305:
296:
1713:
104:
1719:
1724:
1745:
279:(dorsal exoskeleton), all but the posteriormost are laterally expanded and covered the appendages underneath each of them. The boundaries between tergites have a radiated appearance. Dorsal surface of the first tergite have a pair of kidney-shaped eyes. Based on the differentiation of corresponding appendages, the first 2 tergites and the remaining 9 tergites were interpreted as those of
1731:
347:
with spiny surface, situated between the second appendages and the first gnathobasic appendages. The last gnathobasic appendage pair and the remaining 5 appendage pairs anterior to it were interpreted as corresponding to the second tergite and first tergite, respectively. Appendages posterior to the
370:, instead they may have played a tactile role, sensing the property and position of food items. The endopod and exopod of the trunk appendages may have had a locomotory and respiratory function, respectively.
348:
cephalon were 8 pairs of biramous appendages (each compose of a leg-like endopod and a lobe-like exopod) and a pair of furcae, corresponding to the remaining 9 trunk tergites.
366:, using its gnathobases to crush prey while the spiny labrum may have helped lead food items toward the mouth opening. The specialized second appendages are too fragile to be
317:, showing all cephalic appendages and first two trunk appendage pairs. Dark grey (first diagram) and coloured region (second diagram): appendicular structures, Ant (yellow):
283:(head) and trunk, respectively. Contray to the widely-referred reconstruction by Stürmer & Bergström (1978), but as most of the Cheloniellids, There is no evidence of
720:
Ortega-Hernández, Javier; Legg, David A.; Braddy, Simon J. (2013). "The phylogeny of aglaspidid arthropods and the internal relationships within
Artiopoda".
339:, followed by a pair of specialized second appendages that bore spines and esthetasc-like brushes, and 4 pairs of leg-like appendages that have overlapped
837:"A soft-bodied euarthropod from the early Cambrian Xiaoshiba Lagerstätte of China supports a new clade of basal artiopodans with dorsal ecdysial sutures"
656:
Wendruff, Andrew James, et al. "New cheloniellid arthropod with large raptorial appendages from the
Silurian of Wisconsin, USA." BioRxiv (2018): 407379.
1840:
1825:
910:
304:
1835:
668:
Rust, Jes; Bergmann, Alexandra; Bartels, Christoph; Schoenemann, Brigitte; Sedlmeier, Stephanie; KĂĽhl, Gabriele (2016-03-01).
1712:
778:"The Vicissicaudata revisited – insights from a new aglaspidid arthropod with caudal appendages from the Furongian of China"
344:
322:
295:
1830:
280:
1723:
903:
657:
1765:
1215:
275:
range about 20 centimeters in body length (excluding appendages). The flattened, ovoid body compose of 11
1797:
1718:
1730:
1222:
669:
1267:
1048:
181:
1688:
1295:
1246:
1622:
1458:
1350:
1302:
1238:
1169:
1845:
1748:
1631:
1364:
1343:
1274:
1155:
1140:
1032:
896:
874:
753:
634:
555:
400:
340:
119:
1378:
1322:
1230:
1162:
335:
Underneath the tergites were numerous pairs of appendages. The anteriormost appendage pair were
326:
103:
1802:
1447:
1288:
1092:
1659:
1385:
1371:
1357:
866:
858:
817:
799:
745:
737:
697:
689:
336:
318:
257:
1502:
1176:
561:
446:
1258:
1148:
848:
807:
789:
729:
681:
626:
567:
463:
1405:
1208:
1041:
1481:
1433:
1071:
992:
812:
777:
538:
529:
taxon that have a controversial phylogenetic position. It was previously thought to be
254:
155:
1819:
1474:
1124:
946:
733:
546:
522:
386:
168:
61:
878:
757:
638:
1522:
1426:
1313:
670:"The HunsrĂĽck biota: A unique window into the ecology of Lower Devonian arthropods"
241:
1788:
835:
Du, Kun-sheng; Ortega-Hernández, Javier; Yang, Jie; Zhang, Xi-guang (2019–2020).
1419:
1336:
1200:
1187:
1085:
542:
530:
36:
794:
1558:
1549:
1538:
1513:
1495:
1488:
1467:
1440:
1281:
1006:
685:
142:
81:
46:
862:
803:
741:
693:
321:, Pap (green): 2nd cephalic appendage, Est: esthetasc-like brush, Lbr (red):
1677:
1666:
1607:
1600:
1585:
1113:
1099:
1013:
985:
967:
952:
937:
919:
534:
526:
367:
244:
234:
86:
870:
821:
776:
Lerosey-Aubril, Rudy; Zhu, Xuejian; Ortega-Hernández, Javier (2017-09-11).
749:
701:
287:(a medial tail-like terminal structure) in any described fossil materials.
1782:
1652:
1638:
1576:
1531:
1329:
1106:
1055:
1020:
363:
276:
130:
76:
71:
56:
51:
41:
1645:
1565:
999:
630:
360:
261:
91:
66:
853:
836:
931:
284:
1759:
1593:
533:-related, but later studies repeatedly suggests it as a member of
237:
1763:
892:
613:
Stürmer, Wilhelm; Bergström, Jan (1978-06-01). "The arthropod
343:(jaw-like structure on leg base). The mouth was covered by a
888:
329:, Tap (blue): trunk appendage, Ex: exopod, Te: tergite.
1772:
1676:
1621:
1575:
1548:
1521:
1512:
1457:
1403:
1312:
1257:
1186:
1123:
1069:
1031:
965:
904:
313:Ventral structures of the anterior region of
8:
676:. Special Issue: Fossils as Living Beings.
1760:
1518:
1411:
1192:
1077:
977:
973:
962:
911:
897:
889:
102:
20:
852:
811:
793:
325:, Gap (cyan): gnathobasic appendage, Gn:
580:
674:Arthropod Structure & Development
608:
606:
604:
213:
7:
771:
769:
767:
715:
713:
711:
652:
650:
648:
602:
600:
598:
596:
594:
592:
590:
588:
586:
584:
617:from the devonian hunsrĂĽck shale".
438:
416:
392:
382:
14:
1744:
1743:
1729:
1722:
1717:
1711:
734:10.1111/j.1096-0031.2012.00413.x
303:
294:
1:
1841:Fossil taxa described in 1932
247:, known only by one species,
619:Paläontologische Zeitschrift
545:and related genera. Within
16:Extinct genus of arthropods
1862:
1826:Devonian animals of Europe
795:10.1038/s41598-017-11610-5
1739:
1709:
1414:
1195:
1080:
980:
976:
961:
926:
686:10.1016/j.asd.2016.01.004
508:Phylogenetic position of
460:
443:
436:
421:
414:
397:
390:
212:
207:
120:Scientific classification
118:
110:
101:
23:
1836:HunsrĂĽck Slate fossils
253:, discovered from the
1798:Paleobiology Database
537:, forming the clade
315:Cheloniellon calmani
250:Cheloniellon calmani
216:Cheloniellon calmani
113:Cheloniellon calmani
1831:Fossils of Germany
782:Scientific Reports
631:10.1007/BF03006730
111:Reconstruction of
1813:
1812:
1766:Taxon identifiers
1757:
1756:
1707:
1706:
1703:
1702:
1699:
1698:
1617:
1616:
1399:
1398:
1395:
1394:
1065:
1064:
854:10.1111/cla.12344
553:branched next to
516:
515:
502:
501:
493:
492:
484:
483:
475:
474:
226:
225:
220:
202:
1853:
1806:
1805:
1793:
1792:
1791:
1761:
1747:
1746:
1734:
1733:
1726:
1721:
1715:
1519:
1412:
1259:Tremaglaspididae
1193:
1078:
978:
974:
963:
913:
906:
899:
890:
883:
882:
856:
832:
826:
825:
815:
797:
773:
762:
761:
717:
706:
705:
665:
659:
654:
643:
642:
610:
439:
417:
393:
383:
378:
377:
359:may have been a
307:
298:
218:
200:
106:
96:
33:
29:Temporal range:
21:
1861:
1860:
1856:
1855:
1854:
1852:
1851:
1850:
1816:
1815:
1814:
1809:
1801:
1796:
1787:
1786:
1781:
1768:
1758:
1753:
1735:
1728:
1695:
1672:
1613:
1571:
1544:
1508:
1453:
1408:
1406:Trilobitomorpha
1391:
1308:
1253:
1223:Chacharejocaris
1209:Australaglaspis
1182:
1119:
1074:
1061:
1042:Acanthomeridion
1027:
970:
957:
922:
917:
887:
886:
834:
833:
829:
775:
774:
765:
719:
718:
709:
667:
666:
662:
655:
646:
612:
611:
582:
577:
503:
494:
485:
476:
376:
354:
333:
332:
331:
330:
310:
309:
308:
300:
299:
270:
203:
199:
184:
171:
158:
145:
133:
97:
95:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
31:
30:
27:
17:
12:
11:
5:
1859:
1857:
1849:
1848:
1843:
1838:
1833:
1828:
1818:
1817:
1811:
1810:
1808:
1807:
1794:
1778:
1776:
1770:
1769:
1764:
1755:
1754:
1752:
1751:
1740:
1737:
1736:
1710:
1708:
1705:
1704:
1701:
1700:
1697:
1696:
1694:
1693:
1692:
1691:
1682:
1680:
1674:
1673:
1671:
1670:
1663:
1656:
1649:
1642:
1635:
1627:
1625:
1619:
1618:
1615:
1614:
1612:
1611:
1604:
1597:
1590:
1581:
1579:
1573:
1572:
1570:
1569:
1562:
1554:
1552:
1546:
1545:
1543:
1542:
1535:
1527:
1525:
1516:
1510:
1509:
1507:
1506:
1499:
1492:
1485:
1482:Phytophilaspis
1478:
1471:
1463:
1461:
1455:
1454:
1452:
1451:
1444:
1437:
1434:Pygmaclypeatus
1430:
1423:
1415:
1409:
1404:
1401:
1400:
1397:
1396:
1393:
1392:
1390:
1389:
1382:
1375:
1368:
1361:
1354:
1347:
1340:
1333:
1326:
1318:
1316:
1310:
1309:
1307:
1306:
1299:
1292:
1285:
1278:
1271:
1268:Brachyaglaspis
1263:
1261:
1255:
1254:
1252:
1251:
1243:
1235:
1227:
1219:
1212:
1205:
1196:
1190:
1184:
1183:
1181:
1180:
1173:
1166:
1159:
1152:
1145:
1137:
1129:
1127:
1121:
1120:
1118:
1117:
1110:
1103:
1096:
1089:
1081:
1075:
1072:Vicissicaudata
1070:
1067:
1066:
1063:
1062:
1060:
1059:
1052:
1049:Australimicola
1045:
1037:
1035:
1029:
1028:
1026:
1025:
1017:
1010:
1003:
996:
993:Falcatamacaris
989:
981:
971:
966:
959:
958:
956:
955:
949:
940:
934:
927:
924:
923:
918:
916:
915:
908:
901:
893:
885:
884:
847:(3): 269–281.
827:
763:
707:
680:(2): 140–151.
660:
644:
579:
578:
576:
573:
539:Vicissicaudata
521:is a genus of
514:
513:
505:
504:
500:
499:
496:
495:
491:
490:
487:
486:
482:
481:
478:
477:
473:
472:
469:
468:
459:
456:
455:
452:
451:
442:
437:
435:
432:
431:
428:
427:
420:
415:
413:
410:
409:
406:
405:
396:
391:
389:
381:
375:
374:Classification
372:
353:
350:
312:
311:
302:
301:
293:
292:
291:
290:
289:
269:
266:
258:HunsrĂĽck Slate
255:Lower Devonian
224:
223:
222:
221:
210:
209:
205:
204:
192:
190:
186:
185:
182:Cheloniellidae
179:
177:
173:
172:
166:
164:
160:
159:
156:Vicissicaudata
153:
151:
147:
146:
141:
139:
135:
134:
129:
127:
123:
122:
116:
115:
108:
107:
99:
98:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
35:
34:
32:Lower Devonian
28:
15:
13:
10:
9:
6:
4:
3:
2:
1858:
1847:
1844:
1842:
1839:
1837:
1834:
1832:
1829:
1827:
1824:
1823:
1821:
1804:
1799:
1795:
1790:
1784:
1780:
1779:
1777:
1775:
1771:
1767:
1762:
1750:
1742:
1741:
1738:
1732:
1727:
1725:
1720:
1714:
1690:
1686:
1685:
1684:
1683:
1681:
1679:
1675:
1669:
1668:
1664:
1662:
1661:
1657:
1655:
1654:
1650:
1648:
1647:
1643:
1641:
1640:
1636:
1634:
1633:
1629:
1628:
1626:
1624:
1620:
1610:
1609:
1605:
1603:
1602:
1598:
1596:
1595:
1591:
1588:
1587:
1583:
1582:
1580:
1578:
1574:
1568:
1567:
1563:
1561:
1560:
1556:
1555:
1553:
1551:
1547:
1541:
1540:
1536:
1534:
1533:
1529:
1528:
1526:
1524:
1520:
1517:
1515:
1511:
1505:
1504:
1500:
1498:
1497:
1493:
1491:
1490:
1486:
1484:
1483:
1479:
1477:
1476:
1475:Luohuilinella
1472:
1470:
1469:
1465:
1464:
1462:
1460:
1456:
1450:
1449:
1445:
1443:
1442:
1438:
1436:
1435:
1431:
1429:
1428:
1424:
1422:
1421:
1417:
1416:
1413:
1410:
1407:
1402:
1388:
1387:
1383:
1381:
1380:
1376:
1374:
1373:
1369:
1367:
1366:
1362:
1360:
1359:
1355:
1353:
1352:
1348:
1346:
1345:
1341:
1339:
1338:
1334:
1332:
1331:
1327:
1325:
1324:
1320:
1319:
1317:
1315:
1311:
1305:
1304:
1300:
1298:
1297:
1296:Quasimodaspis
1293:
1291:
1290:
1286:
1284:
1283:
1279:
1277:
1276:
1272:
1270:
1269:
1265:
1264:
1262:
1260:
1256:
1249:
1248:
1247:Rozhkovocaris
1244:
1241:
1240:
1236:
1233:
1232:
1228:
1225:
1224:
1220:
1218:
1217:
1213:
1211:
1210:
1206:
1203:
1202:
1198:
1197:
1194:
1191:
1189:
1185:
1179:
1178:
1174:
1172:
1171:
1167:
1165:
1164:
1160:
1158:
1157:
1153:
1151:
1150:
1146:
1143:
1142:
1138:
1136:
1135:
1131:
1130:
1128:
1126:
1125:Cheloniellida
1122:
1116:
1115:
1111:
1109:
1108:
1104:
1102:
1101:
1097:
1095:
1094:
1090:
1088:
1087:
1083:
1082:
1079:
1076:
1073:
1068:
1058:
1057:
1053:
1051:
1050:
1046:
1044:
1043:
1039:
1038:
1036:
1034:
1030:
1023:
1022:
1018:
1016:
1015:
1011:
1009:
1008:
1004:
1002:
1001:
997:
995:
994:
990:
988:
987:
983:
982:
979:
975:
972:
969:
964:
960:
954:
950:
948:
947:Panarthropoda
944:
941:
939:
936:Superphylum:
935:
933:
929:
928:
925:
921:
914:
909:
907:
902:
900:
895:
894:
891:
880:
876:
872:
868:
864:
860:
855:
850:
846:
842:
838:
831:
828:
823:
819:
814:
809:
805:
801:
796:
791:
787:
783:
779:
772:
770:
768:
764:
759:
755:
751:
747:
743:
739:
735:
731:
727:
723:
716:
714:
712:
708:
703:
699:
695:
691:
687:
683:
679:
675:
671:
664:
661:
658:
653:
651:
649:
645:
640:
636:
632:
628:
624:
620:
616:
609:
607:
605:
603:
601:
599:
597:
595:
593:
591:
589:
587:
585:
581:
574:
572:
570:
569:
564:
563:
559:but basal to
558:
557:
552:
548:
547:Cheloniellida
544:
540:
536:
532:
528:
525:, an extinct
524:
523:Cheloniellida
520:
511:
507:
506:
498:
497:
489:
488:
480:
479:
471:
470:
467:
466:
465:
458:
457:
454:
453:
450:
449:
448:
441:
440:
434:
433:
430:
429:
426:
425:
419:
418:
412:
411:
408:
407:
404:
403:
402:
395:
394:
388:
387:Cheloniellida
385:
384:
380:
379:
373:
371:
369:
365:
362:
358:
351:
349:
346:
342:
338:
328:
324:
320:
316:
306:
297:
288:
286:
282:
278:
274:
267:
265:
263:
259:
256:
252:
251:
246:
243:
239:
236:
232:
231:
217:
214:
211:
206:
198:
197:
191:
188:
187:
183:
178:
175:
174:
170:
169:Cheloniellida
165:
162:
161:
157:
152:
149:
148:
144:
140:
137:
136:
132:
128:
125:
124:
121:
117:
114:
109:
105:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
1774:Cheloniellon
1773:
1716:
1665:
1658:
1651:
1644:
1637:
1630:
1623:Conciliterga
1606:
1599:
1592:
1584:
1564:
1557:
1537:
1530:
1523:Emucarididae
1501:
1494:
1487:
1480:
1473:
1466:
1459:Xandarellida
1446:
1439:
1432:
1427:Kwanyinaspis
1425:
1418:
1384:
1377:
1370:
1363:
1356:
1351:Glypharthrus
1349:
1342:
1335:
1328:
1321:
1314:Aglaspididae
1303:Tremaglaspis
1301:
1294:
1287:
1280:
1273:
1266:
1245:
1239:Obrutschewia
1237:
1229:
1221:
1214:
1207:
1199:
1175:
1170:Pseudarthron
1168:
1161:
1154:
1147:
1139:
1134:Cheloniellon
1133:
1132:
1112:
1105:
1098:
1091:
1084:
1054:
1047:
1040:
1019:
1012:
1005:
998:
991:
984:
942:
844:
840:
830:
788:(1): 11117.
785:
781:
728:(1): 15–45.
725:
721:
677:
673:
663:
625:(1): 57–81.
622:
618:
615:Cheloniellon
614:
566:
560:
554:
551:Cheloniellon
550:
519:Cheloniellon
518:
517:
510:Cheloniellon
509:
462:
461:
445:
444:
424:Cheloniellon
423:
422:
399:
398:
357:Cheloniellon
356:
355:
352:Paleoecology
334:
314:
273:Cheloniellon
272:
271:
249:
248:
242:cheloniellid
230:Cheloniellon
229:
228:
227:
219:Broili 1932
215:
196:Cheloniellon
195:
194:
150:Superclass:
112:
25:Cheloniellon
24:
18:
1632:Haifengella
1420:Arthroaspis
1365:Hesselbonia
1344:Chraspedops
1337:Aglaspoides
1275:Chlupacaris
1201:Angarocaris
1188:Aglaspidida
1156:Neostrabops
1141:Drabovaspis
1086:Emeraldella
1033:Protosutura
556:Neostrabops
543:Aglaspidida
531:chelicerate
401:Neostrabops
341:gnathobases
201:Broili 1932
1820:Categories
1559:Misszhouia
1550:Naraoiidae
1539:Kangacaris
1514:Nektaspida
1496:Xandarella
1489:Sinoburius
1468:Cindarella
1441:Retifacies
1379:Tuboculops
1323:Aglaspella
1282:Cyclopites
1231:Girardevia
1216:Beckwithia
1163:Paraduslia
1007:Squamacula
953:Arthropoda
841:Cladistics
722:Cladistics
575:References
327:gnathobase
268:Morphology
143:Arthropoda
1846:Artiopoda
1789:Q96386650
1689:Trilobita
1678:Trilobita
1667:Tegopelte
1608:Tariccoia
1601:Soomaspis
1586:Buenaspis
1448:Tonglaiia
1289:Flobertia
1114:Carimersa
1100:Kodymirus
1093:Eozetetes
1014:Thulaspis
986:Bailongia
968:Artiopoda
938:Ecdysozoa
930:Kingdom:
920:Artiopoda
863:0748-3007
804:2045-2322
742:1096-0031
694:1467-8039
535:Artiopoda
527:arthropod
368:raptorial
245:arthropod
235:monotypic
126:Kingdom:
1783:Wikidata
1749:Category
1660:Skioldia
1653:Saperion
1639:Helmetia
1577:Liwiidae
1532:Emucaris
1386:Uarthrus
1372:Setaspis
1358:Gogglops
1330:Aglaspis
1107:Sidneyia
1056:Zhiwenia
1021:Urokodia
951:Phylum:
932:Animalia
879:89985331
871:34622993
822:28894246
758:85744103
750:34814371
702:26826500
639:87725308
364:predator
337:antennae
281:cephalon
277:tergites
208:Species
176:Family:
138:Phylum:
131:Animalia
1646:Kuamaia
1566:Naraoia
1503:Zhugeia
1177:Triopus
1000:Molaria
813:5593897
562:Triopus
447:Triopus
361:benthic
319:antenna
262:Germany
189:Genus:
163:Order:
1149:Duslia
877:
869:
861:
820:
810:
802:
756:
748:
740:
700:
692:
637:
568:Duslia
464:Duslia
345:labrum
323:labrum
285:telson
1803:19035
1594:Liwia
943:Clade
875:S2CID
754:S2CID
635:S2CID
541:with
238:genus
233:is a
1687:see
867:PMID
859:ISSN
818:PMID
800:ISSN
746:PMID
738:ISSN
698:PMID
690:ISSN
565:and
37:Preęž’
849:doi
808:PMC
790:doi
730:doi
682:doi
627:doi
260:of
240:of
1822::
1800::
1785::
945::
873:.
865:.
857:.
845:35
843:.
839:.
816:.
806:.
798:.
784:.
780:.
766:^
752:.
744:.
736:.
726:29
724:.
710:^
696:.
688:.
678:45
672:.
647:^
633:.
623:52
621:.
583:^
571:.
549:,
512:.
264:.
87:Pg
1589:?
1250:?
1242:?
1234:?
1226:?
1204:?
1144:?
1024:?
912:e
905:t
898:v
881:.
851::
824:.
792::
786:7
760:.
732::
704:.
684::
641:.
629::
193:â€
180:â€
167:â€
154:â€
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:ęž’
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
