124:
102:
236:
274:
in general form and habits, but have short, rounded snouts, without the modifications found in related families. Many species have long, tapering tails, giving them an alternative name of
1481:
1332:
1371:
281:
They are found in temperate and tropical marine waters worldwide. Most species are restricted to depths below 200 m (660 ft), but a few, notably the
289:, can locally be found at relatively shallow depths. They range from 38 to 150 cm (1.25 to 4.92 ft) in maximum total length, depending on species.
1306:
1345:
1105:
1029:
1476:
1132:
sp. nov. (Chimaeriformes: Chimaeridae), a new species of chimaerid from the southeastern
Atlantic and southwestern Indian oceans.
1443:
1147:"A New Genus of Chimaerid Fish (Holocephali, Chimaeridae) from the Upper Oligocene Sooke Formation of British Columbia, Canada"
318:
476:
361:
1350:
1358:
775:
1267:
1254:
123:
579:
871:
1376:
770:
612:
594:
654:
519:
480:
383:
357:
240:
200:
1096:
Roberts, Clive; Stewart, A. L.; Struthers, Carl D.; Barker, Jeremy; Kortet, Salme; Freeborn, Michelle (2015).
278:. Shortnose chimaeras have a venomous spine on their backs, which is sufficiently dangerous to injure humans.
800:
919:
827:
740:
714:
753:
1194:
892:
804:
795:
761:
626:
818:
701:
1407:
925:
862:
856:
784:
731:
682:
673:
669:
537:
1065:
949:
910:
757:
1425:
1272:
749:
1471:
1448:
1386:
1293:
1241:
1146:
906:
841:
832:
813:
705:
696:
608:
588:
468:
463:
375:
349:
344:
472:
379:
353:
940:
901:
634:
441:
745:
1174:
886:
725:
688:
663:
603:
573:
533:
528:
452:
396:
370:
335:
264:
118:
110:
1412:
977:
1394:
1280:
1166:
1111:
1101:
1035:
1025:
954:
934:
877:
649:
621:
514:
430:
326:
1337:
1232:
1435:
1399:
1158:
500:
489:
416:
309:
260:
217:
982:
391:
282:
847:
505:
421:
405:
271:
165:
1465:
1178:
640:
564:
331:
314:
59:
1246:
1259:
1162:
101:
1285:
1019:
1363:
1319:
1226:
998:
286:
175:
34:
1217:
1024:. Fishes of the Gulf of Mexico. Austin: University of Texas Press. p. 38.
559:
244:
224:
79:
44:
1170:
1115:
1039:
986:
135:
84:
1211:
185:
155:
74:
69:
54:
49:
39:
235:
1021:
89:
64:
1324:
145:
1311:
1188:
1145:
Popov, Evgeny V.; Johns, Marjorie J.; Suntok, Stephen (2020-01-02).
1100:. Vol. 2. Wellington, New Zealand: Te Papa Press. p. 40.
1420:
234:
1192:
1298:
1128:
Walovich, K.A., Ebert, D.A. & Kemper, J.M. (2017):
1066:"Family Chimaeridae – Shortnose chimaeras or ratfishes"
297:
The species are grouped into two genera and include:
1201:
8:
550:Last & Stevens, 1994; White et al. 2006
1189:
100:
20:
1010:
1482:Taxa named by Charles Lucien Bonaparte
1018:McEachran, J.; Fechhelm, J.D. (1998).
7:
1426:AC642D14-3A40-FFC1-34F2-F922D80B293A
1387:b410d18a-4350-4b10-af64-f950968a8731
1091:
1089:
1087:
1060:
1058:
1056:
494:Didier, Last & W. T. White, 2008
446:Didier, Last & W. T. White, 2008
410:Didier, Last & W. T. White, 2008
1151:Journal of Vertebrate Paleontology
837:(Eastern Pacific black ghostshark)
14:
981:Popov, Johns & Suntok, 2020
835:, Ebert, Long & Didier, 2009
821:, Didier, Long & Ebert, 2006
629:, Didier, Long & Ebert, 2006
122:
964:sp. D/G (giant black chimaera)
16:Family of cartilaginous fishes
1:
1163:10.1080/02724634.2020.1772275
945:(pointy-nosed blue chimaera)
1477:Cartilaginous fish families
1134:Zootaxa, 4226 (4): 509–520.
970:sp. F (Peruvian rabbitfish)
1498:
872:Hydrolagus novaezealandiae
1098:The fishes of New Zealand
771:Hydrolagus macrophthalmus
213:
208:
119:Scientific classification
117:
108:
99:
23:
920:Hydrolagus purpurescens
852:(large-eyed rabbitfish)
828:Hydrolagus melanophasma
766:(Portuguese rabbitfish)
715:Hydrolagus homonycteris
584:(small-eyed rabbitfish)
239:A Chimaeridae fish and
823:(Galápagos ghostshark)
796:Hydrolagus matallanasi
741:Hydrolagus lusitanicus
437:(Carpenter's chimaera)
248:
1408:Paleobiology Database
897:(Ogilby's ghostshark)
857:Hydrolagus mitsukurii
785:Hydrolagus marmoratus
496:(shortspine chimaera)
338:& Pogonoski, 2008
257:short-nosed chimaeras
238:
1130:Hydrolagus erithacus
861:(D. S. Jordan &
842:Hydrolagus mirabilis
814:Hydrolagus mccoskeri
809:(striped rabbitfish)
791:(marbled ghostshark)
710:(Robin's ghostshark)
697:Hydrolagus erithacus
687:(D. S. Jordan &
589:Hydrolagus africanus
464:Chimaera notafricana
448:(longspine chimaera)
366:(Bahamas ghostshark)
345:Chimaera bahamaensis
270:They resemble other
902:Hydrolagus pallidus
780:(big-eyed chimaera)
750:I. M. R. Figueiredo
645:(ninespot chimaera)
635:Hydrolagus barbouri
510:(Owston's chimaera)
442:Chimaera macrospina
426:(Jordan's chimaera)
412:(southern chimaera)
340:(whitefin chimaera)
887:Hydrolagus ogilbyi
736:(bight ghostshark)
726:Hydrolagus lemures
721:(black ghostshark)
683:Hydrolagus eidolon
664:Hydrolagus colliei
659:(pale ghost shark)
604:Hydrolagus alberti
599:(African chimaera)
574:Hydrolagus affinis
529:Chimaera phantasma
524:(leopard chimaera)
453:Chimaera monstrosa
371:Chimaera carophila
265:cartilaginous fish
249:
111:Hydrolagus alberti
1459:
1458:
1395:Open Tree of Life
1195:Taxon identifiers
1107:978-0-9941041-6-8
1031:978-0-292-75206-1
958:
950:Hydrolagus waitei
944:
935:Hydrolagus trolli
930:(purple chimaera)
929:
914:
896:
882:(dark ghostshark)
881:
866:
851:
836:
822:
808:
790:
779:
765:
735:
720:
709:
692:
678:(spotted ratfish)
677:
658:
650:Hydrolagus bemisi
644:
630:
622:Hydrolagus alphus
616:
598:
583:
569:(Miocene-Recent)
568:
552:(Lombok chimaera)
551:
542:(silver chimaera)
541:
523:
515:Chimaera panthera
509:
495:
484:
458:
447:
436:
431:Chimaera lignaria
425:
411:
400:
386:
365:
339:
327:Chimaera argiloba
321:
241:deepwater mussels
233:
232:
204:
1489:
1452:
1451:
1439:
1438:
1429:
1428:
1416:
1415:
1403:
1402:
1390:
1389:
1380:
1379:
1367:
1366:
1364:NHMSYS0021053520
1354:
1353:
1341:
1340:
1328:
1327:
1315:
1314:
1302:
1301:
1289:
1288:
1276:
1275:
1263:
1262:
1250:
1249:
1237:
1236:
1235:
1222:
1221:
1220:
1190:
1183:
1182:
1142:
1136:
1126:
1120:
1119:
1093:
1082:
1081:
1079:
1077:
1062:
1051:
1050:
1048:
1046:
1015:
953:
938:
923:
905:
890:
875:
860:
845:
831:
819:L. A. K. Barnett
817:
799:
788:
774:
744:
729:
718:
700:
686:
667:
653:
638:
625:
607:
592:
577:
563:
549:
532:
518:
504:
501:Chimaera owstoni
493:
490:Chimaera obscura
467:
456:
445:
434:
420:
417:Chimaera jordani
409:
401:(Cuban chimaera)
395:
387:(brown chimaera)
374:
348:
330:
322:(Eocene-Recent)
313:
199:
127:
126:
104:
94:
31:
27:Temporal range:
21:
1497:
1496:
1492:
1491:
1490:
1488:
1487:
1486:
1462:
1461:
1460:
1455:
1447:
1442:
1434:
1432:
1424:
1419:
1411:
1406:
1398:
1393:
1385:
1383:
1375:
1370:
1362:
1357:
1349:
1344:
1336:
1331:
1323:
1318:
1310:
1305:
1297:
1292:
1284:
1279:
1271:
1266:
1258:
1253:
1245:
1240:
1231:
1230:
1225:
1216:
1215:
1210:
1197:
1187:
1186:
1157:(1): e1772275.
1144:
1143:
1139:
1127:
1123:
1108:
1095:
1094:
1085:
1075:
1073:
1064:
1063:
1054:
1044:
1042:
1032:
1017:
1016:
1012:
1007:
995:
983:Sooke Formation
915:(pale chimaera)
754:Bordalo-Machado
617:(Gulf chimaera)
485:(Cape chimaera)
457:Linnaeus, 1758]
392:Chimaera cubana
382:, Naylor &
295:
283:spotted ratfish
198:
121:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
1495:
1493:
1485:
1484:
1479:
1474:
1464:
1463:
1457:
1456:
1454:
1453:
1440:
1430:
1417:
1404:
1391:
1381:
1368:
1355:
1342:
1329:
1316:
1303:
1290:
1277:
1264:
1251:
1238:
1223:
1207:
1205:
1199:
1198:
1193:
1185:
1184:
1137:
1121:
1106:
1083:
1052:
1030:
1009:
1008:
1006:
1003:
1002:
1001:
994:
991:
990:
989:
973:
972:
971:
965:
959:
946:
931:
916:
898:
883:
868:
853:
838:
824:
810:
792:
781:
767:
737:
722:
711:
704:, Ebert &
693:
679:
660:
646:
631:
618:
600:
585:
555:
554:
553:
543:
525:
511:
497:
486:
460:
449:
438:
427:
413:
406:Chimaera fulva
402:
388:
367:
341:
294:
291:
231:
230:
229:
228:
221:
211:
210:
206:
205:
193:
189:
188:
186:Chimaeriformes
183:
179:
178:
173:
169:
168:
166:Chondrichthyes
163:
159:
158:
153:
149:
148:
143:
139:
138:
133:
129:
128:
115:
114:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
32:
26:
15:
13:
10:
9:
6:
4:
3:
2:
1494:
1483:
1480:
1478:
1475:
1473:
1470:
1469:
1467:
1450:
1445:
1441:
1437:
1431:
1427:
1422:
1418:
1414:
1409:
1405:
1401:
1396:
1392:
1388:
1382:
1378:
1373:
1369:
1365:
1360:
1356:
1352:
1347:
1343:
1339:
1334:
1330:
1326:
1321:
1317:
1313:
1308:
1304:
1300:
1295:
1291:
1287:
1282:
1278:
1274:
1269:
1265:
1261:
1256:
1252:
1248:
1243:
1239:
1234:
1228:
1224:
1219:
1213:
1209:
1208:
1206:
1204:
1200:
1196:
1191:
1180:
1176:
1172:
1168:
1164:
1160:
1156:
1152:
1148:
1141:
1138:
1135:
1131:
1125:
1122:
1117:
1113:
1109:
1103:
1099:
1092:
1090:
1088:
1084:
1071:
1067:
1061:
1059:
1057:
1053:
1041:
1037:
1033:
1027:
1023:
1022:
1014:
1011:
1004:
1000:
997:
996:
992:
988:
984:
980:
979:
974:
969:
966:
963:
960:
956:
952:
951:
947:
942:
939:Didier &
937:
936:
932:
927:
926:C. H. Gilbert
922:
921:
917:
912:
908:
904:
903:
899:
894:
889:
888:
884:
879:
874:
873:
869:
864:
859:
858:
854:
849:
844:
843:
839:
834:
830:
829:
825:
820:
816:
815:
811:
806:
802:
798:
797:
793:
787:
786:
782:
777:
773:
772:
768:
763:
759:
755:
751:
747:
743:
742:
738:
733:
728:
727:
723:
717:
716:
712:
707:
703:
699:
698:
694:
690:
685:
684:
680:
675:
674:E. T. Bennett
671:
666:
665:
661:
656:
652:
651:
647:
642:
637:
636:
632:
628:
624:
623:
619:
614:
610:
606:
605:
601:
596:
591:
590:
586:
581:
580:Brito Capello
576:
575:
571:
570:
566:
562:
561:
556:
547:
544:
539:
535:
531:
530:
526:
521:
517:
516:
512:
507:
506:S. Tanaka (I)
503:
502:
498:
492:
491:
487:
482:
478:
474:
470:
466:
465:
461:
455:
454:
450:
444:
443:
439:
433:
432:
428:
423:
422:S. Tanaka (I)
419:
418:
414:
408:
407:
403:
398:
397:Howell-Rivero
394:
393:
389:
385:
381:
377:
373:
372:
368:
363:
359:
355:
351:
347:
346:
342:
337:
333:
329:
328:
324:
323:
320:
316:
312:
311:
306:
305:
304:
303:
298:
292:
290:
288:
284:
279:
277:
273:
268:
266:
262:
258:
254:
246:
243:at edge of a
242:
237:
227:
226:
222:
220:
219:
215:
214:
212:
207:
202:
197:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
147:
144:
141:
140:
137:
134:
131:
130:
125:
120:
116:
113:
112:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
30:Eocene–Recent
22:
19:
1202:
1154:
1150:
1140:
1133:
1129:
1124:
1097:
1074:. Retrieved
1069:
1043:. Retrieved
1020:
1013:
976:
967:
961:
948:
933:
918:
900:
885:
870:
855:
840:
826:
812:
794:
789:Didier, 2008
783:
769:
739:
724:
719:Didier, 2008
713:
695:
681:
662:
648:
633:
620:
602:
587:
572:
558:
545:
534:D. S. Jordan
527:
513:
499:
488:
462:
459:(Rabbitfish)
451:
440:
435:Didier, 2002
429:
415:
404:
390:
369:
343:
325:
308:
301:
299:
296:
280:
275:
269:
256:
252:
250:
223:
216:
195:
109:
18:
1472:Chimaeridae
1320:iNaturalist
1260:Chimaeridae
1247:Chimaeridae
1233:Chimaeridae
1227:Wikispecies
1203:Chimaeridae
999:Stensioella
867:(spookfish)
833:K. C. James
689:C. L. Hubbs
336:W. T. White
302:Chimaeridae
287:rabbit fish
253:Chimaeridae
196:Chimaeridae
176:Holocephali
24:Chimaeridae
1466:Categories
1076:29 January
1005:References
985:, Canada,
968:Hydrolagus
962:Hydrolagus
776:F. de Buen
758:C. Almeida
560:Hydrolagus
548:cf. sp. E
245:brine pool
225:Hydrolagus
172:Subclass:
1179:221751740
1171:0272-4634
1116:908128805
987:Oligocene
978:Canadodus
613:Schroeder
595:Gilchrist
272:chimaeras
201:Bonaparte
142:Kingdom:
136:Eukaryota
1218:Q1543050
1212:Wikidata
1070:Fishbase
1040:38468784
993:See also
911:Stehmann
746:T. Moura
702:Walovich
627:Quaranta
546:Chimaera
477:Compagno
362:Compagno
315:Linnaeus
310:Chimaera
259:, are a
218:Chimaera
192:Family:
156:Chordata
152:Phylum:
146:Animalia
132:Domain:
1045:12 July
928:, 1905)
895:, 1898)
880:, 1911)
865:, 1904)
850:, 1904)
848:Collett
734:, 1939)
732:Whitley
691:, 1925)
676:, 1839)
643:, 1908)
609:Bigelow
597:, 1922)
582:, 1868)
300:Family
293:Species
276:ratfish
209:Genera
182:Order:
162:Class:
1449:105696
1436:114050
1433:uBio:
1413:132672
1400:550659
1384:NZOR:
1351:161013
1338:114901
1177:
1169:
1114:
1104:
1072:. 2019
1038:
1028:
957:, 1907
955:Fowler
943:, 2002
913:, 1990
909:&
878:Fowler
863:Snyder
807:, 2004
805:Vooren
803:&
778:, 1959
764:, 2005
760:&
708:, 2017
706:Kemper
672:&
657:, 2002
655:Didier
641:Garman
615:, 1951
611:&
567:, 1863
557:Genus
540:, 1900
538:Snyder
536:&
522:, 1998
520:Didier
508:, 1905
483:, 2010
481:Didier
479:&
469:Kemper
424:, 1905
399:, 1936
384:Didier
376:Kemper
364:, 2010
360:&
358:Didier
350:Kemper
307:Genus
261:family
203:, 1831
1444:WoRMS
1421:Plazi
1333:IRMNG
1325:60452
1175:S2CID
941:Séret
907:Hardy
893:Waite
762:Gordo
473:Ebert
380:Ebert
354:Ebert
255:, or
1377:7869
1372:NCBI
1346:ITIS
1312:2181
1307:GBIF
1299:8904
1268:BOLD
1167:ISSN
1112:OCLC
1102:ISBN
1078:2019
1047:2021
1036:OCLC
1026:ISBN
801:Soto
565:Gill
332:Last
319:1758
285:and
251:The
35:PreꞒ
1359:NBN
1294:EoL
1286:84B
1281:CoL
1273:961
1255:AFD
1242:ADW
1159:doi
670:Lay
263:of
1468::
1446::
1423::
1410::
1397::
1374::
1361::
1348::
1335::
1322::
1309::
1296::
1283::
1270::
1257::
1244::
1229::
1214::
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90:N
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50:S
45:O
40:Ꞓ
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