the Dinosaur Park Formation of southern Alberta",
2145:
1931:
1849:
1365:
1254:
622:
538:
521:
47:
4042:
3958:
3808:
3541:
3329:
3263:
3250:
3209:
3088:
2911:
1890:
1464:
1404:
1342:
1302:
968:
862:
659:
491:
239:
121:
92:
57:
4387:
2807:
2740:
2690:
2391:, A New Chasmosaurine Ceratopsid from the Late Campanian of Western Canada".
2840:
2083:
1529:
1525:
1480:
1346:
964:
529:
158:
97:
41:
34:
2948:, gen. nov., with remarks on other genera of Cretaceous horned dinosaurs",
2825:
2758:
2646:
2440:
Campbell, J.A., Ryan, M.J., Holmes, R.B., and Schröder-Adams, C.J. (2016).
501:. Its given name means 'opening lizard', referring to the large openings (
4381:
3843:
3829:
3822:
3385:
3121:
3112:
2287:
1187:
is in a state of flux. For the aforementioned reasons, it is likely that
948:
936:
907:
in 2020, they suggested it should be maintained as a distinct genus from
758:
by Nicholas Longrich, although different teams of researchers have found
682:
613:
487:
191:
178:
87:
82:
67:
62:
52:
1291:
was a medium-size ceratopsid. In 2010 G.S. Paul estimated the length of
1092:
sp. due to the lack of the parietal preserved in the holotypes of both.
782:
sp. due to the lack of the parietal preserved in the holotypes of both.
4429:
4237:
3399:
3343:
2731:
2681:
2266:
Parks, 1924 (Ornithischia: Ankylosauria) and a revision of the genus".
2079:
1377:
is best preserved in the specimen known as NHMUK 4948. The first three
1012:
976:
707:
646:
102:
77:
2524:. George Olshevsky, Publications Requiring Research, San Diego, 157 pp
1114:
Following the original assignment of the holotype and other skulls to
831:, but this has found no acceptance. In 2000, George Olshevsky renamed
3106:
2412:
1203:
Apart from the holotype and paratype several additional specimens of
984:
808:
in 1940, based on specimen NMC 8800 from southwestern Alberta (lower
650:
168:
4358:
1349:
bone. The corner of the frill featured two larger osteoderms on the
1258:; however, the name appeared in the pre-proof of the description of
641:
and his sons found several complete "M. belli" skulls in the middle
2241:
Geological Survey of Canada Contributions to Canadian Palaeontology
1337:
the rear of the frill is V-shaped and its sides are straight. With
2126:
2058:
1505:
1489:
1444:
1364:
1301:
1271:
1174:, sharing features with the contemporaneous derived chasmosaurine
1138:
specimens. This variability, they argued, strongly suggested that
1094:
944:
913:
861:
850:
687:
584:
525:
517:
480:
1428:
specimen NMC 2245 recovered by C.M. Sternberg was accompanied by
4416:
1429:
1236:
In 2015, Nicholas Longrich presented a novel theory that posits
734:
in having very long brow horns. This form is perhaps related to
4362:
3779:
3248:
3099:
3061:
2713:
Fowler, Denver W.; Fowler, Elizabeth A. Freedman (2020-06-05).
3057:
2349:
Brown, B., 1933, "A new longhorned Belly River ceratopsian",
2139:
found in Alberta, Canada by Phil Currie et al., reveals that
2210:
2208:
2206:
2204:
2202:
2200:
2198:
532:)—or anywhere from 2,200 to nearly 5,000 lbs., give or take—
453:
2894:
Maidment, S.C.R.; Barrett, P.M. (2011). "A new specimen of
456:
444:
2191:. Bloomington: Indiana University Press. pp. 312–348.
2189:
Dinosaur Provincial Park: A Spectacular Ecosystem Revealed
1381:
are fused into a unit known as a syncervical, as in other
881:
named in 2001, which stems from the uppermost beds of the
839:
as its fossil material is likely chasmosaurine; this is a
1299:
would have been 4.3 metres long and weighed 1.5 tonnes.
663:"), but that name was preoccupied by the Permian reptile
435:
2577:, M.J. Ryan and K.M. Shepherd, 2001, "A new species of
2522:
An Annotated Checklist of Dinosaur Species by Continent
2442:
A Re-Evaluation of the chasmosaurine ceratopsid genus
2116:, which bore long brow horns, was in fact the male of
3023:
1126:. Campbell and colleagues, in their 2016 analysis of
766:. Campbell and colleagues, in their 2016 analysis of
468:
459:
450:
438:
432:
429:
2537:, sp. nov., a new ceratopsian dinosaur from Texas",
2491:
Sternberg, C.M., 1940, "Ceratopsidae from Alberta",
2372:, Cambridge University Press, Cambridge, pp. 211–229
1471:. The Chasmosaurinae usually have long frills, like
1393:, wider than long, and roughly equal in length. The
891:, in 2010. However, Campbell et al. (2019) referred
340:
321:
304:
4371:
4255:
4213:
4204:
4156:
4122:
4063:
3989:
3853:
3792:
3711:
3672:
3630:
3621:
3517:
3491:
3452:
3357:
3305:
3261:
3208:
3181:
3154:
2478:. Memoirs of the Peabody Museum of Natural History
2436:
2434:
2432:
2430:
2215:Dodson, Peter; et al. (1993). "Chasmosaurus".
447:
441:
1030:, was considered to share features in common with
669:, so he subsequently created the replacement name
2979:"Rare baby dinosaur skeleton unearthed in Canada"
2370:Dinosaur Systematics: Perspectives and Approaches
2143:may have cared for its young, like its relative,
573:come from the lower beds of the formation, while
2476:A revision of the Ceratopsia or horned dinosaurs
2219:. Publications International. pp. 110–111.
790:, "with a short snout". This has been seen as a
1437:from the known fossil skin impression samples.
1264:before being edited out for final publication.
1103:, which has also been considered a specimen of
2511:, Natural History Museum of Los Angeles County
2382:
2380:
2378:
516:, meaning 'lizard'). With a length of 4.3–4.8
512:, meaning 'opening', 'hollow', or 'gulf'; and
3073:
1487:has a basal position in the Chasmosaurinae.S
730:and based on skull AMNH 5401, differing from
653:. Based on these finds, Lambe (1914) erected
8:
2364:
2362:
1221:. The specimen CMN 2245 was referred to the
714:skulls; others are seen as valid species of
2876:
2874:
2872:
2669:Vertebrate Anatomy Morphology Palaeontology
2470:
2468:
1162:was considered doubtful as the holotype of
4497:Late Cretaceous dinosaurs of North America
4359:
4210:
4069:
3859:
3800:
3789:
3776:
3678:
3627:
3499:
3369:
3365:
3273:
3269:
3258:
3245:
3187:
3160:
3151:
3096:
3080:
3066:
3058:
2596:
2594:
1317:are poorly known. Like many ceratopsians,
1015:thought by other researchers to belong to
120:
20:
2860:
2815:
2797:
2748:
2730:
2680:
2636:
2626:
1295:at 4.8 metres, its weight at two tonnes;
1060:. Longrich also regarded the holotype of
754:were thought to probably be exemplars of
2950:Canada Geological Survey Museum Bulletin
2063:Depiction of the mega-herbivores in the
1248:. Because the publication was rejected,
1107:, which itself is probably a synonym of
943: 1983.25.1 consisting of a partial
885:. This species was given its own genus,
4542:Multispecific non-avian dinosaur genera
3030:
2884:, Princeton University Press p. 269–270
2254:
2252:
2179:
1475:itself, whereas their sister-group the
1072:, rather than as the senior synonym of
1056:, rather than as the senior synonym of
877:The most recently described species is
2924:Sternberg, C.M., 1925, "Integument of
2882:The Princeton Field Guide to Dinosaurs
1421:vertebrae, depending on the specimen.
1276:Size comparison of several members of
577:comes from the middle and upper beds.
2772:Fowler, Denver Warwick (2017-11-22).
2708:
2706:
2704:
2702:
2700:
2658:
2656:
991:was named by Nicholas R. Longrich in
922:, sometimes considered its own genus
7:
2509:Dinosaurs Past and Present Volume II
1500:Royal Tyrell Museum of Palaeontology
1479:typically have shorter frills. Most
1357:the outer one was the largest, with
1142:was simply a mature growth stage of
947:including the parietal and from the
1906:
1882:
1841:
1817:
1793:
1743:
1736:
1712:
1662:
1655:
1631:
1607:
1583:
1556:
1543:
1050:was then by Longrich regarded as a
1003:. The generic name is derived from
2583:Canadian Journal of Earth Sciences
2539:Journal of Vertebrate Paleontology
2268:Journal of Vertebrate Paleontology
1134:and, to various degrees, in other
1028:American Museum of Natural History
742:Lambe 1902, had been described as
600:In 1898, at Berry Creek, Alberta,
542:, which were about the size of an
14:
2557:New Mex Mus. Nat. Hist. Sci. Bull
1170:may not even belong to the genus
1064:as probably being an exemplar of
903:likely to be the sister taxon of
4343:
4342:
4318:
4312:
3762:
3756:
3045:
3033:
2168:Timeline of ceratopsian research
1463:was in 1915 by Lambe within the
634:honoured collector Walter Bell.
425:
145:
2112:. In 1933, Lull suggested that
718:or as separate genera. In 1933
2952:12, Geological Series 24: 1–49
2387:Nicholas R. Longrich (2010). "
2096:is suggested by the fact that
1409:, a compound unit composed of
899:. As Fowler and Fowler found
630:to describe his findings. The
549:It was initially to be called
1:
4532:Ceratopsians of North America
4502:Fossil taxa described in 1914
2862:10.12688/f1000research.7573.1
2086:, with successive species of
853:, which has now been renamed
812:). The specific name honours
677:is derived from Greek χάσμα,
626:. He erected the new species
4527:Late Cretaceous ceratopsians
4507:Taxa named by Lawrence Lambe
2799:10.1371/journal.pone.0188426
2628:10.1371/journal.pone.0012292
2461:10.1371/journal.pone.0145805
1146:. Recently, the referral of
608:made the first discovery of
2839:Longrich, Nicholas (2015).
1369:Replica of skin impressions
1217:species closely related to
673:in February 1914. The name
606:Geological Survey of Canada
4558:
2535:Chasmosaurus mariscalensis
2264:Dyoplosaurus acutosquameus
847:Chasmosaurus mariscalensis
845:. Thomas Lehman described
837:Chasmosaurus recurvicornis
639:Charles Hazelius Sternberg
16:Extinct genus of dinosaurs
4332:
4310:
4072:
3862:
3803:
3788:
3775:
3753:
3681:
3502:
3372:
3368:
3276:
3272:
3257:
3244:
3190:
3163:
3150:
3095:
2963:Canadian Field-Naturalist
2930:Canadian Field-Naturalist
2912:10.11646/zootaxa.2963.1.1
2351:American Museum Novitates
2122:Canadian Museum of Nature
1928:
1911:
1904:
1887:
1880:
1863:
1846:
1839:
1822:
1815:
1798:
1791:
1765:
1748:
1741:
1734:
1717:
1710:
1684:
1667:
1660:
1653:
1636:
1629:
1612:
1605:
1588:
1581:
1561:
1554:
1455:Canadian Museum of Nature
1189:Mojoceratops, Eoceratops,
872:Canadian Museum of Nature
833:Monoclonius recurvicornis
802:Charles Mortram Sternberg
788:Chasmosaurus brevirostris
706:, and several additional
373:
366:
301:
296:
142:Scientific classification
140:
128:
119:
23:
1536:according to a study by
1413:, dorsal, and sometimes
1345:, which attached to the
959:were collected from the
569:. Referred specimens of
563:Dinosaur Provincial Park
524:) and a weight of 1.5–2
4537:Paleontology in Alberta
3289:Micropachycephalosaurus
2493:Journal of Paleontology
2393:Journal of Paleontology
2325:Lambe, L.M., 1914, "On
2065:Dinosaur Park Formation
1519:Dinosaur Park Formation
1397:are also amphiplatian.
961:Dinosaur Park Formation
920:Chasmosaurus irvinensis
883:Dinosaur Park Formation
879:Chasmosaurus irvinensis
829:Chasmosaurus sternbergi
810:Dinosaur Park Formation
736:Chasmosaurus canadensis
643:Dinosaur Park Formation
559:Dinosaur Park Formation
2944:L.M. Lambe, 1915, "On
2389:Mojoceratops perifania
2153:. This indicates that
2132:
2072:
1521:
1503:
1457:
1370:
1321:had three main facial
1310:
1285:
1120:Mojoceratops perifania
1111:
1032:Mojoceratops perifania
1001:Mojoceratops perifania
933:Mojoceratops perifania
928:
874:
740:Monoclonius canadensis
699:
597:
557:were collected at the
4464:Paleobiology Database
2946:Eoceratops canadensis
2533:Lehman, T.M., 1989, "
2335:The Ottawa Naturalist
2327:Gryposaurus notabilis
2311:The Ottawa Naturalist
2130:
2062:
1509:
1496:Chasmosaurus russelli
1493:
1448:
1368:
1305:
1275:
1124:Chasmosaurus russelli
1098:
1076:. A 2016 overview of
917:
865:
806:Chasmosaurus russelli
764:Chasmosaurus russelli
744:Eoceratops canadensis
691:
602:Lawrence Morris Lambe
588:
581:Discovery and species
544:African bush elephant
4522:Ornithischian genera
2520:Olshevsky, G. 2000.
2288:10.1671/039.029.0405
2217:The Age of Dinosaurs
2131:Juvenile UALVP 52613
2082:, the east coast of
1252:"priscus" remains a
1048:Chasmosaurus kaiseni
1024:Chasmosaurus kaiseni
752:Chasmosaurus kaiseni
750:and the long-horned
724:Chasmosaurus kaiseni
697:Royal Tyrrell Museum
134:Royal Ontario Museum
4512:Dinosaur Park fauna
3454:Archaeoceratopsidae
2790:2017PLoSO..1288426F
2619:2010PLoSO...512292S
2405:2010JPal...84..681L
2280:2009JVPal..29.1117A
1517:specimens from the
1483:analyses show that
1373:The postcranium of
1183:Today, taxonomy of
1088:to be referable to
814:Loris Shano Russell
778:to be referable to
762:to be a synonym of
590:George F. Sternberg
4174:Crittendenceratops
4124:Albertaceratopsini
2985:. 25 November 2013
2926:Chasmosaurus belli
2896:Chasmosaurus belli
2880:Paul, G.S., 2010,
2732:10.7717/peerj.9251
2682:10.18435/vamp29356
2474:Lull, R.S., 1933,
2331:Chasmosaurus belli
2137:Chasmosaurus belli
2133:
2094:niche partitioning
2073:
1522:
1504:
1458:
1451:Chasmosaurus belli
1379:cervical vertebrae
1371:
1311:
1286:
1112:
929:
875:
784:Richard Swann Lull
746:by Lambe in 1915.
712:Chasmosaurus belli
700:
637:However, in 1913,
598:
4479:
4478:
4451:Open Tree of Life
4365:Taxon identifiers
4356:
4355:
4328:
4327:
4308:
4307:
4304:
4303:
4300:
4299:
4296:
4295:
4257:Pachyrhinosaurini
4195:Yehuecauhceratops
4158:Nasutoceratopsini
4059:
4058:
3771:
3770:
3751:
3750:
3747:
3746:
3743:
3742:
3739:
3738:
3735:
3734:
3731:
3730:
3632:Protoceratopsidae
3519:Leptoceratopsidae
3487:
3486:
3353:
3352:
3240:
3239:
3236:
3235:
3232:
3231:
3228:
3227:
2110:sexual dimorphism
2051:
2050:
2042:
2041:
2033:
2032:
2024:
2023:
2015:
2014:
2006:
2005:
1997:
1996:
1988:
1987:
1979:
1978:
1970:
1969:
1961:
1960:
1952:
1951:
1943:
1942:
1780:
1779:
1699:
1698:
1122:was a synonym of
973:Belly River Group
918:Skull replica of
628:Monoclonius belli
416:
415:
410:
395:
384:
360:
337:
318:
292:
4549:
4517:Campanian genera
4472:
4471:
4459:
4458:
4446:
4445:
4433:
4432:
4420:
4419:
4407:
4406:
4405:
4392:
4391:
4390:
4360:
4346:
4345:
4323:
4322:
4316:
4287:Pachyrhinosaurus
4211:
4085:Machairoceratops
4070:
3910:Coahuilaceratops
3860:
3801:
3790:
3777:
3766:
3761:
3760:
3679:
3628:
3612:Zhuchengceratops
3500:
3370:
3366:
3307:Chaoyangsauridae
3274:
3270:
3259:
3246:
3188:
3161:
3152:
3145:
3144:
3097:
3082:
3075:
3068:
3059:
3050:
3049:
3048:
3038:
3037:
3036:
3029:
3018:
3017:
3015:
3014:
3000:
2994:
2993:
2991:
2990:
2975:
2969:
2959:
2953:
2942:
2936:
2922:
2916:
2915:
2891:
2885:
2878:
2867:
2866:
2864:
2841:"Systematics of
2836:
2830:
2829:
2819:
2801:
2784:(11): e0188426.
2769:
2763:
2762:
2752:
2734:
2710:
2695:
2694:
2684:
2660:
2651:
2650:
2640:
2630:
2598:
2589:
2571:
2565:
2564:
2552:
2546:
2531:
2525:
2518:
2512:
2505:
2499:
2489:
2483:
2472:
2463:
2438:
2425:
2424:
2413:10.1666/09-114.1
2384:
2373:
2366:
2357:
2347:
2341:
2323:
2317:
2306:
2300:
2299:
2274:(4): 1117–1135.
2256:
2247:
2237:
2231:
2230:
2212:
2193:
2192:
2184:
2106:thermoregulation
2104:conceivably for
1907:
1883:
1842:
1818:
1794:
1744:
1737:
1721:Coahuilaceratops
1713:
1663:
1656:
1632:
1608:
1584:
1557:
1544:
1467:assigned to the
1418:
1406:
1395:dorsal vertebrae
1390:
770:specimens found
520:(14.1–15.7
505:) in its frill (
475:
471:
466:
465:
462:
461:
458:
455:
452:
449:
446:
443:
440:
437:
434:
431:
404:
390:
379:
351:
342:
329:
323:
312:
306:
284:
277:
264:
251:
238:
222:
206:
150:
149:
124:
114:
44:
29:Temporal range:
21:
4557:
4556:
4552:
4551:
4550:
4548:
4547:
4546:
4482:
4481:
4480:
4475:
4467:
4462:
4454:
4449:
4441:
4436:
4428:
4423:
4415:
4410:
4401:
4400:
4395:
4386:
4385:
4380:
4367:
4357:
4352:
4324:
4317:
4292:
4251:
4200:
4181:Furcatoceratops
4152:
4133:Albertaceratops
4118:
4092:Menefeeceratops
4055:
4028:Tatankaceratops
3985:
3931:Mercuriceratops
3882:Arrhinoceratops
3849:
3797:
3784:
3767:
3755:
3727:
3707:
3668:
3617:
3584:Montanoceratops
3513:
3508:Graciliceratops
3496:
3483:
3463:Archaeoceratops
3448:
3362:
3349:
3337:Xuanhuaceratops
3323:Hualianceratops
3301:
3282:Albalophosaurus
3266:
3253:
3224:
3204:
3177:
3171:Avemetatarsalia
3156:Avemetatarsalia
3146:
3140:Marginocephalia
3102:
3101:
3091:
3086:
3056:
3046:
3044:
3034:
3032:
3024:
3022:
3021:
3012:
3010:
3002:
3001:
2997:
2988:
2986:
2977:
2976:
2972:
2960:
2956:
2943:
2939:
2923:
2919:
2893:
2892:
2888:
2879:
2870:
2838:
2837:
2833:
2771:
2770:
2766:
2712:
2711:
2698:
2662:
2661:
2654:
2600:
2599:
2592:
2572:
2568:
2554:
2553:
2549:
2532:
2528:
2519:
2515:
2506:
2502:
2490:
2486:
2473:
2466:
2455:(1): e0145805.
2439:
2428:
2386:
2385:
2376:
2367:
2360:
2348:
2344:
2324:
2320:
2307:
2303:
2258:
2257:
2250:
2238:
2234:
2227:
2214:
2213:
2196:
2186:
2185:
2181:
2176:
2164:
2057:
2052:
2043:
2034:
2025:
2016:
2007:
1998:
1989:
1980:
1971:
1962:
1953:
1944:
1826:Arrhinoceratops
1781:
1700:
1443:
1420:
1408:
1392:
1383:neoceratopsians
1306:Restoration of
1270:
835:Cope 1889 into
818:Gregory S. Paul
583:
567:Alberta, Canada
473:
469:
428:
424:
283:
275:
262:
249:
236:
220:
204:
144:
115:
113:
112:
111:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
39:
38:
31:Late Cretaceous
27:
17:
12:
11:
5:
4555:
4553:
4545:
4544:
4539:
4534:
4529:
4524:
4519:
4514:
4509:
4504:
4499:
4494:
4492:Chasmosaurines
4484:
4483:
4477:
4476:
4474:
4473:
4460:
4447:
4434:
4421:
4408:
4393:
4377:
4375:
4369:
4368:
4363:
4354:
4353:
4351:
4350:
4340:
4333:
4330:
4329:
4326:
4325:
4311:
4309:
4306:
4305:
4302:
4301:
4298:
4297:
4294:
4293:
4291:
4290:
4283:
4276:
4269:
4261:
4259:
4253:
4252:
4250:
4249:
4241:
4234:
4227:
4219:
4217:
4208:
4202:
4201:
4199:
4198:
4191:
4188:Nasutoceratops
4184:
4177:
4170:
4162:
4160:
4154:
4153:
4151:
4150:
4147:Medusaceratops
4143:
4136:
4128:
4126:
4120:
4119:
4117:
4116:
4109:
4102:
4095:
4088:
4081:
4078:Diabloceratops
4073:
4067:
4065:Centrosaurinae
4061:
4060:
4057:
4056:
4054:
4053:
4046:
4039:
4036:Titanoceratops
4032:
4024:
4021:Regaliceratops
4017:
4010:
4003:
3995:
3993:
3991:Triceratopsini
3987:
3986:
3984:
3983:
3976:
3969:
3962:
3955:
3952:Sierraceratops
3948:
3941:
3938:Navajoceratops
3934:
3927:
3920:
3913:
3906:
3899:
3892:
3885:
3878:
3871:
3863:
3857:
3855:Chasmosaurinae
3851:
3850:
3848:
3847:
3840:
3833:
3826:
3819:
3816:Brachyceratops
3812:
3804:
3798:
3793:
3786:
3785:
3780:
3773:
3772:
3769:
3768:
3754:
3752:
3749:
3748:
3745:
3744:
3741:
3740:
3737:
3736:
3733:
3732:
3729:
3728:
3726:
3725:
3724:
3723:
3717:
3715:
3709:
3708:
3706:
3705:
3698:
3695:Turanoceratops
3691:
3682:
3676:
3670:
3669:
3667:
3666:
3659:
3652:
3645:
3636:
3634:
3625:
3619:
3618:
3616:
3615:
3608:
3605:Unescoceratops
3601:
3594:
3587:
3580:
3573:
3570:Ischioceratops
3566:
3563:Gryphoceratops
3559:
3552:
3545:
3538:
3531:
3523:
3521:
3515:
3514:
3512:
3511:
3503:
3497:
3492:
3489:
3488:
3485:
3484:
3482:
3481:
3473:
3466:
3458:
3456:
3450:
3449:
3447:
3446:
3443:Sasayamagnomus
3439:
3432:
3425:
3418:
3410:
3403:
3396:
3393:Auroraceratops
3389:
3382:
3373:
3363:
3358:
3355:
3354:
3351:
3350:
3348:
3347:
3340:
3333:
3326:
3319:
3316:Chaoyangsaurus
3311:
3309:
3303:
3302:
3300:
3299:
3296:Psittacosaurus
3292:
3285:
3277:
3267:
3262:
3255:
3254:
3249:
3242:
3241:
3238:
3237:
3234:
3233:
3230:
3229:
3226:
3225:
3223:
3222:
3221:
3220:
3214:
3212:
3206:
3205:
3203:
3202:
3201:
3200:
3191:
3185:
3179:
3178:
3176:
3175:
3174:
3173:
3164:
3158:
3148:
3147:
3143:
3142:
3133:
3124:
3115:
3109:
3100:
3093:
3092:
3087:
3085:
3084:
3077:
3070:
3062:
3055:
3054:
3042:
3020:
3019:
2995:
2970:
2954:
2937:
2917:
2886:
2868:
2831:
2764:
2696:
2652:
2590:
2566:
2547:
2526:
2513:
2500:
2484:
2464:
2426:
2399:(4): 681–694.
2374:
2358:
2342:
2318:
2301:
2248:
2232:
2225:
2194:
2178:
2177:
2175:
2172:
2171:
2170:
2163:
2160:
2056:
2053:
2049:
2048:
2045:
2044:
2040:
2039:
2036:
2035:
2031:
2030:
2027:
2026:
2022:
2021:
2018:
2017:
2013:
2012:
2009:
2008:
2004:
2003:
2000:
1999:
1995:
1994:
1991:
1990:
1986:
1985:
1982:
1981:
1977:
1976:
1973:
1972:
1968:
1967:
1964:
1963:
1959:
1958:
1955:
1954:
1950:
1949:
1946:
1945:
1941:
1940:
1937:
1936:
1927:
1924:
1923:
1920:
1919:
1910:
1905:
1903:
1900:
1899:
1896:
1895:
1886:
1881:
1879:
1876:
1875:
1872:
1871:
1862:
1859:
1858:
1855:
1854:
1845:
1840:
1838:
1835:
1834:
1831:
1830:
1821:
1816:
1814:
1811:
1810:
1807:
1806:
1797:
1792:
1790:
1787:
1786:
1783:
1782:
1778:
1777:
1774:
1773:
1764:
1761:
1760:
1757:
1756:
1747:
1742:
1740:
1735:
1733:
1730:
1729:
1726:
1725:
1716:
1711:
1709:
1706:
1705:
1702:
1701:
1697:
1696:
1693:
1692:
1683:
1680:
1679:
1676:
1675:
1666:
1661:
1659:
1654:
1652:
1649:
1648:
1645:
1644:
1635:
1630:
1628:
1625:
1624:
1621:
1620:
1611:
1606:
1604:
1601:
1600:
1597:
1596:
1587:
1582:
1580:
1577:Chasmosaurinae
1572:
1571:
1568:
1567:
1564:Centrosaurinae
1560:
1555:
1553:
1542:
1540:e.a. in 2010.
1524:The following
1477:Centrosaurinae
1469:Chasmosaurinae
1442:
1441:Classification
1439:
1414:
1402:
1386:
1362:frill corner.
1280:with a human,
1269:
1266:
1261:Sierraceratops
792:junior synonym
582:
579:
490:from the Late
414:
413:
412:
411:
396:
385:
371:
370:
364:
363:
362:
361:
338:
319:
299:
298:
294:
293:
273:
269:
268:
266:Chasmosaurinae
260:
256:
255:
247:
243:
242:
234:
227:
226:
224:Neornithischia
218:
211:
210:
202:
195:
194:
189:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
126:
125:
117:
116:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
28:
15:
13:
10:
9:
6:
4:
3:
2:
4554:
4543:
4540:
4538:
4535:
4533:
4530:
4528:
4525:
4523:
4520:
4518:
4515:
4513:
4510:
4508:
4505:
4503:
4500:
4498:
4495:
4493:
4490:
4489:
4487:
4470:
4465:
4461:
4457:
4452:
4448:
4444:
4439:
4435:
4431:
4426:
4422:
4418:
4413:
4409:
4404:
4398:
4394:
4389:
4383:
4379:
4378:
4376:
4374:
4370:
4366:
4361:
4349:
4341:
4339:
4335:
4334:
4331:
4321:
4315:
4289:
4288:
4284:
4282:
4281:
4277:
4275:
4274:
4270:
4268:
4267:
4266:Achelousaurus
4263:
4262:
4260:
4258:
4254:
4247:
4246:
4245:Styracosaurus
4242:
4240:
4239:
4235:
4233:
4232:
4228:
4226:
4225:
4221:
4220:
4218:
4216:
4215:Centrosaurini
4212:
4209:
4207:
4206:Eucentrosaura
4203:
4197:
4196:
4192:
4190:
4189:
4185:
4183:
4182:
4178:
4176:
4175:
4171:
4169:
4168:
4164:
4163:
4161:
4159:
4155:
4149:
4148:
4144:
4142:
4141:
4137:
4135:
4134:
4130:
4129:
4127:
4125:
4121:
4115:
4114:
4110:
4108:
4107:
4106:Wendiceratops
4103:
4101:
4100:
4096:
4094:
4093:
4089:
4087:
4086:
4082:
4080:
4079:
4075:
4074:
4071:
4068:
4066:
4062:
4052:
4051:
4047:
4045:
4044:
4040:
4038:
4037:
4033:
4030:
4029:
4025:
4023:
4022:
4018:
4016:
4015:
4011:
4009:
4008:
4004:
4002:
4001:
4000:Eotriceratops
3997:
3996:
3994:
3992:
3988:
3982:
3981:
3977:
3975:
3974:
3970:
3968:
3967:
3963:
3961:
3960:
3956:
3954:
3953:
3949:
3947:
3946:
3945:Pentaceratops
3942:
3940:
3939:
3935:
3933:
3932:
3928:
3926:
3925:
3924:Kosmoceratops
3921:
3919:
3918:
3914:
3912:
3911:
3907:
3905:
3904:
3900:
3898:
3897:
3896:Bravoceratops
3893:
3891:
3890:
3889:Bisticeratops
3886:
3884:
3883:
3879:
3877:
3876:
3875:Anchiceratops
3872:
3870:
3869:
3868:Agujaceratops
3865:
3864:
3861:
3858:
3856:
3852:
3846:
3845:
3841:
3839:
3838:
3834:
3832:
3831:
3827:
3825:
3824:
3820:
3818:
3817:
3813:
3811:
3810:
3806:
3805:
3802:
3799:
3796:
3791:
3787:
3783:
3778:
3774:
3765:
3759:
3721:
3720:
3719:
3718:
3716:
3714:
3710:
3704:
3703:
3699:
3697:
3696:
3692:
3689:
3688:
3684:
3683:
3680:
3677:
3675:
3674:Ceratopsoidea
3671:
3665:
3664:
3663:Protoceratops
3660:
3658:
3657:
3656:Breviceratops
3653:
3651:
3650:
3646:
3643:
3642:
3638:
3637:
3635:
3633:
3629:
3626:
3624:
3620:
3614:
3613:
3609:
3607:
3606:
3602:
3600:
3599:
3598:Udanoceratops
3595:
3593:
3592:
3591:Prenoceratops
3588:
3586:
3585:
3581:
3579:
3578:
3577:Leptoceratops
3574:
3572:
3571:
3567:
3565:
3564:
3560:
3558:
3557:
3553:
3551:
3550:
3546:
3544:
3543:
3539:
3537:
3536:
3535:Bainoceratops
3532:
3530:
3529:
3525:
3524:
3522:
3520:
3516:
3510:
3509:
3505:
3504:
3501:
3498:
3495:
3490:
3479:
3478:
3474:
3472:
3471:
3470:Helioceratops
3467:
3465:
3464:
3460:
3459:
3457:
3455:
3451:
3445:
3444:
3440:
3438:
3437:
3436:Mosaiceratops
3433:
3431:
3430:
3426:
3424:
3423:
3419:
3416:
3415:
3411:
3409:
3408:
3407:Koreaceratops
3404:
3402:
3401:
3397:
3395:
3394:
3390:
3388:
3387:
3383:
3380:
3379:
3375:
3374:
3371:
3367:
3364:
3361:
3360:Neoceratopsia
3356:
3346:
3345:
3341:
3339:
3338:
3334:
3332:
3331:
3327:
3325:
3324:
3320:
3318:
3317:
3313:
3312:
3310:
3308:
3304:
3298:
3297:
3293:
3291:
3290:
3286:
3284:
3283:
3279:
3278:
3275:
3271:
3268:
3265:
3260:
3256:
3252:
3247:
3243:
3218:
3217:
3216:
3215:
3213:
3211:
3207:
3199:
3195:
3194:
3193:
3192:
3189:
3186:
3184:
3180:
3172:
3168:
3167:
3166:
3165:
3162:
3159:
3157:
3153:
3149:
3141:
3137:
3134:
3132:
3128:
3125:
3123:
3119:
3116:
3114:
3110:
3108:
3104:
3103:
3098:
3094:
3090:
3083:
3078:
3076:
3071:
3069:
3064:
3063:
3060:
3053:
3043:
3041:
3031:
3027:
3009:
3005:
2999:
2996:
2984:
2980:
2974:
2971:
2967:
2964:
2958:
2955:
2951:
2947:
2941:
2938:
2934:
2931:
2927:
2921:
2918:
2913:
2909:
2905:
2901:
2897:
2890:
2887:
2883:
2877:
2875:
2873:
2869:
2863:
2858:
2854:
2850:
2849:F1000Research
2846:
2844:
2835:
2832:
2827:
2823:
2818:
2813:
2809:
2805:
2800:
2795:
2791:
2787:
2783:
2779:
2775:
2768:
2765:
2760:
2756:
2751:
2746:
2742:
2738:
2733:
2728:
2724:
2720:
2716:
2709:
2707:
2705:
2703:
2701:
2697:
2692:
2688:
2683:
2678:
2674:
2670:
2666:
2659:
2657:
2653:
2648:
2644:
2639:
2634:
2629:
2624:
2620:
2616:
2613:(9): e12292.
2612:
2608:
2604:
2597:
2595:
2591:
2587:
2584:
2580:
2576:
2573:R.B. Holmes,
2570:
2567:
2562:
2558:
2551:
2548:
2544:
2540:
2536:
2530:
2527:
2523:
2517:
2514:
2510:
2504:
2501:
2497:
2494:
2488:
2485:
2481:
2477:
2471:
2469:
2465:
2462:
2458:
2454:
2450:
2447:
2445:
2437:
2435:
2433:
2431:
2427:
2422:
2418:
2414:
2410:
2406:
2402:
2398:
2394:
2390:
2383:
2381:
2379:
2375:
2371:
2365:
2363:
2359:
2355:
2352:
2346:
2343:
2339:
2336:
2332:
2328:
2322:
2319:
2316:(10): 129–135
2315:
2312:
2305:
2302:
2297:
2293:
2289:
2285:
2281:
2277:
2273:
2269:
2265:
2261:
2255:
2253:
2249:
2245:
2242:
2236:
2233:
2228:
2226:0-7853-0443-6
2222:
2218:
2211:
2209:
2207:
2205:
2203:
2201:
2199:
2195:
2190:
2183:
2180:
2173:
2169:
2166:
2165:
2161:
2159:
2156:
2152:
2148:
2147:
2142:
2138:
2129:
2125:
2123:
2119:
2115:
2111:
2107:
2101:
2099:
2095:
2091:
2090:
2085:
2081:
2077:
2070:
2066:
2061:
2054:
2047:
2046:
2038:
2037:
2029:
2028:
2020:
2019:
2011:
2010:
2002:
2001:
1993:
1992:
1984:
1983:
1975:
1974:
1966:
1965:
1957:
1956:
1948:
1947:
1939:
1938:
1935:
1934:
1933:
1926:
1925:
1922:
1921:
1918:
1917:
1916:
1909:
1908:
1902:
1901:
1898:
1897:
1894:
1893:
1892:
1885:
1884:
1878:
1877:
1874:
1873:
1870:
1869:
1868:
1867:Eotriceratops
1861:
1860:
1857:
1856:
1853:
1852:
1851:
1844:
1843:
1837:
1836:
1833:
1832:
1829:
1828:
1827:
1820:
1819:
1813:
1812:
1809:
1808:
1805:
1804:
1803:
1802:Anchiceratops
1796:
1795:
1789:
1788:
1785:
1784:
1776:
1775:
1772:
1771:
1770:
1763:
1762:
1759:
1758:
1755:
1754:
1753:
1752:Kosmoceratops
1746:
1745:
1739:
1738:
1732:
1731:
1728:
1727:
1724:
1723:
1722:
1715:
1714:
1708:
1707:
1704:
1703:
1695:
1694:
1691:
1690:
1689:
1688:Pentaceratops
1682:
1681:
1678:
1677:
1674:
1673:
1672:
1665:
1664:
1658:
1657:
1651:
1650:
1647:
1646:
1643:
1642:
1641:
1640:Agujaceratops
1634:
1633:
1627:
1626:
1623:
1622:
1619:
1618:
1617:
1610:
1609:
1603:
1602:
1599:
1598:
1595:
1594:
1593:
1586:
1585:
1578:
1574:
1573:
1570:
1569:
1566:
1565:
1559:
1558:
1551:
1550:
1546:
1545:
1541:
1539:
1538:Scott Sampson
1535:
1531:
1527:
1520:
1516:
1515:chasmosaurine
1513:positions of
1512:
1511:Stratigraphic
1508:
1501:
1497:
1492:
1488:
1486:
1482:
1478:
1474:
1470:
1466:
1462:
1456:
1452:
1447:
1440:
1438:
1436:
1431:
1427:
1422:
1419:
1412:
1407:
1400:
1396:
1391:
1384:
1380:
1376:
1367:
1363:
1360:
1356:
1352:
1351:parietal bone
1348:
1344:
1340:
1336:
1332:
1328:
1324:
1320:
1316:
1309:
1304:
1300:
1298:
1294:
1290:
1283:
1279:
1274:
1267:
1265:
1263:
1262:
1257:
1256:
1251:
1247:
1243:
1239:
1234:
1232:
1228:
1224:
1223:Vagaceratops-
1220:
1216:
1211:
1206:
1202:
1198:
1194:
1190:
1186:
1181:
1180:
1178:
1173:
1169:
1165:
1161:
1157:
1153:
1149:
1145:
1141:
1137:
1133:
1129:
1125:
1121:
1117:
1110:
1106:
1102:
1097:
1093:
1091:
1087:
1083:
1079:
1075:
1071:
1067:
1063:
1059:
1055:
1054:
1049:
1045:
1041:
1037:
1033:
1029:
1025:
1020:
1018:
1014:
1010:
1009:specific name
1006:
1002:
998:
994:
990:
986:
982:
978:
974:
970:
966:
962:
958:
954:
950:
946:
942:
938:
935:was based on
934:
927:
926:
921:
916:
912:
910:
906:
905:Kosmoceratops
902:
898:
894:
890:
889:
884:
880:
873:
869:
868:C. irvinensis
864:
860:
858:
857:
856:Agujaceratops
852:
849:in 1989 from
848:
844:
843:
838:
834:
830:
826:
824:
823:Pentaceratops
819:
815:
811:
807:
803:
799:
797:
793:
789:
785:
781:
777:
773:
769:
765:
761:
757:
753:
749:
745:
741:
737:
733:
729:
725:
721:
717:
713:
709:
705:
698:
694:
690:
686:
684:
680:
676:
672:
668:
667:
666:Protorosaurus
662:
661:
656:
655:Protorosaurus
652:
648:
644:
640:
635:
633:
632:specific name
629:
625:
624:
619:
618:parietal bone
615:
611:
607:
603:
596:skull in 1914
595:
591:
587:
580:
578:
576:
572:
568:
564:
560:
556:
552:
551:Protorosaurus
547:
545:
541:
540:
535:
531:
527:
523:
519:
515:
511:
508:
504:
500:
499:North America
496:
493:
489:
486:
482:
478:
477:
464:
422:
421:
408:
402:
401:
397:
394:
389:
386:
383:
378:
375:
374:
372:
369:
365:
359:
355:
349:
348:
347:
346:C. irvinensis
339:
336:
332:
328:
327:
320:
316:
311:
310:
303:
302:
300:
295:
291:
287:
282:
281:
274:
271:
270:
267:
261:
258:
257:
254:
248:
245:
244:
241:
235:
232:
229:
228:
225:
219:
216:
213:
212:
209:
203:
200:
197:
196:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
148:
143:
139:
136:specimen 843
135:
131:
127:
123:
118:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
43:
36:
32:
26:
22:
19:
4403:Chasmosaurus
4373:Chasmosaurus
4372:
4285:
4280:Stellasaurus
4278:
4271:
4264:
4243:
4236:
4231:Coronosaurus
4229:
4224:Centrosaurus
4222:
4193:
4186:
4179:
4172:
4165:
4145:
4140:Lokiceratops
4138:
4131:
4113:Xenoceratops
4111:
4104:
4099:Sinoceratops
4097:
4090:
4083:
4076:
4048:
4041:
4034:
4026:
4019:
4012:
4007:Nedoceratops
4005:
3998:
3980:Vagaceratops
3978:
3973:Utahceratops
3971:
3966:Terminocavus
3964:
3957:
3950:
3943:
3936:
3929:
3922:
3917:Judiceratops
3915:
3908:
3903:Chasmosaurus
3902:
3901:
3894:
3887:
3880:
3873:
3866:
3842:
3835:
3828:
3821:
3814:
3807:
3795:Ceratopsidae
3782:Ceratopsidae
3713:Ceratopsidae
3702:Zuniceratops
3700:
3693:
3687:Ajkaceratops
3685:
3661:
3654:
3649:Bagaceratops
3647:
3641:Ajkaceratops
3639:
3623:Coronosauria
3610:
3603:
3596:
3589:
3582:
3575:
3568:
3561:
3554:
3547:
3540:
3533:
3528:Asiaceratops
3526:
3506:
3494:Euceratopsia
3477:Yamaceratops
3475:
3468:
3461:
3441:
3434:
3429:Microceratus
3427:
3422:Liaoceratops
3420:
3412:
3405:
3398:
3391:
3384:
3378:Ajkaceratops
3376:
3342:
3335:
3328:
3321:
3314:
3294:
3287:
3280:
3198:Ornithischia
3183:Ornithischia
3135:
3131:Ornithischia
3126:
3117:
3011:. Retrieved
3008:ScienceDaily
3007:
2998:
2987:. Retrieved
2982:
2973:
2965:
2962:
2957:
2949:
2945:
2940:
2932:
2929:
2925:
2920:
2903:
2899:
2895:
2889:
2881:
2852:
2848:
2843:Chasmosaurus
2842:
2834:
2781:
2777:
2767:
2722:
2718:
2672:
2668:
2610:
2606:
2585:
2582:
2579:Chasmosaurus
2578:
2575:C.A. Forster
2569:
2560:
2556:
2550:
2545:(2): 137–162
2542:
2538:
2534:
2529:
2521:
2516:
2508:
2503:
2498:(5): 468–480
2495:
2492:
2487:
2479:
2475:
2452:
2448:
2444:Chasmosaurus
2443:
2396:
2392:
2388:
2369:
2353:
2350:
2345:
2337:
2334:
2330:
2326:
2321:
2313:
2310:
2304:
2271:
2267:
2263:
2260:Arbour, V.M.
2243:
2240:
2235:
2216:
2188:
2182:
2155:Chasmosaurus
2154:
2151:Chasmosaurus
2150:
2144:
2141:Chasmosaurus
2140:
2136:
2134:
2117:
2113:
2102:
2098:Chasmosaurus
2097:
2092:. A certain
2089:Centrosaurus
2087:
2076:Chasmosaurus
2075:
2074:
2068:
2055:Paleobiology
1930:
1929:
1915:Nedoceratops
1913:
1912:
1889:
1888:
1865:
1864:
1848:
1847:
1824:
1823:
1800:
1799:
1769:Vagaceratops
1767:
1766:
1750:
1749:
1719:
1718:
1686:
1685:
1671:Utahceratops
1669:
1668:
1638:
1637:
1616:Mojoceratops
1614:
1613:
1592:Chasmosaurus
1591:
1590:
1589:
1562:
1549:Ceratopsidae
1547:
1534:Chasmosaurus
1533:
1523:
1495:
1485:Chasmosaurus
1484:
1473:Chasmosaurus
1472:
1461:Chasmosaurus
1460:
1459:
1450:
1435:Chasmosaurus
1434:
1426:Chasmosaurus
1425:
1423:
1401:possessed a
1398:
1389:amphiplatian
1374:
1372:
1358:
1354:
1338:
1334:
1331:Chasmosaurus
1330:
1326:
1319:Chasmosaurus
1318:
1314:
1312:
1307:
1296:
1292:
1289:Chasmosaurus
1288:
1287:
1282:Chasmosaurus
1281:
1278:Ceratopsidae
1259:
1253:
1249:
1245:
1241:
1237:
1235:
1230:
1227:Chasmosaurus
1226:
1222:
1219:Vagaceratops
1218:
1215:Chasmosaurus
1214:
1209:
1204:
1200:
1196:
1192:
1188:
1185:Chasmosaurus
1184:
1182:
1177:Utahceratops
1175:
1172:Chasmosaurus
1171:
1167:
1163:
1159:
1156:Mojoceratops
1155:
1151:
1147:
1143:
1140:Mojoceratops
1139:
1136:Chasmosaurus
1135:
1131:
1128:Chasmosaurus
1127:
1123:
1119:
1116:Mojoceratops
1115:
1113:
1108:
1105:Mojoceratops
1104:
1100:
1099:Holotype of
1090:Chasmosaurus
1089:
1085:
1081:
1078:Chasmosaurus
1077:
1074:M. perifania
1073:
1070:nomen dubium
1069:
1066:Mojoceratops
1065:
1061:
1058:M. perifania
1057:
1053:nomen dubium
1051:
1047:
1044:Mojoceratops
1043:
1040:Chasmosaurus
1039:
1036:Chasmosaurus
1035:
1031:
1023:
1022:The species
1021:
1017:Chasmosaurus
1016:
1000:
997:type species
989:Mojoceratops
988:
981:Saskatchewan
957:Mojoceratops
956:
932:
931:The species
930:
925:Vagaceratops
923:
919:
909:Chasmosaurus
908:
904:
901:Vagaceratops
900:
897:Chasmosaurus
896:
893:Vagaceratops
892:
888:Vagaceratops
886:
878:
876:
867:
866:Holotype of
854:
846:
842:nomen dubium
840:
836:
832:
828:
821:
805:
800:
795:
787:
780:Chasmosaurus
779:
775:
771:
768:Chasmosaurus
767:
763:
760:Mojoceratops
759:
756:Mojoceratops
755:
751:
747:
743:
739:
735:
731:
728:Peter Kaisen
726:, honouring
723:
720:Barnum Brown
716:Chasmosaurus
715:
711:
704:Chasmosaurus
703:
701:
692:
678:
675:Chasmosaurus
674:
671:Chasmosaurus
670:
664:
658:
654:
636:
627:
621:
610:Chasmosaurus
609:
599:
593:
592:preparing a
574:
570:
555:Chasmosaurus
554:
550:
548:
537:
534:Chasmosaurus
533:
513:
509:
420:Chasmosaurus
419:
418:
417:
406:
400:Vagaceratops
398:
388:Mojoceratops
387:
376:
353:
344:
343:
325:
324:
308:
307:
280:Chasmosaurus
279:
278:
253:Ceratopsidae
230:
214:
208:Ornithischia
198:
185:
129:
25:Chasmosaurus
24:
18:
4397:Wikispecies
4273:Einiosaurus
4167:Avaceratops
4050:Triceratops
4014:Ojoceratops
3837:Monoclonius
3549:Ferrisaurus
3414:Kulceratops
2588:: 1423–1438
2146:Triceratops
2135:A juvenile
2078:shared its
2071:on the left
1932:Triceratops
1850:Ojoceratops
1355:C. russelli
1339:C. russelli
1327:C. russelli
1315:C. russelli
1297:C. russelli
1268:Description
1255:nomen nudum
1242:C. russelli
1197:C. russelli
1168:C. russelli
1164:C. russelli
1160:C. russelli
1152:C. kaiseni,
1144:C. russelli
1132:C. russelli
1109:C. russelli
825:sternbergii
816:. In 1987,
693:C. russelli
623:Monoclonius
616:NMC 491, a
571:C. russelli
539:Triceratops
326:C. russelli
259:Subfamily:
40:77–76
4486:Categories
4336:See also:
4043:Torosaurus
3959:Spiclypeus
3809:Agathaumas
3722:see below↓
3542:Cerasinops
3330:Stenopelix
3264:Ceratopsia
3251:Ceratopsia
3219:see below↓
3210:Ceratopsia
3122:Dinosauria
3089:Ceratopsia
3013:2019-08-22
2989:2019-08-22
2675:: 83–100.
2563:: 367–370.
2482:(3): 1–175
2246:(2): 25–81
2174:References
2114:C. kaiseni
1891:Torosaurus
1528:shows the
1465:Ceratopsia
1343:osteoderms
1246:C. priscus
1193:C. kaiseni
1148:Eoceratops
1101:C. kaiseni
1086:Eoceratops
1082:C. kaiseni
1062:Eoceratops
983:, western
967:, 76.5–75
776:C. kaiseni
772:Eoceratops
748:Eoceratops
660:Torosaurus
530:short tons
492:Cretaceous
485:ceratopsid
391:Longrich,
377:Eoceratops
240:Ceratopsia
192:Dinosauria
132:skeleton,
3105:Kingdom:
3040:Dinosaurs
2935:: 108–110
2808:1932-6203
2741:2167-8359
2725:: e9251.
2691:2292-1389
2421:129168541
2340:: 145–155
2084:Laramidia
1530:phylogeny
1526:cladogram
1494:Skull of
1481:cladistic
1449:Skull of
1405:synsacrum
1347:squamosal
1201:C. belli.
971:) of the
965:Campanian
949:paratypes
939:specimen
657:("before
612:remains;
528:(1.7–2.2
503:fenestrae
331:Sternberg
165:Kingdom:
159:Eukaryota
35:Campanian
4382:Wikidata
4348:Category
4338:Timeline
3844:Polyonax
3830:Dysganus
3823:Ceratops
3386:Aquilops
3113:Chordata
3111:Phylum:
3107:Animalia
2983:NBC News
2906:: 1–47.
2855:: 1468.
2826:29166406
2778:PLOS ONE
2759:32547873
2647:20877459
2607:PLOS ONE
2449:PLoS ONE
2296:85665879
2162:See also
2118:C. belli
2069:C. belli
1399:C. belli
1375:C. belli
1359:C. belli
1335:C. belli
1308:C. belli
1293:C. belli
1284:in green
1238:C. belli
1231:C. belli
1210:C. belli
1205:C. belli
1007:and the
995:and the
937:holotype
895:back to
820:renamed
796:C. belli
732:C. belli
683:paratype
614:holotype
594:C. belli
575:C. belli
488:dinosaur
405:Sampson
368:Synonyms
313:(Lambe,
309:C. belli
297:Species
246:Family:
179:Chordata
175:Phylum:
169:Animalia
155:Domain:
130:C. belli
4456:4947070
4443:1039685
4430:3238859
4417:4531609
4388:Q131586
4238:Spinops
3556:Gremlin
3344:Yinlong
3026:Portals
2968:: 67–73
2900:Zootaxa
2817:5699823
2786:Bibcode
2750:7278894
2638:2929175
2615:Bibcode
2401:Bibcode
2276:Bibcode
2080:habitat
1579:
1552:
1353:. With
1013:fossils
977:Alberta
708:species
647:Alberta
604:of the
479:) is a
380:Lambe,
352:Holmes
272:Genus:
109:↓
3052:Canada
2824:
2814:
2806:
2757:
2747:
2739:
2689:
2645:
2635:
2419:
2294:
2223:
1575:
1417:caudal
1411:sacral
1080:found
985:Canada
963:(late
804:added
722:named
679:khasma
651:Canada
526:tonnes
518:metres
514:sauros
510:chasma
495:Period
409:, 2010
407:et al.
354:et al.
4469:38855
4438:IRMNG
3136:Clade
3127:Clade
3118:Clade
2719:PeerJ
2417:S2CID
2356:: 1–3
2292:S2CID
1323:horns
1225:like
945:skull
851:Texas
827:into
507:Greek
481:genus
472:-moh-
286:Lambe
231:Clade
215:Clade
199:Clade
186:Clade
4425:GBIF
3196:see
3169:see
2904:2963
2822:PMID
2804:ISSN
2755:PMID
2737:ISSN
2687:ISSN
2643:PMID
2221:ISBN
1430:skin
1424:The
1240:and
1191:and
1154:and
1084:and
1042:and
1005:mojo
993:2010
979:and
953:AMNH
774:and
393:2010
382:1915
358:2001
335:1940
315:1902
290:1914
48:PreꞒ
4412:EoL
3400:Beg
2928:",
2908:doi
2857:doi
2812:PMC
2794:doi
2745:PMC
2727:doi
2677:doi
2633:PMC
2623:doi
2457:doi
2409:doi
2354:669
2333:",
2284:doi
1532:of
1233:."
1158:to
999:is
975:of
941:TMP
870:at
859:.
794:of
645:of
546:).
497:in
483:of
476:-əs
474:SOR
470:KAZ
454:ɔːr
37:),
4488::
4466::
4453::
4440::
4427::
4414::
4399::
4384::
3138::
3129::
3120::
3006:.
2981:.
2966:41
2933:39
2902:.
2871:^
2851:.
2847:.
2820:.
2810:.
2802:.
2792:.
2782:12
2780:.
2776:.
2753:.
2743:.
2735:.
2721:.
2717:.
2699:^
2685:.
2671:.
2667:.
2655:^
2641:.
2631:.
2621:.
2609:.
2605:.
2593:^
2586:38
2561:35
2559:.
2541:,
2496:14
2467:^
2453:11
2451:,
2429:^
2415:.
2407:.
2397:84
2395:.
2377:^
2361:^
2338:27
2314:27
2290:.
2282:.
2272:29
2270:.
2251:^
2197:^
2067:,
1498:,
1453:,
1250:C.
1150:,
1046:.
1019:.
987:.
969:ma
798:.
695:,
649:,
565:,
561:,
522:ft
445:oʊ
403:?
356:,
350:?
333:,
317:)
288:,
233::
217::
201::
188::
98:Pg
42:Ma
4248:?
4031:?
3690:?
3644:?
3480:?
3417:?
3381:?
3081:e
3074:t
3067:v
3028::
3016:.
2992:.
2914:.
2910::
2865:.
2859::
2853:4
2828:.
2796::
2788::
2761:.
2729::
2723:8
2693:.
2679::
2673:7
2649:.
2625::
2617::
2611:5
2543:9
2480:3
2459::
2423:.
2411::
2403::
2298:.
2286::
2278::
2244:3
2229:.
1502:.
1179:.
463:/
460:s
457:ə
451:s
448:ˈ
442:m
439:z
436:æ
433:k
430:ˌ
427:/
423:(
341:†
322:†
305:†
276:†
263:†
250:†
237:†
221:†
205:†
103:N
93:K
88:J
83:T
78:P
73:C
68:D
63:S
58:O
53:Ꞓ
33:(
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