634:
2077:
804:
2718:
2713:
49:
31:
647:
2750:
209:
2470:
2056:
972:
364:, with only 26 segments (four fewer than the typical number), adult females have only 42 leg pairs, and adult males have only 40 pairs of walking legs (excluding two pairs of gonopods). Adult females with 32 segments (two more than the typical number) have 54 leg pairs (e.g., in the family Peterjohnsiidae), which is the maximum number fixed by species in the class
2740:
2044:
446:
ninth and final stage with 30 segments. The male usually begins to develop gonopods in the seventh stage. Species that produce adults with fewer or more segments than the usual number, however, deviate from the typical pattern by adding a different number of segments at some stage, reaching maturity in a different stage, or both.
429:
Millipedes in this order grow and develop through a series of molts, adding segments until they reach a fixed number in the adult stage, which is usually the same for a given sex in a given species, at which point the molting and the addition of segments and legs stop. This mode of development, known
259:
Chordeumatidans take on a wide variety of forms, including some that are cylindrical and others that are flat-backed. Most species have 26 to 32 body segments (including the telson) behind the head, with the number usually fixed within species. These millipedes range in length from 3.5 mm to 42 mm,
445:
For the typical species in this order, post-embryonic development takes place in nine stages. The juvenile millipede hatches with 6 segments and only 3 pairs of legs in the first stage, then usually goes through stages with 8, 11, 15, 19, 23, 25, and 28 segments, before emerging as adults in the
404:
exhibits the most extensive modifications, including five leg pairs (pairs 7 through 11) in the gonopod complex. Some species also deviate from the usual body plan by reducing or eliminating leg pairs in the adult female. In the family
Chordeumatidae, for example, adult females feature a legless
371:
Many species deviate from the expected number of walking legs, however, because they deviate in terms of sex-linked modifications to their legs. For example, many species involve another leg pair in addition to pairs 8 and 9 in the gonopod complex in adult males. In the family
359:
With these deviations from the usual 30 segments, the number of leg pairs in adults changes, usually with two leg pairs added or subtracted for each segment added to or subtracted from the typical number. For example, in
290:
Most chordeumatidan species have 30 body segments (including the telson) as adults, and adult females in these species have 50 leg pairs. In adult males in this order, two leg pairs (pair 8 and pair 9) are modified into
1890:"Atlas of myriapod biogeography. I. Indigenous ordinal and supra-ordinal distributions in the Diplopoda: Perspectives on taxon origins and ages, and a hypothesis on the origin and early evolution of the class"
1315:. Linnean Society of London, Estuarine and Brackish-water Sciences Association. London: Published for the Linnean Society of London and the Estuarine and Brackish-Water Sciences Association by E.J. Brill.
271:) is relatively narrow, giving the appearance of a distinct "neck" in many species, and the body tapers towards the rear. A dorsal groove runs down the length of the body, and some species feature
376:, leg pair 7 in adult males is modified as part of the gonopod complex. In many species, the gonopod complex instead includes leg pair 10 in addition to pairs 8 and 9 (e.g.,
2007:"Class Diplopoda de Blainville in Gervais, 1844. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness"
260:
although most species are 10 mm to 25 mm long. They are usually drab in color, ranging from various shades of brown to unpigmented, but some feature distinct patterns.
2958:
2997:
1411:
434:, distinguishes this order from most other orders of millipedes. In most other orders, millipedes continue to molt as adults, developing through either
1787:"Golovatchia, new genus, and Golovatchiidae, from the Far East of the Russian Republic, with a comment on Hoffmaneumatidae (Diplopoda: Chordeumatidae)"
1344:
295:, leaving 48 pairs of walking legs in the typical adult male chordeumatidan. Many species in this order, however, deviate from this typical body plan.
1786:
1603:"Craspedosomid Millipedes Discovered in Australia: Reginaterreuma, Neocambrisoma and Peterjohnsia, New Genera (Myriapoda: Diplopoda: Craspedosomida)"
2932:
1721:
1380:
2971:
263:
Species in this order share a set of features that distinguishes them from other millipedes. A key feature is the presence of six large bristles (
2082:
494:, are largely absent from South America. They are abundant in cold, rocky, mountainous areas of Europe and central Asia, and range northward to
251:
feature spinnerets used to build nests of silk. These millipedes produce this silk to create chambers in which to molt or to lay their eggs.
1748:"Branneria bonoculus, n. sp., a second species in the North American milliped family Branneriidae (Diplopoda: Chordeumatida: Brannerioidea)"
2115:
1435:"Myriapodes du Népal. II. Diplopodes Craspedosomides nouveaux de l'Himalaya et de la région indo-malaise (Craspedosomidea et Chordeumidea)"
1911:
Sierwald, Petra; Bond, Jason E. (2007). "Current Status of the
Myriapod Class Diplopoda (Millipedes): Taxonomic Diversity and Phylogeny".
1476:"The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)"
1320:
1232:
1626:"Le cycle biologique de Chamaesoma broelemanni Ribaut et Verhoeff, 1913 (Diplopoda, Craspedosomatida) en forêt d'Orléans (France)"
1181:
3056:
1271:"Anamorphosis in Millipedes (Diplopoda) — The Present State of Knowledge with Some Developmental and Phylogenetic Considerations"
1270:
2060:
1948:"Millipede Taxonomy after 250 Years: Classification and Taxonomic Practices in a Mega-Diverse yet Understudied Arthropod Group"
2976:
2717:
454:
Chordeumatidans have a wide distribution, occurring on all continents except
Antarctica. These millipedes are found in the
2984:
2712:
1813:"Five New Chordeumatidan Millipeds from China: New Species of Vieteuma (Kashmirieumatidae) and Nepalella (Megalotylidae)"
1722:"Japanese chordeumatid millipeds. I. On the genus Speophilosoma Takakuwa (Diplopoda, Chordeumatida, Speophilosomatidae)"
48:
2854:
2831:
306:
344:
in segment number, specifically, adult males with two segments fewer than adult females, for example, in the family
3002:
2911:
2733:
3089:
2924:
298:
Many chordeumatidan species deviate from the usual 30 segments: A few species have 26 segments as adults (e.g.,
2108:
2048:
421:), adult females instead exhibit modifications to the second pair of legs, which are reduced to small nubbins.
633:
1515:"On the Central and East Asian milliped family Diplomaragnidae (Diplopoda, Choredeumatida, Diplomaragnoidea)"
330:
1835:
3084:
2771:
1212:
439:
435:
431:
2859:
1388:
898:
135:
2743:
1666:
1355:
409:(the "platosternite") where a third pair of legs would otherwise be. In other species (e.g., the genus
3038:
1625:
1602:
3061:
3012:
2893:
2818:
2071:
1959:
1812:
1009:
610:
570:
510:
Chordeumatida contains approximately 1200 species, classified in four suborders and approximately 50
384:
158:
803:
2754:
2101:
976:
906:
826:
684:
275:, lateral extensions of the exoskeleton. Paranota are also found in some other millipedes, notably
268:
2469:
2809:
1863:
1702:
1647:
1578:
1025:
764:
699:
638:
544:
487:
373:
240:
182:
120:
43:
873:
279:, from which Chordeumatidans can be distinguished by having a dorsal groove. Unlike most other
3020:
2880:
1987:
1928:
1855:
1767:
1694:
1686:
1570:
1495:
1456:
1326:
1316:
1228:
1211:
Enghoff, Henrik; Golovatch, Sergei; Short, Megan; Stoev, Pavel; Wesener, Thomas (2015-01-01).
1091:
818:
668:
341:
214:
154:
110:
2093:
3048:
2963:
2574:
2021:
1977:
1967:
1920:
1847:
1759:
1678:
1637:
1560:
1522:
1487:
1446:
1282:
1220:
1057:
890:
772:
756:
708:
676:
511:
479:
417:
411:
396:
318:
3025:
30:
2602:
2501:
2341:
2299:
2268:
1924:
1123:
1033:
985:
780:
626:
594:
586:
459:
353:
336:
237:
225:
2076:
2006:
1963:
2919:
2906:
2652:
2592:
2564:
2519:
2514:
2389:
2346:
2287:
2250:
1982:
1947:
1286:
1049:
740:
660:
536:
463:
401:
390:
280:
150:
2823:
646:
3078:
2836:
2662:
2635:
2617:
1851:
1747:
1682:
1651:
1475:
1017:
922:
748:
724:
267:) on the dorsal surface of each body segment, three on each side. The first segment (
1867:
1706:
2885:
2687:
2625:
2509:
2422:
2331:
2318:
1582:
1514:
1156:
1107:
858:
651:
618:
208:
170:
1889:
1972:
1434:
2989:
2945:
2803:
2630:
2533:
2440:
2417:
2407:
2384:
2336:
2212:
1131:
1001:
956:
948:
796:
495:
475:
276:
162:
2794:
2055:
2025:
2640:
2584:
2370:
2241:
2195:
2187:
2087:
1565:
1541:
1491:
1224:
716:
483:
300:
204:
1771:
1763:
1690:
1574:
1499:
1460:
236:, they are found nearly worldwide. Chordeumatida is the largest order in the
2263:
2158:
2152:
2146:
2125:
1330:
1083:
882:
842:
578:
378:
365:
229:
100:
90:
80:
60:
1991:
1932:
1859:
1698:
1451:
971:
2043:
2867:
2846:
2788:
1410:
Golovatch, Sergei I.; Geoffroy, Jean-Jacques; Mauries, Jean-Paul (2006).
1310:
1217:
Treatise on
Zoology - Anatomy, Taxonomy, Biology. The Myriapoda, Volume 2
1115:
471:
406:
345:
324:
312:
272:
2872:
499:
467:
455:
292:
284:
1312:
Millipedes : keys and notes for the identification of the species
2950:
2379:
2140:
1642:
1526:
809:
248:
70:
2937:
2765:
3033:
1354:. The Field Museum, Chicago, IL. 26 September 2006. Archived from
970:
802:
645:
632:
515:
491:
264:
2066:
1836:"Trunk segment numbers and sequential segmentation in myriapods"
1667:"Trunk segment numbers and sequential segmentation in myriapods"
2769:
2485:
2171:
2097:
1542:"Buotidae, a new family for the minute North American milliped
352:(males with 28, females with the usual 30), and in the family
2898:
1720:
Shear, William A.; Tsurusaki, Nobuo; Tanabe, Tsutomu (1994).
1946:
Brewer, Michael S.; Sierwald, Petra; Bond, Jason E. (2012).
1387:. The Field Museum, Chicago. 9 February 2012. Archived from
1269:
Enghoff, Henrik; Dohle, Wolfgang; Blower, J. Gordon (1993).
502:, and in North America up into Canada and southwest Alaska.
1630:
Bulletin du Muséum
National d'Histoire Naturelle, Section A
1549:(Chamberlin) 1940 (Diplopoda, Chordeumatida, Striarioidea)"
328:, in which most species have 29), one genus features 31 (
1412:"Four new Chordeumatida (Diplopoda) from caves in China"
1613:(1): 107–133 – via Biodiversity Heritage Library.
813:(Haaseidae) from Germany. The dorsal groove is visible
1823:(6): 63–72 – via Biodiversity Heritage Library.
514:, although several families contain only one to five
1474:
Antić, Dragan Ž; Makarov, Slobodan E. (2016-12-22).
2778:
2680:
2671:
2649:
2616:
2601:
2583:
2561:
2532:
2500:
2431:
2398:
2368:
2359:
2317:
2298:
2280:
2249:
2240:
2211:
2186:
232:containing more than 1,400 species. Also known as
1817:Proceedings of the California Academy of Sciences
224:(from the Greek word for "sausage") is a large
1888:Shelley, R. M. & Golavatch, S. I. (2011).
2109:
283:(worm-like) millipedes, chordeumatidans lack
8:
1189:. Ohio Division of Wildlife. pp. 3, 24.
356:(males with the usual 30, females with 32).
2766:
2677:
2613:
2558:
2497:
2493:
2482:
2365:
2314:
2310:
2246:
2237:
2183:
2179:
2168:
2116:
2102:
2094:
2075:
1654:– via Biodiversity Heritage Library.
1463:– via Biodiversity Heritage Library.
1345:"Putative apomorphies of millipede clades"
340:). Some species also deviate by featuring
38:Two unidentified species from New Zealand
29:
20:
1981:
1971:
1641:
1564:
1450:
1275:Zoological Journal of the Linnean Society
1148:
482:. Species in this order are present in
1925:10.1146/annurev.ento.52.111805.090210
1596:
1594:
1592:
1264:
980:(Metopidiotrichidae) from New Zealand
348:(males with 26, females with 28), in
7:
3039:2578E112-FFB1-FE20-FF5B-97F9FD70FA7F
3013:0361b0fd-9d56-407e-adaf-aa90525f2a95
2925:3b6a2a08-359d-4225-a20c-d1f1a454e417
2739:
1304:
1302:
1300:
1298:
1296:
1262:
1260:
1258:
1256:
1254:
1252:
1250:
1248:
1246:
1244:
1206:
1204:
1202:
1200:
1198:
1196:
334:), and many species have 32 (e.g.,
1287:10.1111/j.1096-3642.1993.tb00305.x
1157:"Chordeumatida Pocock, 1894 | COL"
995:Superfamily Heterochordeumatoidea
564:Superfamily Anthroleucosomatoidea
14:
1834:Fusco, Giuseppe (December 2005).
1352:Milli-PEET: Millipede Systematics
2749:
2748:
2738:
2716:
2711:
2468:
2054:
2042:
1852:10.1111/j.1525-142X.2005.05064.x
1746:Shear, William A. (2003-07-04).
1683:10.1111/j.1525-142X.2005.05064.x
1607:Memoirs of the Queensland Museum
1540:Shear, William A. (2009-11-16).
1213:"Diplopoda — taxonomic overview"
862:Mikhaljova & Golovatch, 2001
852:Superfamily Neoatractosomatoidea
655:(Craspedosomatidae) from Belgium
642:(Craspedosomatidae) from Germany
207:
47:
1385:Milli-PEET: The class Diplopoda
790:Superfamily Craspedosomatoidea
310:), many species have 28 (e.g.,
934:Suborder Heterochordeumatidea
1:
1624:David, Jean-Francois (1989).
1381:"Millipede Body Organization"
1043:Superfamily Pygmaeosomatoidea
966:Superfamily Diplomaragnoidea
177:c. 50 families, 1200 species
2088:North American Chordeumatida
1973:10.1371/journal.pone.0037240
556:Suborder Craspedosomatidea
530:Superfamily Chordeumatoidea
1913:Annual Review of Entomology
1840:Evolution & Development
1671:Evolution & Development
1601:Mauries, Jean-Paul (1987).
1433:Mauriès, Jean-Paul (1988).
1180:Henen, Derek; Brown, Jeff.
734:Superfamily Cleidogonoidea
307:Opisthocheiron canayerensis
3106:
2026:10.11646/zootaxa.3148.1.32
1811:Shear, William A. (2002).
1785:Shear, William A. (1992).
1513:Shear, William A. (1990).
1309:Blower, J. Gordon (1985).
916:Superfamily Verhoeffioidea
604:Superfamily Brannerioidea
322:), one genus features 29 (
2729:
2709:
2496:
2492:
2481:
2466:
2313:
2182:
2178:
2167:
2135:
2083:Chordeumatida of Tasmania
1774:– via ResearchGate.
1566:10.11646/zootaxa.2290.1.4
1519:American Museum Novitates
1492:10.11646/zootaxa.4211.1.1
1225:10.1163/9789004188273_017
942:Superfamily Conotyloidea
704:Shear & Leonard, 2003
694:Shear & Leonard, 2003
490:and, aside from southern
188:
181:
176:
169:
148:
143:
44:Scientific classification
42:
37:
28:
23:
2443:(flat-backed millipedes)
1764:10.11646/zootaxa.233.1.1
1665:Fusco, Giuseppe (2005).
1439:Revue suisse de zoologie
1101:Superfamily Striarioidea
522:Suborder Chordeumatidea
243:, a group also known as
2271:(giant pill millipedes)
1161:www.catalogueoflife.org
836:Superfamily Haaseoidea
331:Metamastigophorophyllon
2607:(worm-like millipedes)
2322:(suctorial millipedes)
2304:(worm-like millipedes)
1452:10.5962/bhl.part.79638
1077:Superfamily Caseyoidea
981:
814:
656:
643:
470:, and as far south as
362:Chamaesoma broelemanni
301:Chamaesoma broelemanni
195:Craspedosomatida
1069:Suborder Striariidea
974:
899:Mastigophorophyllidae
806:
649:
636:
190:Ascospermophora
2920:Fauna Europaea (new)
2198:(bristle-millipedes)
2124:Orders of the class
2072:Encyclopedia of Life
2051:at Wikimedia Commons
1010:Heterochordeumatidae
611:Brachychaeteumatidae
571:Anthroleucosomatidae
480:Chiloé Island, Chile
385:Neocambrisoma raveni
159:Heterochordeumatidea
1964:2012PLoSO...737240B
1361:on 26 December 2015
907:Neoatractosomatidae
827:Haplobainosomatidae
822:Gray in Jones, 1843
792:Gray in Jones, 1843
685:Macrochaeteumatidae
397:Hoffmaneuma exiguum
350:Xystrosoma beatense
337:Altajosoma kemerovo
245:spinning millipedes
192:Verhoeff, 1900
16:Order of millipedes
2005:Shear, W. (2011).
1391:on 31 October 2013
1183:Millipedes of Ohio
1026:Metopidiotrichidae
982:
815:
765:Lusitaniosomatidae
657:
644:
545:Speophilosomatidae
488:sub-Saharan Africa
379:Branneria carinata
374:Speophilosomatidae
234:sausage millipedes
3072:
3071:
3021:Open Tree of Life
2772:Taxon identifiers
2763:
2762:
2725:
2724:
2707:
2706:
2703:
2702:
2699:
2698:
2695:
2694:
2609:
2528:
2527:
2520:Microdecemplicida
2477:
2476:
2464:
2463:
2460:
2459:
2456:
2455:
2452:
2451:
2444:
2355:
2354:
2324:
2306:
2294:
2293:
2272:
2256:
2254:(pill millipedes)
2207:
2206:
2199:
2129:
2047:Media related to
1136:
1128:
1120:
1112:
1104:
1096:
1092:Urochordeumatidae
1088:
1080:
1072:
1062:
1054:
1046:
1038:
1030:
1022:
1014:
1006:
998:
990:
969:
961:
953:
945:
937:
927:
919:
911:
903:
895:
887:
869:
863:
855:
847:
839:
831:
823:
819:Craspedosomatidae
801:
793:
785:
777:
769:
761:
753:
745:
737:
729:
721:
713:
695:
689:
681:
673:
669:Heterolatzeliidae
665:
631:
623:
615:
607:
599:
591:
583:
575:
567:
559:
549:
541:
540:C. L. Koch, 1847
533:
532:C. L. Koch, 1847
525:
391:Golovatchia magda
342:sexual dimorphism
313:Lipseuma josianae
215:Arthropods portal
203:
202:
198:
197:Jeekel, 1971
193:
155:Craspedosomatidea
139:
3097:
3090:Millipede orders
3065:
3064:
3052:
3051:
3042:
3041:
3029:
3028:
3016:
3015:
3006:
3005:
2993:
2992:
2990:NBNSYS0100002381
2980:
2979:
2967:
2966:
2954:
2953:
2941:
2940:
2928:
2927:
2915:
2914:
2902:
2901:
2889:
2888:
2876:
2875:
2863:
2862:
2850:
2849:
2840:
2839:
2827:
2826:
2814:
2813:
2812:
2799:
2798:
2797:
2767:
2752:
2751:
2742:
2741:
2734:List of families
2720:
2715:
2683:
2678:
2674:
2656:
2614:
2608:
2605:
2568:
2559:
2515:Eoarthropleurida
2498:
2494:
2483:
2472:
2442:
2434:
2401:
2373:
2366:
2362:
2323:
2320:
2315:
2311:
2305:
2302:
2283:
2270:
2269:Sphaerotheriida
2255:
2252:
2247:
2238:
2197:
2184:
2180:
2169:
2127:
2118:
2111:
2104:
2095:
2079:
2059:Data related to
2058:
2046:
2030:
2029:
2011:
2002:
1996:
1995:
1985:
1975:
1943:
1937:
1936:
1908:
1902:
1901:
1885:
1879:
1878:
1876:
1874:
1831:
1825:
1824:
1808:
1802:
1801:
1791:
1782:
1776:
1775:
1743:
1737:
1736:
1726:
1717:
1711:
1710:
1662:
1656:
1655:
1645:
1643:10.5962/p.288286
1621:
1615:
1614:
1598:
1587:
1586:
1568:
1537:
1531:
1530:
1510:
1504:
1503:
1471:
1465:
1464:
1454:
1430:
1424:
1423:
1407:
1401:
1400:
1398:
1396:
1377:
1371:
1370:
1368:
1366:
1360:
1349:
1341:
1335:
1334:
1306:
1291:
1290:
1266:
1239:
1238:
1208:
1191:
1190:
1188:
1177:
1171:
1170:
1168:
1167:
1153:
1134:
1126:
1118:
1110:
1102:
1094:
1086:
1078:
1070:
1060:
1058:Pygmaeosomatidae
1052:
1044:
1036:
1028:
1020:
1012:
1004:
996:
988:
967:
959:
951:
943:
935:
925:
917:
909:
901:
893:
891:Hoffmaneumatidae
885:
879:
867:
861:
853:
845:
837:
829:
821:
799:
791:
783:
775:
773:Opisthocheiridae
767:
759:
757:Entomobielziidae
751:
743:
735:
727:
719:
711:
709:Niponiosomatidae
705:
693:
687:
679:
677:Kashmireumatidae
671:
663:
629:
621:
613:
605:
597:
589:
581:
573:
565:
557:
547:
539:
531:
523:
486:but absent from
440:hemianamorphosis
432:teloanamorphosis
415:and the species
319:Haasea hungarica
217:
212:
211:
196:
191:
134:
52:
51:
33:
21:
3105:
3104:
3100:
3099:
3098:
3096:
3095:
3094:
3075:
3074:
3073:
3068:
3060:
3055:
3047:
3045:
3037:
3032:
3024:
3019:
3011:
3009:
3001:
2996:
2988:
2983:
2975:
2970:
2962:
2957:
2949:
2944:
2936:
2931:
2923:
2918:
2910:
2905:
2897:
2892:
2884:
2879:
2871:
2866:
2858:
2853:
2845:
2843:
2835:
2830:
2822:
2817:
2808:
2807:
2802:
2793:
2792:
2787:
2774:
2764:
2759:
2721:
2691:
2681:
2672:
2667:
2650:
2645:
2606:
2604:
2603:Helminthomorpha
2597:
2579:
2575:Zosterogrammida
2562:
2553:
2551:
2549:
2547:
2545:
2543:
2541:
2539:
2537:
2535:
2524:
2502:Arthropleuridea
2488:
2473:
2448:
2432:
2427:
2399:
2394:
2369:
2360:
2351:
2342:Siphonocryptida
2321:
2303:
2301:
2300:Helminthomorpha
2290:
2281:
2276:
2253:
2232:
2230:
2228:
2226:
2224:
2222:
2220:
2218:
2216:
2214:
2203:
2174:
2163:
2131:
2122:
2090:on BugGuide.net
2067:"Chordeumatida"
2039:
2034:
2033:
2009:
2004:
2003:
1999:
1945:
1944:
1940:
1910:
1909:
1905:
1887:
1886:
1882:
1872:
1870:
1833:
1832:
1828:
1810:
1809:
1805:
1794:Myriapodologica
1789:
1784:
1783:
1779:
1745:
1744:
1740:
1729:Myriapodologica
1724:
1719:
1718:
1714:
1664:
1663:
1659:
1623:
1622:
1618:
1600:
1599:
1590:
1539:
1538:
1534:
1512:
1511:
1507:
1473:
1472:
1468:
1432:
1431:
1427:
1409:
1408:
1404:
1394:
1392:
1379:
1378:
1374:
1364:
1362:
1358:
1347:
1343:
1342:
1338:
1323:
1308:
1307:
1294:
1268:
1267:
1242:
1235:
1210:
1209:
1194:
1186:
1179:
1178:
1174:
1165:
1163:
1155:
1154:
1150:
1145:
1127:Silvestri, 1909
1124:Rhiscosomididae
1095:Silvestri, 1909
1034:Peterjohnsiidae
1021:Golovatch, 1978
986:Diplomaragnidae
894:Golovatch, 1978
877:
870:
866:Cyrnosomatidae
781:Trichopetalidae
703:
696:
692:Microlympiidae
627:Chamaesomatidae
595:Vandeleumatidae
587:Origmatogonidae
508:
460:Central America
452:
427:
354:Peterjohnsiidae
257:
213:
206:
194:
161:
157:
153:
133:
46:
17:
12:
11:
5:
3103:
3101:
3093:
3092:
3087:
3077:
3076:
3070:
3069:
3067:
3066:
3053:
3043:
3030:
3017:
3007:
2994:
2981:
2968:
2955:
2942:
2929:
2916:
2907:Fauna Europaea
2903:
2890:
2877:
2864:
2851:
2841:
2828:
2815:
2800:
2784:
2782:
2776:
2775:
2770:
2761:
2760:
2758:
2757:
2746:
2736:
2730:
2727:
2726:
2723:
2722:
2710:
2708:
2705:
2704:
2701:
2700:
2697:
2696:
2693:
2692:
2686:
2684:
2675:
2669:
2668:
2666:
2665:
2659:
2657:
2653:incertae sedis
2647:
2646:
2644:
2643:
2641:Palaeosomatida
2638:
2633:
2628:
2622:
2620:
2611:
2599:
2598:
2596:
2595:
2593:Amynilyspedida
2589:
2587:
2581:
2580:
2578:
2577:
2571:
2569:
2565:incertae sedis
2556:
2530:
2529:
2526:
2525:
2523:
2522:
2517:
2512:
2510:Arthropleurida
2506:
2504:
2490:
2489:
2486:
2479:
2478:
2475:
2474:
2467:
2465:
2462:
2461:
2458:
2457:
2454:
2453:
2450:
2449:
2447:
2446:
2437:
2435:
2429:
2428:
2426:
2425:
2420:
2415:
2410:
2404:
2402:
2396:
2395:
2393:
2392:
2390:Spirostreptida
2387:
2382:
2376:
2374:
2363:
2357:
2356:
2353:
2352:
2350:
2349:
2347:Siphonophorida
2344:
2339:
2334:
2328:
2326:
2308:
2296:
2295:
2292:
2291:
2288:Glomeridesmida
2286:
2284:
2278:
2277:
2275:
2274:
2266:
2260:
2258:
2244:
2235:
2209:
2208:
2205:
2204:
2202:
2201:
2192:
2190:
2176:
2175:
2172:
2165:
2164:
2162:
2161:
2155:
2149:
2143:
2136:
2133:
2132:
2123:
2121:
2120:
2113:
2106:
2098:
2092:
2091:
2085:
2080:
2064:
2063:at Wikispecies
2052:
2038:
2037:External links
2035:
2032:
2031:
1997:
1938:
1919:(1): 401–420.
1903:
1880:
1846:(6): 608–617.
1826:
1803:
1777:
1738:
1712:
1677:(6): 608–617.
1657:
1636:(3): 639–647.
1616:
1588:
1548:
1545:
1532:
1505:
1466:
1425:
1402:
1372:
1336:
1321:
1292:
1281:(2): 103–234.
1240:
1233:
1192:
1172:
1147:
1146:
1144:
1141:
1140:
1139:
1138:
1137:
1129:
1121:
1113:
1099:
1098:
1097:
1089:
1087:Verhoeff, 1909
1079:Verhoeff, 1909
1074:
1073:
1066:
1065:
1064:
1063:
1055:
1050:Lankasomatidae
1041:
1040:
1039:
1031:
1023:
1015:
1007:
1005:Verhoeff, 1914
993:
992:
991:
964:
963:
962:
954:
939:
938:
931:
930:
929:
928:
926:Verhoeff, 1899
918:Verhoeff, 1899
914:
913:
912:
910:Verhoeff, 1901
904:
902:Verhoeff, 1899
896:
888:
880:
864:
854:Verhoeff, 1901
850:
849:
848:
834:
833:
832:
830:Verhoeff, 1909
824:
816:
800:Verhoeff, 1899
788:
787:
786:
784:Verhoeff, 1914
778:
770:
768:Schubart, 1953
762:
760:Verhoeff, 1899
754:
746:
741:Biokoviellidae
732:
731:
730:
722:
714:
712:Verhoeff, 1941
706:
690:
688:Verhoeff, 1914
682:
674:
666:
661:Golovatchiidae
658:
630:Verhoeff, 1913
624:
616:
614:Verhoeff, 1910
602:
601:
600:
598:Mauriès, 1970
592:
584:
582:Hoffman, 1980
576:
574:Verhoeff 1899
561:
560:
553:
552:
551:
550:
548:Takakuwa, 1949
542:
537:Chordeumatidae
527:
526:
507:
506:Classification
504:
464:Southeast Asia
451:
448:
436:euanamorphosis
426:
423:
418:Vieteuma longi
402:Chordeumatidae
400:). The family
281:helminthomorph
256:
253:
247:because their
219:
218:
201:
200:
186:
185:
179:
178:
174:
173:
167:
166:
151:Chordeumatidea
146:
145:
141:
140:
128:
124:
123:
118:
114:
113:
108:
104:
103:
98:
94:
93:
88:
84:
83:
78:
74:
73:
68:
64:
63:
58:
54:
53:
40:
39:
35:
34:
26:
25:
24:Chordeumatida
15:
13:
10:
9:
6:
4:
3:
2:
3102:
3091:
3088:
3086:
3085:Chordeumatida
3083:
3082:
3080:
3063:
3058:
3054:
3050:
3044:
3040:
3035:
3031:
3027:
3022:
3018:
3014:
3008:
3004:
2999:
2995:
2991:
2986:
2982:
2978:
2973:
2969:
2965:
2960:
2956:
2952:
2947:
2943:
2939:
2934:
2930:
2926:
2921:
2917:
2913:
2908:
2904:
2900:
2895:
2891:
2887:
2882:
2878:
2874:
2869:
2865:
2861:
2856:
2852:
2848:
2842:
2838:
2837:Chordeumatida
2833:
2829:
2825:
2824:Chordeumatida
2820:
2816:
2811:
2810:Chordeumatida
2805:
2801:
2796:
2790:
2786:
2785:
2783:
2781:
2780:Chordeumatida
2777:
2773:
2768:
2756:
2747:
2745:
2737:
2735:
2732:
2731:
2728:
2719:
2714:
2690:(superfamily)
2689:
2685:
2679:
2676:
2670:
2664:
2661:
2660:
2658:
2655:
2654:
2648:
2642:
2639:
2637:
2634:
2632:
2629:
2627:
2624:
2623:
2621:
2619:
2618:Archipolypoda
2615:
2612:
2610:
2600:
2594:
2591:
2590:
2588:
2586:
2582:
2576:
2573:
2572:
2570:
2567:
2566:
2560:
2557:
2555:
2531:
2521:
2518:
2516:
2513:
2511:
2508:
2507:
2505:
2503:
2499:
2495:
2491:
2484:
2480:
2471:
2445:
2439:
2438:
2436:
2430:
2424:
2421:
2419:
2416:
2414:
2413:Chordeumatida
2411:
2409:
2406:
2405:
2403:
2397:
2391:
2388:
2386:
2383:
2381:
2378:
2377:
2375:
2372:
2367:
2364:
2358:
2348:
2345:
2343:
2340:
2338:
2335:
2333:
2330:
2329:
2327:
2325:
2319:Colobognatha
2316:
2312:
2309:
2307:
2297:
2289:
2285:
2279:
2273:
2267:
2265:
2262:
2261:
2259:
2257:
2251:Oniscomorpha
2248:
2245:
2243:
2239:
2236:
2234:
2210:
2200:
2194:
2193:
2191:
2189:
2185:
2181:
2177:
2170:
2166:
2160:
2156:
2154:
2150:
2148:
2144:
2142:
2138:
2137:
2134:
2130:
2119:
2114:
2112:
2107:
2105:
2100:
2099:
2096:
2089:
2086:
2084:
2081:
2078:
2074:
2073:
2068:
2065:
2062:
2061:Chordeumatida
2057:
2053:
2050:
2049:Chordeumatida
2045:
2041:
2040:
2036:
2027:
2023:
2019:
2015:
2008:
2001:
1998:
1993:
1989:
1984:
1979:
1974:
1969:
1965:
1961:
1958:(5): e37240.
1957:
1953:
1949:
1942:
1939:
1934:
1930:
1926:
1922:
1918:
1914:
1907:
1904:
1899:
1895:
1894:Insecta Mundi
1891:
1884:
1881:
1869:
1865:
1861:
1857:
1853:
1849:
1845:
1841:
1837:
1830:
1827:
1822:
1818:
1814:
1807:
1804:
1799:
1795:
1788:
1781:
1778:
1773:
1769:
1765:
1761:
1757:
1753:
1749:
1742:
1739:
1734:
1730:
1723:
1716:
1713:
1708:
1704:
1700:
1696:
1692:
1688:
1684:
1680:
1676:
1672:
1668:
1661:
1658:
1653:
1649:
1644:
1639:
1635:
1632:(in French).
1631:
1627:
1620:
1617:
1612:
1608:
1604:
1597:
1595:
1593:
1589:
1584:
1580:
1576:
1572:
1567:
1562:
1558:
1554:
1550:
1546:
1543:
1536:
1533:
1528:
1524:
1520:
1516:
1509:
1506:
1501:
1497:
1493:
1489:
1485:
1481:
1477:
1470:
1467:
1462:
1458:
1453:
1448:
1444:
1441:(in French).
1440:
1436:
1429:
1426:
1421:
1417:
1413:
1406:
1403:
1390:
1386:
1382:
1376:
1373:
1357:
1353:
1346:
1340:
1337:
1332:
1328:
1324:
1322:90-04-07698-0
1318:
1314:
1313:
1305:
1303:
1301:
1299:
1297:
1293:
1288:
1284:
1280:
1276:
1272:
1265:
1263:
1261:
1259:
1257:
1255:
1253:
1251:
1249:
1247:
1245:
1241:
1236:
1234:9789004156128
1230:
1226:
1222:
1218:
1214:
1207:
1205:
1203:
1201:
1199:
1197:
1193:
1185:
1184:
1176:
1173:
1162:
1158:
1152:
1149:
1142:
1135:Bollman, 1893
1133:
1130:
1125:
1122:
1117:
1114:
1109:
1106:
1105:
1103:Bollman, 1893
1100:
1093:
1090:
1085:
1082:
1081:
1076:
1075:
1068:
1067:
1059:
1056:
1051:
1048:
1047:
1042:
1037:Mauriès, 1987
1035:
1032:
1027:
1024:
1019:
1018:Megalotylidae
1016:
1011:
1008:
1003:
1000:
999:
994:
987:
984:
983:
979:
978:
977:Schedotrigona
973:
968:Attems, 1907
965:
958:
955:
950:
947:
946:
941:
940:
933:
932:
924:
923:Verhoeffiidae
921:
920:
915:
908:
905:
900:
897:
892:
889:
884:
881:
878:Mauriès, 1969
876:
875:
868:Mauriès, 2003
865:
860:
857:
856:
851:
844:
841:
840:
835:
828:
825:
820:
817:
812:
811:
805:
798:
795:
794:
789:
782:
779:
774:
771:
766:
763:
758:
755:
750:
749:Cleidogonidae
747:
742:
739:
738:
733:
726:
725:Trachygonidae
723:
720:Loomis, 1966
718:
715:
710:
707:
702:
701:
691:
686:
683:
680:Mauriès, 1982
678:
675:
672:Verhoeff 1899
670:
667:
662:
659:
654:
653:
648:
641:
640:
635:
628:
625:
620:
617:
612:
609:
608:
603:
596:
593:
590:Verhoeff 1914
588:
585:
580:
577:
572:
569:
568:
566:Verhoeff 1899
563:
562:
555:
554:
546:
543:
538:
535:
534:
529:
528:
521:
520:
519:
517:
513:
505:
503:
501:
497:
493:
489:
485:
481:
477:
473:
469:
465:
461:
457:
449:
447:
443:
441:
437:
433:
424:
422:
420:
419:
414:
413:
408:
403:
399:
398:
393:
392:
388:
386:
381:
380:
375:
369:
367:
363:
357:
355:
351:
347:
343:
339:
338:
333:
332:
327:
326:
321:
320:
315:
314:
309:
308:
303:
302:
296:
294:
288:
286:
282:
278:
274:
270:
266:
261:
254:
252:
250:
246:
242:
239:
235:
231:
227:
223:
222:Chordeumatida
216:
210:
205:
199:
187:
184:
180:
175:
172:
168:
165:
164:
160:
156:
152:
147:
142:
137:
132:
131:Chordeumatida
129:
126:
125:
122:
119:
116:
115:
112:
109:
107:Subterclass:
106:
105:
102:
99:
96:
95:
92:
89:
86:
85:
82:
79:
76:
75:
72:
69:
66:
65:
62:
59:
56:
55:
50:
45:
41:
36:
32:
27:
22:
19:
2779:
2688:Xyloiuloidea
2663:Pleurojulida
2651:
2636:Euphoberiida
2631:Cowiedesmida
2626:Archidesmida
2563:
2441:Polydesmida
2418:Siphoniulida
2412:
2332:Platydesmida
2282:Limacomorpha
2145:Superphylum
2128:(millipedes)
2070:
2017:
2013:
2000:
1955:
1951:
1941:
1916:
1912:
1906:
1897:
1893:
1883:
1871:. Retrieved
1843:
1839:
1829:
1820:
1816:
1806:
1797:
1793:
1780:
1755:
1751:
1741:
1732:
1728:
1715:
1674:
1670:
1660:
1633:
1629:
1619:
1610:
1606:
1559:(1): 41–49.
1556:
1552:
1535:
1518:
1508:
1486:(1): 1–205.
1483:
1479:
1469:
1442:
1438:
1428:
1419:
1415:
1405:
1393:. Retrieved
1389:the original
1384:
1375:
1363:. Retrieved
1356:the original
1351:
1339:
1311:
1278:
1274:
1216:
1182:
1175:
1164:. Retrieved
1160:
1151:
1111:Loomis, 1966
1108:Apterouridae
1053:Mauriès 1978
1029:Attems, 1907
1013:Pocock, 1894
997:Pocock, 1894
989:Attems, 1907
975:
886:Attems, 1926
872:
859:Altajellidae
846:Attems, 1899
838:Attems, 1899
808:
776:Ribaut, 1913
698:
652:Craspedosoma
650:
637:
619:Branneriidae
524:Pocock 1894
509:
453:
450:Distribution
444:
428:
416:
410:
395:
389:
383:
377:
370:
361:
358:
349:
335:
329:
323:
317:
311:
305:
299:
297:
289:
262:
258:
244:
233:
221:
220:
189:
149:
130:
117:Superorder:
18:
2946:iNaturalist
2804:Wikispecies
2423:Stemmiulida
2408:Callipodida
2400:Nematophora
2385:Spirobolida
2337:Polyzoniida
2196:Polyxenida
2188:Penicillata
2020:: 159–164.
1800:(4): 25–36.
1735:(4): 25–36.
1422:(1): 75–92.
1219:: 363–453.
1132:Striariidae
1119:Shear, 2009
1002:Eudigonidae
957:Conotylidae
952:Shear, 1971
949:Adritylidae
936:Shear, 2000
871:monotypic:
797:Attemsiidae
744:Mrsic, 1992
700:Microlympia
697:monotypic:
664:Shear, 1992
639:Atractosoma
496:Scandinavia
476:New Zealand
425:Development
412:Kashmireuma
277:Polydesmida
255:Description
241:Nematophora
163:Striariidea
138:, 1894
121:Nematophora
87:Subphylum:
3079:Categories
2755:Millipedes
2682:Juliformia
2585:Pentazonia
2371:Juliformia
2242:Pentazonia
2157:Subphylum
2153:Arthropoda
2126:Diplopoda
1758:(1): 1–7.
1547:carolinus
1416:Zoosystema
1166:2024-07-04
1143:References
1071:Cook, 1896
1061:Carl, 1941
1045:Carl, 1941
960:Cook, 1896
944:Cook, 1896
807:A pair of
752:Cook, 1896
736:Cook, 1896
728:Cook, 1896
717:Tingupidae
622:Cook, 1896
606:Cook, 1896
558:Cook, 1895
484:Madagascar
238:superorder
230:millipedes
144:Suborders
81:Arthropoda
2744:Diplopoda
2433:Merocheta
2264:Glomerida
2159:Myriapoda
2147:Ecdysozoa
1873:25 August
1772:1175-5334
1691:1520-541X
1652:132547501
1575:1175-5334
1527:2246/5072
1500:1175-5334
1461:0035-418X
1084:Caseyidae
883:Faginidae
874:Cyrnosoma
843:Haaseidae
579:Haasiidae
366:Diplopoda
171:Diversity
101:Diplopoda
91:Myriapoda
67:Kingdom:
61:Eukaryota
2868:BugGuide
2844:BioLib:
2795:Q4300743
2789:Wikidata
2673:Eugnatha
2361:Eugnatha
2141:Animalia
2139:Kingdom
1992:22615951
1952:PLOS ONE
1933:17163800
1900:: 1–134.
1868:21401688
1860:16336414
1707:21401688
1699:16336414
1521:(2977).
1445:: 3–49.
1395:18 March
1365:20 March
1331:13439686
1116:Buotidae
512:families
472:Tasmania
407:sternite
346:Buotidae
325:Tianella
293:gonopods
285:ozopores
273:paranota
183:Synonyms
111:Eugnatha
77:Phylum:
71:Animalia
57:Domain:
2487:Extinct
2151:Phylum
2069:at the
2014:Zootaxa
1983:3352885
1960:Bibcode
1752:Zootaxa
1583:4833397
1553:Zootaxa
1480:Zootaxa
500:Siberia
468:Oceania
456:tropics
394:, and
249:telsons
127:Order:
97:Class:
3062:887894
3049:582683
3046:uBio:
3010:NZOR:
3003:118457
2977:154417
2860:259567
2753:
2380:Julida
2173:Living
1990:
1980:
1931:
1866:
1858:
1770:
1705:
1697:
1689:
1650:
1581:
1573:
1544:Buotus
1498:
1459:
1329:
1319:
1231:
810:Haasea
516:genera
478:, and
466:, and
269:collum
136:Pocock
3057:WoRMS
3034:Plazi
3026:77633
2964:11642
2959:IRMNG
2951:50425
2912:11526
2886:7NF8X
2873:97644
2847:19684
2010:(PDF)
1864:S2CID
1790:(PDF)
1725:(PDF)
1703:S2CID
1648:S2CID
1579:S2CID
1359:(PDF)
1348:(PDF)
1187:(PDF)
492:Chile
265:setae
226:order
2998:NCBI
2972:ITIS
2938:1017
2933:GBIF
2899:6397
2855:BOLD
2018:3148
1988:PMID
1929:PMID
1875:2020
1856:PMID
1768:ISSN
1695:PMID
1687:ISSN
1634:4:11
1571:ISSN
1557:2290
1496:ISSN
1484:4211
1457:ISSN
1397:2014
1367:2014
1327:OCLC
1317:ISBN
1229:ISBN
316:and
304:and
2985:NBN
2894:EoL
2881:CoL
2832:AFD
2819:ADW
2022:doi
1978:PMC
1968:doi
1921:doi
1898:158
1848:doi
1760:doi
1756:233
1679:doi
1638:doi
1561:doi
1523:hdl
1488:doi
1447:doi
1283:doi
1279:109
1221:doi
458:of
438:or
430:as
228:of
3081::
3059::
3036::
3023::
3000::
2987::
2974::
2961::
2948::
2935::
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2896::
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