Knowledge (XXG)

Cittarium pica

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generally does not live at great depths, though this has occasionally been reported. Most individuals are found at the water's edge, and have little tendency to disperse. Minimum recorded depth is 0 m. Maximum recorded depth is 7 m.
859:. During the reproductive season, which normally occurs from June to November in the field, male individuals release their sperm into the water, as females simultaneously release their green colored unfertilised 681:. Those spots have a tendency to become axial lines in older, larger individuals. The upper surface is often entirely black. The aperture is commonly white, with an inner iridescence because of the nacre. 692:
On some old, empty shells of large individuals, the black colored parts become slightly higher in relief, compared to the white areas surrounding it. This unusual morphology may be due to the action of
439:, where this is a popular food item. The word "will" or "wilks" can be used both as a singular form and a plural. This species is however not at all closely related to the species that are known as 684:
Young shells, or well-preserved adults, have the spire whorls sculptured by oblique folds, cut by a few spiral sulci. The periphery and the base in the half-grown shells are spirally lirate.
520:. The vetigastropods are considered to be among the most primitive living neo-gastropods, and are widely distributed in all oceans of the world. It is also part of the superfamily 992:
These large sea snails are boiled and eaten in a variety of different local recipes. Because of their popularity as a food item and the problem of overfishing, in the
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there are territorial regulations to protect these snails: no collecting is allowed during their reproductive season, and there is a minimum harvest size regulation.
1703: 1252: 1850: 1755: 879:. These trochophore larvae do not spend much time in the plankton, because settlement occurs relatively soon, after 3.5 to 4.5 days. Individuals usually reach 1167:
The living marine resources of the Western Central Atlantic. Volume 1: Introduction, molluscs, crustaceans, hagfishes, sharks, batoid fishes, and chimaeras.
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FAO Species Identification Guide for Fishery Purposes and American Society of Ichthyologists and Herpetologists Special Publication No. 5. 1600p.
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at shell lengths of 32–34 mm. The life span of this species is still unknown, but estimates for other top shells reach 30 years.
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is arcuate, produced above in a heavy porcellanous callous deposit, half-surrounding the umbilicus and deeply notched in the middle.
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individuals possess shells ornamented by spiral lines and strong cords, in contrast to the nearly smooth, homogeneous surface of
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of this species can be up to 137 mm in maximum dimension. It is very thick and heavy, having an outline that is between
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This snail is known as "wilk" or "wilks" (or sometimes as "whelks") in the English-speaking Caribbean islands of the
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Field guide to the commercial marine and brackish-water resources of the northern coast of South America
632: 272: 123: 1312:(Mollusca: Gastropoda: Trochidae) From The Oligocene of Europe and its Paleobiogeographic Implications" 443:
in the U.S. and in Europe. In some Spanish-speaking parts of the Caribbean, when used as a food source
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The lusterless color pattern is rather distinct, overall white with black zigzag flammules on each
667: 663: 643: 1107: 646:, which is deep and devoid of sculpture, but spirally bicostate inside. The semicircular, oblique 1835: 1331: 1276: 978:
In the wild, the shell of this species is used extensively by the large land hermit crab species
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over time. Feeding commonly occurs during the nocturnal period, when the snails are most active.
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This large snail is found on or under rocks, in exposed and moderately sheltered shores, both in
756: 647: 302: 152: 1799: 1786: 1708: 1651: 1534: 1389: 1139: 993: 1524: 1514: 1379: 1369: 1323: 1268: 1106:. Department of Planning and Natural Resources Division of Fish and Wildlife. Archived from 694: 502: 493: 422: 1493:"Population Genetics of a Trochid Gastropod Broadens Picture of Caribbean Sea Connectivity" 880: 764: 678: 671: 659: 628: 620: 605: 581: 483: 478: 429: 141: 1510: 1529: 1492: 1384: 1355: 744: 517: 412: 209: 1406: 110: 1829: 860: 852: 371: 348: 324: 289: 61: 1280: 1133: 787: 1427: 712: 1656: 1138:. FAO species identification sheets for fishery purposes. Rome: FAO. p. 513. 1519: 1374: 1067: 1794: 1742: 1690: 1623: 1407:
Tryon (1889), Manual of Conchology XI, Academy of Natural Sciences, Philadelphia
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is a legally protected species in Bermuda, where its collection is forbidden.
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fertilised eggs, which will further develop into lecitotrophic (yolk feeding)
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shells (or some other similarly large shell) to use as shelter. Nowadays,
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Ministry of Environment and Sports: Department of Conservation Services.
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is simple. The lip is edged inside by black, or black and white. The
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Leal, J.H. (2002). Gastropods. p. 99-147. In: Carpenter, K.E. (ed.).
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in shape, with rounded shoulders and a somewhat low conical form.
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is glossy and delicate, and has a node that projects towards the
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by Philippi (1847), Rosenberg (2005) and Hendy et al. (2008).
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Díaz-Ferguson, E.; Haney, R.; Wares, J.; Silliman, B. (2010).
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Proceedings of the Academy of Natural Sciences of Philadelphia
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A small shallow seawater pool found on St. Croix's north shore
1267:(1). The Academy of Natural Sciences of Philadelphia: 27–47. 1132:
Cervigón, F. (1993). "Gastropods". In Carpenter, K.E. (ed.).
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10.1666/0022-3360(2002)076<0767:frotcg>2.0.co;2
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10.1635/0097-3157(2003)153[0027:TEWIWC]2.0.CO;2
875:. They emerge from the egg capsules as shell-cap-bearing 497:). It has gone locally extinct in some habitats due to 962:(also known as ringed wormsnail) and the tube dwelling 824:
Rocky shore on St. Croix US Virgin Islands, habitat of
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which has been polished to make it look more attractive
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as the inner shell layer, and its subordinated family
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More information and recipes from the Cayman Islands
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The spire is conoidal. It contains about six convex
1598: 589:is very distinctive. This shell has been polished. 1479:view from below; the umbilicus is clearly visible 1011:populations, because these arthropods need empty 555:. For a long time, only one species in the genus 1034: 1032: 455:(snails, in Spanish). In Venezuela it is called 1179:Sartwell, T. Wood, J. B.; Valdivia, A. (eds.). 1576:Photo of live animal and a lot of information 551:This is the only living species in the genus 8: 1349: 1347: 1345: 1246: 1244: 1242: 1240: 1238: 1236: 1234: 1232: 1230: 1228: 1226: 1224: 1222: 891:The west Indian top shell is known to be an 559:was known, however in 2002 a fossil species 1220: 1218: 1216: 1214: 1212: 1210: 1208: 1206: 1204: 1202: 1062: 1060: 428:. This species has a large black and white 1586: 131: 109: 20: 1528: 1518: 1383: 1373: 1044:Cittarium pica Linnaeus, 1758 (Top Snail) 1094: 1092: 1090: 1088: 1086: 1084: 1445: 1426:. Government of Bermuda. Archived from 1322:(4). Paleontological Society: 767–770. 1028: 775:. The species has been reintroduced to 1185:West Indian Top Shell (Cittarium pica) 1161: 1159: 1157: 1155: 1101:"U.S.V.I. Animal Fact Sheet #15 Whelk" 1007:. This caused a serious impact on the 473:important invertebrate species in the 1424:West Indian Topshell (Cittarium pica) 1308:"First Record of The Caribbean Genus 7: 1851:Arthropods of the Dominican Republic 565:Lozouet, 2002 was discovered in the 536:. (1924) recombined this species as 544:, and it was finally recombined as 540:. Weisbord (1962) recombined it as 122:, showing the inner surface of the 971:may live attached to the shell of 711:This species occurs rarely in the 14: 1181:"Marine Invertebrates of Bermuda" 118:A colored illustration of a live 1472: 1460: 1448: 1253:"The edible West Indian "whelk" 895:, feeding on a large variety of 715:, and in the Caribbean coast of 627:is depressed-globose. The outer 156: 654:inside as is the case in other 585:The wide and deep umbilicus of 1: 469:is considered the third most 1841:Gastropods described in 1758 1520:10.1371/journal.pone.0012675 1375:10.1371/journal.pone.0008776 1846:Taxa named by Carl Linnaeus 1068:"Malacolog (Version 4.1.1)" 1040:"The Paleobiology Database" 1867: 1003:became locally extinct in 863:. The encounter of those 569:deposits of southwestern 308: 301: 278: 271: 153:Scientific classification 151: 139: 130: 117: 108: 23: 925:, often uses a shell of 899:, and sometimes also on 735:. It also occurs in the 459:; in Cuba it is called 1316:Journal of Paleontology 911:Biological interactions 855:in this species occurs 658:, and is circular. The 642:presents a rather wide 1251:Robertson, R. (2003). 999:For a similar reason, 929: 919:The land hermit crab, 836: 828: 796: 601: 597:Drawing of a shell of 590: 1782:Paleobiology Database 1354:Welch, J. J. (2010). 918: 834: 823: 790: 596: 584: 397:West Indian top shell 1306:Lozouet, P. (2002). 959:Dendropoma corrodens 945:Pinnotheres barbatus 700:Plectonema terebrans 562:Cittarium maestratii 516:is within the clade 16:Species of gastropod 1511:2010PLoSO...512675D 1099:Toller, W. (2003). 981:Coenobita clypeatus 922:Coenobita clypeatus 773:Trinidad and Tobago 650:is distinguishably 1430:on 7 February 2011 948:is mentioned as a 939:Lottia leucopleura 930: 837: 829: 797: 791:An empty shell of 757:Dominican Republic 602: 591: 312:Cittarium picoides 32:Pleistocene–Recent 1823: 1822: 1592:Taxon identifiers 1420:"Bermuda Species" 1113:on 16 August 2011 994:US Virgin Islands 577:Shell description 384: 383: 378: 364: 356: 340: 332: 316: 315:Gould, A.A., 1853 1858: 1816: 1815: 1803: 1802: 1790: 1789: 1777: 1776: 1764: 1763: 1751: 1750: 1738: 1737: 1725: 1724: 1712: 1711: 1699: 1698: 1686: 1685: 1673: 1672: 1660: 1659: 1647: 1646: 1634: 1633: 1632: 1619: 1618: 1617: 1587: 1572: 1570: 1568: 1542: 1532: 1522: 1476: 1467:Abapertural view 1464: 1452: 1440: 1439: 1437: 1435: 1415: 1409: 1404: 1398: 1397: 1387: 1377: 1351: 1340: 1339: 1303: 1297: 1296: 1294: 1292: 1283:. Archived from 1248: 1197: 1196: 1194: 1192: 1176: 1170: 1163: 1150: 1149: 1129: 1123: 1122: 1120: 1118: 1112: 1105: 1096: 1079: 1078: 1076: 1074: 1064: 1055: 1054: 1052: 1050: 1036: 1009:land hermit crab 767:as far south as 695:blue-green algae 503:overexploitation 494:Eustrombus gigas 407:of large edible 370: 362: 360:Trochus picoides 346: 338: 322: 314: 284: 161: 160: 140:Five views of a 135: 113: 103: 33: 29:Temporal range: 21: 1866: 1865: 1861: 1860: 1859: 1857: 1856: 1855: 1826: 1825: 1824: 1819: 1811: 1806: 1798: 1793: 1785: 1780: 1772: 1767: 1759: 1754: 1746: 1741: 1733: 1728: 1720: 1715: 1707: 1702: 1694: 1689: 1681: 1676: 1668: 1663: 1655: 1650: 1642: 1637: 1628: 1627: 1622: 1613: 1612: 1607: 1594: 1566: 1564: 1552: 1549: 1490: 1487: 1485:Further reading 1480: 1477: 1468: 1465: 1456: 1453: 1444: 1443: 1433: 1431: 1417: 1416: 1412: 1405: 1401: 1353: 1352: 1343: 1305: 1304: 1300: 1290: 1288: 1287:on 2 March 2012 1250: 1249: 1200: 1190: 1188: 1178: 1177: 1173: 1164: 1153: 1146: 1131: 1130: 1126: 1116: 1114: 1110: 1103: 1098: 1097: 1082: 1072: 1070: 1066: 1065: 1058: 1048: 1046: 1038: 1037: 1030: 1025: 990: 913: 889: 881:sexual maturity 842: 802: 785: 765:Lesser Antilles 709: 688:Unusual erosion 660:parietal callus 579: 538:Cittarium picum 511: 484:Panulirus argus 451:, or simply as 336:Livona picoides 297: 286: 280: 267: 155: 104: 102: 101: 100: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 39: 31: 30: 27: 17: 12: 11: 5: 1864: 1862: 1854: 1853: 1848: 1843: 1838: 1828: 1827: 1821: 1820: 1818: 1817: 1804: 1791: 1778: 1765: 1752: 1739: 1726: 1713: 1700: 1687: 1674: 1661: 1648: 1635: 1630:Cittarium pica 1620: 1604: 1602: 1600:Cittarium pica 1596: 1595: 1590: 1584: 1583: 1578: 1573: 1562:Gastropods.com 1556:Cittarium pica 1548: 1547:External links 1545: 1544: 1543: 1486: 1483: 1482: 1481: 1478: 1471: 1469: 1466: 1459: 1457: 1455:Apertural view 1454: 1447: 1442: 1441: 1410: 1399: 1341: 1298: 1255:Cittarium pica 1198: 1171: 1151: 1144: 1124: 1080: 1056: 1027: 1026: 1024: 1021: 1017:Cittarium pica 1013:Cittarium pica 1001:Cittarium pica 989: 986: 973:Cittarium pica 952:. The sessile 927:Cittarium pica 912: 909: 888: 887:Feeding habits 885: 845:Cittarium pica 841: 838: 826:Cittarium pica 814:Cittarium pica 801: 798: 793:Cittarium pica 784: 781: 745:Cayman Islands 708: 705: 690: 689: 640:Cittarium pica 599:Cittarium pica 587:Cittarium pica 578: 575: 546:Cittarium pica 518:Vetigastropoda 514:Cittarium pica 510: 507: 467:Cittarium pica 445:Cittarium pica 388:Cittarium pica 382: 381: 380: 379: 365: 357: 344:Meleagris pica 341: 333: 317: 306: 305: 299: 298: 287: 282:Cittarium pica 276: 275: 269: 268: 261: 259: 255: 254: 247: 243: 242: 237: 233: 232: 227: 223: 222: 217: 213: 212: 210:Vetigastropoda 207: 203: 202: 197: 193: 192: 187: 183: 182: 177: 173: 172: 167: 163: 162: 149: 148: 146:Cittarium pica 137: 136: 128: 127: 120:Cittarium pica 115: 114: 106: 105: 97: 96: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 40: 35: 34: 28: 25:Cittarium pica 15: 13: 10: 9: 6: 4: 3: 2: 1863: 1852: 1849: 1847: 1844: 1842: 1839: 1837: 1834: 1833: 1831: 1814: 1809: 1805: 1801: 1796: 1792: 1788: 1783: 1779: 1775: 1770: 1766: 1762: 1757: 1753: 1749: 1744: 1740: 1736: 1731: 1727: 1723: 1718: 1714: 1710: 1705: 1701: 1697: 1692: 1688: 1684: 1679: 1675: 1671: 1666: 1662: 1658: 1653: 1649: 1645: 1640: 1636: 1631: 1625: 1621: 1616: 1610: 1606: 1605: 1603: 1601: 1597: 1593: 1588: 1582: 1579: 1577: 1574: 1563: 1559: 1557: 1551: 1550: 1546: 1540: 1536: 1531: 1526: 1521: 1516: 1512: 1508: 1505:(9): e12675. 1504: 1500: 1499: 1494: 1489: 1488: 1484: 1475: 1470: 1463: 1458: 1451: 1446: 1429: 1425: 1421: 1414: 1411: 1408: 1403: 1400: 1395: 1391: 1386: 1381: 1376: 1371: 1367: 1363: 1362: 1357: 1350: 1348: 1346: 1342: 1337: 1333: 1329: 1325: 1321: 1317: 1313: 1311: 1302: 1299: 1286: 1282: 1278: 1274: 1270: 1266: 1262: 1258: 1256: 1247: 1245: 1243: 1241: 1239: 1237: 1235: 1233: 1231: 1229: 1227: 1225: 1223: 1221: 1219: 1217: 1215: 1213: 1211: 1209: 1207: 1205: 1203: 1199: 1186: 1182: 1175: 1172: 1168: 1162: 1160: 1158: 1156: 1152: 1147: 1145:92-5-103129-0 1141: 1137: 1136: 1128: 1125: 1109: 1102: 1095: 1093: 1091: 1089: 1087: 1085: 1081: 1069: 1063: 1061: 1057: 1045: 1041: 1035: 1033: 1029: 1022: 1020: 1018: 1014: 1010: 1006: 1002: 997: 995: 987: 985: 983: 982: 976: 974: 970: 969: 965: 961: 960: 955: 951: 947: 946: 941: 940: 935: 928: 924: 923: 917: 910: 908: 906: 902: 898: 894: 886: 884: 882: 878: 874: 870: 866: 862: 858: 854: 853:fertilisation 850: 846: 839: 833: 827: 822: 818: 815: 811: 807: 799: 794: 789: 782: 780: 778: 774: 770: 766: 762: 758: 754: 750: 746: 742: 738: 734: 730: 726: 722: 718: 714: 706: 704: 702: 701: 696: 687: 686: 685: 682: 680: 675: 673: 669: 665: 661: 657: 653: 649: 645: 641: 638:The shell of 636: 634: 630: 626: 622: 617: 615: 611: 607: 600: 595: 588: 583: 576: 574: 572: 568: 564: 563: 558: 554: 549: 547: 543: 539: 535: 531: 527: 524:, presenting 523: 519: 515: 508: 506: 504: 500: 496: 495: 490: 486: 485: 480: 479:spiny lobster 476: 472: 468: 464: 462: 458: 454: 450: 446: 442: 438: 433: 431: 427: 424: 420: 417: 414: 410: 406: 402: 398: 394: 390: 389: 377: 373: 369: 366: 361: 358: 354: 350: 345: 342: 339:(Gould, 1853) 337: 334: 330: 326: 321: 318: 313: 310: 309: 307: 304: 300: 295: 291: 285: 283: 277: 274: 273:Binomial name 270: 266: 265: 260: 257: 256: 253: 252: 248: 245: 244: 241: 238: 235: 234: 231: 228: 226:Superfamily: 225: 224: 221: 218: 215: 214: 211: 208: 205: 204: 201: 198: 195: 194: 191: 188: 185: 184: 181: 178: 175: 174: 171: 168: 165: 164: 159: 154: 150: 147: 143: 138: 134: 129: 125: 121: 116: 112: 107: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 38: 26: 22: 19: 1599: 1565:. Retrieved 1561: 1555: 1502: 1496: 1432:. Retrieved 1428:the original 1423: 1413: 1402: 1368:(1): e8776. 1365: 1359: 1319: 1315: 1309: 1301: 1289:. Retrieved 1285:the original 1264: 1260: 1254: 1189:. Retrieved 1184: 1174: 1134: 1127: 1115:. Retrieved 1108:the original 1071:. Retrieved 1047:. Retrieved 1043: 1016: 1012: 1000: 998: 991: 979: 977: 972: 966: 957: 943: 937: 931: 926: 920: 890: 877:trochophores 844: 843: 825: 813: 808:and shallow 803: 792: 713:Florida Keys 710: 707:Distribution 698: 691: 683: 676: 639: 637: 623:. The large 618: 603: 598: 586: 560: 556: 552: 550: 545: 541: 537: 533: 513: 512: 492: 482: 477:, after the 471:economically 466: 465: 460: 456: 452: 448: 447:is known as 444: 434: 401:magpie shell 400: 396: 387: 386: 385: 367: 359: 343: 335: 319: 311: 281: 279: 264:C. pica 263: 262: 250: 145: 119: 24: 18: 1795:SeaLifeBase 1743:NatureServe 1691:iNaturalist 1624:Wikispecies 761:Puerto Rico 674:specimens. 614:turbiniform 573:and named. 542:Livona pica 532:. Woodring 499:overfishing 489:queen conch 437:West Indies 393:common name 363:Gould, 1853 320:Livona pica 126:at the top 1830:Categories 1567:15 January 1291:13 October 1117:13 October 1073:13 October 1023:References 964:polychaete 956:gastropod 857:externally 840:Life cycle 806:intertidal 763:, and the 721:Costa Rica 697:, such as 656:Trochoidea 625:body whorl 610:trochiform 522:Trochoidea 487:) and the 368:Turbo pica 230:Trochoidea 206:Subclass: 200:Gastropoda 1836:Cittarium 1748:2.1256165 1434:6 October 1310:Cittarium 988:Human use 968:Spirorbis 950:commensal 893:herbivore 849:dioecious 733:Venezuela 664:umbilicus 644:umbilicus 633:columella 567:Oligocene 557:Cittarium 553:Cittarium 530:Tegulidae 475:Caribbean 453:caracoles 426:Tegulidae 416:gastropod 409:sea snail 258:Species: 251:Cittarium 240:Tegulidae 176:Kingdom: 170:Eukaryota 124:operculum 1709:11253878 1615:Q3171877 1609:Wikidata 1539:20844767 1498:PLoS ONE 1394:20098740 1361:PLoS ONE 1281:86363886 1191:19 April 1049:14 April 954:vermetid 932:A small 901:detritus 867:produce 810:subtidal 769:Barbados 729:Colombia 668:Juvenile 652:nacreous 648:aperture 509:Taxonomy 372:Linnaeus 349:Linnaeus 325:Linnaeus 303:Synonyms 290:Linnaeus 236:Family: 220:Trochida 190:Mollusca 186:Phylum: 180:Animalia 166:Domain: 1683:2293038 1530:2937038 1507:Bibcode 1385:2808249 1336:1307109 1005:Bermuda 905:erosion 865:gametes 812:zones. 800:Habitat 783:Ecology 777:Bermuda 749:Jamaica 737:Bahamas 421:in the 419:mollusk 405:species 403:, is a 246:Genus: 216:Order: 196:Class: 98:↓ 1813:419449 1774:419449 1761:164121 1722:113340 1696:133677 1670:482715 1644:201583 1537:  1527:  1392:  1382:  1334:  1279:  1187:. BBSR 1142:  934:limpet 873:larvae 755:, the 743:, the 725:Panama 717:Mexico 672:mature 621:whorls 571:France 457:quigua 449:bulgao 441:whelks 423:family 413:marine 1808:WoRMS 1787:90935 1735:70011 1704:IRMNG 1332:JSTOR 1277:S2CID 1111:(PDF) 1104:(PDF) 897:algae 869:yolky 753:Haiti 679:whorl 606:shell 534:et al 526:nacre 461:cigua 430:shell 142:shell 1800:2879 1769:OBIS 1756:NCBI 1730:ITIS 1678:GBIF 1657:VMPF 1639:BOLD 1569:2019 1535:PMID 1436:2010 1390:PMID 1293:2009 1193:2010 1140:ISBN 1119:2009 1075:2009 1051:2010 861:eggs 771:and 741:Cuba 731:and 612:and 604:The 501:and 411:, a 395:the 376:1758 353:1758 329:1758 294:1758 37:PreꞒ 1717:ISC 1665:EoL 1652:CoL 1525:PMC 1515:doi 1380:PMC 1370:doi 1324:doi 1269:doi 1265:153 847:is 629:lip 399:or 144:of 1832:: 1810:: 1797:: 1784:: 1771:: 1758:: 1745:: 1732:: 1719:: 1706:: 1693:: 1680:: 1667:: 1654:: 1641:: 1626:: 1611:: 1560:. 1533:. 1523:. 1513:. 1501:. 1495:. 1422:. 1388:. 1378:. 1364:. 1358:. 1344:^ 1330:. 1320:76 1318:. 1314:. 1275:. 1263:. 1259:. 1201:^ 1183:. 1154:^ 1083:^ 1059:^ 1042:. 1031:^ 984:. 936:, 779:. 759:, 751:, 747:, 739:, 727:, 723:, 719:, 666:. 505:. 463:. 432:. 391:, 374:, 351:, 327:, 292:, 87:Pg 1571:. 1558:" 1554:" 1541:. 1517:: 1509:: 1503:5 1438:. 1396:. 1372:: 1366:5 1338:. 1326:: 1295:. 1271:: 1195:. 1148:. 1121:. 1077:. 1053:. 491:( 481:( 355:) 347:( 331:) 323:( 296:) 288:( 92:N 82:K 77:J 72:T 67:P 62:C 57:D 52:S 47:O 42:Ꞓ

Index

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operculum

shell
Scientific classification
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Eukaryota
Animalia
Mollusca
Gastropoda
Vetigastropoda
Trochida
Trochoidea
Tegulidae
Cittarium
Binomial name
Linnaeus
1758

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