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Chriacus

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124: 393:. This difference mirrors the history of the mammals classified as arctocyonids. They were first considered creodonts (imagined to be the ancestors of modern carnivores), and then "condylarths," (imagined to be the ancestors of hoofed mammals). Modern studies suggest the confusion is due to the fact that ungulates, carnivores, and creodonts are related groups, and flesh-eating lineages and adaptations evolved within each of them. 104: 355:
body size) and the neocortex was less developed than later mammals. By the standards of modern mammals, they would have been neither especially quick nor intelligent, but their brains were comparable to many mammals of their time. Derived features of the inner ear were shared with fossils that are assigned to
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suggest these animals depended more on their sense of smell than sight, may have been able to hear about as well as a modern aardvark, and were slow-moving to moderately agile. The encephalization quotient (EQ) had a range of 0.12–0.41 (1.0=average brain size compared with modern mammals of a similar
323:, though they were not closely related to any living mammal. Well preserved fossils allow clear information on what they looked like. They were about 1 metre (3.3 ft) long including a long, robust tail, which may or may not have been 707: 619: 556: 499:"Virtual endocranial and inner ear endocasts of the Paleocene 'condylarth' Chriacus: new insight into the neurosensory system and evolution of early placental mammals" 796: 327:. Other features include a light build, weighing approximately 7 kg (15 lb), and many adaptations typical of animals that live in trees. These include 497:
Bertrand, Ornella C., Sarah L. Shelley John R. Wible Thomas E. Williamson Luke T. Holbrook Stephen G.B. Chester Ian B. Butler, and Stephen L. Brusatte (2019).
335:. The powerfully built limbs had flexible joints, especially the ankles, an adaptation that allows an animal to turn its hind feet behind it, like modern tree 842: 862: 359:(artiodactyls+perissodactyls), suggesting the genus may be close to the origin of ungulates, though it is too different in form to be a direct ancestor. 847: 783: 852: 857: 481: 724: 371:
in relation to other groups is disputed. Halliday et al. (2015) consider it a member of the family Oxyclaenidae, a sister group to
635:"A structural intermediate between triisodontids and mesonychians (Mammalia, Acreodi) from the earliest Eocene of Portugal" 367:
There are nine species currently recognized in the genus. Like most early placental mammals, the classification of
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epoch and died out after the early Eocene. In life, members of the genus would have looked something like a
690:"Resolving_the_higher-level_phylogenetic_relationships_of_Triisodontidae_'Condylarthra'_within_Placentalia" 751: 809: 613: 550: 837: 646: 430: 211: 111: 670: 538: 118: 814: 801: 701: 662: 601: 530: 477: 454: 446: 654: 591: 583: 520: 510: 438: 419:"Climbing adaptations in the early eocene mammal Chriacus and the origin of artiodactyla" 650: 434: 596: 571: 525: 498: 372: 418: 831: 689: 542: 191: 61: 674: 634: 379:, while Tabuce et al. (2011) classify it as an arctocyonid, most closely related to 220: 719: 328: 178: 36: 774: 736: 658: 390: 324: 81: 46: 450: 442: 385: 376: 320: 312: 135: 86: 666: 605: 534: 474:
The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals
458: 103: 768: 356: 340: 336: 316: 155: 76: 71: 56: 51: 41: 788: 633:
Tabuce, Rodolphe; Clavel, Julien; Telles Antunes, Miguel (2011-02-01).
91: 66: 688:
Shelley, Sarah L, Stephen L Brusatte, and Thomas E Williamson (2015).
587: 515: 308: 165: 145: 745: 570:
Halliday, Thomas J. D.; Upchurch, Paul; Goswami, Anjali (2015).
332: 749: 572:""Resolving the relationships of Paleocene placental mammals"" 331:
of their five-toed feet, and having long, curved, compressed
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lies somewhere within the range of their early relatives.
758: 311:that lived in what is now North America during the 725:Technical information in the Paleobiology database 346:Analysis of casts of the brain and inner ear from 343:, eating fruit, eggs, insects and small mammals. 339:, in order to climb downward. They were probably 8: 706:: CS1 maint: multiple names: authors list ( 618:: CS1 maint: multiple names: authors list ( 555:: CS1 maint: multiple names: authors list ( 746: 476:. London: Marshall Editions. p. 234. 102: 20: 595: 524: 514: 406: 699: 611: 548: 7: 412: 410: 843:Paleocene mammals of North America 14: 863:Taxa named by Edward Drinker Cope 307:is an extinct genus of placental 122: 848:Eocene mammals of North America 1: 853:Fossil taxa described in 1883 858:Prehistoric placental genera 741:reconstructed from skeletons 720:Article on Paleocene mammals 32:Early Paleocene–Early Eocene 879: 417:Rose, K. D. (1987-04-17). 659:10.1007/s00114-010-0747-y 240: 235: 226: 219: 119:Scientific classification 117: 110: 101: 23: 472:Palmer, D., ed. (1999). 16:Extinct genus of mammals 639:Die Naturwissenschaften 443:10.1126/science.3426662 810:Paleobiology Database 389:, and allied to the 329:walking on the soles 651:2011NW.....98..145T 435:1987Sci...236..314R 112:Life reconstruction 576:Biological Reviews 503:Journal of Anatomy 348:Chriacus pelvidens 229:Chriacus pelvidens 825: 824: 752:Taxon identifiers 588:10.1111/brv.12242 516:10.1111/joa.13084 429:(4799): 314–316. 300: 299: 215: 870: 818: 817: 805: 804: 792: 791: 779: 778: 777: 747: 712: 711: 705: 697: 685: 679: 678: 630: 624: 623: 617: 609: 599: 567: 561: 560: 554: 546: 528: 518: 494: 488: 487: 469: 463: 462: 414: 210: 203: 190: 177: 127: 126: 106: 96: 33: 29:Temporal range: 21: 878: 877: 873: 872: 871: 869: 868: 867: 828: 827: 826: 821: 813: 808: 800: 795: 787: 782: 773: 772: 767: 754: 733: 716: 715: 698: 687: 686: 682: 632: 631: 627: 610: 569: 568: 564: 547: 496: 495: 491: 484: 471: 470: 466: 416: 415: 408: 403: 365: 231: 209: 201: 188: 175: 121: 97: 95: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 39: 31: 30: 27: 17: 12: 11: 5: 876: 874: 866: 865: 860: 855: 850: 845: 840: 830: 829: 823: 822: 820: 819: 806: 793: 780: 764: 762: 756: 755: 750: 744: 743: 732: 731:External links 729: 728: 727: 722: 714: 713: 680: 625: 582:(1): 521–550. 562: 489: 482: 464: 405: 404: 402: 399: 364: 363:Classification 361: 298: 297: 296: 295: 289: 283: 281:C. oconostotae 277: 271: 265: 259: 253: 247: 238: 237: 233: 232: 224: 223: 217: 216: 199: 195: 194: 186: 182: 181: 173: 169: 168: 163: 159: 158: 153: 149: 148: 143: 139: 138: 133: 129: 128: 115: 114: 108: 107: 99: 98: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 40: 35: 34: 28: 15: 13: 10: 9: 6: 4: 3: 2: 875: 864: 861: 859: 856: 854: 851: 849: 846: 844: 841: 839: 836: 835: 833: 816: 811: 807: 803: 798: 794: 790: 785: 781: 776: 770: 766: 765: 763: 761: 757: 753: 748: 742: 740: 737:Picture of a 735: 734: 730: 726: 723: 721: 718: 717: 709: 703: 695: 691: 684: 681: 676: 672: 668: 664: 660: 656: 652: 648: 645:(2): 145–55. 644: 640: 636: 629: 626: 621: 615: 607: 603: 598: 593: 589: 585: 581: 577: 573: 566: 563: 558: 552: 544: 540: 536: 532: 527: 522: 517: 512: 508: 504: 500: 493: 490: 485: 483:1-84028-152-9 479: 475: 468: 465: 460: 456: 452: 448: 444: 440: 436: 432: 428: 424: 420: 413: 411: 407: 400: 398: 396: 392: 388: 387: 382: 378: 374: 373:Palaeoryctids 370: 362: 360: 358: 353: 349: 344: 342: 338: 334: 330: 326: 322: 318: 314: 310: 306: 305: 294: 290: 288: 284: 282: 278: 276: 275:C. metocometi 272: 270: 266: 264: 260: 258: 257:C. calenancus 254: 252: 248: 246: 242: 241: 239: 234: 230: 225: 222: 218: 213: 208: 207: 200: 197: 196: 193: 192:Arctocyonidae 187: 184: 183: 180: 174: 171: 170: 167: 164: 161: 160: 157: 154: 151: 150: 147: 144: 141: 140: 137: 134: 131: 130: 125: 120: 116: 113: 109: 105: 100: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 38: 26: 22: 19: 759: 738: 694:ResearchGate 693: 683: 642: 638: 628: 614:cite journal 579: 575: 565: 551:cite journal 509:(1): 21–49. 506: 502: 492: 473: 467: 426: 422: 394: 384: 380: 368: 366: 351: 347: 345: 303: 302: 301: 292: 287:C. pelvidens 286: 280: 274: 268: 262: 256: 250: 244: 228: 221:Type species 205: 204: 24: 18: 838:Condylarths 381:Loxolophus, 352:C. baldwini 269:C. katrinae 263:C. gallinae 251:C. baldwini 245:C. badgleyi 179:Arctocyonia 832:Categories 401:References 391:Mesonychia 357:Euungulata 325:prehensile 293:C. punitor 543:204969130 451:0036-8075 386:Arctocyon 377:Creodonts 341:omnivores 337:squirrels 321:binturong 313:Paleocene 142:Kingdom: 136:Eukaryota 775:Q2398857 769:Wikidata 760:Chriacus 739:Chriacus 702:cite web 675:22526630 667:21181109 606:28075073 535:31667836 395:Chriacus 369:Chriacus 317:kinkajou 304:Chriacus 236:Species 206:Chriacus 185:Family: 166:Mammalia 156:Chordata 152:Phylum: 146:Animalia 132:Domain: 25:Chriacus 802:1207752 789:4826585 647:Bibcode 597:6849585 526:6904649 459:3426662 431:Bibcode 423:Science 309:mammals 198:Genus: 172:Order: 162:Class: 673:  665:  604:  594:  541:  533:  523:  480:  457:  449:  214:, 1883 815:42203 797:IRMNG 671:S2CID 539:S2CID 383:then 333:claws 784:GBIF 708:link 663:PMID 620:link 602:PMID 557:link 531:PMID 478:ISBN 455:PMID 447:ISSN 375:and 350:and 212:Cope 37:PreꞒ 655:doi 592:PMC 584:doi 521:PMC 511:doi 507:236 439:doi 427:236 319:or 834:: 812:: 799:: 786:: 771:: 704:}} 700:{{ 692:. 669:. 661:. 653:. 643:98 641:. 637:. 616:}} 612:{{ 600:. 590:. 580:92 578:. 574:. 553:}} 549:{{ 537:. 529:. 519:. 505:. 501:. 453:. 445:. 437:. 425:. 421:. 409:^ 87:Pg 710:) 696:. 677:. 657:: 649:: 622:) 608:. 586:: 559:) 545:. 513:: 486:. 461:. 441:: 433:: 291:† 285:† 279:† 273:† 267:† 261:† 255:† 249:† 243:† 227:† 202:† 189:† 176:† 92:N 82:K 77:J 72:T 67:P 62:C 57:D 52:S 47:O 42:Ꞓ

Index

PreꞒ

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D
C
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Life reconstruction
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Mammalia
Arctocyonia
Arctocyonidae
Chriacus
Cope
Type species
mammals
Paleocene
kinkajou
binturong
prehensile

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