Knowledge (XXG)

Chordeumatida

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634: 2077: 804: 2718: 2713: 49: 31: 647: 2750: 209: 2470: 2056: 972: 364:, with only 26 segments (four fewer than the typical number), adult females have only 42 leg pairs, and adult males have only 40 pairs of walking legs (excluding two pairs of gonopods). Adult females with 32 segments (two more than the typical number) have 54 leg pairs (e.g., in the family Peterjohnsiidae), which is the maximum number fixed by species in the class 2740: 2044: 446:
ninth and final stage with 30 segments. The male usually begins to develop gonopods in the seventh stage. Species that produce adults with fewer or more segments than the usual number, however, deviate from the typical pattern by adding a different number of segments at some stage, reaching maturity in a different stage, or both.
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Millipedes in this order grow and develop through a series of molts, adding segments until they reach a fixed number in the adult stage, which is usually the same for a given sex in a given species, at which point the molting and the addition of segments and legs stop. This mode of development, known
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Chordeumatidans take on a wide variety of forms, including some that are cylindrical and others that are flat-backed. Most species have 26 to 32 body segments (including the telson) behind the head, with the number usually fixed within species. These millipedes range in length from 3.5 mm to 42 mm,
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For the typical species in this order, post-embryonic development takes place in nine stages. The juvenile millipede hatches with 6 segments and only 3 pairs of legs in the first stage, then usually goes through stages with 8, 11, 15, 19, 23, 25, and 28 segments, before emerging as adults in the
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exhibits the most extensive modifications, including five leg pairs (pairs 7 through 11) in the gonopod complex. Some species also deviate from the usual body plan by reducing or eliminating leg pairs in the adult female. In the family Chordeumatidae, for example, adult females feature a legless
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Many species deviate from the expected number of walking legs, however, because they deviate in terms of sex-linked modifications to their legs. For example, many species involve another leg pair in addition to pairs 8 and 9 in the gonopod complex in adult males. In the family
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With these deviations from the usual 30 segments, the number of leg pairs in adults changes, usually with two leg pairs added or subtracted for each segment added to or subtracted from the typical number. For example, in
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Most chordeumatidan species have 30 body segments (including the telson) as adults, and adult females in these species have 50 leg pairs. In adult males in this order, two leg pairs (pair 8 and pair 9) are modified into
1890:"Atlas of myriapod biogeography. I. Indigenous ordinal and supra-ordinal distributions in the Diplopoda: Perspectives on taxon origins and ages, and a hypothesis on the origin and early evolution of the class" 1315:. Linnean Society of London, Estuarine and Brackish-water Sciences Association. London: Published for the Linnean Society of London and the Estuarine and Brackish-Water Sciences Association by E.J. Brill. 271:) is relatively narrow, giving the appearance of a distinct "neck" in many species, and the body tapers towards the rear. A dorsal groove runs down the length of the body, and some species feature 376:, leg pair 7 in adult males is modified as part of the gonopod complex. In many species, the gonopod complex instead includes leg pair 10 in addition to pairs 8 and 9 (e.g., 2007:"Class Diplopoda de Blainville in Gervais, 1844. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness" 260:
although most species are 10 mm to 25 mm long. They are usually drab in color, ranging from various shades of brown to unpigmented, but some feature distinct patterns.
2958: 2997: 1411: 434:, distinguishes this order from most other orders of millipedes. In most other orders, millipedes continue to molt as adults, developing through either 1787:"Golovatchia, new genus, and Golovatchiidae, from the Far East of the Russian Republic, with a comment on Hoffmaneumatidae (Diplopoda: Chordeumatidae)" 1344: 295:, leaving 48 pairs of walking legs in the typical adult male chordeumatidan. Many species in this order, however, deviate from this typical body plan. 1786: 1603:"Craspedosomid Millipedes Discovered in Australia: Reginaterreuma, Neocambrisoma and Peterjohnsia, New Genera (Myriapoda: Diplopoda: Craspedosomida)" 2932: 1721: 1380: 2971: 263:
Species in this order share a set of features that distinguishes them from other millipedes. A key feature is the presence of six large bristles (
2082: 494:, are largely absent from South America. They are abundant in cold, rocky, mountainous areas of Europe and central Asia, and range northward to 251:
feature spinnerets used to build nests of silk. These millipedes produce this silk to create chambers in which to molt or to lay their eggs.
1748:"Branneria bonoculus, n. sp., a second species in the North American milliped family Branneriidae (Diplopoda: Chordeumatida: Brannerioidea)" 2115: 1435:"Myriapodes du Népal. II. Diplopodes Craspedosomides nouveaux de l'Himalaya et de la région indo-malaise (Craspedosomidea et Chordeumidea)" 1911:
Sierwald, Petra; Bond, Jason E. (2007). "Current Status of the Myriapod Class Diplopoda (Millipedes): Taxonomic Diversity and Phylogeny".
1476:"The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" 1320: 1232: 1626:"Le cycle biologique de Chamaesoma broelemanni Ribaut et Verhoeff, 1913 (Diplopoda, Craspedosomatida) en forêt d'Orléans (France)" 1181: 3056: 1271:"Anamorphosis in Millipedes (Diplopoda) — The Present State of Knowledge with Some Developmental and Phylogenetic Considerations" 1270: 2060: 1948:"Millipede Taxonomy after 250 Years: Classification and Taxonomic Practices in a Mega-Diverse yet Understudied Arthropod Group" 2976: 2717: 454:
Chordeumatidans have a wide distribution, occurring on all continents except Antarctica. These millipedes are found in the
2984: 2712: 1813:"Five New Chordeumatidan Millipeds from China: New Species of Vieteuma (Kashmirieumatidae) and Nepalella (Megalotylidae)" 1722:"Japanese chordeumatid millipeds. I. On the genus Speophilosoma Takakuwa (Diplopoda, Chordeumatida, Speophilosomatidae)" 48: 2854: 2831: 306: 344:
in segment number, specifically, adult males with two segments fewer than adult females, for example, in the family
3002: 2911: 2733: 3089: 2924: 298:
Many chordeumatidan species deviate from the usual 30 segments: A few species have 26 segments as adults (e.g.,
2108: 2048: 421:), adult females instead exhibit modifications to the second pair of legs, which are reduced to small nubbins. 633: 1515:"On the Central and East Asian milliped family Diplomaragnidae (Diplopoda, Choredeumatida, Diplomaragnoidea)" 330: 1835: 3084: 2771: 1212: 439: 435: 431: 2859: 1388: 898: 135: 2743: 1666: 1355: 409:(the "platosternite") where a third pair of legs would otherwise be. In other species (e.g., the genus 3038: 1625: 1602: 3061: 3012: 2893: 2818: 2071: 1959: 1812: 1009: 610: 570: 510:
Chordeumatida contains approximately 1200 species, classified in four suborders and approximately 50
384: 158: 803: 2754: 2101: 976: 906: 826: 684: 275:, lateral extensions of the exoskeleton. Paranota are also found in some other millipedes, notably 268: 2469: 2809: 1863: 1702: 1647: 1578: 1025: 764: 699: 638: 544: 487: 373: 240: 182: 120: 43: 873: 279:, from which Chordeumatidans can be distinguished by having a dorsal groove. Unlike most other 3020: 2880: 1987: 1928: 1855: 1767: 1694: 1686: 1570: 1495: 1456: 1326: 1316: 1228: 1211:
Enghoff, Henrik; Golovatch, Sergei; Short, Megan; Stoev, Pavel; Wesener, Thomas (2015-01-01).
1091: 818: 668: 341: 214: 154: 110: 2093: 3048: 2963: 2574: 2021: 1977: 1967: 1920: 1847: 1759: 1678: 1637: 1560: 1522: 1487: 1446: 1282: 1220: 1057: 890: 772: 756: 708: 676: 511: 479: 417: 411: 396: 318: 3025: 30: 2602: 2501: 2341: 2299: 2268: 1924: 1123: 1033: 985: 780: 626: 594: 586: 459: 353: 336: 237: 225: 2076: 2006: 1963: 2919: 2906: 2652: 2592: 2564: 2519: 2514: 2389: 2346: 2287: 2250: 1982: 1947: 1286: 1049: 740: 660: 536: 463: 401: 390: 280: 150: 2823: 646: 3078: 2836: 2662: 2635: 2617: 1851: 1747: 1682: 1651: 1475: 1017: 922: 748: 724: 267:) on the dorsal surface of each body segment, three on each side. The first segment ( 1867: 1706: 2885: 2687: 2625: 2509: 2422: 2331: 2318: 1582: 1514: 1156: 1107: 858: 651: 618: 208: 170: 1889: 1972: 1434: 2989: 2945: 2803: 2630: 2533: 2440: 2417: 2407: 2384: 2336: 2212: 1131: 1001: 956: 948: 796: 495: 475: 276: 162: 2794: 2055: 2025: 2640: 2584: 2370: 2241: 2195: 2187: 2087: 1565: 1541: 1491: 1224: 716: 483: 300: 204: 1771: 1763: 1690: 1574: 1499: 1460: 236:, they are found nearly worldwide. Chordeumatida is the largest order in the 2263: 2158: 2152: 2146: 2125: 1330: 1083: 882: 842: 578: 378: 365: 229: 100: 90: 80: 60: 1991: 1932: 1859: 1698: 1451: 971: 2043: 2867: 2846: 2788: 1410:
Golovatch, Sergei I.; Geoffroy, Jean-Jacques; Mauries, Jean-Paul (2006).
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Treatise on Zoology - Anatomy, Taxonomy, Biology. The Myriapoda, Volume 2
1115: 471: 406: 345: 324: 312: 272: 2872: 499: 467: 455: 292: 284: 1312:
Millipedes : keys and notes for the identification of the species
2950: 2379: 2140: 1642: 1526: 809: 248: 70: 2937: 2765: 3033: 1354:. The Field Museum, Chicago, IL. 26 September 2006. Archived from 970: 802: 645: 632: 515: 491: 264: 2066: 1836:"Trunk segment numbers and sequential segmentation in myriapods" 1667:"Trunk segment numbers and sequential segmentation in myriapods" 2769: 2485: 2171: 2097: 1542:"Buotidae, a new family for the minute North American milliped 352:(males with 28, females with the usual 30), and in the family 2898: 1720:
Shear, William A.; Tsurusaki, Nobuo; Tanabe, Tsutomu (1994).
1946:
Brewer, Michael S.; Sierwald, Petra; Bond, Jason E. (2012).
1387:. The Field Museum, Chicago. 9 February 2012. Archived from 1269:
Enghoff, Henrik; Dohle, Wolfgang; Blower, J. Gordon (1993).
502:, and in North America up into Canada and southwest Alaska. 1630:
Bulletin du Muséum National d'Histoire Naturelle, Section A
1549:(Chamberlin) 1940 (Diplopoda, Chordeumatida, Striarioidea)" 328:, in which most species have 29), one genus features 31 ( 1412:"Four new Chordeumatida (Diplopoda) from caves in China" 1613:(1): 107–133 – via Biodiversity Heritage Library. 813:(Haaseidae) from Germany. The dorsal groove is visible 1823:(6): 63–72 – via Biodiversity Heritage Library. 514:, although several families contain only one to five 1474:
Antić, Dragan Ž; Makarov, Slobodan E. (2016-12-22).
2778: 2680: 2671: 2649: 2616: 2601: 2583: 2561: 2532: 2500: 2431: 2398: 2368: 2359: 2317: 2298: 2280: 2249: 2240: 2211: 2186: 232:containing more than 1,400 species. Also known as 1817:Proceedings of the California Academy of Sciences 224:(from the Greek word for "sausage") is a large 1888:Shelley, R. M. & Golavatch, S. I. (2011). 2109: 283:(worm-like) millipedes, chordeumatidans lack 8: 1189:. Ohio Division of Wildlife. pp. 3, 24. 356:(males with the usual 30, females with 32). 2766: 2677: 2613: 2558: 2497: 2493: 2482: 2365: 2314: 2310: 2246: 2237: 2183: 2179: 2168: 2116: 2102: 2094: 2075: 1654:– via Biodiversity Heritage Library. 1463:– via Biodiversity Heritage Library. 1345:"Putative apomorphies of millipede clades" 340:). Some species also deviate by featuring 38:Two unidentified species from New Zealand 29: 20: 1981: 1971: 1641: 1564: 1450: 1275:Zoological Journal of the Linnean Society 1148: 482:. Species in this order are present in 1925:10.1146/annurev.ento.52.111805.090210 1596: 1594: 1592: 1264: 980:(Metopidiotrichidae) from New Zealand 348:(males with 26, females with 28), in 7: 3039:2578E112-FFB1-FE20-FF5B-97F9FD70FA7F 3013:0361b0fd-9d56-407e-adaf-aa90525f2a95 2925:3b6a2a08-359d-4225-a20c-d1f1a454e417 2739: 1304: 1302: 1300: 1298: 1296: 1262: 1260: 1258: 1256: 1254: 1252: 1250: 1248: 1246: 1244: 1206: 1204: 1202: 1200: 1198: 1196: 334:), and many species have 32 (e.g., 1287:10.1111/j.1096-3642.1993.tb00305.x 1157:"Chordeumatida Pocock, 1894 | COL" 995:Superfamily Heterochordeumatoidea 564:Superfamily Anthroleucosomatoidea 14: 1834:Fusco, Giuseppe (December 2005). 1352:Milli-PEET: Millipede Systematics 2749: 2748: 2738: 2716: 2711: 2468: 2054: 2042: 1852:10.1111/j.1525-142X.2005.05064.x 1746:Shear, William A. (2003-07-04). 1683:10.1111/j.1525-142X.2005.05064.x 1607:Memoirs of the Queensland Museum 1540:Shear, William A. (2009-11-16). 1213:"Diplopoda — taxonomic overview" 862:Mikhaljova & Golovatch, 2001 852:Superfamily Neoatractosomatoidea 655:(Craspedosomatidae) from Belgium 642:(Craspedosomatidae) from Germany 207: 47: 1385:Milli-PEET: The class Diplopoda 790:Superfamily Craspedosomatoidea 310:), many species have 28 (e.g., 934:Suborder Heterochordeumatidea 1: 1624:David, Jean-Francois (1989). 1381:"Millipede Body Organization" 1043:Superfamily Pygmaeosomatoidea 966:Superfamily Diplomaragnoidea 177:c. 50 families, 1200 species 2088:North American Chordeumatida 1973:10.1371/journal.pone.0037240 556:Suborder Craspedosomatidea 530:Superfamily Chordeumatoidea 1913:Annual Review of Entomology 1840:Evolution & Development 1671:Evolution & Development 1601:Mauries, Jean-Paul (1987). 1433:Mauriès, Jean-Paul (1988). 1180:Henen, Derek; Brown, Jeff. 734:Superfamily Cleidogonoidea 307:Opisthocheiron canayerensis 3106: 2026:10.11646/zootaxa.3148.1.32 1811:Shear, William A. (2002). 1785:Shear, William A. (1992). 1513:Shear, William A. (1990). 1309:Blower, J. Gordon (1985). 916:Superfamily Verhoeffioidea 604:Superfamily Brannerioidea 322:), one genus features 29 ( 2729: 2709: 2496: 2492: 2481: 2466: 2313: 2182: 2178: 2167: 2135: 2083:Chordeumatida of Tasmania 1774:– via ResearchGate. 1566:10.11646/zootaxa.2290.1.4 1519:American Museum Novitates 1492:10.11646/zootaxa.4211.1.1 1225:10.1163/9789004188273_017 942:Superfamily Conotyloidea 704:Shear & Leonard, 2003 694:Shear & Leonard, 2003 490:and, aside from southern 188: 181: 176: 169: 148: 143: 44:Scientific classification 42: 37: 28: 23: 2443:(flat-backed millipedes) 1764:10.11646/zootaxa.233.1.1 1665:Fusco, Giuseppe (2005). 1439:Revue suisse de zoologie 1101:Superfamily Striarioidea 522:Suborder Chordeumatidea 243:, a group also known as 2271:(giant pill millipedes) 1161:www.catalogueoflife.org 836:Superfamily Haaseoidea 331:Metamastigophorophyllon 2607:(worm-like millipedes) 2322:(suctorial millipedes) 2304:(worm-like millipedes) 1452:10.5962/bhl.part.79638 1077:Superfamily Caseyoidea 981: 814: 656: 643: 470:, and as far south as 362:Chamaesoma broelemanni 301:Chamaesoma broelemanni 195:Craspedosomatida  1069:Suborder Striariidea 974: 899:Mastigophorophyllidae 806: 649: 636: 190:Ascospermophora  2920:Fauna Europaea (new) 2198:(bristle-millipedes) 2124:Orders of the class 2072:Encyclopedia of Life 2051:at Wikimedia Commons 1010:Heterochordeumatidae 611:Brachychaeteumatidae 571:Anthroleucosomatidae 480:Chiloé Island, Chile 385:Neocambrisoma raveni 159:Heterochordeumatidea 1964:2012PLoSO...737240B 1361:on 26 December 2015 907:Neoatractosomatidae 827:Haplobainosomatidae 822:Gray in Jones, 1843 792:Gray in Jones, 1843 685:Macrochaeteumatidae 397:Hoffmaneuma exiguum 350:Xystrosoma beatense 337:Altajosoma kemerovo 245:spinning millipedes 192:Verhoeff, 1900 16:Order of millipedes 2005:Shear, W. (2011). 1391:on 31 October 2013 1183:Millipedes of Ohio 1026:Metopidiotrichidae 982: 815: 765:Lusitaniosomatidae 657: 644: 545:Speophilosomatidae 488:sub-Saharan Africa 379:Branneria carinata 374:Speophilosomatidae 234:sausage millipedes 3072: 3071: 3021:Open Tree of Life 2772:Taxon identifiers 2763: 2762: 2725: 2724: 2707: 2706: 2703: 2702: 2699: 2698: 2695: 2694: 2609: 2528: 2527: 2520:Microdecemplicida 2477: 2476: 2464: 2463: 2460: 2459: 2456: 2455: 2452: 2451: 2444: 2355: 2354: 2324: 2306: 2294: 2293: 2272: 2256: 2254:(pill millipedes) 2207: 2206: 2199: 2129: 2047:Media related to 1136: 1128: 1120: 1112: 1104: 1096: 1092:Urochordeumatidae 1088: 1080: 1072: 1062: 1054: 1046: 1038: 1030: 1022: 1014: 1006: 998: 990: 969: 961: 953: 945: 937: 927: 919: 911: 903: 895: 887: 869: 863: 855: 847: 839: 831: 823: 819:Craspedosomatidae 801: 793: 785: 777: 769: 761: 753: 745: 737: 729: 721: 713: 695: 689: 681: 673: 669:Heterolatzeliidae 665: 631: 623: 615: 607: 599: 591: 583: 575: 567: 559: 549: 541: 540:C. L. Koch, 1847 533: 532:C. L. Koch, 1847 525: 391:Golovatchia magda 342:sexual dimorphism 313:Lipseuma josianae 215:Arthropods portal 203: 202: 198: 197:Jeekel, 1971 193: 155:Craspedosomatidea 139: 3097: 3090:Millipede orders 3065: 3064: 3052: 3051: 3042: 3041: 3029: 3028: 3016: 3015: 3006: 3005: 2993: 2992: 2990:NBNSYS0100002381 2980: 2979: 2967: 2966: 2954: 2953: 2941: 2940: 2928: 2927: 2915: 2914: 2902: 2901: 2889: 2888: 2876: 2875: 2863: 2862: 2850: 2849: 2840: 2839: 2827: 2826: 2814: 2813: 2812: 2799: 2798: 2797: 2767: 2752: 2751: 2742: 2741: 2734:List of families 2720: 2715: 2683: 2678: 2674: 2656: 2614: 2608: 2605: 2568: 2559: 2515:Eoarthropleurida 2498: 2494: 2483: 2472: 2442: 2434: 2401: 2373: 2366: 2362: 2323: 2320: 2315: 2311: 2305: 2302: 2283: 2270: 2269:Sphaerotheriida 2255: 2252: 2247: 2238: 2197: 2184: 2180: 2169: 2127: 2118: 2111: 2104: 2095: 2079: 2059:Data related to 2058: 2046: 2030: 2029: 2011: 2002: 1996: 1995: 1985: 1975: 1943: 1937: 1936: 1908: 1902: 1901: 1885: 1879: 1878: 1876: 1874: 1831: 1825: 1824: 1808: 1802: 1801: 1791: 1782: 1776: 1775: 1743: 1737: 1736: 1726: 1717: 1711: 1710: 1662: 1656: 1655: 1645: 1643:10.5962/p.288286 1621: 1615: 1614: 1598: 1587: 1586: 1568: 1537: 1531: 1530: 1510: 1504: 1503: 1471: 1465: 1464: 1454: 1430: 1424: 1423: 1407: 1401: 1400: 1398: 1396: 1377: 1371: 1370: 1368: 1366: 1360: 1349: 1341: 1335: 1334: 1306: 1291: 1290: 1266: 1239: 1238: 1208: 1191: 1190: 1188: 1177: 1171: 1170: 1168: 1167: 1153: 1134: 1126: 1118: 1110: 1102: 1094: 1086: 1078: 1070: 1060: 1058:Pygmaeosomatidae 1052: 1044: 1036: 1028: 1020: 1012: 1004: 996: 988: 967: 959: 951: 943: 935: 925: 917: 909: 901: 893: 891:Hoffmaneumatidae 885: 879: 867: 861: 853: 845: 837: 829: 821: 799: 791: 783: 775: 773:Opisthocheiridae 767: 759: 757:Entomobielziidae 751: 743: 735: 727: 719: 711: 709:Niponiosomatidae 705: 693: 687: 679: 677:Kashmireumatidae 671: 663: 629: 621: 613: 605: 597: 589: 581: 573: 565: 557: 547: 539: 531: 523: 486:but absent from 440:hemianamorphosis 432:teloanamorphosis 415:and the species 319:Haasea hungarica 217: 212: 211: 196: 191: 134: 52: 51: 33: 21: 3105: 3104: 3100: 3099: 3098: 3096: 3095: 3094: 3075: 3074: 3073: 3068: 3060: 3055: 3047: 3045: 3037: 3032: 3024: 3019: 3011: 3009: 3001: 2996: 2988: 2983: 2975: 2970: 2962: 2957: 2949: 2944: 2936: 2931: 2923: 2918: 2910: 2905: 2897: 2892: 2884: 2879: 2871: 2866: 2858: 2853: 2845: 2843: 2835: 2830: 2822: 2817: 2808: 2807: 2802: 2793: 2792: 2787: 2774: 2764: 2759: 2721: 2691: 2681: 2672: 2667: 2650: 2645: 2606: 2604: 2603:Helminthomorpha 2597: 2579: 2575:Zosterogrammida 2562: 2553: 2551: 2549: 2547: 2545: 2543: 2541: 2539: 2537: 2535: 2524: 2502:Arthropleuridea 2488: 2473: 2448: 2432: 2427: 2399: 2394: 2369: 2360: 2351: 2342:Siphonocryptida 2321: 2303: 2301: 2300:Helminthomorpha 2290: 2281: 2276: 2253: 2232: 2230: 2228: 2226: 2224: 2222: 2220: 2218: 2216: 2214: 2203: 2174: 2163: 2131: 2122: 2090:on BugGuide.net 2067:"Chordeumatida" 2039: 2034: 2033: 2009: 2004: 2003: 1999: 1945: 1944: 1940: 1910: 1909: 1905: 1887: 1886: 1882: 1872: 1870: 1833: 1832: 1828: 1810: 1809: 1805: 1794:Myriapodologica 1789: 1784: 1783: 1779: 1745: 1744: 1740: 1729:Myriapodologica 1724: 1719: 1718: 1714: 1664: 1663: 1659: 1623: 1622: 1618: 1600: 1599: 1590: 1539: 1538: 1534: 1512: 1511: 1507: 1473: 1472: 1468: 1432: 1431: 1427: 1409: 1408: 1404: 1394: 1392: 1379: 1378: 1374: 1364: 1362: 1358: 1347: 1343: 1342: 1338: 1323: 1308: 1307: 1294: 1268: 1267: 1242: 1235: 1210: 1209: 1194: 1186: 1179: 1178: 1174: 1165: 1163: 1155: 1154: 1150: 1145: 1127:Silvestri, 1909 1124:Rhiscosomididae 1095:Silvestri, 1909 1034:Peterjohnsiidae 1021:Golovatch, 1978 986:Diplomaragnidae 894:Golovatch, 1978 877: 870: 866:Cyrnosomatidae 781:Trichopetalidae 703: 696: 692:Microlympiidae 627:Chamaesomatidae 595:Vandeleumatidae 587:Origmatogonidae 508: 460:Central America 452: 427: 354:Peterjohnsiidae 257: 213: 206: 194: 161: 157: 153: 133: 46: 17: 12: 11: 5: 3103: 3101: 3093: 3092: 3087: 3077: 3076: 3070: 3069: 3067: 3066: 3053: 3043: 3030: 3017: 3007: 2994: 2981: 2968: 2955: 2942: 2929: 2916: 2907:Fauna Europaea 2903: 2890: 2877: 2864: 2851: 2841: 2828: 2815: 2800: 2784: 2782: 2776: 2775: 2770: 2761: 2760: 2758: 2757: 2746: 2736: 2730: 2727: 2726: 2723: 2722: 2710: 2708: 2705: 2704: 2701: 2700: 2697: 2696: 2693: 2692: 2686: 2684: 2675: 2669: 2668: 2666: 2665: 2659: 2657: 2653:incertae sedis 2647: 2646: 2644: 2643: 2641:Palaeosomatida 2638: 2633: 2628: 2622: 2620: 2611: 2599: 2598: 2596: 2595: 2593:Amynilyspedida 2589: 2587: 2581: 2580: 2578: 2577: 2571: 2569: 2565:incertae sedis 2556: 2530: 2529: 2526: 2525: 2523: 2522: 2517: 2512: 2510:Arthropleurida 2506: 2504: 2490: 2489: 2486: 2479: 2478: 2475: 2474: 2467: 2465: 2462: 2461: 2458: 2457: 2454: 2453: 2450: 2449: 2447: 2446: 2437: 2435: 2429: 2428: 2426: 2425: 2420: 2415: 2410: 2404: 2402: 2396: 2395: 2393: 2392: 2390:Spirostreptida 2387: 2382: 2376: 2374: 2363: 2357: 2356: 2353: 2352: 2350: 2349: 2347:Siphonophorida 2344: 2339: 2334: 2328: 2326: 2308: 2296: 2295: 2292: 2291: 2288:Glomeridesmida 2286: 2284: 2278: 2277: 2275: 2274: 2266: 2260: 2258: 2244: 2235: 2209: 2208: 2205: 2204: 2202: 2201: 2192: 2190: 2176: 2175: 2172: 2165: 2164: 2162: 2161: 2155: 2149: 2143: 2136: 2133: 2132: 2123: 2121: 2120: 2113: 2106: 2098: 2092: 2091: 2085: 2080: 2064: 2063:at Wikispecies 2052: 2038: 2037:External links 2035: 2032: 2031: 1997: 1938: 1919:(1): 401–420. 1903: 1880: 1846:(6): 608–617. 1826: 1803: 1777: 1738: 1712: 1677:(6): 608–617. 1657: 1636:(3): 639–647. 1616: 1588: 1548: 1545: 1532: 1505: 1466: 1425: 1402: 1372: 1336: 1321: 1292: 1281:(2): 103–234. 1240: 1233: 1192: 1172: 1147: 1146: 1144: 1141: 1140: 1139: 1138: 1137: 1129: 1121: 1113: 1099: 1098: 1097: 1089: 1087:Verhoeff, 1909 1079:Verhoeff, 1909 1074: 1073: 1066: 1065: 1064: 1063: 1055: 1050:Lankasomatidae 1041: 1040: 1039: 1031: 1023: 1015: 1007: 1005:Verhoeff, 1914 993: 992: 991: 964: 963: 962: 954: 939: 938: 931: 930: 929: 928: 926:Verhoeff, 1899 918:Verhoeff, 1899 914: 913: 912: 910:Verhoeff, 1901 904: 902:Verhoeff, 1899 896: 888: 880: 864: 854:Verhoeff, 1901 850: 849: 848: 834: 833: 832: 830:Verhoeff, 1909 824: 816: 800:Verhoeff, 1899 788: 787: 786: 784:Verhoeff, 1914 778: 770: 768:Schubart, 1953 762: 760:Verhoeff, 1899 754: 746: 741:Biokoviellidae 732: 731: 730: 722: 714: 712:Verhoeff, 1941 706: 690: 688:Verhoeff, 1914 682: 674: 666: 661:Golovatchiidae 658: 630:Verhoeff, 1913 624: 616: 614:Verhoeff, 1910 602: 601: 600: 598:Mauriès, 1970 592: 584: 582:Hoffman, 1980 576: 574:Verhoeff 1899 561: 560: 553: 552: 551: 550: 548:Takakuwa, 1949 542: 537:Chordeumatidae 527: 526: 507: 506:Classification 504: 464:Southeast Asia 451: 448: 436:euanamorphosis 426: 423: 418:Vieteuma longi 402:Chordeumatidae 400:). The family 281:helminthomorph 256: 253: 247:because their 219: 218: 201: 200: 186: 185: 179: 178: 174: 173: 167: 166: 151:Chordeumatidea 146: 145: 141: 140: 128: 124: 123: 118: 114: 113: 108: 104: 103: 98: 94: 93: 88: 84: 83: 78: 74: 73: 68: 64: 63: 58: 54: 53: 40: 39: 35: 34: 26: 25: 24:Chordeumatida 15: 13: 10: 9: 6: 4: 3: 2: 3102: 3091: 3088: 3086: 3085:Chordeumatida 3083: 3082: 3080: 3063: 3058: 3054: 3050: 3044: 3040: 3035: 3031: 3027: 3022: 3018: 3014: 3008: 3004: 2999: 2995: 2991: 2986: 2982: 2978: 2973: 2969: 2965: 2960: 2956: 2952: 2947: 2943: 2939: 2934: 2930: 2926: 2921: 2917: 2913: 2908: 2904: 2900: 2895: 2891: 2887: 2882: 2878: 2874: 2869: 2865: 2861: 2856: 2852: 2848: 2842: 2838: 2837:Chordeumatida 2833: 2829: 2825: 2824:Chordeumatida 2820: 2816: 2811: 2810:Chordeumatida 2805: 2801: 2796: 2790: 2786: 2785: 2783: 2781: 2780:Chordeumatida 2777: 2773: 2768: 2756: 2747: 2745: 2737: 2735: 2732: 2731: 2728: 2719: 2714: 2690:(superfamily) 2689: 2685: 2679: 2676: 2670: 2664: 2661: 2660: 2658: 2655: 2654: 2648: 2642: 2639: 2637: 2634: 2632: 2629: 2627: 2624: 2623: 2621: 2619: 2618:Archipolypoda 2615: 2612: 2610: 2600: 2594: 2591: 2590: 2588: 2586: 2582: 2576: 2573: 2572: 2570: 2567: 2566: 2560: 2557: 2555: 2531: 2521: 2518: 2516: 2513: 2511: 2508: 2507: 2505: 2503: 2499: 2495: 2491: 2484: 2480: 2471: 2445: 2439: 2438: 2436: 2430: 2424: 2421: 2419: 2416: 2414: 2413:Chordeumatida 2411: 2409: 2406: 2405: 2403: 2397: 2391: 2388: 2386: 2383: 2381: 2378: 2377: 2375: 2372: 2367: 2364: 2358: 2348: 2345: 2343: 2340: 2338: 2335: 2333: 2330: 2329: 2327: 2325: 2319:Colobognatha 2316: 2312: 2309: 2307: 2297: 2289: 2285: 2279: 2273: 2267: 2265: 2262: 2261: 2259: 2257: 2251:Oniscomorpha 2248: 2245: 2243: 2239: 2236: 2234: 2210: 2200: 2194: 2193: 2191: 2189: 2185: 2181: 2177: 2170: 2166: 2160: 2156: 2154: 2150: 2148: 2144: 2142: 2138: 2137: 2134: 2130: 2119: 2114: 2112: 2107: 2105: 2100: 2099: 2096: 2089: 2086: 2084: 2081: 2078: 2074: 2073: 2068: 2065: 2062: 2061:Chordeumatida 2057: 2053: 2050: 2049:Chordeumatida 2045: 2041: 2040: 2036: 2027: 2023: 2019: 2015: 2008: 2001: 1998: 1993: 1989: 1984: 1979: 1974: 1969: 1965: 1961: 1958:(5): e37240. 1957: 1953: 1949: 1942: 1939: 1934: 1930: 1926: 1922: 1918: 1914: 1907: 1904: 1899: 1895: 1894:Insecta Mundi 1891: 1884: 1881: 1869: 1865: 1861: 1857: 1853: 1849: 1845: 1841: 1837: 1830: 1827: 1822: 1818: 1814: 1807: 1804: 1799: 1795: 1788: 1781: 1778: 1773: 1769: 1765: 1761: 1757: 1753: 1749: 1742: 1739: 1734: 1730: 1723: 1716: 1713: 1708: 1704: 1700: 1696: 1692: 1688: 1684: 1680: 1676: 1672: 1668: 1661: 1658: 1653: 1649: 1644: 1639: 1635: 1632:(in French). 1631: 1627: 1620: 1617: 1612: 1608: 1604: 1597: 1595: 1593: 1589: 1584: 1580: 1576: 1572: 1567: 1562: 1558: 1554: 1550: 1546: 1543: 1536: 1533: 1528: 1524: 1520: 1516: 1509: 1506: 1501: 1497: 1493: 1489: 1485: 1481: 1477: 1470: 1467: 1462: 1458: 1453: 1448: 1444: 1441:(in French). 1440: 1436: 1429: 1426: 1421: 1417: 1413: 1406: 1403: 1390: 1386: 1382: 1376: 1373: 1357: 1353: 1346: 1340: 1337: 1332: 1328: 1324: 1322:90-04-07698-0 1318: 1314: 1313: 1305: 1303: 1301: 1299: 1297: 1293: 1288: 1284: 1280: 1276: 1272: 1265: 1263: 1261: 1259: 1257: 1255: 1253: 1251: 1249: 1247: 1245: 1241: 1236: 1234:9789004156128 1230: 1226: 1222: 1218: 1214: 1207: 1205: 1203: 1201: 1199: 1197: 1193: 1185: 1184: 1176: 1173: 1162: 1158: 1152: 1149: 1142: 1135:Bollman, 1893 1133: 1130: 1125: 1122: 1117: 1114: 1109: 1106: 1105: 1103:Bollman, 1893 1100: 1093: 1090: 1085: 1082: 1081: 1076: 1075: 1068: 1067: 1059: 1056: 1051: 1048: 1047: 1042: 1037:Mauriès, 1987 1035: 1032: 1027: 1024: 1019: 1018:Megalotylidae 1016: 1011: 1008: 1003: 1000: 999: 994: 987: 984: 983: 979: 978: 977:Schedotrigona 973: 968:Attems, 1907 965: 958: 955: 950: 947: 946: 941: 940: 933: 932: 924: 923:Verhoeffiidae 921: 920: 915: 908: 905: 900: 897: 892: 889: 884: 881: 878:Mauriès, 1969 876: 875: 868:Mauriès, 2003 865: 860: 857: 856: 851: 844: 841: 840: 835: 828: 825: 820: 817: 812: 811: 805: 798: 795: 794: 789: 782: 779: 774: 771: 766: 763: 758: 755: 750: 749:Cleidogonidae 747: 742: 739: 738: 733: 726: 725:Trachygonidae 723: 720:Loomis, 1966 718: 715: 710: 707: 702: 701: 691: 686: 683: 680:Mauriès, 1982 678: 675: 672:Verhoeff 1899 670: 667: 662: 659: 654: 653: 648: 641: 640: 635: 628: 625: 620: 617: 612: 609: 608: 603: 596: 593: 590:Verhoeff 1914 588: 585: 580: 577: 572: 569: 568: 566:Verhoeff 1899 563: 562: 555: 554: 546: 543: 538: 535: 534: 529: 528: 521: 520: 519: 517: 513: 505: 503: 501: 497: 493: 489: 485: 481: 477: 473: 469: 465: 461: 457: 449: 447: 443: 441: 437: 433: 424: 422: 420: 419: 414: 413: 408: 403: 399: 398: 393: 392: 388: 386: 381: 380: 375: 369: 367: 363: 357: 355: 351: 347: 343: 339: 338: 333: 332: 327: 326: 321: 320: 315: 314: 309: 308: 303: 302: 296: 294: 288: 286: 282: 278: 274: 270: 266: 261: 254: 252: 250: 246: 242: 239: 235: 231: 227: 223: 222:Chordeumatida 216: 210: 205: 199: 187: 184: 180: 175: 172: 168: 165: 164: 160: 156: 152: 147: 142: 137: 132: 131:Chordeumatida 129: 126: 125: 122: 119: 116: 115: 112: 109: 107:Subterclass: 106: 105: 102: 99: 96: 95: 92: 89: 86: 85: 82: 79: 76: 75: 72: 69: 66: 65: 62: 59: 56: 55: 50: 45: 41: 36: 32: 27: 22: 19: 2779: 2688:Xyloiuloidea 2663:Pleurojulida 2651: 2636:Euphoberiida 2631:Cowiedesmida 2626:Archidesmida 2563: 2441:Polydesmida 2418:Siphoniulida 2412: 2332:Platydesmida 2282:Limacomorpha 2145:Superphylum 2128:(millipedes) 2070: 2017: 2013: 2000: 1955: 1951: 1941: 1916: 1912: 1906: 1897: 1893: 1883: 1871:. Retrieved 1843: 1839: 1829: 1820: 1816: 1806: 1797: 1793: 1780: 1755: 1751: 1741: 1732: 1728: 1715: 1674: 1670: 1660: 1633: 1629: 1619: 1610: 1606: 1559:(1): 41–49. 1556: 1552: 1535: 1518: 1508: 1486:(1): 1–205. 1483: 1479: 1469: 1442: 1438: 1428: 1419: 1415: 1405: 1393:. Retrieved 1389:the original 1384: 1375: 1363:. Retrieved 1356:the original 1351: 1339: 1311: 1278: 1274: 1216: 1182: 1175: 1164:. Retrieved 1160: 1151: 1111:Loomis, 1966 1108:Apterouridae 1053:Mauriès 1978 1029:Attems, 1907 1013:Pocock, 1894 997:Pocock, 1894 989:Attems, 1907 975: 886:Attems, 1926 872: 859:Altajellidae 846:Attems, 1899 838:Attems, 1899 808: 776:Ribaut, 1913 698: 652:Craspedosoma 650: 637: 619:Branneriidae 524:Pocock 1894 509: 453: 450:Distribution 444: 428: 416: 410: 395: 389: 383: 377: 370: 361: 358: 349: 335: 329: 323: 317: 311: 305: 299: 297: 289: 262: 258: 244: 233: 221: 220: 189: 149: 130: 117:Superorder: 18: 2946:iNaturalist 2804:Wikispecies 2423:Stemmiulida 2408:Callipodida 2400:Nematophora 2385:Spirobolida 2337:Polyzoniida 2196:Polyxenida 2188:Penicillata 2020:: 159–164. 1800:(4): 25–36. 1735:(4): 25–36. 1422:(1): 75–92. 1219:: 363–453. 1132:Striariidae 1119:Shear, 2009 1002:Eudigonidae 957:Conotylidae 952:Shear, 1971 949:Adritylidae 936:Shear, 2000 871:monotypic: 797:Attemsiidae 744:Mrsic, 1992 700:Microlympia 697:monotypic: 664:Shear, 1992 639:Atractosoma 496:Scandinavia 476:New Zealand 425:Development 412:Kashmireuma 277:Polydesmida 255:Description 241:Nematophora 163:Striariidea 138:, 1894 121:Nematophora 87:Subphylum: 3079:Categories 2755:Millipedes 2682:Juliformia 2585:Pentazonia 2371:Juliformia 2242:Pentazonia 2157:Subphylum 2153:Arthropoda 2126:Diplopoda 1758:(1): 1–7. 1547:carolinus 1416:Zoosystema 1166:2024-07-04 1143:References 1071:Cook, 1896 1061:Carl, 1941 1045:Carl, 1941 960:Cook, 1896 944:Cook, 1896 807:A pair of 752:Cook, 1896 736:Cook, 1896 728:Cook, 1896 717:Tingupidae 622:Cook, 1896 606:Cook, 1896 558:Cook, 1895 484:Madagascar 238:superorder 230:millipedes 144:Suborders 81:Arthropoda 2744:Diplopoda 2433:Merocheta 2264:Glomerida 2159:Myriapoda 2147:Ecdysozoa 1873:25 August 1772:1175-5334 1691:1520-541X 1652:132547501 1575:1175-5334 1527:2246/5072 1500:1175-5334 1461:0035-418X 1084:Caseyidae 883:Faginidae 874:Cyrnosoma 843:Haaseidae 579:Haasiidae 366:Diplopoda 171:Diversity 101:Diplopoda 91:Myriapoda 67:Kingdom: 61:Eukaryota 2868:BugGuide 2844:BioLib: 2795:Q4300743 2789:Wikidata 2673:Eugnatha 2361:Eugnatha 2141:Animalia 2139:Kingdom 1992:22615951 1952:PLOS ONE 1933:17163800 1900:: 1–134. 1868:21401688 1860:16336414 1707:21401688 1699:16336414 1521:(2977). 1445:: 3–49. 1395:18 March 1365:20 March 1331:13439686 1116:Buotidae 512:families 472:Tasmania 407:sternite 346:Buotidae 325:Tianella 293:gonopods 285:ozopores 273:paranota 183:Synonyms 111:Eugnatha 77:Phylum: 71:Animalia 57:Domain: 2487:Extinct 2151:Phylum 2069:at the 2014:Zootaxa 1983:3352885 1960:Bibcode 1752:Zootaxa 1583:4833397 1553:Zootaxa 1480:Zootaxa 500:Siberia 468:Oceania 456:tropics 394:, and 249:telsons 127:Order: 97:Class: 3062:887894 3049:582683 3046:uBio: 3010:NZOR: 3003:118457 2977:154417 2860:259567 2753:  2380:Julida 2173:Living 1990:  1980:  1931:  1866:  1858:  1770:  1705:  1697:  1689:  1650:  1581:  1573:  1544:Buotus 1498:  1459:  1329:  1319:  1231:  810:Haasea 516:genera 478:, and 466:, and 269:collum 136:Pocock 3057:WoRMS 3034:Plazi 3026:77633 2964:11642 2959:IRMNG 2951:50425 2912:11526 2886:7NF8X 2873:97644 2847:19684 2010:(PDF) 1864:S2CID 1790:(PDF) 1725:(PDF) 1703:S2CID 1648:S2CID 1579:S2CID 1359:(PDF) 1348:(PDF) 1187:(PDF) 492:Chile 265:setae 226:order 2998:NCBI 2972:ITIS 2938:1017 2933:GBIF 2899:6397 2855:BOLD 2018:3148 1988:PMID 1929:PMID 1875:2020 1856:PMID 1768:ISSN 1695:PMID 1687:ISSN 1634:4:11 1571:ISSN 1557:2290 1496:ISSN 1484:4211 1457:ISSN 1397:2014 1367:2014 1327:OCLC 1317:ISBN 1229:ISBN 316:and 304:and 2985:NBN 2894:EoL 2881:CoL 2832:AFD 2819:ADW 2022:doi 1978:PMC 1968:doi 1921:doi 1898:158 1848:doi 1760:doi 1756:233 1679:doi 1638:doi 1561:doi 1523:hdl 1488:doi 1447:doi 1283:doi 1279:109 1221:doi 458:of 438:or 430:as 228:of 3081:: 3059:: 3036:: 3023:: 3000:: 2987:: 2974:: 2961:: 2948:: 2935:: 2922:: 2909:: 2896:: 2883:: 2870:: 2857:: 2834:: 2821:: 2806:: 2791:: 2016:. 2012:. 1986:. 1976:. 1966:. 1954:. 1950:. 1927:. 1917:52 1915:. 1896:. 1892:. 1862:. 1854:. 1842:. 1838:. 1821:53 1819:. 1815:. 1796:. 1792:. 1766:. 1754:. 1750:. 1731:. 1727:. 1701:. 1693:. 1685:. 1673:. 1669:. 1646:. 1628:. 1611:25 1609:. 1605:. 1591:^ 1577:. 1569:. 1555:. 1551:. 1517:. 1494:. 1482:. 1478:. 1455:. 1443:95 1437:. 1420:28 1418:. 1414:. 1383:. 1350:. 1325:. 1295:^ 1277:. 1273:. 1243:^ 1227:. 1215:. 1195:^ 1159:. 518:. 498:, 474:, 462:, 442:. 382:, 368:. 287:. 2554:a 2552:h 2550:t 2548:a 2546:n 2544:g 2542:o 2540:l 2538:i 2536:h 2534:C 2233:a 2231:h 2229:t 2227:a 2225:n 2223:g 2221:o 2219:l 2217:i 2215:h 2213:C 2117:e 2110:t 2103:v 2028:. 2024:: 1994:. 1970:: 1962:: 1956:7 1935:. 1923:: 1877:. 1850:: 1844:7 1798:3 1762:: 1733:3 1709:. 1681:: 1675:7 1640:: 1585:. 1563:: 1529:. 1525:: 1502:. 1490:: 1449:: 1399:. 1369:. 1333:. 1289:. 1285:: 1237:. 1223:: 1169:. 387:,

Index


Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Myriapoda
Diplopoda
Eugnatha
Nematophora
Chordeumatida
Pocock
Chordeumatidea
Craspedosomatidea
Heterochordeumatidea
Striariidea
Diversity
Synonyms
icon
Arthropods portal
order
millipedes
superorder
Nematophora
telsons
setae
collum
paranota
Polydesmida
helminthomorph

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