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Chimaerasuchidae

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Marinho, T.S. & Carvalho, I.S. (2007). Revision of the Sphagesauridae KUHN, 1968 (Crocodyliformes, Mesoeucrocodylia). In: Carvalho, I.S.; Cassab, R.C.T.; Schwanke, C.; Carvalho, M.A.; Fernandes, A.C.S.; Rodrigues, M.A.; Carvalho, M.S.S.; Arai, M.; Oliveira, M.E.Q. (Eds.). Paleontologia: Cenários
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does not have any of the synapomorphies of this family that was originally erected by Kuhn (1968). In this new paper, the authors propose an emended diagnosis for the Sphagesauridae, pointing new synapomorphies for this taxon and propose that the Chimaerasuchidae would be a valid family in spite of
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Candeiro, C.R.A. & Martinelli, A.G. (2006). A review of paleogeographical and chronostratigraphical distribution of mesoeucrocodylian species from the upper Cretaceous beds from the Baurus (Brazil) and Neuquén (Argentina) groups, Southern South America.
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in the analysis. Neosuchians and notosuchians are the two major clades of mesoeucrocodylians. When neosuchians are included in analyses with
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In a recent contribution, Marinho & Carvalho (2007) did a revision of Sphagesauridae and noticed that
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of Sphagesauridae. This is because Sphagesauridae was named before Chimaerasuchidae, and under
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Kuhn, O. 1968. Die Vortzeitlichen Krokodile. Verlag Oeben, Krailing, München, 124 p.
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Nature "A Pugnosed Crocodyliform from the Late Cretaceous of Madagascar"
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placed Chimaerasuchidae within a new clade of mesoeucrocodylians called
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of small (~1 metre long), possibly herbivorous crocodyliforms from the
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Carvahlo, I.S., Ribeiro, L.C.B. and Avilla, L.S. (2004).
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belongs to Shagesauridae, Chimaerasuchidae would be a
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Rio de Janeiro: Interciência, 431:, another newly erected clade that contained 8: 946:the poor phylogenetic analysis of Carvalho 2317: 2287: 2137: 2048: 1927: 1872: 1863: 1746: 1742: 1731: 1718: 1650: 1427: 1250: 1219: 1215: 1204: 1191: 1137: 1128: 1079: 1063: 1049: 1041: 114: 20: 912:has been found to be the sister taxon of 399:In the phylogenetic analysis of Carvalho 990:Journal of South American Earth Sciences 938:rules, the oldest name has priority. 7: 1023:Ancient Crocrodile Chomped On Plants 710: 698: 669: 644: 617: 588: 561: 532: 477: 465: 454: 2437:Early Cretaceous first appearances 965:Journal of Vertebrate Paleontology 922:is often assigned its own family, 14: 2422:Early Cretaceous crocodylomorphs 2272: 2264: 2257: 2250: 1704: 1697: 1691: 1685: 136: 2432:Late Cretaceous crocodylomorphs 975:Uberasuchus terrificus sp. nov. 959:Pol, D. (2003). New remains of 1: 427:. It was the sister taxon of 2442:Late Cretaceous extinctions 2427:Terrestrial crocodylomorphs 2302:Terrestrial crocodylomorphs 2463: 16:Extinct family of reptiles 2297: 2286: 2248: 2140: 2051: 1930: 1875: 1749: 1745: 1730: 1717: 1683: 1649: 1430: 1253: 1222: 1218: 1203: 1190: 1140: 1127: 1078: 758: 732: 715: 708: 696: 674: 667: 649: 642: 622: 615: 593: 586: 566: 559: 537: 530: 499: 482: 475: 463: 300: 293: 133:Scientific classification 131: 122: 113: 23: 125:Chimaerasuchus paradoxus 403:, the family contained 2447:Pseudosuchian families 443:. Below is a modified 396: 344:. It was erected as a 2394:Paleobiology Database 389: 348:in 2004 by Carvalho 961:Sphagesaurus huenei 390:The possible genus 2292:Related categories 2002:Pissarrachampsinae 1028:2012-09-25 at the 397: 364:and possibly also 342:mesoeucrocodylians 336:crocodiles") is a 2409: 2408: 2381:Open Tree of Life 2323:Taxon identifiers 2314: 2313: 2310: 2309: 2282: 2281: 2246: 2245: 2242: 2241: 2238: 2237: 2234: 2233: 2230: 2229: 2038: 2037: 2034: 2033: 1851: 1850: 1713: 1712: 1681: 1680: 1677: 1676: 1673: 1672: 1645: 1644: 1641: 1640: 1562:Colhuehuapisuchus 1292:Mahajangasuchidae 1238: 1237: 1186: 1185: 1182: 1181: 1178: 1177: 979:Gondwana Research 883: 882: 834: 833: 825: 824: 816: 815: 807: 806: 798: 797: 789: 788: 780: 779: 771: 770: 747: 746: 514: 513: 327: 326: 289: 2454: 2402: 2401: 2389: 2388: 2376: 2375: 2363: 2362: 2350: 2349: 2348: 2331:Chimaerasuchidae 2318: 2288: 2277: 2276: 2269: 2268: 2262: 2261: 2254: 2146:Adamantinasuchus 2138: 2049: 1928: 1873: 1864: 1747: 1743: 1732: 1719: 1708: 1702: 1701: 1695: 1689: 1651: 1428: 1251: 1220: 1216: 1205: 1192: 1138: 1129: 1122: 1121: 1108:Mesoeucrocodylia 1080: 1065: 1058: 1051: 1042: 894:did not include 873:Peirosauriformes 864:Peirosaurimorpha 711: 699: 670: 645: 618: 589: 562: 533: 478: 471:Chimaerasuchidae 466: 455: 358:Early Cretaceous 330:Chimaerasuchidae 315: 305: 280: 277:Chimaerasuchidae 275: 262: 246: 141: 140: 118: 108: 38: 29:Early Cretaceous 27:Temporal range: 24:Chimaerasuchidae 21: 2462: 2461: 2457: 2456: 2455: 2453: 2452: 2451: 2412: 2411: 2410: 2405: 2397: 2392: 2384: 2379: 2371: 2366: 2358: 2353: 2344: 2343: 2338: 2325: 2315: 2306: 2293: 2278: 2271: 2263: 2256: 2226: 2205:Caipirasuchinae 2199: 2180:Armadillosuchus 2165: 2127: 2084: 2030: 2018:Pissarrachampsa 1996: 1991:Stratiotosuchus 1955: 1936:Cynodontosuchus 1917: 1911:Razanandrongobe 1904:Pehuenchesuchus 1860: 1858:Xenodontosuchia 1847: 1811:Candidodontidae 1805: 1785:Razanandrongobe 1739: 1726: 1709: 1696: 1669: 1637: 1597:Montealtosuchus 1569:Gasparinisuchus 1519: 1464: 1417: 1313: 1308:Mahajangasuchus 1286: 1247: 1234: 1212: 1199: 1174: 1154: 1123: 1085: 1084: 1074: 1069: 1030:Wayback Machine 1016:Taxon Feedback 1013: 956: 884: 875: 868: 867: 852: 851: 848:Notosuchiformes 841:Notosuchimorpha 835: 826: 817: 808: 799: 790: 781: 772: 748: 652:Candidodontidae 569:Uruguaysuchidae 515: 429:Notosuchimorpha 384: 372:Late Cretaceous 279: 273: 264:Xenodontosuchia 260: 244: 232:Crocodyliformes 219:Crocodylomorpha 135: 109: 107: 106: 105: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 33: 32: 25: 17: 12: 11: 5: 2460: 2458: 2450: 2449: 2444: 2439: 2434: 2429: 2424: 2414: 2413: 2407: 2406: 2404: 2403: 2390: 2377: 2364: 2351: 2335: 2333: 2327: 2326: 2321: 2312: 2311: 2308: 2307: 2305: 2304: 2298: 2295: 2294: 2291: 2284: 2283: 2280: 2279: 2249: 2247: 2244: 2243: 2240: 2239: 2236: 2235: 2232: 2231: 2228: 2227: 2225: 2224: 2221:Morrinhosuchus 2217: 2209: 2207: 2201: 2200: 2198: 2197: 2190: 2183: 2175: 2173: 2171:Sphagesaurinae 2167: 2166: 2164: 2163: 2156: 2149: 2141: 2135: 2133:Sphagesauridae 2129: 2128: 2126: 2125: 2118: 2110: 2103: 2094: 2092: 2086: 2085: 2083: 2082: 2079:Eptalofosuchus 2075: 2068: 2061: 2057:Brasileosaurus 2052: 2046: 2040: 2039: 2036: 2035: 2032: 2031: 2029: 2028: 2021: 2014: 2006: 2004: 1998: 1997: 1995: 1994: 1987: 1980: 1973: 1965: 1963: 1957: 1956: 1954: 1953: 1946: 1943:Gondwanasuchus 1939: 1931: 1925: 1919: 1918: 1916: 1915: 1907: 1900: 1892: 1884: 1881:Chimaerasuchus 1876: 1870: 1861: 1856: 1853: 1852: 1849: 1848: 1846: 1845: 1838: 1831: 1828:Lavocatchampsa 1824: 1816: 1814: 1807: 1806: 1804: 1803: 1796: 1789: 1781: 1774: 1766: 1759: 1750: 1740: 1735: 1728: 1727: 1722: 1715: 1714: 1711: 1710: 1684: 1682: 1679: 1678: 1675: 1674: 1671: 1670: 1668: 1667: 1666: 1665: 1659: 1657: 1647: 1646: 1643: 1642: 1639: 1638: 1636: 1635: 1628: 1621: 1614: 1607: 1600: 1593: 1586: 1579: 1572: 1565: 1558: 1555:Bayomesasuchus 1551: 1544: 1537: 1529: 1527: 1521: 1520: 1518: 1517: 1510: 1503: 1496: 1489: 1482: 1479:Barreirosuchus 1474: 1472: 1466: 1465: 1463: 1462: 1458:Stolokrosuchus 1454: 1447: 1439: 1431: 1425: 1419: 1418: 1416: 1415: 1408: 1401: 1394: 1387: 1380: 1373: 1366: 1359: 1356:Dentaneosuchus 1352: 1345: 1338: 1331: 1323: 1321: 1315: 1314: 1312: 1311: 1304: 1296: 1294: 1288: 1287: 1285: 1284: 1277: 1273:Trematochampsa 1269: 1262: 1254: 1248: 1243: 1240: 1239: 1236: 1235: 1233: 1232: 1223: 1213: 1208: 1201: 1200: 1195: 1188: 1187: 1184: 1183: 1180: 1179: 1176: 1175: 1173: 1172: 1171: 1170: 1164: 1162: 1156: 1155: 1153: 1152: 1151: 1150: 1141: 1135: 1125: 1124: 1120: 1119: 1110: 1104: 1098: 1092: 1083: 1076: 1075: 1070: 1068: 1067: 1060: 1053: 1045: 1039: 1038: 1033: 1021:Science News " 1019: 1012: 1011:External links 1009: 1008: 1007: 999: 996: 985: 971: 955: 952: 950:(2004) paper. 943:Chimaerasuchus 932:junior synonym 928:Chimaerasuchus 924:Sphagesauridae 909:Chimaerasuchus 904:Chimaerasuchus 881: 880: 877: 876: 836: 832: 831: 828: 827: 823: 822: 819: 818: 814: 813: 810: 809: 805: 804: 801: 800: 796: 795: 792: 791: 787: 786: 783: 782: 778: 777: 774: 773: 769: 768: 765: 764: 757: 754: 753: 750: 749: 745: 744: 741: 740: 731: 728: 727: 724: 723: 714: 709: 707: 697: 695: 692:Peirosauroidea 687: 686: 683: 682: 673: 668: 666: 660: 659: 656: 655: 648: 643: 641: 633: 632: 629: 628: 621: 616: 614: 606: 605: 602: 601: 592: 587: 585: 577: 576: 573: 572: 565: 560: 558: 550: 549: 546: 545: 541:Stolokrosuchus 536: 531: 529: 521: 520: 517: 516: 512: 511: 508: 507: 498: 495: 494: 491: 490: 486:Chimaerasuchus 481: 476: 474: 464: 462: 459:Gondwanasuchia 453: 447:from Carvalho 437:sebecosuchians 425:Gondwanasuchia 405:Chimaerasuchus 383: 380: 354:Chimaerasuchus 325: 324: 323: 322: 311: 308:Chimaerasuchus 298: 297: 291: 290: 271: 267: 266: 258: 251: 250: 242: 235: 234: 229: 222: 221: 216: 209: 208: 203: 196: 195: 190: 183: 182: 177: 173: 172: 167: 163: 162: 157: 153: 152: 147: 143: 142: 129: 128: 120: 119: 111: 110: 102: 101: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 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1986: 1985: 1981: 1979: 1978: 1977:Aplestosuchus 1974: 1972: 1971: 1970:Aphaurosuchus 1967: 1966: 1964: 1962: 1961:Baurusuchinae 1958: 1952: 1951: 1947: 1945: 1944: 1940: 1938: 1937: 1933: 1932: 1929: 1926: 1924: 1923:Baurusuchidae 1920: 1913: 1912: 1908: 1906: 1905: 1901: 1898: 1897: 1893: 1890: 1889: 1888:Comahuesuchus 1885: 1883: 1882: 1878: 1877: 1874: 1871: 1869: 1865: 1862: 1859: 1854: 1844: 1843: 1839: 1837: 1836: 1832: 1830: 1829: 1825: 1823: 1822: 1818: 1817: 1815: 1812: 1808: 1802: 1801: 1800:Uruguaysuchus 1797: 1795: 1794: 1790: 1787: 1786: 1782: 1780: 1779: 1775: 1772: 1771: 1770:Comahuesuchus 1767: 1765: 1764: 1763:Araripesuchus 1760: 1757: 1756: 1752: 1751: 1748: 1744: 1741: 1738: 1733: 1729: 1725: 1720: 1716: 1707: 1703: 1700: 1694: 1688: 1663: 1662: 1661: 1660: 1658: 1656: 1652: 1648: 1634: 1633: 1632:Uberabasuchus 1629: 1627: 1626: 1622: 1620: 1619: 1615: 1613: 1612: 1608: 1606: 1605: 1601: 1599: 1598: 1594: 1592: 1591: 1587: 1585: 1584: 1580: 1578: 1577: 1573: 1571: 1570: 1566: 1564: 1563: 1559: 1557: 1556: 1552: 1550: 1549: 1548:Barrosasuchus 1545: 1543: 1542: 1541:Barcinosuchus 1538: 1536: 1535: 1531: 1530: 1528: 1526: 1525:Peirosaurinae 1522: 1516: 1515: 1511: 1509: 1508: 1504: 1502: 1501: 1497: 1495: 1494: 1490: 1488: 1487: 1483: 1481: 1480: 1476: 1475: 1473: 1471: 1467: 1460: 1459: 1455: 1453: 1452: 1451:Miadanasuchus 1448: 1445: 1444: 1440: 1438: 1437: 1433: 1432: 1429: 1426: 1424: 1423:Peirosauridae 1420: 1414: 1413: 1409: 1407: 1406: 1402: 1400: 1399: 1395: 1393: 1392: 1388: 1386: 1385: 1381: 1379: 1378: 1374: 1372: 1371: 1367: 1365: 1364: 1360: 1358: 1357: 1353: 1351: 1350: 1346: 1344: 1343: 1339: 1337: 1336: 1332: 1330: 1329: 1325: 1324: 1322: 1320: 1316: 1310: 1309: 1305: 1303: 1302: 1298: 1297: 1295: 1293: 1289: 1283: 1282: 1278: 1275: 1274: 1270: 1268: 1267: 1263: 1261: 1260: 1256: 1255: 1252: 1249: 1246: 1241: 1230: 1229: 1225: 1224: 1221: 1217: 1214: 1211: 1206: 1202: 1198: 1193: 1189: 1168: 1167: 1166: 1165: 1163: 1161: 1157: 1149: 1145: 1144: 1143: 1142: 1139: 1136: 1134: 1130: 1126: 1118: 1114: 1111: 1109: 1105: 1103: 1099: 1097: 1093: 1091: 1087: 1086: 1081: 1077: 1073: 1066: 1061: 1059: 1054: 1052: 1047: 1046: 1043: 1037: 1034: 1031: 1027: 1024: 1020: 1018: 1015: 1014: 1010: 1005: 1000: 997: 994: 991: 986: 983: 980: 976: 972: 969: 966: 962: 958: 957: 953: 951: 949: 944: 939: 937: 933: 929: 925: 921: 917: 916: 911: 910: 905: 901: 897: 893: 890:(2004) paper 889: 886:The Carvalho 879: 878: 874: 871: 865: 862: 858: 855: 849: 846: 842: 839: 830: 829: 821: 820: 812: 811: 803: 802: 794: 793: 785: 784: 776: 775: 767: 766: 763: 762: 761:Peirosauridae 756: 755: 752: 751: 743: 742: 739: 738: 737: 730: 729: 726: 725: 722: 721: 720: 713: 712: 705: 701: 700: 693: 689: 688: 685: 684: 681: 680: 679: 678:Araripesuchus 672: 671: 665: 662: 661: 658: 657: 654: 653: 647: 646: 639: 635: 634: 631: 630: 627: 626: 620: 619: 612: 608: 607: 604: 603: 600: 599: 598: 597:Comahuesuchus 591: 590: 583: 579: 578: 575: 574: 571: 570: 564: 563: 556: 552: 551: 548: 547: 544: 543: 542: 535: 534: 527: 523: 522: 519: 518: 510: 509: 506: 505: 504: 497: 496: 493: 492: 489: 488: 487: 480: 479: 472: 468: 467: 460: 457: 456: 452: 450: 446: 442: 438: 434: 430: 426: 422: 418: 414: 410: 406: 402: 395: 394: 388: 381: 379: 377: 373: 369: 368: 363: 359: 355: 352:and included 351: 347: 343: 339: 335: 331: 320: 319: 312: 310: 309: 302: 301: 299: 296: 292: 288: 284: 278: 272: 269: 268: 265: 259: 256: 253: 252: 249: 243: 240: 237: 236: 233: 230: 227: 224: 223: 220: 217: 214: 211: 210: 207: 204: 201: 198: 197: 194: 191: 188: 185: 184: 181: 178: 175: 174: 171: 168: 165: 164: 161: 158: 155: 154: 151: 148: 145: 144: 139: 134: 130: 127: 126: 121: 117: 112: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 37: 30: 22: 19: 2330: 2255: 2219: 2212: 2194:Sphagesaurus 2192: 2185: 2178: 2158: 2151: 2144: 2120: 2112: 2105: 2097: 2090:Notosuchidae 2077: 2070: 2063: 2055: 2044:Sphagesauria 2023: 2016: 2009: 1989: 1982: 1975: 1968: 1948: 1941: 1934: 1909: 1902: 1894: 1886: 1879: 1840: 1835:Malawisuchus 1833: 1826: 1819: 1798: 1791: 1783: 1778:Libycosuchus 1776: 1768: 1761: 1753: 1690: 1630: 1623: 1616: 1609: 1604:Patagosuchus 1602: 1595: 1588: 1581: 1576:Hamadasuchus 1574: 1567: 1560: 1553: 1546: 1539: 1534:Antaeusuchus 1532: 1512: 1505: 1498: 1493:Epoidesuchus 1491: 1486:Caririsuchus 1484: 1477: 1470:Pepesuchinae 1456: 1449: 1443:Fortignathus 1441: 1436:Amargasuchus 1434: 1410: 1403: 1398:Sahitisuchus 1396: 1389: 1382: 1375: 1368: 1361: 1354: 1347: 1340: 1335:Barinasuchus 1333: 1326: 1306: 1299: 1279: 1271: 1264: 1257: 1226: 1148:Pseudosuchia 1133:Pseudosuchia 1112: 1003: 992: 989: 981: 978: 974: 967: 964: 960: 947: 942: 940: 927: 920:Sphagesaurus 919: 915:Sphagesaurus 913: 907: 903: 899: 887: 885: 869: 860: 857:Terriasuchia 853: 844: 837: 759: 734: 733: 719:Malawisuchus 717: 716: 676: 675: 663: 650: 637: 623: 595: 594: 581: 567: 554: 539: 538: 525: 501: 500: 484: 483: 470: 448: 441:peirosaurids 433:notosuchians 420: 408: 404: 400: 398: 391: 365: 353: 349: 329: 328: 316: 306: 282: 276: 254: 238: 225: 212: 206:Pseudosuchia 199: 186: 123: 18: 2107:Llanosuchus 2025:Wargosuchus 1984:Baurusuchus 1896:Eremosuchus 1868:Baurusuchia 1755:Anatosuchus 1625:Rukwasuchus 1611:Peirosaurus 1514:Roxochampsa 1412:Zulmasuchus 1363:Iberosuchus 1349:Bretesuchus 1342:Bergisuchus 1328:Ayllusuchus 1301:Kaprosuchus 1259:Baharijodon 1228:Microsuchus 1210:Ziphosuchia 1197:Ziphosuchia 1160:Ziphosuchia 1072:Ziphosuchia 896:neosuchians 704:Itasuchidae 625:Ziphosuchia 193:Archosauria 2416:Categories 2160:Yacarerani 2122:Notosuchus 1842:Pakasuchus 1821:Candidodon 1793:Simosuchus 1737:Notosuchia 1724:Notosuchia 1664:see below↓ 1655:Notosuchia 1618:Pepesuchus 1590:Lomasuchus 1583:Kinesuchus 1507:Pepesuchus 1391:Ogresuchus 1384:Lorosuchus 1377:Langstonia 1281:Wanosuchus 1169:see below↓ 1117:Metasuchia 1102:Sauropsida 1006:: 589-599. 984:: 975-1002 954:References 900:Simosuchus 611:Notosuchia 503:Simosuchus 417:Cretaceous 409:Simosuchus 393:Simosuchus 376:Madagascar 367:Simosuchus 318:Simosuchus 248:Notosuchia 1950:Pabwehshi 1500:Itasuchus 1370:Ilchunaia 1319:Sebecidae 1266:Doratodon 1088:Kingdom: 970:: 817-831 892:phylogeny 736:Itasuchus 445:cladogram 382:Phylogeny 370:from the 356:from the 281:Carvalho 156:Kingdom: 150:Eukaryota 34:120  2346:Q5099282 2340:Wikidata 1096:Chordata 1094:Phylum: 1090:Animalia 1026:Archived 995::116-129 451:(2004): 270:Family: 180:Reptilia 170:Chordata 166:Phylum: 160:Animalia 146:Domain: 2386:3605945 2360:4822290 1405:Sebecus 1245:Sebecia 1106:Order: 1100:Class: 706:  694:  640:  613:  584:  557:  528:  473:  461:  334:Chimera 176:Class: 103:↓ 2399:114379 2373:117579 948:et al. 888:et al. 702:  690:  636:  609:  580:  553:  524:  469:  439:, and 413:genera 411:, two 338:family 295:Genera 283:et al. 2368:IRMNG 1113:Clade 926:. If 449:et al 421:et al 401:et al 362:China 350:et al 346:clade 255:Clade 239:Clade 226:Clade 213:Clade 200:Clade 187:Clade 2355:GBIF 1146:see 936:ICZN 902:and 407:and 287:2004 42:PreꞒ 374:of 360:of 340:of 2418:: 2396:: 2383:: 2370:: 2357:: 2342:: 1115:: 993:22 968:23 918:. 866:, 859:, 850:, 843:, 435:, 332:(" 285:, 257:: 241:: 228:: 215:: 202:: 189:: 92:Pg 36:Ma 31:, 2117:? 2102:? 2060:? 1914:? 1899:? 1891:? 1813:? 1788:? 1773:? 1758:? 1461:? 1446:? 1276:? 1231:? 1064:e 1057:t 1050:v 1032:" 1004:1 982:4 870:5 861:4 854:3 845:2 838:1 664:5 638:4 582:3 555:2 526:1 321:? 314:† 304:† 274:† 261:† 245:† 97:N 87:K 82:J 77:T 72:P 67:C 62:D 57:S 52:O 47:Ꞓ

Index

Early Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Chimaerasuchus paradoxus
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Archosauria
Pseudosuchia
Crocodylomorpha
Crocodyliformes
Notosuchia
Xenodontosuchia
Chimaerasuchidae
2004

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