825:
1015:
509:
122:
689:
102:
463:
964:
392:
1032:
998:
593:
981:
669:
each stage of development. The warmer the temperature is, the quicker the life cycle, and the cooler the temperature is the slower the life cycle. If temperatures dip below a threshold temperature for an extended period of time, clerids and most other insects will have growth and developmental progress arrested. Like all beetles, Cleridae follow a
1049:
733:
stage can last from 6 weeks to one year depending on the need to overwinter, and how short the overall life cycle is for a particular species. A majority of clerid species pupate in earthen cells which are made from soil and certain enzymes secreted from their mouths. The rest remain in pupal cells.
418:
in shape and range from 3–24 millimetres (0.12–0.94 in) in length. Their entire bodies are covered with bristly hairs and many display an ornate body color pattern. These often brightly color patterns can be red, yellow, orange, or blue. The antennae are clubbed at the tip for most species, but
876:
agent for these pests. Researchers and forestry officials have used bark beetle aggregation pheromones to attract the checkered beetle to specific trees. This causes the bark beetles to be overwhelmed, extensively preyed upon by the clerid beetles, and typically eliminated. There is also additional
704:
takes place while the female feeds, because females need a large amount of food for egg development. The female lays her eggs 36–72 hours after copulation. The eggs are laid in between pieces of bark on wood-borer-infested trees or under stones in the soil. She may lay 28–42 eggs at a time. For the
484:
formula is 5–5–5, meaning that on each of the front, middle and hind legs there are 5 tarsomeres (individual subsegments of the feet/tarsi). One or more of these subsegments on each leg is typically lobed, and the 4th tarsomere is normally difficult to distinguish. Furthermore, an important feature
668:
The general life cycle of clerids has been known to last anywhere from 35 days to more than 3 years, and is strongly dependent on the life cycle of their prey. While the life cycle can vary in length between genus and species, temperature is also a major determinant in the length of time spent in
563:
into the bark. Nest robbing species typically hunt termite, bee, and wasp larvae, and one particular species has been noted to prey primarily on grasshopper egg masses. Not all nest robbing species actively hunt live prey, some species for example prefer to feed only on dead honey bee larvae and
728:
area of the last body segment. Immediately after birth, they start searching for food close to where they hatched. They feed on wood-borer insects on trees, or feed on their species' substrate or prey of choice. Feeding is the main purpose of the larval stage to prepare for pupation. Once their
576:
In general, the bulk of adult
Cleridae feed mainly on other adult beetles while the larvae stage feed on other beetle larvae. Some checkered beetles are known to have an extremely voracious appetite with some larvae able to consume "several times their own body weight" in a day.
947:
580:
Although most species of checkered beetles are predaceous in nature, some are scavengers and others have been found feeding on flower pollen. Because of the checkered beetles predaceous nature and insatiable appetite, they are often key players in the
525:
Cleridae can be found in the
Americas, Africa, Europe, the Middle East and even in Australia. There are approximately 3,500 species in the world and about 500 species in North America. Due to this wide distribution there are many different
723:
When larvae hatch from their eggs, they are either red or yellow. Their bodies have a slender and flat appearance with short legs due to their minimal movement. The larvae are covered in hair and have two horn-type projections on the
820:
or PMI. Although the checkered beetle is not the most significant insect on carrion, the beetle's predaceous nature and its ability to reproduce in carrion that is exposed to the environment provides some forensic importance.
572:
The
Cleridae contains many species of predaceous beetles that feed on other beetles and beetle larvae in their natural habitat. The most common prey item for checkered beetles are bark beetles and wood boring beetles.
552:. These "tree living species" are found in forests across the world with various climates and an array of easily preyed upon insects. They seek protection under the bark and hunt for other insects above and below the
379:) and are found infesting various food products. Research efforts related to Cleridae have focused primarily on using certain species as biological controls. This is a very effective technique for controlling
780:, especially those products that are stored unwrapped for long periods of time. Adults feed on the surface of the products, while the larvae damage the meat by boring down usually in the fatty parts.
488:
The second defining characteristic of the family
Cleridae is that clerids never have eversible vesicles (small usually hidden balloon-like structures thought to be scent glands) on their
4674:"Attraction of bark beetle predator, Thanasimus undatulus (Coleoptera: Cleridae), to pheromones of the spruce beetle and two secondary bark beetles (Coleoptera: Scolytidae)"
4591:
7086:
6960:
559:
The third type of clerid beetles is the "nest robbing species" which live in shrubbery and in trees. Unlike the tree living species, these species do not actually
6999:
4349:
485:
that eliminates many other families of beetles is that clerids' front coxae (base of the leg) expose the second segment of the legs known as the trochanter.
6758:
6934:
681:, and the pupa shedding its skin to emerge as an adult. The larvae of the majority of the known species of Cleridae feed upon the eggs and young of
919:. The following list of tribes and selected genera is thus preliminary. Some notable species are also listed. The oldest members of the family are
459:
suture (found under the pronotal shield) is not present. To further identify Clerid beetles, a few additional characteristics need to be examined.
6973:
842:
There is ongoing research with some clerid species. Forensic research is limited because of their late arrival on carrion, but members such as
4178:
784:
has been recorded feeding upon a large variety of items ranging from hides and dried figs to
Egyptian mummies. In addition, products such as
4871:
345:. Checkered beetles range in length between 3 and 24 millimetres (0.12 and 0.94 in). Cleridae can be identified based on their 5–5–5
4673:
419:
others can be "clubbed, saw-tooth, or thread-like." The pronotum region is nearly cylindrical and characteristically narrower than the
4622:
4539:
4266:
4217:
4146:
4599:
816:
is attracted towards carrion in the later stages of decomposition, so its arrival on carrion can help provide an estimate for the
4294:. Misc. Publ. 1426. U.S. Forestry Service. Washington D.C: U.S. Department of Agriculture, Forest Service. 1985. pp. 274–275
585:
of other insects. The checkered beetles have also developed a unique adaptation to aid in their quest for prey. The beetles use
500:
which sometimes has these glands. This trait is very important in correctly differentiating checkered beetles from
Melyridae.
4473:
6978:
6986:
4715:"Ecological notes on species of Cleridae (Insecta: Coleoptera) associated with the prairie flora of central North America"
618:
are attracted to dry carrion and other decomposing animal matter such as bones and skin as well as various meat products.
4746:"The oldest known clerid fossils from the Middle Jurassic of China, with a review of Cleridae systematics (Coleoptera)"
7081:
6843:
6820:
4039:
3776:
3753:
3710:
3487:
3474:
2897:
2817:
1340:
4256:
4623:"Diptera and Coleoptera of potential forensic importance in southeastern Brazil: relative abundance and seasonality"
800:
Since clerids are predaceous in nature, they have been found feeding on fly larvae as well as the skin and bones of
7004:
6913:
4356:
4313:
849:
4785:"The dating of the fourth volume of Guillaume-Antoine Olivier's "Entomologie, ou histoire naturelle des insectes""
1048:
6926:
3002:
2163:
1053:
769:
376:
218:
121:
4864:
1019:
1137:
968:
729:
larval stage is complete the tree dwelling species make their way to the bottom of the tree to pupate. The
6773:
2144:
2097:
397:
364:
of trees. Larvae are predaceous and feed vigorously before pupation and subsequently emergence as adults.
3761:
609:
The diversity of checkered beetle's feeding habits is quite evident when different species are examined.
341:
feeders. Clerids have elongated bodies with bristly hairs, are usually bright colored, and have variable
7035:
6848:
3013:
2875:
3718:
3535:
2675:
2282:
1797:
7053:
3853:
3823:
3565:
3459:
3339:
3269:
2752:
2715:
2705:
1947:
1737:
1687:
904:
660:, in which larvae feed on the pollen of flowering plants and adults prey upon grasshoppers and wasps.
545:. These species are found in moist, sunny environments where flowering plants are found in abundance.
7014:
6882:
6010:
3883:
3299:
2905:
2892:
2812:
2168:
1847:
1697:
1547:
1235:
940:
861:
857:
844:
817:
701:
3813:
3705:
3685:
3665:
3615:
3482:
3469:
3111:
2725:
2377:
1967:
1807:
1617:
1418:
1408:
985:
824:
734:
Adult beetles emerge from pupation and spend a variable time of their life maturing, and eventually
6229:
4857:
4327:
4059:
3923:
3863:
3833:
3803:
3793:
3738:
3728:
3575:
3555:
3545:
3515:
3379:
3219:
3121:
3071:
3041:
2997:
2954:
2742:
2503:
2442:
2193:
2176:
2130:
2067:
1837:
1567:
1537:
1468:
1378:
1285:
1092:
1036:
368:
4017:
4006:
3843:
3695:
3635:
3505:
3495:
3449:
3369:
3329:
3259:
3141:
2658:
2580:
2422:
2357:
2213:
2153:
2047:
1987:
1977:
1917:
1757:
1707:
1637:
1627:
1577:
1448:
1315:
1275:
1132:
1122:
1014:
451:(base of the leg), do not divide the first and second abdominal/ventral plates which are known as
4765:
4650:
4563:
4307:
4242:
Checkered
Beetles: Illustrated Key to the Cleridae and Thanerocleridae of the Western Palaearctic
4034:
3984:
3933:
3913:
3748:
3645:
3625:
3595:
3525:
3359:
3319:
3279:
3101:
2959:
2926:
2802:
2772:
2651:
2644:
2617:
2560:
2553:
2523:
2473:
2456:
2367:
2299:
2137:
2017:
1927:
1867:
1767:
1727:
1717:
1677:
1657:
1647:
1597:
1557:
1438:
1398:
1358:
1335:
1305:
1215:
1145:
1007:
951:
927:
921:
894:
873:
682:
602:
582:
261:
116:
4658:
4571:
4085:
3963:
3903:
3893:
3873:
3785:
3771:
3181:
2792:
2627:
2533:
2493:
2483:
2402:
2333:
2245:
2183:
2057:
1957:
1897:
1857:
1747:
1667:
1607:
1458:
1245:
1112:
508:
7040:
4074:
3605:
3349:
3239:
3151:
2885:
2835:
2695:
2607:
2600:
2313:
2110:
2007:
1777:
1518:
1428:
1388:
1295:
1265:
1205:
7076:
7022:
6869:
6498:
5915:
4814:
4642:
4443:
4262:
4213:
4174:
4142:
3389:
3249:
3209:
3161:
2685:
2570:
2463:
2449:
2387:
2323:
2306:
2289:
2252:
2027:
1528:
1368:
1348:
1255:
1168:
1156:
1002:
829:
694:
513:
342:
6965:
4688:
3675:
2855:
2845:
2825:
2762:
1887:
1877:
1478:
1225:
907:, or splitting off the Thaneroclerinae as distinct family, or circumscribing the Korynetinae
7027:
6595:
6374:
6369:
6292:
6050:
5066:
4804:
4794:
4757:
4634:
4555:
4433:
3429:
3419:
3289:
3008:
2412:
1997:
1488:
1198:
764:
725:
597:
468:
432:
326:
4173:(illustrated and revised ed.). New York: Houghton Mifflin Harcourt. pp. 208–209.
1188:
6284:
6237:
6082:
6066:
5742:
5502:
5471:
5386:
5304:
5195:
5163:
4844:
4391:
4048:
3092:
932:
747:
275:
480:
Clerid beetles have unique legs that help to distinguish them from other families. Their
688:
462:
101:
6921:
6908:
6811:
6687:
6638:
6493:
6300:
5968:
5952:
5891:
5786:
5757:
5685:
5649:
5633:
5549:
5494:
5399:
5296:
5145:
5107:
5087:
4983:
4834:
4809:
4784:
4638:
4559:
4206:
3975:
1059:
963:
670:
448:
391:
346:
7070:
6669:
6473:
6464:
6205:
6176:
6168:
6160:
6134:
6109:
6005:
5992:
5931:
5849:
5752:
5747:
5628:
5562:
5533:
5517:
5481:
5451:
5404:
5358:
5325:
5241:
5233:
5204:
5042:
3409:
3171:
3052:
3021:
1024:
909:
899:
553:
481:
372:
361:
357:
268:
59:
4769:
4654:
4567:
4438:
946:
6646:
6570:
6316:
6308:
6263:
6247:
6197:
6118:
6074:
6026:
5944:
5857:
5726:
5713:
5693:
5654:
5554:
5541:
5512:
5446:
5434:
5330:
5287:
5270:
5262:
5212:
5050:
5020:
4951:
4940:
4924:
4355:. Colorado State University Department of Entomology. pp. 1–15. Archived from
3031:
2865:
2782:
2666:
2634:
2433:
2347:
2262:
2077:
1031:
990:
877:
research being done pertaining to the impact of clerids on pollination in flowers.
777:
634:
254:
247:
193:
4485:
4414:
4289:
6874:
4168:
6991:
6947:
6825:
6805:
6732:
6703:
6677:
6664:
6604:
6562:
6546:
6538:
6530:
6514:
6506:
6442:
6400:
6392:
6361:
6276:
6255:
6221:
6213:
6126:
6058:
6042:
6034:
5997:
5984:
5960:
5878:
5870:
5820:
5812:
5770:
5734:
5705:
5677:
5672:
5588:
5583:
5575:
5525:
5461:
5456:
5370:
5335:
5320:
5312:
5228:
5220:
5037:
3439:
3309:
3229:
3081:
2223:
2203:
1907:
1102:
997:
890:
735:
647:
456:
380:
34:
592:
6659:
6620:
6612:
6554:
6417:
6379:
6332:
6324:
6144:
6092:
5976:
5907:
5778:
5762:
5718:
5659:
5623:
5603:
5567:
5476:
5466:
5340:
5074:
5058:
4799:
4714:
4026:
3995:
3655:
3585:
3399:
3061:
2967:
2732:
2590:
2543:
2236:
2102:
2087:
2037:
973:
915:
773:
768:, commonly known as the red-legged ham beetle, is of particular importance in
717:
707:
639:
620:
586:
334:
330:
240:
79:
44:
6796:
4849:
868:
industry because in large numbers they can cause damage and kill live trees.
427:. Their elytra have tiny pits or depressions, and never expose more than two
6895:
6724:
6716:
6708:
6695:
6651:
6450:
6425:
6353:
6181:
6152:
5836:
5667:
5641:
5593:
5507:
5409:
5378:
5350:
5183:
5158:
5153:
5134:
5099:
4988:
4971:
4963:
4912:
4900:
3943:
3188:
3131:
2944:
2934:
2915:
2120:
1508:
1498:
980:
936:
805:
656:
556:. The primary source of prey for these bark living hunters is bark beetles.
497:
452:
444:
424:
423:(special hardened front wings), while the head is as wide or wider than the
371:. Some species are occasionally found on carrion in the later dry stages of
353:, and the absence of a special type of vesicle. Female Cleridae lay between
311:
203:
183:
153:
133:
84:
4818:
4447:
4745:
4646:
4419:(De Geer, 1775) (Coleoptera; Cleridae) associated with pet food in Brazil"
6856:
6790:
6628:
6578:
6522:
6481:
6430:
6087:
6018:
5899:
5611:
5417:
5249:
5082:
5029:
5008:
4918:
3201:
2987:
2977:
2340:
1937:
1827:
1074:
1041:
956:
625:
611:
527:
518:
493:
473:
428:
402:
350:
286:
233:
74:
69:
54:
49:
39:
6835:
856:
is a predator of bark beetles. Some species of bark beetles such as the
746:
overwinter inside the wood-borer-infested trees and oviposit during the
6345:
6337:
6189:
5923:
5844:
5828:
4789:
4761:
1787:
1587:
1178:
1082:
801:
643:
615:
538:
533:
Many of the species are known as "flower visitors", that prey on other
489:
319:
89:
64:
4839:
4208:
Forensic
Entomology: The Utility of Arthropods in Legal Investigations
6952:
6861:
5936:
5796:
5429:
4906:
4894:
4881:
3953:
2513:
2272:
1325:
1161:
886:
865:
651:
560:
542:
534:
420:
338:
318:. The family Cleridae has a worldwide distribution, and a variety of
307:
173:
163:
143:
6939:
6767:
6900:
4524:(9). Berkeley, California: University of California Press: 249–262.
7048:
1817:
1047:
1030:
1013:
996:
979:
962:
945:
823:
739:
713:
687:
674:
591:
549:
507:
461:
390:
4141:(7th ed.). Belmont: Brooks/Cole. pp. 365–400, 428–429.
789:
785:
730:
678:
629:
628:
areas where bark beetle species constitute their main source of
415:
6771:
5180:
5131:
5005:
4937:
4853:
4554:(1). Institute of Zoology, Chinese Academy of Sciences: 55–62.
496:. This characteristic distinguishes them from a similar family
333:
and feed on other beetles and larvae; however other genera are
4255:
Bellows, Thomas S.; Fisher, T. W.; Caltagirone, L. E. (1999).
6887:
893:, though some genera still defy easy classification. Several
1068:
5252:(furniture beetles, death watch beetles, spider beetles)
4681:
4139:
548:
Another habitat commonly inhabited by clerid beetles is
4244:. Weikersheim, Germany: Margraf Verlag. pp. 12–15.
808:
because of their food choice, but some species such as
897:
schemes exist, recognizing for example a group around
383:
due to the voracious appetite of many clerid species.
4672:
Poland, Therese M.; Borden, John H. (December 1997).
712:
this occurs in late summer or early fall to give the
677:, which grows and feeds, changing its skin to form a
447:. Key characteristics of Polyphaga are that the hind
4598:. University of California Riverside. Archived from
4484:(1). University of Wisconsin-Madison. Archived from
6780:
6686:
6637:
6603:
6594:
6472:
6463:
6416:
6391:
6275:
6246:
6143:
6117:
6108:
5890:
5869:
5795:
5602:
5493:
5349:
5295:
5286:
5261:
5203:
5194:
5019:
4518:
University of California Publications in Entomology
4137:Johnson, Norman F.; Triplehorn, Charles A. (2004).
685:, while the adults feed on the adult bark beetles.
589:to help them locate, kill, and consume their prey.
4205:
4621:Souza, Adriana; Linhares, Aricio (28 June 2008).
4338:. Agriculture Canada Publication 1861/E. 208–211.
6129:(jewel beetles, or metallic wood-boring beetles)
4512:Linsley, Earle G. (1936). "Studies in the genus
4204:Byrd, Jason H. (2001). James L., Castner (ed.).
4835:Atlas of checkered beetles (Cleridae) of Russia
4261:. San Diego: Academic Press. pp. 429–430.
716:enough time for proper growth before having to
414:Generally, checkered beetles are elongated and
5852:(true weevils, bark beetles, ambrosia beetles)
4865:
4744:Kolibáč, JiĹ™Ă; Huang, Diying (October 2016).
4336:Checklist of the Beetles of Canada and Alaska
4170:A Field Guide to the Beetles of North America
530:in which the checkered beetles can be found.
8:
6095:(ironclad beetles, cylindrical bark beetles)
4392:"A guide to the Cleridae of Atlantic Canada"
4235:
4233:
4231:
4229:
4413:Gredilha, R.; Lima, A. F. (February 2007).
848:have been researched as a possible role in
6768:
6600:
6469:
6114:
5292:
5200:
5191:
5177:
5128:
5016:
5002:
4934:
4872:
4858:
4850:
4840:Roland Gerstmeier publications on Cleridae
4585:
4583:
4581:
4544:(Coleoptera: Cleridae) in Southern Norway"
100:
20:
6759:List of subgroups of the order Coleoptera
5839:(straight snout weevils, New York weevil)
4808:
4798:
4592:"Chapter 7 Pests of Stored Food Products"
4507:
4505:
4503:
4437:
4132:
4130:
4128:
4126:
4124:
4122:
4120:
4118:
6266:(cicada beetle, cicada parasite beetles)
4633:(1). Royal Entomological Society: 8–12.
792:can become infested, but not destroyed.
4467:
4465:
4463:
4461:
4459:
4457:
4385:
4383:
4381:
4379:
4377:
4212:(2nd ed.). Boca Raton: CRC Press.
4162:
4160:
4158:
4114:
443:Clerid beetles fall under the suborder
6356:(false fireflies, long-lipped beetles)
4305:
4199:
4197:
889:of Cleridae are divided among several
4533:
4531:
4474:"Know Your Friends Checkered Beetles"
4350:"Clerid Beetles- Voracious Predators"
367:Clerids have a minor significance in
7:
7087:Taxa named by Pierre André Latreille
7054:396787A7-5C40-1B48-FF1D-FD7FFD62F14C
7015:c93ca142-b3f9-46f4-a408-3911cdf55287
6927:960dc211-c8d1-4463-a368-4664a3a8697e
6382:(false metallic wood-boring beetles)
4328:"Family Cleridae: checkered beetles"
329:and feeding habits. Most genera are
4390:Majka, Christopher (20 June 2006).
1513:Chevrolat in Guérin-Ménéville, 1839
673:life cycle: the egg hatches into a
4639:10.1111/j.1365-2915.1997.tb00284.x
4590:Ebeling, Walter (23 August 2002).
4560:10.1111/j.1744-7917.1998.tb00298.x
872:has been researched as a possible
360:at a time predominately under the
111:described in the mid-19th century
14:
4627:Medical and Veterinary Entomology
2939:Perty in Spix & Martius, 1830
804:. Most clerids are not useful in
632:. The primary source of prey for
272:Jacquelin du Val, 1861 (disputed)
5993:Perimylopidae, or Promecheilidae
5578:(minute brown scavenger beetles)
4516:Spinola (Coleoptera-Cleridae)".
650:larvae. One of the more diverse
375:. Also, some species are pests (
294:
280:
120:
6208:(variegated mud-loving beetles)
4439:10.1590/S1519-69842007000100026
4713:Mawdsley, Jonathan R. (2002).
4258:Handbook of biological control
705:longer-living species such as
1:
6037:(narrow-waisted bark beetles)
5729:(parasitic flat bark beetles)
5680:(short-winged flower beetles)
5536:(lady beetles, or God's cows)
5273:(tooth-necked fungus beetles)
2267:Fairmaire & Germain, 1861
314:. They are commonly known as
6549:(sand-loving scarab beetles)
5939:(minute tree-fungus beetles)
5765:(silvanid flat bark beetles)
4719:The Great Lakes Entomologist
4426:Brazilian Journal of Biology
1300:Schenkling in Sjöstedt, 1908
1150:Sedlacek & Winkler, 1975
738:. Sexually mature adults or
6698:(primitive carrion beetles)
6654:(minute mud-loving beetles)
6517:(bumble bee scarab beetles)
6287:(soft-bodied plant beetles)
6258:(soft bodied plant beetles)
5061:(predaceous diving beetles)
4845:Cleridae of Atlantic Canada
4240:Gerstmeier, Ronald (1998).
3540:Gerstmeier & Kuff, 1992
3530:Gerstmeier & Kuff, 1992
3056:(sometimes in Korynetinae)
2670:(including Phyllobaeninae)
2437:(sometimes in Korynetinae)
2240:(sometimes in Korynetinae)
1054:Dermestoides sanguinicollis
7103:
6533:(scavenger scarab beetles)
6525:(enigmatic scarab beetles)
5698:Lamingtonium binnaberrense
5412:(soft-wing flower beetles)
4291:Insects of eastern forests
4167:White, Richard E. (1998).
3948:Laporte de Castelnau, 1832
3723:Laporte de Castelnau, 1836
3590:Wolcott & Chapin, 1918
3434:Fischer von Waldheim, 1829
3096:(tentatively placed here)
2910:Laporte de Castelnau, 1836
2518:Laporte de Castelnau, 1836
2417:Laporte de Castelnau, 1836
1792:Laporte de Castelnau, 1836
1742:Laporte de Castelnau, 1836
943:in Inner Mongolia, China.
850:integrated pest management
833:attacking bark beetle prey
6756:
6672:(water scavenger beetles)
6405:Rhinorhipus tamborinensis
6184:(long-toed water beetles)
6077:(polypore fungus beetles)
5979:(palm and flower beetles)
5963:(tumbling flower beetles)
5910:(ant-like flower beetles)
5688:(lined flat bark beetles)
5662:(pleasing fungus beetles)
5557:(handsome fungus beetles)
5190:
5176:
5141:
5127:
5102:(burrowing water beetles)
5015:
5001:
4947:
4933:
4889:
4800:10.3897/zookeys.734.22901
4348:Costello, Sheryl (2003).
4044:(Clerinae? Hydnocerinae?)
4013:(Clerinae? Hydnocerinae?)
3003:red-shouldered ham beetle
2427:Wolcott & Dybas, 1947
2164:Trogodendron fasciculatum
770:stored product entomology
759:Stored product entomology
466:5 rear leg tarsomeres of
377:stored product entomology
322:and feeding preferences.
231:
226:
117:Scientific classification
115:
108:
99:
23:
6719:(feather-winged beetles)
6053:(false longhorn beetles)
6000:(jugular-horned beetles)
5947:(false darkling beetles)
5918:(cryptic fungus beetles)
5737:(shining flower beetles)
5077:(crawling water beetles)
4976:Sikhotealinia zhiltzovae
4538:Hue, Yi (28 June 2008).
4472:Mahr, Susan (May 2000).
4394:. Chebucto Community Net
3713:& van der Wiel, 1949
3477:& van der Wiel, 1949
864:can become pests to the
504:Distribution and ecology
6200:(forest stream beetles)
6061:(synchroa bark beetles)
6021:(dead log bark beetles)
6013:(fire-coloured beetles)
5902:(ant-like leaf beetles)
5636:(silken fungus beetles)
5570:(well polished beetles)
5544:(minute fungus beetles)
5363:Acanthocnemus nigricans
4687:: 35–41. Archived from
4478:Biological Control News
4055:(Clerinae? Enopliinae?)
4002:(Clerinae? Enopliinae?)
1260:Dupont in Spinola, 1844
1020:Tarsostenus univittatus
796:Medico-legal entomology
251:Gistel, 1856 (disputed)
244:Gistel, 1856 (disputed)
6711:(round fungus beetles)
6045:(false flower beetles)
6029:(wedge-shaped beetles)
5987:(false blister beetle)
5971:(hairy fungus beetles)
5926:(conifer bark beetles)
5823:(leaf-rolling weevils)
5484:(bark-gnawing beetles)
5244:(wounded-tree beetles)
5110:(false ground beetles)
5032:(trout-stream beetles)
4542:Thanasimus formicarius
4334:: Bousquet, Y. (Ed.).
4312:: CS1 maint: others (
3404:Gray in Griffith, 1832
2318:Gray in Griffith, 1832
2145:Trichodes leucopsideus
2098:Thanasimus formicarius
2042:Gray in Griffith, 1832
1087:Jacquelin du Val, 1860
1064:
1045:
1028:
1011:
994:
977:
969:Phymatophaea guttigera
960:
903:as distinct subfamily
834:
698:
606:
522:
477:
406:
398:Enoclerus ichneumoneus
279:Chapin, 1921 (but see
109:Some checkered beetles
30:Middle Jurassic–Recent
7036:Paleobiology Database
6623:(false clown beetles)
6445:(plate-thigh beetles)
6327:(false click beetles)
6232:(water-penny beetles)
5881:(ship-timber beetles)
5860:(pine flower weevils)
5708:(root-eating beetles)
5528:(minute bark beetles)
5439:Phloiophilus edwardsi
4966:(reticulated beetles)
4783:Bousquet, Y. (2018).
4750:Systematic Entomology
4081:(Clerinae? Tillinae?)
3991:(Clerinae? Tillinae?)
3014:red-legged ham beetle
1051:
1034:
1017:
1000:
983:
966:
949:
827:
691:
595:
511:
465:
394:
6922:Fauna Europaea (new)
6501:(false stag beetles)
6486:Belohina inexpectata
6348:(net-winged beetles)
6311:(rare click beetles)
6224:(travertine beetles)
6216:(minute mud beetles)
5781:(dry-fungus beetles)
5616:Agapytho foveicollis
5373:(false skin beetles)
5236:(Jacobson's beetles)
4956:Crowsoniella relicta
4326:McNamara, J. (1991)
2700:G.R.Waterhouse, 1839
2690:C.O.Waterhouse, 1879
2169:Yellow-horned Clerid
1652:C.O.Waterhouse, 1879
870:Thanasimus undatulus
862:mountain pine beetle
858:southern pine beetle
854:Thanasimus undatulus
845:Thanasimus undatulus
818:post-mortem interval
6499:Diphyllostomatidae
5831:(primitive weevils)
5644:(flat bark beetles)
5394:(checkered beetles)
5381:(fruitworm beetles)
5069:(whirligig beetles)
2998:Necrobia ruficollis
2955:Korynetes caeruleus
2131:Trichodes alvearius
1037:Diplocladus kuwerti
683:wood-boring beetles
395:Narrow pronotum in
369:forensic entomology
325:Cleridae have many
310:of the superfamily
7082:Polyphaga families
6364:(glowworm beetles)
6069:(darkling beetles)
5916:Archeocrypticidae
5773:(palmetto beetles)
5520:(dry bark beetles)
5307:(longhorn beetles)
4762:10.1111/syen.12192
4661:on 5 January 2013.
4602:on 8 November 2015
4574:on 5 January 2013.
2960:steely blue beetle
2138:Trichodes apiarius
1065:
1046:
1029:
1012:
995:
978:
961:
952:Clerus mutillarius
874:biological control
835:
754:Forensic relevance
699:
624:spp. are found in
607:
583:biological control
523:
478:
407:
7064:
7063:
7023:Open Tree of Life
6774:Taxon identifiers
6765:
6764:
6752:
6751:
6748:
6747:
6744:
6743:
6736:
6728:
6727:(carrion beetles)
6720:
6712:
6699:
6673:
6655:
6624:
6616:
6590:
6589:
6582:
6574:
6566:
6558:
6550:
6542:
6534:
6526:
6518:
6510:
6502:
6489:
6459:
6458:
6446:
6438:
6408:
6383:
6365:
6357:
6349:
6341:
6328:
6320:
6312:
6304:
6303:(soldier beetles)
6296:
6293:Brachypsectridae
6288:
6267:
6259:
6233:
6225:
6217:
6209:
6201:
6193:
6185:
6172:
6164:
6156:
6130:
6104:
6103:
6096:
6078:
6070:
6062:
6054:
6051:Stenotrachelidae
6046:
6038:
6030:
6022:
6014:
6001:
5988:
5980:
5972:
5964:
5956:
5955:(blister beetles)
5948:
5940:
5927:
5919:
5911:
5903:
5882:
5861:
5853:
5840:
5832:
5824:
5816:
5806:
5782:
5774:
5766:
5738:
5730:
5722:
5709:
5701:
5689:
5681:
5663:
5645:
5637:
5619:
5579:
5571:
5558:
5545:
5537:
5529:
5521:
5503:Akalyptoischiidae
5485:
5442:
5425:
5413:
5395:
5382:
5374:
5366:
5316:
5308:
5282:
5281:
5274:
5253:
5245:
5237:
5224:
5216:
5172:
5171:
5149:
5123:
5122:
5119:
5118:
5111:
5103:
5095:
5078:
5070:
5062:
5054:
5046:
5033:
4997:
4996:
4979:
4967:
4959:
4415:"First record of
4180:978-0-395-91089-4
4105:
4104:
4091:
4080:
4065:
4054:
4043:
4023:
4012:
4001:
3990:
3969:
3959:
3949:
3939:
3929:
3919:
3909:
3899:
3889:
3879:
3869:
3859:
3849:
3839:
3829:
3819:
3809:
3799:
3780:
3767:
3757:
3744:
3734:
3724:
3714:
3701:
3691:
3681:
3671:
3661:
3651:
3641:
3631:
3621:
3611:
3601:
3591:
3581:
3571:
3561:
3551:
3541:
3536:Flabellotilloidea
3531:
3521:
3511:
3501:
3491:
3478:
3465:
3455:
3445:
3435:
3425:
3415:
3405:
3395:
3385:
3375:
3365:
3355:
3345:
3335:
3334:Solervicens, 1976
3325:
3315:
3305:
3295:
3285:
3275:
3265:
3255:
3245:
3235:
3225:
3215:
3194:
3177:
3167:
3157:
3147:
3137:
3127:
3117:
3107:
3087:
3077:
3067:
3047:
3037:
3027:
2993:
2983:
2973:
2950:
2940:
2921:
2911:
2901:
2881:
2880:Solervicens, 1987
2871:
2861:
2851:
2841:
2831:
2821:
2808:
2798:
2788:
2778:
2768:
2767:Motschulsky, 1845
2758:
2748:
2738:
2721:
2711:
2701:
2691:
2681:
2640:
2623:
2613:
2596:
2586:
2576:
2566:
2549:
2539:
2529:
2519:
2509:
2499:
2489:
2479:
2469:
2428:
2418:
2408:
2393:
2383:
2373:
2363:
2353:
2329:
2328:Solervicens, 1989
2319:
2295:
2278:
2268:
2258:
2229:
2219:
2209:
2199:
2189:
2159:
2126:
2116:
2093:
2083:
2073:
2063:
2053:
2043:
2033:
2023:
2013:
2003:
1993:
1983:
1973:
1963:
1953:
1943:
1933:
1923:
1913:
1903:
1893:
1883:
1873:
1863:
1853:
1843:
1833:
1823:
1813:
1803:
1798:Oodontophlogistus
1793:
1783:
1773:
1763:
1753:
1743:
1733:
1723:
1713:
1703:
1693:
1683:
1673:
1663:
1653:
1643:
1633:
1623:
1613:
1603:
1593:
1583:
1573:
1563:
1553:
1543:
1529:Eunatalis porcata
1524:
1514:
1504:
1494:
1484:
1474:
1464:
1454:
1444:
1434:
1424:
1414:
1404:
1403:Solervicens, 1997
1394:
1384:
1374:
1364:
1354:
1344:
1331:
1321:
1311:
1301:
1291:
1290:Quedenfeldt, 1885
1281:
1271:
1261:
1251:
1241:
1231:
1221:
1211:
1194:
1184:
1174:
1151:
1141:
1128:
1118:
1108:
1107:Quedenfeldt, 1885
1098:
1088:
1003:Necrobia violacea
830:Thanasimus dubius
695:Thanasimus dubius
541:and also feed on
514:Trichodes ornatus
316:checkered beetles
301:
300:
291:
284:
273:
266:
259:
252:
245:
238:
222:
16:Checkered beetles
7094:
7057:
7056:
7044:
7043:
7031:
7030:
7018:
7017:
7008:
7007:
6995:
6994:
6992:NHMSYS0001717699
6982:
6981:
6969:
6968:
6956:
6955:
6943:
6942:
6930:
6929:
6917:
6916:
6904:
6903:
6891:
6890:
6878:
6877:
6865:
6864:
6852:
6851:
6839:
6838:
6829:
6828:
6816:
6815:
6814:
6801:
6800:
6799:
6769:
6734:
6726:
6718:
6710:
6697:
6671:
6653:
6622:
6614:
6601:
6596:Staphyliniformia
6580:
6572:
6564:
6556:
6548:
6540:
6532:
6524:
6516:
6508:
6500:
6483:
6470:
6444:
6435:Declinia relicta
6432:
6402:
6381:
6375:Sinopyrophoridae
6370:Rhagophthalmidae
6363:
6355:
6347:
6339:
6326:
6318:
6310:
6302:
6294:
6286:
6285:Artematopodidae
6265:
6257:
6231:
6223:
6215:
6207:
6199:
6192:(riffle beetles)
6191:
6183:
6171:(turtle beetles)
6170:
6162:
6154:
6128:
6115:
6094:
6076:
6068:
6060:
6052:
6044:
6036:
6028:
6020:
6012:
5999:
5986:
5978:
5970:
5962:
5954:
5946:
5938:
5925:
5917:
5909:
5901:
5880:
5859:
5851:
5838:
5830:
5822:
5815:(fungus weevils)
5814:
5805:
5804:
5800:
5787:Tasmosalpingidae
5780:
5772:
5764:
5736:
5728:
5720:
5707:
5695:
5687:
5679:
5661:
5643:
5635:
5613:
5577:
5569:
5556:
5543:
5535:
5527:
5519:
5483:
5436:
5419:
5411:
5393:
5380:
5372:
5360:
5359:Acanthocnemidae
5314:
5306:
5293:
5272:
5251:
5243:
5235:
5222:
5214:
5201:
5192:
5178:
5147:
5129:
5109:
5101:
5089:
5076:
5068:
5060:
5052:
5045:(ground beetles)
5044:
5031:
5017:
5003:
4973:
4965:
4953:
4952:Crowsoniellidae
4935:
4874:
4867:
4860:
4851:
4823:
4822:
4812:
4802:
4793:(734): 137–148.
4780:
4774:
4773:
4741:
4735:
4734:
4732:
4730:
4710:
4704:
4703:
4701:
4699:
4693:
4678:
4669:
4663:
4662:
4657:. Archived from
4618:
4612:
4611:
4609:
4607:
4596:Urban Entomology
4587:
4576:
4575:
4570:. Archived from
4535:
4526:
4525:
4509:
4498:
4497:
4495:
4493:
4469:
4452:
4451:
4441:
4423:
4417:Necrobia rufipes
4410:
4404:
4403:
4401:
4399:
4387:
4372:
4371:
4369:
4367:
4361:
4354:
4345:
4339:
4324:
4318:
4317:
4311:
4303:
4301:
4299:
4286:
4280:
4279:
4277:
4275:
4252:
4246:
4245:
4237:
4224:
4223:
4211:
4201:
4192:
4191:
4189:
4187:
4164:
4153:
4152:
4134:
4089:
4078:
4063:
4052:
4038:
4021:
4010:
3999:
3988:
3967:
3957:
3947:
3937:
3927:
3917:
3908:Schenkling, 1903
3907:
3897:
3887:
3878:Schenkling, 1902
3877:
3867:
3857:
3847:
3837:
3827:
3824:Pseudoteloclerus
3817:
3807:
3797:
3775:
3765:
3752:
3742:
3732:
3722:
3709:
3699:
3690:Schenkling, 1907
3689:
3679:
3669:
3659:
3649:
3639:
3629:
3619:
3609:
3599:
3589:
3579:
3569:
3566:Impressopallenis
3559:
3549:
3539:
3529:
3519:
3510:Schenkling, 1899
3509:
3499:
3486:
3473:
3463:
3460:Eburneocladiscus
3453:
3443:
3433:
3423:
3413:
3403:
3393:
3383:
3373:
3363:
3353:
3343:
3340:Cladiscopallenis
3333:
3323:
3313:
3303:
3293:
3283:
3273:
3270:Biflabellotillus
3263:
3254:Schenkling, 1906
3253:
3243:
3233:
3223:
3213:
3192:
3175:
3165:
3155:
3145:
3135:
3125:
3115:
3105:
3085:
3075:
3065:
3045:
3035:
3025:
3009:Necrobia rufipes
2991:
2981:
2971:
2948:
2938:
2919:
2909:
2896:
2879:
2869:
2859:
2849:
2839:
2829:
2816:
2806:
2796:
2786:
2776:
2766:
2756:
2753:Cucujocallimerus
2746:
2736:
2719:
2716:Brachycallimerus
2710:Schenkling, 1903
2709:
2706:Blaesiophthalmus
2699:
2689:
2679:
2638:
2621:
2611:
2594:
2584:
2574:
2564:
2547:
2537:
2527:
2517:
2507:
2497:
2487:
2477:
2467:
2426:
2416:
2406:
2391:
2381:
2372:Schenkling, 1900
2371:
2361:
2351:
2327:
2317:
2293:
2276:
2266:
2256:
2227:
2217:
2207:
2197:
2187:
2157:
2124:
2114:
2091:
2081:
2071:
2061:
2051:
2041:
2031:
2021:
2011:
2001:
1991:
1981:
1971:
1961:
1951:
1948:Priocleromorphus
1941:
1931:
1922:Schenkling, 1902
1921:
1911:
1901:
1891:
1881:
1871:
1861:
1851:
1841:
1831:
1821:
1811:
1801:
1791:
1781:
1771:
1761:
1752:Schenkling, 1906
1751:
1741:
1731:
1721:
1712:Gerstmeier, 1991
1711:
1701:
1691:
1688:Micropteroclerus
1681:
1672:Schenkling, 1899
1671:
1661:
1651:
1641:
1631:
1621:
1612:Schenkling, 1915
1611:
1601:
1591:
1581:
1571:
1562:Gerstmeier, 2002
1561:
1551:
1541:
1523:Schenkling, 1909
1522:
1512:
1502:
1492:
1482:
1472:
1462:
1452:
1442:
1432:
1422:
1412:
1402:
1393:Schenkling, 1906
1392:
1383:Schenkling, 1899
1382:
1372:
1362:
1352:
1339:
1329:
1319:
1309:
1299:
1289:
1279:
1270:Lacordaire, 1857
1269:
1259:
1249:
1239:
1229:
1219:
1209:
1192:
1182:
1172:
1149:
1136:
1126:
1117:Schenkling, 1903
1116:
1106:
1097:Schenkling, 1906
1096:
1086:
1069:
905:Neorthopleurinae
838:Ongoing research
814:Necrobia rufipes
810:Necrobia rufipes
782:Necrobia rufipes
765:Necrobia rufipes
598:Necrobia rufipes
469:Tillus elongatus
356:
306:are a family of
289:
278:
271:
264:
257:
250:
243:
236:
217:
125:
124:
104:
94:
31:
27:Temporal range:
21:
7102:
7101:
7097:
7096:
7095:
7093:
7092:
7091:
7067:
7066:
7065:
7060:
7052:
7047:
7039:
7034:
7026:
7021:
7013:
7011:
7003:
6998:
6990:
6985:
6977:
6972:
6964:
6959:
6951:
6946:
6938:
6933:
6925:
6920:
6912:
6907:
6899:
6894:
6886:
6881:
6873:
6868:
6860:
6855:
6847:
6842:
6834:
6832:
6824:
6819:
6810:
6809:
6804:
6795:
6794:
6789:
6776:
6766:
6761:
6740:
6682:
6633:
6615:(clown beetles)
6586:
6557:(betsy beetles)
6465:Scarabaeiformia
6455:
6412:
6387:
6319:(click beetles)
6295:(Texas beetles)
6271:
6242:
6238:Ptilodactylidae
6163:(cedar beetles)
6139:
6100:
6083:Trictenotomidae
5969:Mycetophagidae
5886:
5865:
5802:
5801:
5799:
5791:
5743:Phloeostichidae
5694:Lamingtoniidae
5686:Laemophloeidae
5634:Cryptophagidae
5598:
5489:
5472:Thanerocleridae
5435:Phloiophilidae
5422:Metaxina ornata
5387:Chaetosomatidae
5345:
5278:
5257:
5215:(auger beetles)
5196:Bostrichiformia
5186:
5168:
5164:Torridincolidae
5148:(skiff beetles)
5146:Hydroscaphidae
5137:
5115:
5108:Trachypachidae
5053:(tiger beetles)
5011:
4993:
4943:
4929:
4885:
4878:
4831:
4826:
4782:
4781:
4777:
4743:
4742:
4738:
4728:
4726:
4712:
4711:
4707:
4697:
4695:
4694:on 27 July 2011
4691:
4676:
4671:
4670:
4666:
4620:
4619:
4615:
4605:
4603:
4589:
4588:
4579:
4540:"Life Cycle of
4537:
4536:
4529:
4511:
4510:
4501:
4491:
4489:
4488:on 29 June 2008
4471:
4470:
4455:
4421:
4412:
4411:
4407:
4397:
4395:
4389:
4388:
4375:
4365:
4363:
4362:on 24 July 2011
4359:
4352:
4347:
4346:
4342:
4325:
4321:
4304:
4297:
4295:
4288:
4287:
4283:
4273:
4271:
4269:
4254:
4253:
4249:
4239:
4238:
4227:
4220:
4203:
4202:
4195:
4185:
4183:
4181:
4166:
4165:
4156:
4149:
4136:
4135:
4116:
4112:
4107:
4106:
4022:Schaeffer, 1771
4011:Fairmaire, 1889
3898:Fairmaire, 1903
3884:Synellapotillus
3838:Fairmaire, 1886
3610:Fairmaire, 1886
3444:Fairmaire, 1885
3424:Fairmaire, 1888
3394:Latreille, 1825
3374:Fairmaire, 1889
3354:Chevrolat, 1843
3300:Bostrichoclerus
3244:Fairmaire, 1903
3224:Fairmaire, 1902
3214:Fairmaire, 1903
3093:Thaneroclerinae
3076:Blackburn, 1900
2992:Latreille, 1797
2900:& Pic, 1940
2893:Stenocallimerus
2813:Lasiocallimerus
2777:Blanchard, 1842
2680:Fairmaire, 1902
2362:Chevrolat, 1834
2294:Latreille, 1802
2092:Latreille, 1806
2002:Chevrolat, 1874
1882:Chevrolat, 1843
1848:Phlogistomorpha
1822:Latreille, 1802
1762:Blackburn, 1900
1732:Fairmaire, 1886
1698:Microstigmatium
1682:Wollaston, 1867
1548:Eurymetomorphon
1453:Chevrolat, 1842
1330:Fabricius, 1775
1320:Fairmaire, 1889
1236:Bousquetoclerus
1127:Wollaston, 1867
933:Middle Jurassic
883:
840:
812:can be useful.
798:
761:
756:
666:
570:
506:
441:
412:
389:
354:
292:
285:
276:Thaneroclerinae
274:
267:
260:
253:
246:
239:
237:Latreille, 1802
216:
119:
110:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
7100:
7098:
7090:
7089:
7084:
7079:
7069:
7068:
7062:
7061:
7059:
7058:
7045:
7032:
7019:
7009:
6996:
6983:
6970:
6957:
6944:
6931:
6918:
6909:Fauna Europaea
6905:
6892:
6879:
6866:
6853:
6840:
6830:
6817:
6802:
6786:
6784:
6778:
6777:
6772:
6763:
6762:
6757:
6754:
6753:
6750:
6749:
6746:
6745:
6742:
6741:
6739:
6738:
6735:(rove beetles)
6733:Staphylinidae
6730:
6722:
6714:
6706:
6701:
6692:
6690:
6688:Staphylinoidea
6684:
6683:
6681:
6680:
6675:
6670:Hydrophilidae
6667:
6662:
6657:
6649:
6643:
6641:
6639:Hydrophiloidea
6635:
6634:
6632:
6631:
6626:
6618:
6609:
6607:
6598:
6592:
6591:
6588:
6587:
6585:
6584:
6581:(hide beetles)
6576:
6568:
6565:(rain beetles)
6560:
6552:
6544:
6541:(stag beetles)
6536:
6528:
6520:
6512:
6504:
6496:
6494:Bolboceratidae
6491:
6478:
6476:
6467:
6461:
6460:
6457:
6456:
6454:
6453:
6448:
6440:
6428:
6422:
6420:
6414:
6413:
6411:
6410:
6401:Rhinorhipidae
6397:
6395:
6393:Rhinorhipoidea
6389:
6388:
6386:
6385:
6377:
6372:
6367:
6359:
6351:
6343:
6335:
6330:
6322:
6314:
6306:
6298:
6290:
6281:
6279:
6273:
6272:
6270:
6269:
6261:
6252:
6250:
6244:
6243:
6241:
6240:
6235:
6227:
6219:
6211:
6206:Heteroceridae
6203:
6195:
6187:
6179:
6174:
6169:Chelonariidae
6166:
6161:Callirhipidae
6158:
6155:(pill beetles)
6149:
6147:
6141:
6140:
6138:
6137:
6132:
6123:
6121:
6112:
6106:
6105:
6102:
6101:
6099:
6098:
6090:
6085:
6080:
6072:
6067:Tenebrionidae
6064:
6056:
6048:
6040:
6032:
6024:
6016:
6008:
6003:
5995:
5990:
5982:
5974:
5966:
5958:
5950:
5942:
5934:
5929:
5921:
5913:
5905:
5896:
5894:
5892:Tenebrionoidea
5888:
5887:
5885:
5884:
5875:
5873:
5867:
5866:
5864:
5863:
5855:
5850:Curculionidae
5847:
5842:
5834:
5826:
5818:
5809:
5807:
5797:Curculionoidea
5793:
5792:
5790:
5789:
5784:
5776:
5768:
5760:
5758:Protocucujidae
5755:
5750:
5745:
5740:
5732:
5724:
5716:
5711:
5703:
5691:
5683:
5675:
5670:
5665:
5657:
5652:
5650:Cybocephalidae
5647:
5639:
5631:
5626:
5621:
5608:
5606:
5600:
5599:
5597:
5596:
5591:
5586:
5581:
5573:
5565:
5560:
5552:
5550:Discolomatidae
5547:
5539:
5534:Coccinellidae
5531:
5523:
5518:Bothrideridae
5515:
5510:
5505:
5499:
5497:
5495:Coccinelloidea
5491:
5490:
5488:
5487:
5482:Trogossitidae
5479:
5474:
5469:
5464:
5459:
5454:
5449:
5444:
5432:
5427:
5415:
5407:
5402:
5400:Lophocateridae
5397:
5389:
5384:
5376:
5368:
5355:
5353:
5347:
5346:
5344:
5343:
5338:
5333:
5328:
5323:
5318:
5315:(leaf beetles)
5313:Chrysomelidae
5310:
5301:
5299:
5297:Chrysomeloidea
5290:
5284:
5283:
5280:
5279:
5277:
5276:
5267:
5265:
5259:
5258:
5256:
5255:
5247:
5242:Nosodendridae
5239:
5234:Jacobsoniidae
5231:
5226:
5223:(skin beetles)
5218:
5209:
5207:
5198:
5188:
5187:
5181:
5174:
5173:
5170:
5169:
5167:
5166:
5161:
5156:
5151:
5142:
5139:
5138:
5132:
5125:
5124:
5121:
5120:
5117:
5116:
5114:
5113:
5105:
5097:
5092:Meru phyllisae
5085:
5080:
5072:
5064:
5056:
5048:
5040:
5035:
5026:
5024:
5013:
5012:
5006:
4999:
4998:
4995:
4994:
4992:
4991:
4986:
4984:Micromalthidae
4981:
4969:
4961:
4948:
4945:
4944:
4938:
4931:
4930:
4928:
4927:
4921:
4915:
4909:
4903:
4897:
4890:
4887:
4886:
4879:
4877:
4876:
4869:
4862:
4854:
4848:
4847:
4842:
4837:
4830:
4829:External links
4827:
4825:
4824:
4775:
4756:(4): 808–823.
4736:
4705:
4664:
4613:
4577:
4548:Insect Science
4527:
4499:
4453:
4405:
4373:
4340:
4319:
4281:
4267:
4247:
4225:
4218:
4193:
4179:
4154:
4147:
4113:
4111:
4108:
4103:
4102:
4098:
4097:
4082:
4071:
4056:
4045:
4031:
4014:
4003:
3992:
3976:Incertae sedis
3971:
3970:
3960:
3950:
3940:
3930:
3920:
3910:
3900:
3890:
3880:
3870:
3860:
3850:
3840:
3830:
3820:
3814:Pseudopallenis
3810:
3800:
3798:Duvivier, 1892
3790:
3781:
3768:
3758:
3745:
3735:
3733:Miyatake, 1965
3725:
3715:
3706:Orthocladiscus
3702:
3692:
3686:Notocymatodera
3682:
3672:
3666:Neocallotillus
3662:
3652:
3642:
3632:
3622:
3616:Macroliostylus
3612:
3602:
3592:
3582:
3572:
3562:
3552:
3542:
3532:
3522:
3512:
3502:
3492:
3483:Elasmocylidrus
3479:
3470:Egenocladiscus
3466:
3456:
3446:
3436:
3426:
3416:
3406:
3396:
3386:
3376:
3366:
3356:
3346:
3336:
3326:
3316:
3306:
3304:Van Dyke, 1938
3296:
3286:
3276:
3266:
3256:
3246:
3236:
3226:
3216:
3197:
3196:
3195:
3185:
3178:
3176:Lefebvre, 1838
3168:
3158:
3148:
3138:
3128:
3118:
3112:Compactoclerus
3108:
3106:Champion, 1913
3089:
3088:
3078:
3068:
3049:
3048:
3038:
3028:
3018:
3017:
3016:
3005:
2984:
2974:
2964:
2963:
2962:
2941:
2923:
2922:
2912:
2902:
2889:
2882:
2872:
2862:
2852:
2842:
2832:
2822:
2809:
2799:
2789:
2779:
2769:
2759:
2749:
2739:
2729:
2726:Brachyptevenus
2722:
2712:
2702:
2692:
2682:
2663:
2662:
2655:
2648:
2641:
2631:
2624:
2614:
2604:
2597:
2587:
2577:
2567:
2557:
2550:
2540:
2530:
2520:
2510:
2500:
2490:
2480:
2470:
2460:
2453:
2446:
2430:
2429:
2419:
2409:
2399:
2384:
2374:
2364:
2354:
2344:
2337:
2330:
2320:
2310:
2303:
2296:
2286:
2279:
2269:
2259:
2249:
2232:
2231:
2230:
2220:
2210:
2200:
2190:
2180:
2173:
2172:
2171:
2150:
2149:
2148:
2141:
2134:
2117:
2107:
2106:
2105:
2084:
2074:
2064:
2054:
2044:
2034:
2024:
2014:
2004:
1994:
1984:
1974:
1972:Miyatake, 1968
1968:Pseudolesterus
1964:
1954:
1944:
1934:
1924:
1914:
1904:
1894:
1884:
1874:
1864:
1854:
1844:
1834:
1824:
1814:
1808:Operculiphorus
1804:
1794:
1784:
1774:
1764:
1754:
1744:
1734:
1724:
1714:
1704:
1694:
1684:
1674:
1664:
1654:
1644:
1634:
1624:
1618:Inhumeroclerus
1614:
1604:
1594:
1584:
1574:
1564:
1554:
1544:
1534:
1533:
1532:
1515:
1505:
1495:
1485:
1475:
1465:
1455:
1445:
1435:
1425:
1419:Dasyteneclines
1415:
1409:Dasyceroclerus
1405:
1395:
1385:
1375:
1365:
1355:
1345:
1332:
1322:
1312:
1302:
1292:
1282:
1272:
1262:
1252:
1242:
1232:
1222:
1212:
1202:
1195:
1185:
1175:
1165:
1152:
1142:
1129:
1119:
1109:
1099:
1089:
1067:
1066:
1060:incertae sedis
986:Lemidia aptera
882:
879:
839:
836:
797:
794:
760:
757:
755:
752:
671:holometabolous
665:
662:
569:
568:Feeding habits
566:
505:
502:
440:
439:Identification
437:
411:
408:
388:
385:
349:, division of
347:tarsal formula
299:
298:
229:
228:
224:
223:
211:
207:
206:
201:
197:
196:
191:
187:
186:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
131:
127:
126:
113:
112:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
32:
26:
15:
13:
10:
9:
6:
4:
3:
2:
7099:
7088:
7085:
7083:
7080:
7078:
7075:
7074:
7072:
7055:
7050:
7046:
7042:
7037:
7033:
7029:
7024:
7020:
7016:
7010:
7006:
7001:
6997:
6993:
6988:
6984:
6980:
6975:
6971:
6967:
6962:
6958:
6954:
6949:
6945:
6941:
6936:
6932:
6928:
6923:
6919:
6915:
6910:
6906:
6902:
6897:
6893:
6889:
6884:
6880:
6876:
6871:
6867:
6863:
6858:
6854:
6850:
6845:
6841:
6837:
6831:
6827:
6822:
6818:
6813:
6807:
6803:
6798:
6792:
6788:
6787:
6785:
6783:
6779:
6775:
6770:
6760:
6755:
6737:
6731:
6729:
6723:
6721:
6715:
6713:
6707:
6705:
6702:
6700:
6694:
6693:
6691:
6689:
6685:
6679:
6676:
6674:
6668:
6666:
6663:
6661:
6658:
6656:
6650:
6648:
6645:
6644:
6642:
6640:
6636:
6630:
6627:
6625:
6621:Sphaeritidae
6619:
6617:
6611:
6610:
6608:
6606:
6602:
6599:
6597:
6593:
6583:
6577:
6575:
6571:Scarabaeidae
6569:
6567:
6561:
6559:
6553:
6551:
6545:
6543:
6537:
6535:
6529:
6527:
6521:
6519:
6513:
6511:
6509:(dor beetles)
6505:
6503:
6497:
6495:
6492:
6490:
6487:
6480:
6479:
6477:
6475:
6474:Scarabaeoidea
6471:
6468:
6466:
6462:
6452:
6449:
6447:
6441:
6439:
6436:
6429:
6427:
6424:
6423:
6421:
6419:
6415:
6409:
6406:
6399:
6398:
6396:
6394:
6390:
6384:
6378:
6376:
6373:
6371:
6368:
6366:
6360:
6358:
6352:
6350:
6344:
6342:
6336:
6334:
6331:
6329:
6323:
6321:
6315:
6313:
6309:Cerophytidae
6307:
6305:
6299:
6297:
6291:
6289:
6283:
6282:
6280:
6278:
6274:
6268:
6264:Rhipiceridae
6262:
6260:
6254:
6253:
6251:
6249:
6245:
6239:
6236:
6234:
6228:
6226:
6220:
6218:
6212:
6210:
6204:
6202:
6198:Eulichadidae
6196:
6194:
6188:
6186:
6180:
6178:
6177:Cneoglossidae
6175:
6173:
6167:
6165:
6159:
6157:
6151:
6150:
6148:
6146:
6142:
6136:
6135:Schizopodidae
6133:
6131:
6125:
6124:
6122:
6120:
6116:
6113:
6111:
6110:Elateriformia
6107:
6097:
6091:
6089:
6086:
6084:
6081:
6079:
6075:Tetratomidae
6073:
6071:
6065:
6063:
6057:
6055:
6049:
6047:
6041:
6039:
6033:
6031:
6027:Ripiphoridae
6025:
6023:
6017:
6015:
6011:Pyrochroidae
6009:
6007:
6006:Pterogeniidae
6004:
6002:
5996:
5994:
5991:
5989:
5983:
5981:
5975:
5973:
5967:
5965:
5959:
5957:
5951:
5949:
5945:Melandryidae
5943:
5941:
5935:
5933:
5932:Chalcodryidae
5930:
5928:
5922:
5920:
5914:
5912:
5906:
5904:
5898:
5897:
5895:
5893:
5889:
5883:
5877:
5876:
5874:
5872:
5868:
5862:
5858:Nemonychidae
5856:
5854:
5848:
5846:
5843:
5841:
5835:
5833:
5827:
5825:
5819:
5817:
5811:
5810:
5808:
5798:
5794:
5788:
5785:
5783:
5777:
5775:
5769:
5767:
5761:
5759:
5756:
5754:
5753:Propalticidae
5751:
5749:
5748:Priasilphidae
5746:
5744:
5741:
5739:
5733:
5731:
5727:Passandridae
5725:
5723:
5721:(sap beetles)
5717:
5715:
5712:
5710:
5704:
5702:
5699:
5692:
5690:
5684:
5682:
5676:
5674:
5671:
5669:
5666:
5664:
5658:
5656:
5653:
5651:
5648:
5646:
5640:
5638:
5632:
5630:
5629:Cavognathidae
5627:
5625:
5622:
5620:
5617:
5610:
5609:
5607:
5605:
5601:
5595:
5592:
5590:
5587:
5585:
5582:
5580:
5574:
5572:
5566:
5564:
5563:Eupsilobiidae
5561:
5559:
5555:Endomychidae
5553:
5551:
5548:
5546:
5542:Corylophidae
5540:
5538:
5532:
5530:
5524:
5522:
5516:
5514:
5511:
5509:
5506:
5504:
5501:
5500:
5498:
5496:
5492:
5486:
5480:
5478:
5475:
5473:
5470:
5468:
5465:
5463:
5460:
5458:
5457:Protopeltidae
5455:
5453:
5452:Prionoceridae
5450:
5448:
5445:
5443:
5440:
5433:
5431:
5428:
5426:
5423:
5416:
5414:
5408:
5406:
5405:Mauroniscidae
5403:
5401:
5398:
5396:
5390:
5388:
5385:
5383:
5377:
5375:
5369:
5367:
5364:
5357:
5356:
5354:
5352:
5348:
5342:
5339:
5337:
5334:
5332:
5329:
5327:
5326:Megalopodidae
5324:
5322:
5319:
5317:
5311:
5309:
5305:Cerambycidae
5303:
5302:
5300:
5298:
5294:
5291:
5289:
5285:
5275:
5271:Derodontidae
5269:
5268:
5266:
5264:
5263:Derodontoidea
5260:
5254:
5248:
5246:
5240:
5238:
5232:
5230:
5229:Endecatomidae
5227:
5225:
5219:
5217:
5213:Bostrichidae
5211:
5210:
5208:
5206:
5205:Bostrichoidea
5202:
5199:
5197:
5193:
5189:
5185:
5179:
5175:
5165:
5162:
5160:
5159:Sphaeriusidae
5157:
5155:
5152:
5150:
5144:
5143:
5140:
5136:
5130:
5126:
5112:
5106:
5104:
5098:
5096:
5093:
5086:
5084:
5081:
5079:
5073:
5071:
5065:
5063:
5057:
5055:
5051:Cicindelidae
5049:
5047:
5041:
5039:
5036:
5034:
5028:
5027:
5025:
5022:
5018:
5014:
5010:
5004:
5000:
4990:
4987:
4985:
4982:
4980:
4977:
4970:
4968:
4962:
4960:
4957:
4950:
4949:
4946:
4942:
4936:
4932:
4926:
4922:
4920:
4916:
4914:
4910:
4908:
4904:
4902:
4898:
4896:
4892:
4891:
4888:
4883:
4875:
4870:
4868:
4863:
4861:
4856:
4855:
4852:
4846:
4843:
4841:
4838:
4836:
4833:
4832:
4828:
4820:
4816:
4811:
4806:
4801:
4796:
4792:
4791:
4786:
4779:
4776:
4771:
4767:
4763:
4759:
4755:
4751:
4747:
4740:
4737:
4724:
4720:
4716:
4709:
4706:
4690:
4686:
4682:
4675:
4668:
4665:
4660:
4656:
4652:
4648:
4644:
4640:
4636:
4632:
4628:
4624:
4617:
4614:
4601:
4597:
4593:
4586:
4584:
4582:
4578:
4573:
4569:
4565:
4561:
4557:
4553:
4549:
4545:
4543:
4534:
4532:
4528:
4523:
4519:
4515:
4508:
4506:
4504:
4500:
4487:
4483:
4479:
4475:
4468:
4466:
4464:
4462:
4460:
4458:
4454:
4449:
4445:
4440:
4435:
4431:
4427:
4420:
4418:
4409:
4406:
4393:
4386:
4384:
4382:
4380:
4378:
4374:
4358:
4351:
4344:
4341:
4337:
4333:
4329:
4323:
4320:
4315:
4309:
4293:
4292:
4285:
4282:
4270:
4268:0-12-257305-6
4264:
4260:
4259:
4251:
4248:
4243:
4236:
4234:
4232:
4230:
4226:
4221:
4219:0-8493-8120-7
4215:
4210:
4209:
4200:
4198:
4194:
4182:
4176:
4172:
4171:
4163:
4161:
4159:
4155:
4150:
4148:0-03-096835-6
4144:
4140:
4133:
4131:
4129:
4127:
4125:
4123:
4121:
4119:
4115:
4109:
4101:
4095:
4092:(Korynetinae
4090:Winkler, 1984
4088:
4087:
4083:
4079:Spinola, 1841
4077:
4076:
4072:
4069:
4066:(Korynetinae
4062:
4061:
4060:Parapelonides
4057:
4053:Spinola, 1844
4051:
4050:
4046:
4041:
4037:
4036:
4032:
4029:
4028:
4024:(Korynetinae
4020:
4019:
4015:
4009:
4008:
4004:
3998:
3997:
3993:
3987:
3986:
3982:
3981:
3980:
3979:
3978:
3977:
3966:
3965:
3961:
3958:Olivier, 1790
3956:
3955:
3951:
3946:
3945:
3941:
3936:
3935:
3931:
3926:
3925:
3924:Tillodadiscus
3921:
3916:
3915:
3911:
3906:
3905:
3901:
3896:
3895:
3891:
3886:
3885:
3881:
3876:
3875:
3871:
3868:Spinola, 1844
3866:
3865:
3864:Stenocylidrus
3861:
3856:
3855:
3851:
3848:Kolibac, 1997
3846:
3845:
3841:
3836:
3835:
3834:Rhopaloclerus
3831:
3826:
3825:
3821:
3816:
3815:
3811:
3806:
3805:
3804:Pseudogyponix
3801:
3796:
3795:
3794:Pseudachlamys
3791:
3788:
3787:
3782:
3778:
3774:
3773:
3769:
3764:
3763:
3759:
3755:
3751:
3750:
3746:
3741:
3740:
3739:Paradoxocerus
3736:
3731:
3730:
3729:Paracladiscus
3726:
3721:
3720:
3716:
3712:
3708:
3707:
3703:
3698:
3697:
3693:
3688:
3687:
3683:
3678:
3677:
3673:
3668:
3667:
3663:
3660:Spinola, 1841
3658:
3657:
3653:
3648:
3647:
3643:
3638:
3637:
3633:
3628:
3627:
3623:
3618:
3617:
3613:
3608:
3607:
3603:
3598:
3597:
3593:
3588:
3587:
3583:
3578:
3577:
3576:Isocymatodera
3573:
3568:
3567:
3563:
3558:
3557:
3556:Gracilotillus
3553:
3548:
3547:
3546:Gastrocentrum
3543:
3538:
3537:
3533:
3528:
3527:
3523:
3518:
3517:
3516:Falsopallenis
3513:
3508:
3507:
3503:
3498:
3497:
3493:
3489:
3485:
3484:
3480:
3476:
3472:
3471:
3467:
3462:
3461:
3457:
3452:
3451:
3447:
3442:
3441:
3437:
3432:
3431:
3427:
3422:
3421:
3417:
3412:
3411:
3410:Cymatoderella
3407:
3402:
3401:
3397:
3392:
3391:
3387:
3382:
3381:
3380:Cylidroctenus
3377:
3372:
3371:
3367:
3362:
3361:
3357:
3352:
3351:
3347:
3342:
3341:
3337:
3332:
3331:
3327:
3322:
3321:
3317:
3314:Wolcott, 1911
3312:
3311:
3307:
3302:
3301:
3297:
3292:
3291:
3287:
3284:Kolibac, 1997
3282:
3281:
3277:
3272:
3271:
3267:
3262:
3261:
3257:
3252:
3251:
3247:
3242:
3241:
3237:
3232:
3231:
3227:
3222:
3221:
3220:Arachnoclerus
3217:
3212:
3211:
3207:
3206:
3205:
3204:
3203:
3198:
3193:Wolcott, 1910
3191:
3190:
3186:
3184:
3183:
3179:
3174:
3173:
3172:Thaneroclerus
3169:
3164:
3163:
3159:
3154:
3153:
3149:
3146:Kolibáč, 1992
3144:
3143:
3139:
3134:
3133:
3129:
3124:
3123:
3122:Cyrtinoclerus
3119:
3114:
3113:
3109:
3104:
3103:
3099:
3098:
3097:
3095:
3094:
3086:Spinola, 1844
3084:
3083:
3079:
3074:
3073:
3072:Tarsostenodes
3069:
3064:
3063:
3059:
3058:
3057:
3055:
3054:
3053:Tarsosteninae
3046:Winkler, 1960
3044:
3043:
3042:Romanaeclerus
3039:
3036:Spinola, 1844
3034:
3033:
3029:
3024:
3023:
3022:Neorthopleura
3019:
3015:
3011:
3010:
3006:
3004:
3000:
2999:
2995:
2994:
2990:
2989:
2985:
2982:R.Lucas, 1918
2980:
2979:
2975:
2972:Spinola, 1844
2970:
2969:
2965:
2961:
2957:
2956:
2952:
2951:
2947:
2946:
2942:
2937:
2936:
2932:
2931:
2930:
2929:
2928:
2918:
2917:
2913:
2908:
2907:
2903:
2899:
2895:
2894:
2890:
2888:
2887:
2883:
2878:
2877:
2873:
2868:
2867:
2863:
2858:
2857:
2853:
2848:
2847:
2843:
2838:
2837:
2833:
2830:Spinola, 1841
2828:
2827:
2823:
2819:
2815:
2814:
2810:
2805:
2804:
2800:
2795:
2794:
2790:
2785:
2784:
2780:
2775:
2774:
2770:
2765:
2764:
2760:
2755:
2754:
2750:
2745:
2744:
2743:Cephaloclerus
2740:
2735:
2734:
2730:
2728:
2727:
2723:
2718:
2717:
2713:
2708:
2707:
2703:
2698:
2697:
2693:
2688:
2687:
2683:
2678:
2677:
2673:
2672:
2671:
2669:
2668:
2661:
2660:
2656:
2654:
2653:
2649:
2647:
2646:
2642:
2637:
2636:
2632:
2630:
2629:
2625:
2622:Spinola, 1844
2620:
2619:
2615:
2610:
2609:
2605:
2603:
2602:
2598:
2593:
2592:
2588:
2583:
2582:
2578:
2573:
2572:
2568:
2563:
2562:
2558:
2556:
2555:
2551:
2546:
2545:
2541:
2536:
2535:
2531:
2526:
2525:
2521:
2516:
2515:
2511:
2506:
2505:
2504:Hapsidopteris
2501:
2498:Spinola, 1841
2496:
2495:
2491:
2488:Spinola, 1844
2486:
2485:
2481:
2476:
2475:
2471:
2466:
2465:
2461:
2459:
2458:
2454:
2452:
2451:
2447:
2445:
2444:
2443:Acanthocollum
2440:
2439:
2438:
2436:
2435:
2425:
2424:
2420:
2415:
2414:
2410:
2405:
2404:
2400:
2397:
2392:Spinola, 1841
2390:
2389:
2385:
2380:
2379:
2375:
2370:
2369:
2365:
2360:
2359:
2355:
2350:
2349:
2345:
2343:
2342:
2338:
2336:
2335:
2331:
2326:
2325:
2321:
2316:
2315:
2311:
2309:
2308:
2304:
2302:
2301:
2297:
2292:
2291:
2287:
2285:
2284:
2280:
2277:LeConte, 1861
2275:
2274:
2270:
2265:
2264:
2260:
2257:Spinola, 1841
2255:
2254:
2250:
2248:
2247:
2243:
2242:
2241:
2239:
2238:
2233:
2228:Spinola, 1841
2226:
2225:
2221:
2218:Winkler, 1992
2216:
2215:
2211:
2206:
2205:
2201:
2198:Winkler, 1981
2196:
2195:
2194:Wittmeridecus
2191:
2186:
2185:
2181:
2179:
2178:
2177:Wilsonoclerus
2174:
2170:
2166:
2165:
2161:
2160:
2158:Spinola, 1841
2156:
2155:
2151:
2147:
2146:
2142:
2140:
2139:
2135:
2133:
2132:
2128:
2127:
2123:
2122:
2118:
2115:Spinola, 1841
2113:
2112:
2108:
2104:
2100:
2099:
2095:
2094:
2090:
2089:
2085:
2080:
2079:
2075:
2070:
2069:
2068:Thalerocnemis
2065:
2060:
2059:
2055:
2052:Spinola, 1841
2050:
2049:
2045:
2040:
2039:
2035:
2030:
2029:
2025:
2022:Winkler, 1972
2020:
2019:
2015:
2012:Spinola, 1841
2010:
2009:
2005:
2000:
1999:
1995:
1990:
1989:
1985:
1980:
1979:
1975:
1970:
1969:
1965:
1960:
1959:
1955:
1950:
1949:
1945:
1940:
1939:
1935:
1932:Spinola, 1841
1930:
1929:
1925:
1920:
1919:
1915:
1912:Wolcott, 1910
1910:
1909:
1905:
1900:
1899:
1895:
1890:
1889:
1885:
1880:
1879:
1875:
1872:Spinola, 1841
1870:
1869:
1865:
1860:
1859:
1855:
1850:
1849:
1845:
1840:
1839:
1838:Oxystigmatium
1835:
1830:
1829:
1825:
1820:
1819:
1815:
1810:
1809:
1805:
1800:
1799:
1795:
1790:
1789:
1785:
1782:Spinola, 1841
1780:
1779:
1775:
1772:Rifkind, 2008
1770:
1769:
1765:
1760:
1759:
1755:
1750:
1749:
1745:
1740:
1739:
1735:
1730:
1729:
1725:
1722:Kolibac, 1997
1720:
1719:
1715:
1710:
1709:
1705:
1700:
1699:
1695:
1690:
1689:
1685:
1680:
1679:
1675:
1670:
1669:
1665:
1660:
1659:
1655:
1650:
1649:
1645:
1640:
1639:
1635:
1630:
1629:
1625:
1620:
1619:
1615:
1610:
1609:
1605:
1600:
1599:
1595:
1590:
1589:
1585:
1580:
1579:
1575:
1570:
1569:
1568:Falsoorthrius
1565:
1560:
1559:
1555:
1550:
1549:
1545:
1542:Spinola, 1841
1540:
1539:
1535:
1531:
1530:
1526:
1525:
1521:
1520:
1516:
1511:
1510:
1506:
1501:
1500:
1496:
1491:
1490:
1486:
1483:Winkler, 1987
1481:
1480:
1476:
1471:
1470:
1469:Eburneoclerus
1466:
1463:Spinola, 1841
1461:
1460:
1456:
1451:
1450:
1446:
1441:
1440:
1436:
1431:
1430:
1426:
1421:
1420:
1416:
1411:
1410:
1406:
1401:
1400:
1396:
1391:
1390:
1386:
1381:
1380:
1379:Corynommadius
1376:
1371:
1370:
1366:
1361:
1360:
1356:
1353:Spinola, 1841
1351:
1350:
1346:
1342:
1338:
1337:
1333:
1328:
1327:
1323:
1318:
1317:
1313:
1308:
1307:
1303:
1298:
1297:
1293:
1288:
1287:
1286:Cardiostichus
1283:
1280:Winkler, 1982
1278:
1277:
1273:
1268:
1267:
1263:
1258:
1257:
1253:
1248:
1247:
1243:
1238:
1237:
1233:
1228:
1227:
1223:
1220:Rifkind, 1996
1218:
1217:
1213:
1208:
1207:
1203:
1201:
1200:
1196:
1191:
1190:
1186:
1183:Spinola, 1841
1181:
1180:
1176:
1171:
1170:
1166:
1163:
1159:
1158:
1153:
1148:
1147:
1143:
1139:
1135:
1134:
1130:
1125:
1124:
1120:
1115:
1114:
1110:
1105:
1104:
1100:
1095:
1094:
1093:Anthicoclerus
1090:
1085:
1084:
1080:
1079:
1078:
1077:
1076:
1071:
1070:
1062:
1061:
1056:
1055:
1050:
1043:
1039:
1038:
1033:
1026:
1025:Tarsosteninae
1022:
1021:
1016:
1009:
1005:
1004:
999:
992:
988:
987:
982:
975:
971:
970:
965:
958:
954:
953:
948:
944:
942:
938:
934:
930:
929:
924:
923:
918:
917:
912:
911:
910:sensu stricto
906:
902:
901:
900:Neorthopleura
896:
892:
888:
880:
878:
875:
871:
867:
863:
859:
855:
851:
847:
846:
837:
832:
831:
826:
822:
819:
815:
811:
807:
803:
795:
793:
791:
787:
783:
779:
775:
772:. It infests
771:
767:
766:
758:
753:
751:
749:
745:
741:
737:
732:
727:
721:
719:
715:
711:
709:
703:
697:
696:
690:
686:
684:
680:
676:
672:
663:
661:
659:
658:
653:
649:
645:
641:
637:
636:
631:
627:
623:
622:
617:
614:
613:
604:
600:
599:
594:
590:
588:
584:
578:
574:
567:
565:
562:
557:
555:
551:
546:
544:
540:
536:
531:
529:
521:) on a flower
520:
516:
515:
510:
503:
501:
499:
495:
491:
486:
483:
475:
471:
470:
464:
460:
458:
454:
450:
446:
438:
436:
434:
430:
426:
422:
417:
409:
404:
400:
399:
393:
386:
384:
382:
378:
374:
370:
365:
363:
359:
352:
348:
344:
340:
336:
332:
328:
323:
321:
317:
313:
309:
305:
297:
296:
288:
282:
277:
270:
269:Tarsosteninae
265:Laporte, 1836
263:
258:Spinola, 1844
256:
249:
242:
235:
230:
225:
220:
215:
212:
209:
208:
205:
202:
200:Superfamily:
199:
198:
195:
192:
189:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
123:
118:
114:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
6781:
6660:Helophoridae
6652:Georissidae
6647:Epimetopidae
6563:Pleocomidae
6547:Ochodaeidae
6531:Hybosoridae
6515:Glaphyridae
6507:Geotrupidae
6485:
6482:Belohinidae
6443:Eucinetidae
6434:
6431:Decliniidae
6404:
6362:Phengodidae
6301:Cantharidae
6256:Dascillidae
6248:Dascilloidea
6230:Psephenidae
6222:Lutrochidae
6214:Limnichidae
6127:Buprestidae
6119:Buprestoidea
6059:Synchroidae
6043:Scraptiidae
6035:Salpingidae
5998:Prostomidae
5985:Oedemeridae
5961:Mordellidae
5879:Lymexylidae
5871:Lymexyloidea
5821:Attelabidae
5813:Anthribidae
5771:Smicripidae
5735:Phalacridae
5719:Nitidulidae
5714:Myraboliidae
5706:Monotomidae
5697:
5678:Kateretidae
5655:Cyclaxyridae
5615:
5612:Agapythidae
5576:Latridiidae
5526:Cerylonidae
5513:Anamorphidae
5447:Phycosecidae
5438:
5421:
5418:Metaxinidae
5391:
5371:Biphyllidae
5362:
5331:Orsodacnidae
5288:Cucujiformia
5221:Dermestidae
5091:
5030:Amphizoidae
4975:
4955:
4941:Archostemata
4925:Holometabola
4923:Superorder:
4917:Infraclass:
4788:
4778:
4753:
4749:
4739:
4727:. Retrieved
4722:
4718:
4708:
4696:. Retrieved
4689:the original
4684:
4680:
4667:
4659:the original
4630:
4626:
4616:
4604:. Retrieved
4600:the original
4595:
4572:the original
4551:
4547:
4541:
4521:
4517:
4513:
4490:. Retrieved
4486:the original
4481:
4477:
4429:
4425:
4416:
4408:
4396:. Retrieved
4364:. Retrieved
4357:the original
4343:
4335:
4331:
4322:
4296:. Retrieved
4290:
4284:
4272:. Retrieved
4257:
4250:
4241:
4207:
4184:. Retrieved
4169:
4138:
4099:
4093:
4084:
4073:
4067:
4058:
4047:
4033:
4025:
4018:Dermestoides
4016:
4007:Cleropiestus
4005:
3994:
3989:Kirsch, 1871
3983:
3974:
3973:
3972:
3962:
3952:
3942:
3932:
3922:
3912:
3902:
3892:
3882:
3872:
3862:
3852:
3844:Smudlotillus
3842:
3832:
3822:
3818:Kuwert, 1893
3812:
3802:
3792:
3784:
3770:
3760:
3747:
3743:Kraatz, 1899
3737:
3727:
3717:
3704:
3700:Chapin, 1945
3696:Onychotillus
3694:
3684:
3680:Gorham, 1892
3674:
3664:
3654:
3644:
3640:Chapin, 1919
3636:Melanoclerus
3634:
3624:
3614:
3604:
3600:Kraatz, 1899
3594:
3584:
3574:
3564:
3554:
3550:Gorham, 1876
3544:
3534:
3524:
3514:
3506:Eucymatodera
3504:
3496:Enoploclerus
3494:
3481:
3468:
3458:
3454:Heller, 1921
3450:Diplopherusa
3448:
3438:
3428:
3418:
3408:
3398:
3388:
3384:Kraatz, 1899
3378:
3370:Cteniopachys
3368:
3358:
3348:
3338:
3330:Chilioclerus
3328:
3318:
3308:
3298:
3288:
3278:
3268:
3260:Basilewskyus
3258:
3248:
3238:
3228:
3218:
3208:
3200:
3199:
3187:
3180:
3170:
3160:
3150:
3142:Meprinogenus
3140:
3130:
3126:Chapin, 1924
3120:
3110:
3100:
3091:
3090:
3080:
3070:
3066:Gorham, 1876
3060:
3051:
3050:
3040:
3032:Opetiopalpus
3030:
3020:
3007:
2996:
2986:
2976:
2966:
2953:
2949:Herbst, 1792
2943:
2933:
2925:
2924:
2920:Chapin, 1917
2914:
2904:
2891:
2884:
2874:
2870:Dejean, 1837
2866:Phyllobaenus
2864:
2854:
2844:
2840:Gorham, 1892
2834:
2824:
2811:
2801:
2797:Gorham, 1877
2791:
2787:Chapin, 1917
2783:Isohydnocera
2781:
2771:
2761:
2751:
2747:Kuwert, 1893
2741:
2737:Gorham, 1876
2731:
2724:
2720:Chapin, 1924
2714:
2704:
2694:
2684:
2674:
2667:Hydnocerinae
2665:
2664:
2659:Turbophloeus
2657:
2650:
2643:
2639:Kuwert, 1894
2635:Pyticeroides
2633:
2626:
2616:
2606:
2599:
2589:
2581:Parvochaetus
2579:
2569:
2559:
2552:
2542:
2532:
2522:
2512:
2502:
2492:
2482:
2472:
2462:
2455:
2448:
2441:
2434:Epiphloeinae
2432:
2431:
2423:Thriocerodes
2421:
2411:
2401:
2395:
2386:
2382:Newman, 1840
2376:
2366:
2358:Platynoptera
2356:
2352:Pascoe, 1876
2348:Phymatophaea
2346:
2339:
2332:
2322:
2312:
2305:
2298:
2288:
2281:
2271:
2263:Corinthiscus
2261:
2251:
2244:
2235:
2234:
2222:
2214:Zahradnikius
2212:
2208:Gorham, 1892
2202:
2192:
2188:Menier, 1986
2182:
2175:
2162:
2154:Trogodendron
2152:
2143:
2136:
2129:
2125:Herbst, 1792
2119:
2109:
2096:
2086:
2082:Murray, 1867
2078:Thanasimodes
2076:
2066:
2056:
2048:Systenoderes
2046:
2036:
2032:Kuwert, 1893
2026:
2016:
2006:
1996:
1988:Pujoliclerus
1986:
1982:Chapin, 1924
1978:Pseudomadius
1976:
1966:
1956:
1946:
1936:
1926:
1918:Plathanocera
1916:
1906:
1896:
1886:
1876:
1866:
1862:Gorham, 1876
1856:
1846:
1842:Kraatz, 1899
1836:
1832:Gorham, 1876
1826:
1816:
1812:Kuwert, 1894
1806:
1802:Elston, 1923
1796:
1786:
1776:
1766:
1758:Neoscrobiger
1756:
1746:
1736:
1726:
1716:
1708:Mimolesterus
1706:
1702:Kraatz, 1899
1696:
1692:Chapin, 1920
1686:
1676:
1666:
1656:
1646:
1638:Languropilus
1636:
1632:Fursov, 1936
1628:Jenjouristia
1626:
1616:
1606:
1602:Gorham, 1876
1596:
1592:Gorham, 1883
1586:
1582:Gorham, 1901
1578:Graptoclerus
1576:
1566:
1556:
1546:
1536:
1527:
1517:
1507:
1497:
1493:Newman, 1840
1487:
1477:
1467:
1457:
1449:Dozocolletus
1447:
1437:
1427:
1417:
1413:Kuwert, 1894
1407:
1397:
1387:
1377:
1373:Pascoe, 1860
1367:
1363:Chapin, 1924
1357:
1347:
1334:
1324:
1316:Cleropiestus
1314:
1310:Gorham, 1876
1304:
1294:
1284:
1276:Canariclerus
1274:
1264:
1254:
1244:
1240:Menier, 1997
1234:
1230:Gorham, 1882
1224:
1214:
1204:
1197:
1187:
1177:
1173:Kuwert, 1894
1167:
1155:
1144:
1133:Aptinoclerus
1131:
1123:Apteroclerus
1121:
1111:
1101:
1091:
1081:
1073:
1072:
1058:
1052:
1035:
1018:
1001:
991:Hydnocerinae
984:
967:
950:
941:Daohugou bed
926:
920:
914:
908:
898:
884:
869:
853:
843:
841:
828:
813:
809:
799:
781:
778:smoked meats
763:
762:
743:
722:
706:
700:
693:
667:
655:
635:Phyllobaenus
633:
619:
610:
608:
596:
579:
575:
571:
558:
547:
532:
524:
512:
487:
479:
467:
455:. Also, the
442:
413:
396:
381:bark beetles
366:
324:
315:
303:
302:
255:Hydnocerinae
248:Epiphloeinae
232:
227:Subfamilies
213:
194:Cucujiformia
190:Infraorder:
18:
6948:iNaturalist
6806:Wikispecies
6704:Hydraenidae
6678:Spercheidae
6665:Hydrochidae
6629:Synteliidae
6613:Histeridae
6605:Histeroidea
6555:Passalidae
6523:Glaresidae
6380:Throscidae
6340:(fireflies)
6338:Lampyridae
6325:Eucnemidae
6317:Elateridae
6277:Elateroidea
6093:Zopheridae
5977:Mycteridae
5908:Anthicidae
5779:Sphindidae
5763:Silvanidae
5673:Hobartiidae
5660:Erotylidae
5589:Mycetaeidae
5584:Murmidiidae
5568:Euxestidae
5462:Rentoniidae
5336:Oxypeltidae
5321:Disteniidae
5154:Lepiceridae
5083:Hygrobiidae
5075:Haliplidae
5059:Dytiscidae
5038:Aspidytidae
4035:Evenoclerus
3985:Aphelocerus
3968:Hintz, 1902
3938:Hintz, 1905
3934:Tillodenops
3918:White, 1849
3914:Tilloclerus
3858:Lewis, 1892
3749:Paraspinoza
3670:Burke, 2016
3646:Microtillus
3626:Magnotillus
3596:Leptoclerus
3580:Hintz, 1902
3526:Falsotillus
3500:Hintz, 1902
3440:Diplocladus
3360:Cladomorpha
3324:Hintz, 1902
3320:Ceratocopus
3310:Callotillus
3280:Bilbotillus
3230:Araeodontia
3156:Lewis, 1892
3136:Lewis, 1892
3102:Cleridopsis
3082:Tarsostenus
2927:Korynetinae
2860:Sharp, 1877
2850:Sharp, 1877
2803:Laiomorphus
2773:Eurymetopum
2652:Stegnoclava
2645:Silveirasia
2618:Plocamocera
2612:Opitz, 1997
2595:Opitz, 2008
2585:Opitz, 2006
2565:Opitz, 1997
2561:Megatrachys
2554:Megaphloeus
2538:Opitz, 1997
2528:Opitz, 1997
2524:Iontoclerus
2508:Opitz, 1997
2478:Opitz, 1997
2474:Diapromeces
2468:Opitz, 2008
2457:Decaphloeus
2407:Gahan, 1910
2394:(including
2368:Pseudichnea
2300:Exochonotus
2224:Zenithicola
2204:Xenorthrius
2072:Lohde, 1900
2018:Sedlacekius
1962:Hintz, 1902
1942:Kirby, 1818
1928:Platyclerus
1908:Placopterus
1868:Phloiocopus
1852:Hintz, 1908
1768:Ohanlonella
1728:Myrmecomaea
1718:Mitrandiria
1678:Microclerus
1658:Memorthrius
1648:Lissaulicus
1598:Hemitrachys
1558:Falsomadius
1503:Gahan, 1910
1443:Opitz, 2009
1439:Dologenitus
1433:Gahan, 1908
1399:Ctenoclerus
1359:Coptoclerus
1336:Clytomadius
1306:Cleromorpha
1216:Barrotillus
1210:Opitz, 2003
1193:Kirby, 1818
1146:Aradamicula
1103:Aphelochroa
1008:Korynetinae
928:Wangweiella
922:Protoclerus
891:subfamilies
881:Systematics
744:Thanasiumus
648:hymenoptera
642:, immature
640:wood borers
603:Korynetinae
457:notopleural
387:Description
290:Leach, 1815
262:Korynetinae
7071:Categories
6725:Silphidae
6717:Ptiliidae
6709:Leiodidae
6696:Agyrtidae
6539:Lucanidae
6418:Scirtoidea
6354:Omethidae
6333:Jurasaidae
6182:Dryopidae
6153:Byrrhidae
6145:Byrrhoidea
5837:Brentidae
5642:Cucujidae
5624:Boganiidae
5604:Cucujoidea
5477:Thymalidae
5467:Rhadalidae
5410:Melyridae
5379:Byturidae
5341:Vesperidae
5250:Ptiniidae
5100:Noteridae
5067:Gyrinidae
5043:Carabidae
4972:Jurodidae
4964:Cupedidae
4911:Subclass:
4901:Arthropoda
4882:Coleoptera
4725:(1): 15–22
4432:(1): 187.
4110:References
4086:Syriopelta
4064:Barr, 1980
3996:Apteropilo
3964:Tylotosoma
3904:Teloclerus
3894:Synellapus
3874:Strotocera
3786:Prospinoza
3772:Picoclerus
3766:Klug, 1842
3762:Philocalus
3656:Monophylla
3586:Lecontella
3414:Barr, 1962
3400:Cymatodera
3294:Barr, 1978
3234:Barr, 1952
3182:Viticlerus
3062:Paratillus
3026:Barr, 1976
2968:Lebasiella
2793:Isolemidia
2733:Callimerus
2628:Pteroferus
2591:Pennasolis
2575:Barr, 2006
2544:Madoniella
2534:Katamyurus
2494:Epiphloeus
2484:Ellipotoma
2403:Teneroides
2334:Paracregya
2246:Antygodera
2237:Enopliinae
2184:Winklerius
2103:Ant Beetle
2088:Thanasimus
2062:Hong, 2002
2058:Tanocleria
2038:Stigmatium
1958:Prioclerus
1902:Klug, 1837
1898:Placocerus
1858:Phlogistus
1748:Neogyponyx
1668:Metademius
1608:Homalopilo
1538:Erymanthus
1459:Eburiphora
1246:Burgeoneus
1113:Apopempsis
974:Enopliinae
916:sensu lato
718:overwinter
708:Thanasimus
702:Copulation
664:Life cycle
621:Thanasimus
587:pheromones
410:Appearance
335:scavengers
331:predaceous
241:Enopliinae
180:Suborder:
174:Coleoptera
154:Arthropoda
6579:Trogidae
6573:(scarabs)
6451:Scirtidae
6426:Clambidae
6019:Pythidae
5953:Meloidae
5900:Aderidae
5803:(weevils)
5668:Helotidae
5594:Teredidae
5508:Alexiidae
5392:Cleridae
5351:Cleroidea
5184:Polyphaga
5182:Suborder
5135:Myxophaga
5133:Suborder
5088:Meruidae
5007:Suborder
4989:Ommatidae
4939:Suborder
4913:Pterygota
4893:Kingdom:
4308:cite book
4075:Perilypus
4040:Corporaal
4000:Lea, 1908
3944:Tilloidea
3928:Pic, 1953
3888:Pic, 1939
3828:Pic, 1932
3808:Pic, 1939
3777:Corporaal
3754:Corporaal
3711:Corporaal
3650:Pic, 1950
3630:Pic, 1936
3620:Pic, 1939
3606:Liostylus
3570:Pic, 1953
3560:Pic, 1933
3520:Pic, 1926
3488:Corporaal
3475:Corporaal
3464:Pic, 1955
3364:Pic, 1949
3350:Cladiscus
3344:Pic, 1949
3274:Pic, 1949
3264:Pic, 1950
3240:Archalius
3189:Zenodosus
3152:Neoclerus
3132:Isoclerus
3116:Pic, 1939
2945:Korynetes
2935:Chariessa
2916:Wolcottia
2898:Corporaal
2886:Solemidia
2876:Silviella
2836:Neohydnus
2818:Corporaal
2807:Pic, 1927
2757:Pic, 1929
2696:Allelidea
2608:Pilosirus
2601:Pericales
2548:Pic, 1935
2396:Pelonides
2314:Lasiodera
2121:Trichodes
2111:Tillicera
2008:Scrobiger
1992:Pic, 1947
1952:Pic, 1950
1892:Pic, 1940
1778:Olesterus
1662:Pic, 1940
1642:Pic, 1940
1622:Pic, 1955
1572:Pic, 1940
1552:Pic, 1950
1519:Eunatalis
1509:Epiclines
1499:Enoclerus
1473:Pic, 1950
1429:Dieropsis
1423:Pic, 1941
1389:Ctenaxina
1341:Corporaal
1296:Caridopus
1266:Calendyma
1250:Pic, 1950
1206:Barriella
937:Callovian
931:the late
895:taxonomic
806:forensics
692:Larva of
657:Trichodes
638:spp. are
537:visiting
498:Melyridae
453:sternites
445:Polyphaga
435:plates).
351:sternites
312:Cleroidea
219:Latreille
204:Cleroidea
184:Polyphaga
140:Kingdom:
134:Eukaryota
7077:Cleridae
6857:BugGuide
6833:BioLib:
6826:Cleridae
6812:Cleridae
6791:Wikidata
6782:Cleridae
6346:Lycidae
6190:Elmidae
6088:Ulodidae
5924:Boridae
5829:Belidae
5430:Peltidae
5023:families
5009:Adephaga
4919:Neoptera
4899:Phylum:
4895:Animalia
4884:families
4819:29674858
4770:89399958
4729:20 March
4698:20 March
4655:30551315
4606:22 March
4568:85273860
4492:20 March
4448:17505768
4398:19 March
4366:19 March
4298:22 March
4274:19 March
4186:22 March
3789:(fossil)
3719:Pallenis
3390:Cylidrus
3250:Aroterus
3210:Antenius
3202:Tillinae
3162:Onerunka
3012:–
3001:–
2988:Necrobia
2978:Loedelia
2958:–
2686:Achlamys
2676:Abrosius
2571:Opitzius
2464:Decorosa
2450:Amboakis
2388:Pyticara
2341:Pelonium
2324:Neopylus
2307:Hublella
2290:Enoplium
2283:Curacavi
2253:Apolopha
2167:–
2101:–
2028:Sikorius
1938:Priocera
1828:Orthrius
1369:Cormodes
1349:Colyphus
1256:Caestron
1169:Astigmus
1157:Arawakis
1075:Clerinae
1042:Tillinae
957:Clerinae
860:and the
852:or IPM.
736:oviposit
626:woodland
612:Necrobia
564:adults.
528:habitats
519:Clerinae
494:pronotum
474:Tillinae
429:tergites
425:pronotum
403:Clerinae
343:antennae
320:habitats
304:Cleridae
293:and see
287:Tillinae
234:Clerinae
214:Cleridae
210:Family:
150:Phylum:
144:Animalia
130:Domain:
24:Cleridae
6797:Q377988
5937:Ciidae
5845:Caridae
4907:Insecta
4905:Class:
4880:Extant
4810:5904342
4790:ZooKeys
4647:9061672
4514:Aulicus
4330:(PDF).
3854:Spinoza
3676:Nodepus
3166:Kolibáč
2856:Paupris
2846:Parmius
2826:Lemidia
2763:Emmepus
2413:Tenerus
1888:Pieleus
1878:Phonius
1788:Omadius
1738:Natalis
1588:Gyponyx
1479:Ekisius
1226:Blaxima
1179:Aulicus
1083:Allonyx
802:carrion
644:weevils
539:insects
490:abdomen
308:beetles
170:Order:
164:Insecta
160:Class:
7028:192254
7012:NZOR:
7005:186087
6979:113852
6966:115923
6901:1CLERF
5021:Extant
4817:
4807:
4768:
4653:
4645:
4566:
4446:
4265:
4216:
4177:
4145:
4100:
4049:Muisca
4042:, 1950
3954:Tillus
3779:, 1936
3756:, 1942
3490:, 1939
3430:Denops
3420:Dedana
3290:Bogcia
2906:Theano
2820:, 1939
2514:Ichnea
2273:Cregya
1998:Sallea
1489:Eleale
1343:, 1949
1326:Clerus
1199:Balcus
1162:fossil
1140:, 1893
1138:Kuwert
887:genera
866:lumber
748:spring
740:imagos
726:dorsal
714:larvae
652:genera
646:, and
561:burrow
543:pollen
535:flower
482:tarsal
433:dorsal
421:elytra
339:pollen
327:niches
221:, 1802
7049:Plazi
7041:69399
6961:IRMNG
6953:55051
6914:11033
6849:24402
4766:S2CID
4692:(PDF)
4677:(PDF)
4651:S2CID
4564:S2CID
4428:. 1.
4422:(PDF)
4360:(PDF)
4353:(PDF)
2378:Pylus
1818:Opilo
1189:Axina
774:dried
731:pupal
675:larva
550:trees
373:decay
355:28–42
295:below
7000:NCBI
6974:ITIS
6940:3795
6935:GBIF
6896:EPPO
6888:7460
6844:BOLD
6836:9979
4815:PMID
4731:2009
4700:2009
4643:PMID
4608:2009
4494:2009
4444:PMID
4400:2009
4368:2009
4314:link
4300:2009
4276:2009
4263:ISBN
4214:ISBN
4188:2009
4175:ISBN
4143:ISBN
4094:s.l.
4068:s.l.
4027:s.l.
925:and
885:The
790:silk
788:and
786:wool
679:pupa
630:prey
616:spp.
554:bark
492:and
449:coxa
416:oval
362:bark
358:eggs
281:text
35:Preęž’
6987:NBN
6883:EoL
6875:8BP
6870:CoL
6862:113
6821:AFD
4805:PMC
4795:doi
4758:doi
4635:doi
4556:doi
4434:doi
913:or
776:or
742:of
654:is
337:or
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