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Clownfish

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meals and occasional dead anemone tentacles, and functions as a safe nest site. In return, the anemonefish defends the anemone from its predators and parasites. The anemone also picks up nutrients from the anemonefish's excrement. The nitrogen excreted from anemonefish increases the number of algae incorporated into the tissue of their hosts, which aids the anemone in tissue growth and regeneration. The activity of the anemonefish results in greater water circulation around the sea anemone, and it has been suggested that their bright coloring might lure small fish to the anemone, which then catches them. Studies on anemonefish have found that they alter the flow of water around sea anemone tentacles by certain behaviors and movements such as "wedging" and "switching". Aeration of the host anemone tentacles allows for benefits to the metabolism of both partners, mainly by increasing anemone body size and both anemonefish and anemone respiration.
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responsibility for the eggs, with males expending most of the time and effort. Male anemonefish care for their eggs by fanning and guarding them for 6 to 10 days until they hatch. In general, eggs develop more rapidly in a clutch when males fan properly, and fanning represents a crucial mechanism for successfully developing eggs. This suggests that males can control the success of hatching an egg clutch by investing different amounts of time and energy toward the eggs. For example, a male could choose to fan less in times of scarcity or fan more in times of abundance. Furthermore, males display increased alertness when guarding more valuable broods, or eggs in which paternity is guaranteed. Females, though, display generally less preference for parental behavior than males. All these suggest that males have increased parental investment towards eggs compared to females.
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Clownfish hatchlings undergo development after hatching in regards to both their body size and fins. If maintained at the demanded thermal regulation, clownfish undergo proper development of their fins. Clownfish follow the ensuing order in their fin development "Pectorals < caudal < dorsal
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in anemonefish may rest on the case that nonbreeders modulate their phenotype in a way that causes breeders to tolerate them. This strategy prevents conflict by reducing competition between males for one female. For example, by purposefully modifying their growth rate to remain small and submissive,
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Bleaching of the host anemone can occur when warm temperatures cause a reduction in algal symbionts within the anemone. Bleaching of the host can cause a short-term increase in the metabolic rate of resident anemonefish, probably as a result of acute stress. Over time, however, there appears to be a
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Anemonefish and sea anemones have a symbiotic, mutualistic relationship, each providing many benefits to the other. The individual species are generally highly host specific. The sea anemone protects the anemonefish from predators, as well as providing food through the scraps left from the anemone's
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Clownfish sold from captivity make up a very small account (10%) of the total trade of these fishes. Designer Clownfish, scientifically named A. ocellaris are much costlier and obtaining them has disrupted their coral reefs. Their attractive allure, color, and patterning have made them out to be an
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Anemonefish make up approximately 43% of the global marine ornamental trade, and approximately 25% of the global trade comes from fish bred in captivity, while the majority is captured from the wild, accounting for decreased densities in exploited areas. Public aquaria and captive-breeding programs
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does not occur and only the adult pair exhibits reproductive behavior. However, if the female dies, the social hierarchy shifts with the breeding male exhibiting protandrous sex reversal to become the breeding female. The largest juvenile then becomes the new breeding male after a period of rapid
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Anemonefish lay eggs on any flat surface close to their host anemones. In the wild, anemonefish spawn around the time of the full moon. Depending on the species, they can lay hundreds or thousands of eggs. The male parent guards the eggs until they hatch about 6–10 days later, typically two hours
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means that the eggs hatch around the full moon or new moon periods. One explanation for this lunar clock is that spring tides produce the highest tides during full or new moons. Nocturnal hatching during high tide may reduce predation by allowing for a greater capacity for escape. Namely, the
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Before making the clutch, the parents often clear an oval-shaped clutch varying in diameter for the spawn. Fecundity, or reproductive rate, of the females, usually ranges from 600 to 1,500 eggs depending on her size. In contrast to most animal species, the female only occasionally takes
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Vargas-Abúndez, Arturo Jorge; Randazzo, Basilio; Foddai, Marco; Sanchini, Lorenzo; Truzzi, Cristina; Giorgini, Elisabetta; Gasco, Laura; Olivotto, Ike (January 2019). "Insect meal based diets for clownfish: Biometric, histological, spectroscopic, biochemical and molecular implications".
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stronger currents and greater water volume during high tide protect the hatchlings by effectively sweeping them to safety. Before spawning, anemonefish exhibit increased rates of anemone and substrate biting, which help prepare and clean the nest for the spawn.
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has been in closed captivity. Members of some anemonefish species, such as the maroon clownfish, become aggressive in captivity; others, like the false percula clownfish, can be kept successfully with other individuals of the same species.
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Dhaneesh, K. V.; Vinoth, R.; Ghosh, Swagat; Gopi, M.; Kumar, T. T. Ajith; Balasubramanian, T. (2013). "Hatchery Production of Marine Ornamental Fishes: An Alternate Livelihood Option for the Island Community at Lakshadweep".
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down-regulation of metabolism and a reduced growth rate for fish associated with bleached anemones. These effects may stem from reduced food availability (e.g. anemone waste products, symbiotic algae) for the anemonefish.
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was thought to be the key innovation that allowed anemonefish to radiate rapidly, with rapid and convergent morphological changes correlated with the ecological niches offered by the host anemones. The complexity of
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Anemonefish colonies usually consist of the reproductive male and female and a few male juveniles, which help tend the colony. Although multiple males cohabit an environment with a single female,
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have never been sequenced. The authors hypothetically placed this species in the Indian clade because it is the most parsimonious solution regarding the biogeography of anemonefish species.
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The reproductive cycle of anemonefish is often correlated with the lunar cycle. Rates of spawning for anemonefish peak around the first and third quarters of the moon. The timing of this
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Shuman, Craig S.; Hodgson, Gregor; Ambrose, Richard F. (December 2005). "Population impacts of collecting sea anemones and anemonefish for the marine aquarium trade in the Philippines".
253:. Depending on the species, anemonefish are overall yellow, orange, or a reddish or blackish color, and many show white bars or patches. The largest can reach a length of 17 cm ( 2776:"Historic hybridization and introgression between two iconic Australian anemonefish and contemporary patterns of population connectivity: Historic Hybridization between Anemonefish" 3026: 1820:. The popularity of anemonefish for aquaria increased following the film's release; it is the first film associated with an increase in the numbers of those captured in the wild. 2483:
Buston, Peter (November 2004). "Does the presence of non-breeders enhance the fitness of breeders? An experimental analysis in the clown anemonefish Amphiprion percula".
2686: 1769:. Once an anemone or coral has been adopted, the anemonefish will defend it. Anemonefish, however, are not obligately tied to hosts, and can survive alone in captivity. 1749:
are essential to sustain their trade as marine ornamentals, and has recently become economically feasible. It is one of a handful of marine ornamentals whose complete
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of certain species of anemonefish with specific anemone host species may have allowed the fish to evolve an immunity to the nematocysts and toxins of their hosts.
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Porat, D.; Chadwick-Furman, N.E. (March 2005). "Effects of anemonefish on giant sea anemones: Ammonium uptake, zooxanthella content and tissue regeneration".
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Anemonefish are the best known example of fish that are able to live among the venomous sea anemone tentacles, but several others occur, including juvenile
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in the evolutionary past. The two evolutionary groups had individuals of both species detected, thus the species lacked reciprocal monophyly. No shared
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the juveniles in a colony present no threat to the fitness of the adult male, thereby protecting themselves from being evicted by the dominant fish.
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Roux, Natacha; Lami, Raphaël; Salis, Pauline; Magré, Kévin; Romans, Pascal; Masanet, Patrick; Lecchini, David; Laudet, Vincent (December 2019).
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exists. The largest and most aggressive female is found at the top. Only two anemonefish, a male and a female, in a group reproduce – through
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Buston, P. (2004). "Does the Presence of Non-Breeders Enhance the Fitness of Breeders? An Experimental Analysis in the Clown Anemonefish
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Watson, Craig A.; Hill, Jeffrey E. (May 2006). "Design criteria for recirculating, marine ornamental production systems".
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Mebs, D. (September 1994). "Anemonefish symbiosis: Vulnerability and resistance of fish to the toxin of the sea anemone".
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of the head, tooth shape, and body proportions are used. These features have been used to group species into six
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Casas, Laura; Saborido-Rey, Fran; Ryu, Taewoo; Michell, Craig; Ravasi, Timothy; Irigoien, Xabier (17 October 2016).
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Litsios, Glenn; Sims, Carrie A; Wüest, Rafael O; Pearman, Peter B; Zimmermann, Niklaus E; Salamin, Nicolas (2012).
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structure shown by genetic analysis of the Australian clade suggested evolutionary connectivity among samples of
3159: 1120: 1067: 480: 182: 2528:"Determining the level of parental care relating fanning behavior of five species of clownfishes in captivity" 942: 2597: 1485: 1456: 923: 888: 510: 404:. This would mean that anemones fail to recognize the fish as a potential food source and do not fire their 1059: 3538: 3437: 2827: 1713: 1421: 1402: 1147: 1092: 951: 896: 761: 585: 501: 476: 3137: 3128: 2012:
Growth, reproduction and survival of a tropical sea anemone (Actiniaria): benefits of hosting anemonefish
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Cortese, Daphne; Norin, Tommy; Beldade, Ricardo; Crespel, Amelie; Killen, Shaun; Mills, Suzanne (2021).
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Norin, Tommy; Mills, Suzanne; Crespel, Amelie; Cortese, Daphne; Beldade, Ricardo; Killen, Shaun (2018).
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matter from the anemonefish provides nutrients to the sea anemone. Anemonefish primarily feed on small
552:= anal < pelvic". The early larval stage is crucial to ensure a healthy progression of growth. 3515: 3358: 3310: 3251: 3000: 2957: 2728: 2636: 2492: 2361: 2253: 2216: 1997: 1929: 1696: 1355: 1328: 1303: 1278: 1253: 1199: 1009: 869: 787: 589: 561: 444: 427: 368: 335: 178: 3299:"Sea anemone and clownfish microbiota diversity and variation during the initial steps of symbiosis" 3475: 1816: 1731: 1672: 1594: 1539: 1429: 1228: 1112: 1038: 779: 691: 472: 436: 331: 246: 49: 596:. As can be seen from the gallery, each of the fish in these complexes has a similar appearance. 3543: 3384: 2973: 2508: 2465: 2448:
Ross, Robert M. (1978). "Reproductive Behavior of the Anemonefish Amphiprion melanopus on Guam".
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Anikuttan Kuttan Kuravamparambu; Rameshkumar Palsamy; Nazar Abdul Khudus; et al. (2022).
2290: 2269: 2185: 2126: 2051: 1893: 1862:"Clown Anemonefish, Clown Anemonefish Pictures, Clown Anemonefish Facts – National Geographic" 1477: 1320: 696: 452: 448: 3502: 1814:
main characters Nemo, his father Marlin, and his mother Coral are clownfish from the species
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features and color pattern in the field, while in a laboratory, other features such as
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Calado, Ricardo; Olivotto, Ike; Oliver, Miquel Planas; Holt, G. Joan (6 March 2017).
2922:. UNEP world conservation and monitoring centre (WCMC). pp. 1–64. Archived from 2851: 2823: 2311: 2265: 1788: 712: 573: 223: 141: 3388: 2977: 2512: 2396: 2381: 1949: 1347: 1030: 805: 366:
larvae, with a small portion of their diet coming from algae, with the exception of
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van der Meer, M. H.; Jones, G. P.; Hobbs, J.-P. A.; van Herwerden, L. (July 2012).
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Szczebak, J. T.; Henry, R. P.; Al-Horani, F. A.; Chadwick, N. E. (15 March 2013).
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Szczebak, J. T.; Henry, R. P.; Al-Horani, F. A.; Chadwick, N. E. (15 March 2013).
1903: 1448: 861: 3055: 2889: 3484: 3469: 3183: 3029:; Douglas Warmolts (2003). "23". In James C. Cato; Christopher L. Brown (eds.). 2016: 1531: 495: 463: 416: 355: 303:
Anemonefish are endemic to the warmer waters of the Indian Ocean, including the
250: 131: 121: 3322: 3097: 915: 2969: 2649: 2573: 2556: 2504: 2373: 2011: 1941: 1762: 565: 238: 173: 3460: 3271: 2741: 2717:"Mutualism with sea anemones triggered the adaptive radiation of clownfishes" 2582: 392:
Several theories are given about how they can survive the sea anemone venom:
2625:"Hybridisation and diversification in the adaptive radiation of clownfishes" 2229: 2204: 1781: 1681: 716: 601: 515: 409: 405: 243: 78: 3340: 3289: 2809: 2760: 2668: 2189: 2170: 2154:"Anemone bleaching increases the metabolic demands of symbiont anemonefish" 2130: 2055: 2273: 17: 3454: 3240:"Sex Change in Clownfish: Molecular Insights from Transcriptome Analysis" 2864: 2836: 2081: 1758: 1677: 1639: 363: 359: 347: 98: 3379: 2526:
Ghosh, Swagat; Kumar, T. T. Ajith; Balasubramanian, T. (October 2012).
2469: 2121: 2104: 2046: 2029: 1394: 401: 304: 227: 190: 3489: 3263: 3060:(illustrated ed.). Barron's Educational Series. pp. 11–22. 2791: 1992: 1796: 316: 88: 3431: 2461: 2916:
From ocean to aquarium: A global trade in marine ornamental species
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than to the saddleback fish with which it was previously grouped.
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and can feed on undigested food from their host anemones, and the
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Marine Ornamental Species: Collection, Culture and Conservation
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Biodiversity of the Central Coast. Retrieved 29 January 2015.
3121:"The anemonefish symbiosis: what is known and what is not" 1970:. National Geographic Society. 10 May 2011. Archived from 267: in), while the smallest barely achieve 7–8 cm ( 1885:
Field Guide to Anemone Fishes and Their Host Sea Anemones
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that the authors theorize was the result of historical
2687:"What we really know about the diversity of Clownfish" 2105:"Anemonefish oxygenate their anemone hosts at night" 2080:. The MarineBio Conservation Society. Archived from 2030:"Anemonefish oxygenate their anemone hosts at night" 3444: 3090:"Aquarium Fish: On the Clownfishes' Range of Hosts" 2882:
Climate Change and Island and Coastal Vulnerability
2557:"Designer clown fishes: Unraveling the ambiguities" 2422:. Florida Museum of Natural History. Archived from 2623:Litsios, Glenn; Salamin, Nicolas (December 2014). 1761:, the anemonefish may settle in some varieties of 560:Historically, anemonefish have been identified by 1638:(clown anemonefish) in a 'normal' orange and a 2913:Taylor, M.; Razak, T. & Green, E. (2003). 1922:Marine and Freshwater Behaviour and Physiology 396:The mucus coating of the fish may be based on 230:of clownfish are recognized: one in the genus 8: 3422:Monterey Bay Aquarium: Video and information 2710: 2708: 2680: 2678: 1882:Fautin, Daphne G.; Allen, Gerald R. (1997). 1877: 1875: 1860:Society, National Geographic (10 May 2011). 319:. No anemonefish are found in the Atlantic. 2550: 2548: 3432: 1915: 1913: 1757:When a sea anemone is not available in an 726: 40: 31: 3427:Clown Fish underwater photography gallery 3378: 3330: 3279: 2799: 2750: 2740: 2658: 2648: 2618: 2616: 2614: 2572: 2228: 2179: 2169: 2120: 2045: 663:. Other significant differences are that 425:may develop resistance to the toxin from 2332:Fretwell, K.; and B. Starzomski (2014). 1780: 671:is in the Indian clade rather than with 1852: 1829: 1623: 1591: 1582: 600:has shown that these complexes are not 236:, while the remaining are in the genus 3214:. John Wiley & Sons. p. 179. 604:, particularly the 11 species in the 3211:Marine Ornamental Species Aquaculture 3033:. Wiley-Blackwell. pp. 303–326. 2535:Indian Journal of Geo-Marine Sciences 7: 471:In a group of anemonefish, a strict 3162:: T.F.H. Publications. p. 83. 2485:Behavioral Ecology and Sociobiology 2354:Behavioral Ecology and Sociobiology 2009:Holbrook, S. J. and Schmitt, R. J. 1773:attractive target in wild trading. 749: 667:also has monospecific lineage, and 2397:"Clownfish breeding for beginners" 2158:Proceedings of the Royal Society B 1620:Morphological diversity by complex 25: 3371:10.1016/j.aquaculture.2018.08.018 2286:Lieske, E.; and R. Myers (1999). 1791:featuring clownfish are popular. 1722: 1705: 1688: 1664: 1647: 1626: 1609: 1575: 1530: 1503: 1476: 1447: 1420: 1393: 1346: 1319: 1294: 1269: 1244: 1219: 1190: 1163: 1138: 1111: 1083: 1058: 1029: 1000: 971: 941: 914: 887: 860: 831: 804: 778: 65: 2109:Journal of Experimental Biology 2034:Journal of Experimental Biology 358:from the water column, such as 479:. Anemonefish are protandrous 1: 3013:10.1016/j.aquaeng.2005.07.002 2306:Patzner, R.A. (5 July 2017). 719:were found between species. 659:in the Australian clade with 242:. In the wild, they all form 2890:10.1007/978-94-007-6016-5_17 2266:10.1016/0041-0101(94)90390-5 1700:(red saddleback anemonefish) 499:A pair of pink anemonefish ( 467:Clownfish swimming movements 3549:Ray-finned fish subfamilies 3119:Daphne Gail Fautin (1991). 2561:Frontiers in Marine Science 1676:(saddleback clownfish) off 679:is more closely related to 449:incognito (or anemone) goby 372:, which primarily feeds on 3570: 3323:10.1038/s41598-019-55756-w 3057:Clownfish and Sea Anemones 1016:Red saddleback anemonefish 723:Phylogenetic relationships 647:being in an Indian clade, 535:swimming around an anemone 3224:– via Google Books. 3154:Ronald L. Shimek (2004). 2970:10.1007/s00338-005-0027-z 2650:10.1186/s12862-014-0245-5 2574:10.3389/fmars.2022.907362 2505:10.1007/s00265-004-0833-2 2374:10.1007/s00265-004-0833-2 1942:10.1080/10236240500057929 1890:Western Australian Museum 1583: 1206:Red and black anemonefish 750: 745: 742: 737: 734: 731: 675:, the skunk anemonefish. 514:growth. The existence of 481:sequential hermaphrodites 307:, and Pacific Ocean, the 169: 164: 62:Scientific classification 60: 48: 39: 34: 2993:Aquacultural Engineering 2868:. December 2011 version. 2854:; Pauly, Daniel (eds.). 2840:. December 2011 version. 2826:; Pauly, Daniel (eds.). 2742:10.1186/1471-2148-12-212 2721:BMC Evolutionary Biology 2629:BMC Evolutionary Biology 1735:(maroon anemonefish) in 1717:(pink skunk anemonefish) 1154:White-bonnet anemonefish 299:Distribution and habitat 3136:: 23–46. Archived from 2230:10.1111/1365-2435.13729 1765:, or large polyp stony 1260:False clown anemonefish 505:) in their anemone home 380:Symbiosis and mutualism 336:magnificent sea anemone 3054:Tullock, John (1998). 2171:10.1098/rspb.2018.0282 2070:"Clown Anemonefishes, 1792: 1566:Three-band anemonefish 1463:Australian anemonefish 1436:Saddleback anemonefish 1409:Pink skunk anemonefish 1310:Orange Storm Clownfish 1179:Whitesnout anemonefish 1127:Madagascar anemonefish 1074:Seychelles anemonefish 957:Orange-fin anemonefish 762:Amphiprion akallopisos 536: 506: 502:Amphiprion perideraion 477:external fertilization 468: 369:Amphiprion perideraion 343: 3415:13 March 2021 at the 3184:"Finding Nemo (2003)" 2780:Ecology and Evolution 1968:Nat Geo Wild: Animals 1784: 1659:(Clark's anemonefish) 1546:Thielle's anemonefish 1285:Black Storm Clownfish 1099:Wide-band anemonefish 930:Mauritian anemonefish 530: 498: 466: 330: 3554:Fish of Saudi Arabia 3408:Amphiprion ocellaris 3156:Marine Invertebrates 3088:Fatherree, James W. 2884:. pp. 253–265. 2693:on 25 September 2015 2072:Amphiprion ocellaris 1998:Encyclopedia of Life 1808:and its 2016 sequel 876:Two-band anemonefish 847:Barber's anemonefish 820:Allard's anemonefish 445:Banggai cardinalfish 428:Heteractis magnifica 340:Heteractis magnifica 54:Amphiprion ocellaris 3363:2019Aquac.498....1V 3315:2019NatSR...919491R 3256:2016NatSR...635461C 3005:2006AqEng..34..157W 2962:2005CorRe..24..564S 2733:2012BMCEE..12..212L 2641:2014BMCEE..14..245L 2497:2004BEcoS..57...23B 2399:. Mad Hatter's Reef 2366:2004BEcoS..57...23B 2308:"Gobius incognitus" 2258:1994Txcn...32.1059M 2221:2021FuEco..35..663C 1964:"Clown Anemonefish" 1934:2005MFBP...38...43P 1868:on 13 January 2010. 1595:Premnas biaculeatus 1362:Pacific anemonefish 1235:Maldive anemonefish 987:Clark's anemonefish 728: 685:Premnas biaculeatus 620:, with six species, 602:monophyletic groups 491:Parental investment 473:dominance hierarchy 437:threespot dascyllus 332:Ocellaris clownfish 218:from the subfamily 50:Ocellaris clownfish 27:Subfamily of fishes 3303:Scientific Reports 3244:Scientific Reports 2856:"Species in genus 2828:"Species in genus 2426:on 28 October 2005 2350:Amphiprion percula 2335:Painted greenling. 2288:Coral Reef Fishes. 2209:Functional Ecology 2122:10.1242/jeb.075648 2084:on 27 October 2011 2047:10.1242/jeb.075648 1974:on 13 January 2010 1793: 1777:In popular culture 1600:Maroon anemonefish 1492:Orange anemonefish 1045:Tomato anemonefish 903:Chagos anemonefish 727: 608:group, where only 537: 533:cinnamon clownfish 507: 469: 422:Amphiprion percula 344: 309:Great Barrier Reef 3526: 3525: 3498:Open Tree of Life 3438:Taxon identifiers 3406:Photo Gallery of 3264:10.1038/srep35461 3169:978-1-890087-66-1 3094:Advanced Aquarist 3040:978-0-8138-2987-6 2899:978-94-007-6015-8 2420:"Clown Anemofish" 2418:Stephanie Boyer. 2015:, 2005, cited in 1906:on 14 April 2015. 1617: 1616: 1519:Sebae anemonefish 1384:Clown anemonefish 767:Skunk anemonefish 697:mitochondrial DNA 641:A. latifasciatus 637:A. fuscocaudatus 528: 453:painted greenling 205: 204: 199: 186: 160: 16:(Redirected from 3561: 3519: 3518: 3506: 3505: 3493: 3492: 3480: 3479: 3478: 3465: 3464: 3463: 3433: 3404: 3392: 3382: 3344: 3334: 3293: 3283: 3226: 3225: 3205: 3199: 3198: 3196: 3194: 3180: 3174: 3173: 3160:Neptune City, NJ 3151: 3145: 3144: 3142: 3125: 3116: 3110: 3109: 3107: 3105: 3100:on 22 March 2014 3096:. Archived from 3085: 3079: 3078: 3076: 3074: 3051: 3045: 3044: 3023: 3017: 3016: 2988: 2982: 2981: 2945: 2939: 2938: 2936: 2934: 2928: 2921: 2910: 2904: 2903: 2876: 2870: 2869: 2848: 2842: 2841: 2820: 2814: 2813: 2803: 2792:10.1002/ece3.251 2786:(7): 1592–1604. 2771: 2765: 2764: 2754: 2744: 2712: 2703: 2702: 2700: 2698: 2689:. Archived from 2682: 2673: 2672: 2662: 2652: 2620: 2609: 2608: 2606: 2604: 2593: 2587: 2586: 2576: 2552: 2543: 2542: 2532: 2523: 2517: 2516: 2480: 2474: 2473: 2445: 2436: 2435: 2433: 2431: 2415: 2409: 2408: 2406: 2404: 2392: 2386: 2385: 2345: 2339: 2330: 2324: 2323: 2321: 2319: 2314:on 7 August 2020 2310:. Archived from 2303: 2297: 2284: 2278: 2277: 2252:(9): 1059–1068. 2241: 2235: 2234: 2232: 2200: 2194: 2193: 2183: 2173: 2149: 2143: 2142: 2124: 2100: 2094: 2093: 2091: 2089: 2066: 2060: 2059: 2049: 2025: 2019: 2007: 2001: 1990: 1984: 1983: 1981: 1979: 1960: 1954: 1953: 1917: 1908: 1907: 1902:. Archived from 1879: 1870: 1869: 1864:. Archived from 1857: 1841: 1838:A. fuscocaudatus 1834: 1737:Papua New Guinea 1726: 1709: 1692: 1668: 1651: 1642:blackish variant 1630: 1613: 1579: 1534: 1507: 1480: 1451: 1424: 1397: 1350: 1335:Oman anemonefish 1323: 1298: 1273: 1248: 1223: 1194: 1167: 1142: 1122:A. latifasciatus 1115: 1087: 1069:A. fuscocaudatus 1062: 1033: 1004: 975: 945: 918: 891: 864: 835: 808: 782: 732:Scientific name 729: 616:are in the same 598:Genetic analysis 529: 346:Anemonefish are 294: 293: 289: 286: 280: 279: 275: 272: 266: 265: 261: 258: 194: 177: 155: 70: 69: 44: 32: 21: 3569: 3568: 3564: 3563: 3562: 3560: 3559: 3558: 3529: 3528: 3527: 3522: 3514: 3509: 3501: 3496: 3488: 3483: 3474: 3473: 3468: 3459: 3458: 3453: 3440: 3417:Wayback Machine 3402: 3399: 3347: 3296: 3237: 3234: 3232:Further reading 3229: 3222: 3207: 3206: 3202: 3192: 3190: 3188:Rotten Tomatoes 3182: 3181: 3177: 3170: 3153: 3152: 3148: 3143:on 25 May 2012. 3140: 3123: 3118: 3117: 3113: 3103: 3101: 3087: 3086: 3082: 3072: 3070: 3068: 3053: 3052: 3048: 3041: 3025: 3024: 3020: 2990: 2989: 2985: 2947: 2946: 2942: 2932: 2930: 2926: 2919: 2912: 2911: 2907: 2900: 2878: 2877: 2873: 2850: 2849: 2845: 2822: 2821: 2817: 2773: 2772: 2768: 2714: 2713: 2706: 2696: 2694: 2684: 2683: 2676: 2622: 2621: 2612: 2602: 2600: 2598:"Anemonefishes" 2595: 2594: 2590: 2554: 2553: 2546: 2530: 2525: 2524: 2520: 2482: 2481: 2477: 2462:10.2307/1443829 2447: 2446: 2439: 2429: 2427: 2417: 2416: 2412: 2402: 2400: 2394: 2393: 2389: 2347: 2346: 2342: 2331: 2327: 2317: 2315: 2305: 2304: 2300: 2285: 2281: 2243: 2242: 2238: 2202: 2201: 2197: 2151: 2150: 2146: 2102: 2101: 2097: 2087: 2085: 2068: 2067: 2063: 2027: 2026: 2022: 2008: 2004: 1991: 1987: 1977: 1975: 1962: 1961: 1957: 1919: 1918: 1911: 1900: 1881: 1880: 1873: 1859: 1858: 1854: 1850: 1845: 1844: 1835: 1831: 1826: 1779: 1746: 1744:In the aquarium 1739: 1727: 1718: 1710: 1701: 1693: 1684: 1669: 1660: 1652: 1643: 1631: 1622: 1487:A. sandaracinos 1458:A. rubrocinctus 952:A. chrysopterus 925:A. chrysogaster 725: 649:A. chrysopterus 633:A. chrosgaster 558: 522: 493: 461: 451:, and juvenile 382: 325: 301: 291: 287: 284: 282: 277: 273: 270: 268: 263: 259: 256: 254: 154: 64: 28: 23: 22: 15: 12: 11: 5: 3567: 3565: 3557: 3556: 3551: 3546: 3541: 3531: 3530: 3524: 3523: 3521: 3520: 3507: 3494: 3481: 3476:Amphiprioninae 3466: 3450: 3448: 3446:Amphiprioninae 3442: 3441: 3436: 3430: 3429: 3424: 3419: 3410:and their eggs 3398: 3397:External links 3395: 3394: 3393: 3345: 3294: 3233: 3230: 3228: 3227: 3220: 3200: 3175: 3168: 3146: 3111: 3080: 3066: 3046: 3039: 3018: 2999:(3): 157–162. 2983: 2956:(4): 564–573. 2940: 2929:on 1 July 2004 2905: 2898: 2871: 2852:Froese, Rainer 2843: 2824:Froese, Rainer 2815: 2766: 2704: 2685:DeAngelis, R. 2674: 2610: 2588: 2544: 2518: 2475: 2456:(1): 103–107. 2437: 2410: 2395:Jeff Hesketh. 2387: 2340: 2325: 2298: 2279: 2236: 2215:(3): 663–674. 2195: 2144: 2115:(6): 970–976. 2095: 2061: 2040:(6): 970–976. 2020: 2002: 1985: 1955: 1909: 1898: 1871: 1851: 1849: 1846: 1843: 1842: 1828: 1827: 1825: 1822: 1789:birthday cakes 1778: 1775: 1745: 1742: 1741: 1740: 1732:P. biaculeatus 1728: 1721: 1719: 1714:A. perideraion 1711: 1704: 1702: 1694: 1687: 1685: 1670: 1663: 1661: 1653: 1646: 1644: 1632: 1625: 1621: 1618: 1615: 1614: 1607: 1604: 1601: 1598: 1590: 1589: 1581: 1580: 1573: 1570: 1567: 1564: 1556: 1555: 1553: 1550: 1547: 1544: 1536: 1535: 1528: 1525: 1520: 1517: 1509: 1508: 1501: 1498: 1496:A. akallopisos 1493: 1490: 1482: 1481: 1474: 1469: 1464: 1461: 1453: 1452: 1445: 1442: 1437: 1434: 1426: 1425: 1418: 1415: 1413:A. akallopisos 1410: 1407: 1404:A. perideraion 1399: 1398: 1391: 1388: 1385: 1382: 1374: 1373: 1371: 1368: 1366:A. akallopisos 1363: 1360: 1352: 1351: 1344: 1339: 1336: 1333: 1325: 1324: 1317: 1314: 1311: 1308: 1300: 1299: 1292: 1289: 1286: 1283: 1275: 1274: 1267: 1264: 1261: 1258: 1250: 1249: 1242: 1239: 1236: 1233: 1225: 1224: 1217: 1212: 1207: 1204: 1196: 1195: 1188: 1183: 1180: 1177: 1169: 1168: 1161: 1158: 1155: 1152: 1149:A. leucokranos 1144: 1143: 1136: 1131: 1128: 1125: 1117: 1116: 1109: 1106: 1100: 1097: 1094:A. latezonatus 1089: 1088: 1081: 1078: 1075: 1072: 1064: 1063: 1056: 1051: 1046: 1043: 1035: 1034: 1027: 1022: 1017: 1014: 1006: 1005: 998: 993: 988: 985: 977: 976: 969: 964: 958: 955: 947: 946: 939: 934: 931: 928: 920: 919: 912: 907: 904: 901: 898:A. chagosensis 893: 892: 885: 880: 877: 874: 866: 865: 858: 853: 848: 845: 837: 836: 829: 824: 821: 818: 810: 809: 802: 797: 794: 792: 784: 783: 776: 773: 771:A. akallopisos 768: 765: 757: 756: 748: 747: 744: 741: 736: 733: 724: 721: 677:A. latezonatus 673:A. akallopisos 665:A. latezonatus 629:A. chagosensis 557: 554: 492: 489: 460: 457: 433: 432: 413: 381: 378: 324: 321: 315:or in shallow 300: 297: 222:in the family 220:Amphiprioninae 203: 202: 201: 200: 187: 167: 166: 162: 161: 152:Amphiprioninae 149: 145: 144: 139: 135: 134: 129: 125: 124: 119: 112: 111: 109:Actinopterygii 106: 102: 101: 96: 92: 91: 86: 82: 81: 76: 72: 71: 58: 57: 46: 45: 37: 36: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 3566: 3555: 3552: 3550: 3547: 3545: 3542: 3540: 3539:Pomacentridae 3537: 3536: 3534: 3517: 3512: 3508: 3504: 3499: 3495: 3491: 3486: 3482: 3477: 3471: 3467: 3462: 3456: 3452: 3451: 3449: 3447: 3443: 3439: 3434: 3428: 3425: 3423: 3420: 3418: 3414: 3411: 3409: 3401: 3400: 3396: 3390: 3386: 3381: 3376: 3372: 3368: 3364: 3360: 3356: 3352: 3346: 3342: 3338: 3333: 3328: 3324: 3320: 3316: 3312: 3308: 3304: 3300: 3295: 3291: 3287: 3282: 3277: 3273: 3269: 3265: 3261: 3257: 3253: 3249: 3245: 3241: 3236: 3235: 3231: 3223: 3221:9780470673904 3217: 3213: 3212: 3204: 3201: 3189: 3185: 3179: 3176: 3171: 3165: 3161: 3157: 3150: 3147: 3139: 3135: 3131: 3130: 3122: 3115: 3112: 3099: 3095: 3091: 3084: 3081: 3069: 3067:9780764105111 3063: 3059: 3058: 3050: 3047: 3042: 3036: 3032: 3028: 3027:Hall, Heather 3022: 3019: 3014: 3010: 3006: 3002: 2998: 2994: 2987: 2984: 2979: 2975: 2971: 2967: 2963: 2959: 2955: 2951: 2944: 2941: 2925: 2918: 2917: 2909: 2906: 2901: 2895: 2891: 2887: 2883: 2875: 2872: 2867: 2866: 2861: 2859: 2853: 2847: 2844: 2839: 2838: 2833: 2831: 2825: 2819: 2816: 2811: 2807: 2802: 2797: 2793: 2789: 2785: 2781: 2777: 2770: 2767: 2762: 2758: 2753: 2748: 2743: 2738: 2734: 2730: 2726: 2722: 2718: 2711: 2709: 2705: 2692: 2688: 2681: 2679: 2675: 2670: 2666: 2661: 2656: 2651: 2646: 2642: 2638: 2634: 2630: 2626: 2619: 2617: 2615: 2611: 2599: 2592: 2589: 2584: 2580: 2575: 2570: 2566: 2562: 2558: 2551: 2549: 2545: 2541:(5): 430–441. 2540: 2536: 2529: 2522: 2519: 2514: 2510: 2506: 2502: 2498: 2494: 2490: 2486: 2479: 2476: 2471: 2467: 2463: 2459: 2455: 2451: 2444: 2442: 2438: 2425: 2421: 2414: 2411: 2398: 2391: 2388: 2383: 2379: 2375: 2371: 2367: 2363: 2359: 2355: 2351: 2344: 2341: 2337: 2336: 2329: 2326: 2313: 2309: 2302: 2299: 2296: 2295:0-691-00481-1 2292: 2289: 2283: 2280: 2275: 2271: 2267: 2263: 2259: 2255: 2251: 2247: 2240: 2237: 2231: 2226: 2222: 2218: 2214: 2210: 2206: 2199: 2196: 2191: 2187: 2182: 2177: 2172: 2167: 2163: 2159: 2155: 2148: 2145: 2140: 2136: 2132: 2128: 2123: 2118: 2114: 2110: 2106: 2099: 2096: 2083: 2079: 2075: 2073: 2065: 2062: 2057: 2053: 2048: 2043: 2039: 2035: 2031: 2024: 2021: 2018: 2014: 2013: 2006: 2003: 2000: 1999: 1994: 1989: 1986: 1973: 1969: 1965: 1959: 1956: 1951: 1947: 1943: 1939: 1935: 1931: 1927: 1923: 1916: 1914: 1910: 1905: 1901: 1899:9780730983651 1895: 1891: 1887: 1886: 1878: 1876: 1872: 1867: 1863: 1856: 1853: 1847: 1839: 1836:Exemplars of 1833: 1830: 1823: 1821: 1819: 1818: 1813: 1812: 1807: 1806: 1802:'s 2003 film 1801: 1798: 1790: 1787: 1783: 1776: 1774: 1770: 1768: 1764: 1760: 1755: 1752: 1743: 1738: 1734: 1733: 1725: 1720: 1716: 1715: 1708: 1703: 1699: 1698: 1691: 1686: 1683: 1679: 1675: 1674: 1667: 1662: 1658: 1657: 1650: 1645: 1641: 1637: 1636: 1629: 1624: 1619: 1612: 1608: 1605: 1602: 1599: 1597: 1596: 1592: 1587: 1578: 1574: 1571: 1568: 1565: 1563: 1562: 1561:A. tricinctus 1558: 1557: 1554: 1551: 1549:Likely hybrid 1548: 1545: 1543: 1542: 1538: 1537: 1533: 1529: 1526: 1524: 1521: 1518: 1516: 1515: 1511: 1510: 1506: 1502: 1499: 1497: 1494: 1491: 1489: 1488: 1484: 1483: 1479: 1475: 1473: 1470: 1468: 1465: 1462: 1460: 1459: 1455: 1454: 1450: 1446: 1443: 1441: 1438: 1435: 1433: 1432: 1428: 1427: 1423: 1419: 1416: 1414: 1411: 1408: 1406: 1405: 1401: 1400: 1396: 1392: 1389: 1386: 1383: 1381: 1380: 1376: 1375: 1372: 1369: 1367: 1364: 1361: 1359: 1358: 1354: 1353: 1349: 1345: 1343: 1340: 1337: 1334: 1332: 1331: 1327: 1326: 1322: 1318: 1315: 1312: 1309: 1307: 1306: 1302: 1301: 1297: 1293: 1290: 1287: 1284: 1282: 1281: 1277: 1276: 1272: 1268: 1265: 1262: 1259: 1257: 1256: 1252: 1251: 1247: 1243: 1240: 1237: 1234: 1232: 1231: 1227: 1226: 1222: 1218: 1216: 1213: 1211: 1210:A. ephippium 1208: 1205: 1203: 1202: 1198: 1197: 1193: 1189: 1187: 1184: 1181: 1178: 1176: 1175: 1174:A. mccullochi 1171: 1170: 1166: 1162: 1159: 1157:Likely hybrid 1156: 1153: 1151: 1150: 1146: 1145: 1141: 1137: 1135: 1132: 1129: 1126: 1124: 1123: 1119: 1118: 1114: 1110: 1107: 1104: 1101: 1098: 1096: 1095: 1091: 1090: 1086: 1082: 1079: 1076: 1073: 1071: 1070: 1066: 1065: 1061: 1057: 1055: 1052: 1050: 1047: 1044: 1042: 1041: 1037: 1036: 1032: 1028: 1026: 1023: 1021: 1018: 1015: 1013: 1012: 1008: 1007: 1003: 999: 997: 994: 992: 989: 986: 984: 983: 979: 978: 974: 970: 968: 965: 962: 959: 956: 954: 953: 949: 948: 944: 940: 938: 935: 932: 929: 927: 926: 922: 921: 917: 913: 911: 908: 905: 902: 900: 899: 895: 894: 890: 886: 884: 881: 878: 875: 873: 872: 868: 867: 863: 859: 857: 854: 852: 851:A. ephippium 849: 846: 844: 843: 839: 838: 834: 830: 828: 825: 822: 819: 817: 816: 812: 811: 807: 803: 801: 798: 795: 793: 791: 790: 786: 785: 781: 777: 774: 772: 769: 766: 764: 763: 759: 758: 754: 740: 730: 722: 720: 718: 714: 713:introgression 710: 709:hybridization 706: 705:A. mccullochi 702: 698: 693: 688: 686: 682: 678: 674: 670: 666: 662: 661:A. mccullochi 658: 655:lineage, and 654: 650: 646: 642: 638: 634: 630: 626: 623: 619: 615: 611: 607: 603: 599: 595: 591: 587: 583: 579: 575: 571: 567: 563: 562:morphological 555: 553: 549: 545: 542: 534: 520: 517: 512: 504: 503: 497: 490: 488: 484: 482: 478: 474: 465: 458: 456: 454: 450: 446: 442: 438: 430: 429: 424: 423: 418: 414: 411: 407: 403: 399: 395: 394: 393: 390: 386: 379: 377: 375: 371: 370: 365: 361: 357: 353: 349: 341: 337: 334:nestled in a 333: 329: 322: 320: 318: 314: 310: 306: 298: 296: 252: 248: 245: 241: 240: 235: 234: 229: 225: 224:Pomacentridae 221: 217: 213: 209: 197: 193: 192: 188: 184: 180: 176: 175: 171: 170: 168: 163: 158: 153: 150: 147: 146: 143: 142:Pomacentridae 140: 137: 136: 133: 130: 127: 126: 123: 120: 117: 114: 113: 110: 107: 104: 103: 100: 97: 94: 93: 90: 87: 84: 83: 80: 77: 74: 73: 68: 63: 59: 55: 51: 47: 43: 38: 33: 30: 19: 3445: 3407: 3380:2318/1674109 3354: 3350: 3309:(1): 19491. 3306: 3302: 3247: 3243: 3210: 3203: 3191:. Retrieved 3187: 3178: 3155: 3149: 3138:the original 3133: 3127: 3114: 3102:. Retrieved 3098:the original 3093: 3083: 3071:. Retrieved 3056: 3049: 3030: 3021: 2996: 2992: 2986: 2953: 2949: 2943: 2931:. Retrieved 2924:the original 2915: 2908: 2881: 2874: 2863: 2857: 2846: 2835: 2829: 2818: 2783: 2779: 2769: 2724: 2720: 2697:20 September 2695:. Retrieved 2691:the original 2632: 2628: 2603:20 September 2601:. Retrieved 2596:Goemans, B. 2591: 2564: 2560: 2538: 2534: 2521: 2491:(1): 23–31. 2488: 2484: 2478: 2453: 2449: 2430:15 September 2428:. Retrieved 2424:the original 2413: 2401:. Retrieved 2390: 2360:(1): 23–31. 2357: 2353: 2349: 2343: 2333: 2328: 2316:. Retrieved 2312:the original 2301: 2287: 2282: 2249: 2245: 2239: 2212: 2208: 2198: 2161: 2157: 2147: 2112: 2108: 2098: 2086:. Retrieved 2082:the original 2077: 2071: 2064: 2037: 2033: 2023: 2017:blogspot.com 2010: 2005: 1996: 1988: 1976:. Retrieved 1972:the original 1967: 1958: 1928:(1): 43–51. 1925: 1921: 1904:the original 1884: 1866:the original 1855: 1837: 1832: 1817:A. ocellaris 1815: 1811:Finding Dory 1809: 1805:Finding Nemo 1803: 1794: 1786:Finding Nemo 1785: 1771: 1756: 1747: 1730: 1712: 1697:A. ephippium 1695: 1671: 1654: 1633: 1593: 1585: 1560: 1540: 1522: 1513: 1495: 1486: 1472:A. ephippium 1471: 1467:A. ephippium 1466: 1457: 1439: 1430: 1412: 1403: 1378: 1365: 1357:A. pacificus 1356: 1341: 1330:A. omanensis 1329: 1305:A. ocellaris 1304: 1280:A. ocellaris 1279: 1255:A. ocellaris 1254: 1229: 1215:A. ephippium 1214: 1209: 1201:A. melanopus 1200: 1186:A. ephippium 1185: 1173: 1148: 1133: 1121: 1103:Monospecific 1093: 1068: 1054:A. ephippium 1053: 1049:A. ephippium 1048: 1039: 1025:A. ephippium 1024: 1020:A. ephippium 1019: 1011:A. ephippium 1010: 995: 990: 981: 966: 961:Monospecific 950: 936: 924: 909: 897: 882: 871:A. bicinctus 870: 856:A. ephippium 855: 850: 841: 826: 814: 799: 789:A. akindynos 788: 770: 760: 752: 735:Common name 704: 701:A. akindynos 700: 689: 684: 680: 676: 672: 668: 664: 660: 657:A. akindynos 656: 653:monospecific 648: 645:A. omanensis 644: 640: 636: 632: 628: 625:A. bicinctus 624: 621: 614:A. tricintus 613: 609: 605: 559: 550: 546: 538: 508: 500: 487:after dusk. 485: 470: 459:Reproduction 441:cardinalfish 434: 426: 420: 400:rather than 391: 387: 383: 367: 345: 339: 302: 251:sea anemones 237: 231: 219: 211: 207: 206: 189: 172: 151: 128:(unranked): 115: 53: 29: 3485:iNaturalist 3470:Wikispecies 3403:(in German) 3351:Aquaculture 3104:31 December 2950:Coral Reefs 2088:19 December 1993:"Clownfish" 1978:19 December 1763:soft corals 1673:A. polymnus 1541:A. thiellei 1523:A. polymnus 1440:A. polymnus 1431:A. polymnus 1230:A. nigripes 1040:A. frenatus 669:A. nigripes 622:A . allardi 531:Video of a 417:coevolution 408:, or sting 406:nematocysts 356:zooplankton 295: in). 212:anemonefish 148:Subfamily: 132:Ovalentaria 122:Percomorpha 3533:Categories 2830:Amphiprion 2727:(1): 212. 2635:(1): 245. 2403:19 October 2318:10 January 1848:References 1656:A. clarkii 1640:melanistic 1635:A. percula 1527:Saddleback 1444:Saddleback 1379:A. percula 1342:A. clarkii 1182:Australian 1134:A. clarkii 1108:Saddleback 1080:A. clarkii 996:A. clarkii 991:A. clarkii 982:A. clarkii 967:A. clarkii 937:A. clarkii 910:A. clarkii 883:A. clarkii 842:A. barberi 827:A. clarkii 815:A. allardi 800:A. clarkii 796:Australian 753:Amphiprion 717:haplotypes 681:A. percula 610:A. clarkii 606:A. clarkii 590:saddleback 439:, certain 410:organelles 348:omnivorous 247:mutualisms 239:Amphiprion 174:Amphiprion 35:Clownfish 18:Clown fish 3544:Symbiosis 3272:2045-2322 3250:: 35461. 3129:Symbiosis 2583:2296-7745 2078:Marinebio 1751:lifecycle 1682:Indonesia 1390:Clownfish 1316:Clownfish 1291:Clownfish 1266:Clownfish 692:mutualism 690:Obligate 570:complexes 566:scalation 516:protandry 443:(such as 244:symbiotic 226:. Thirty 208:Clownfish 183:Schneider 85:Kingdom: 79:Eukaryota 3455:Wikidata 3413:Archived 3389:92357750 3357:: 1–11. 3341:31862916 3290:27748421 2978:25027153 2933:18 April 2865:FishBase 2837:FishBase 2810:22957165 2761:23122007 2669:25433367 2513:24516887 2382:24516887 2190:29643214 2164:(1876). 2131:23447664 2056:23447664 1950:53051081 1759:aquarium 1678:Sulawesi 1514:A. sebae 743:Complex 556:Taxonomy 511:polygamy 402:proteins 364:tunicate 360:copepods 138:Family: 99:Chordata 95:Phylum: 89:Animalia 75:Domain: 3503:5846181 3461:Q472616 3359:Bibcode 3332:6925283 3311:Bibcode 3281:5066260 3252:Bibcode 3193:5 April 3001:Bibcode 2958:Bibcode 2858:Premnas 2801:3434915 2752:3532366 2729:Bibcode 2660:4264551 2637:Bibcode 2493:Bibcode 2470:1443829 2362:Bibcode 2274:7801342 2254:Bibcode 2246:Toxicon 2217:Bibcode 2181:5904320 1995:at the 1930:Bibcode 1603:Percula 1586:Premnas 1572:Clarkii 1569:Clarkii 1387:Percula 1313:Percula 1288:Percula 1263:Percula 1105:lineage 1077:Indian 963:lineage 651:having 586:clarkii 574:percula 317:lagoons 305:Red Sea 290:⁄ 276:⁄ 262:⁄ 233:Premnas 228:species 191:Premnas 165:Genera 105:Class: 3516:714652 3490:343197 3387:  3339:  3329:  3288:  3278:  3270:  3218:  3166:  3073:11 May 3064:  3037:  2976:  2896:  2808:  2798:  2759:  2749:  2667:  2657:  2581:  2511:  2468:  2450:Copeia 2380:  2293:  2272:  2188:  2178:  2139:205352 2137:  2129:  2054:  1948:  1896:  1797:Disney 1767:corals 1606:Maroon 1584:Genus 1338:Indian 1238:Indian 1130:Indian 933:Indian 906:Indian 879:Indian 823:Indian 751:Genus 746:image 643:, and 594:maroon 592:, and 578:tomato 398:sugars 216:fishes 198:, 1816 196:Cuvier 185:, 1801 181:& 159:, 1975 3511:WoRMS 3385:S2CID 3141:(PDF) 3124:(PDF) 2974:S2CID 2927:(PDF) 2920:(PDF) 2531:(PDF) 2509:S2CID 2466:JSTOR 2378:S2CID 2135:S2CID 1946:S2CID 1824:Notes 1800:Pixar 1729:Male 1552:Skunk 1500:Skunk 1417:Skunk 1370:Skunk 1241:Skunk 1160:Skunk 775:Skunk 739:Clade 618:clade 582:skunk 541:spawn 374:algae 352:fecal 313:reefs 249:with 179:Bloch 157:Allen 116:Clade 3337:PMID 3286:PMID 3268:ISSN 3216:ISBN 3195:2016 3164:ISBN 3106:2016 3075:2015 3062:ISBN 3035:ISBN 2935:2013 2894:ISBN 2806:PMID 2757:PMID 2699:2015 2665:PMID 2605:2015 2579:ISSN 2454:1978 2432:2013 2405:2023 2320:2018 2291:ISBN 2270:PMID 2186:PMID 2127:PMID 2090:2011 2052:PMID 1980:2011 1894:ISBN 711:and 703:and 683:and 612:and 415:The 362:and 323:Diet 214:are 3375:hdl 3367:doi 3355:498 3327:PMC 3319:doi 3276:PMC 3260:doi 3009:doi 2966:doi 2886:doi 2796:PMC 2788:doi 2747:PMC 2737:doi 2655:PMC 2645:doi 2569:doi 2501:doi 2458:doi 2370:doi 2352:". 2262:doi 2225:doi 2176:PMC 2166:doi 2162:285 2117:doi 2113:216 2042:doi 2038:216 1938:doi 1795:In 447:), 210:or 3535:: 3513:: 3500:: 3487:: 3472:: 3457:: 3383:. 3373:. 3365:. 3353:. 3335:. 3325:. 3317:. 3305:. 3301:. 3284:. 3274:. 3266:. 3258:. 3246:. 3242:. 3186:. 3158:. 3134:10 3132:. 3126:. 3092:. 3007:. 2997:34 2995:. 2972:. 2964:. 2954:24 2952:. 2892:. 2862:. 2834:. 2804:. 2794:. 2782:. 2778:. 2755:. 2745:. 2735:. 2725:12 2723:. 2719:. 2707:^ 2677:^ 2663:. 2653:. 2643:. 2633:14 2631:. 2627:. 2613:^ 2577:. 2567:. 2563:. 2559:. 2547:^ 2539:41 2537:. 2533:. 2507:. 2499:. 2489:57 2487:. 2464:. 2452:. 2440:^ 2376:. 2368:. 2358:57 2356:. 2268:. 2260:. 2250:32 2248:. 2223:. 2213:35 2211:. 2207:. 2184:. 2174:. 2160:. 2156:. 2133:. 2125:. 2111:. 2107:. 2076:. 2050:. 2036:. 2032:. 1966:. 1944:. 1936:. 1926:38 1924:. 1912:^ 1892:. 1888:. 1874:^ 1680:, 1588:: 755:: 639:, 635:, 631:, 627:, 588:, 584:, 580:, 576:, 572:: 455:. 376:. 118:: 56:) 3391:. 3377:: 3369:: 3361:: 3343:. 3321:: 3313:: 3307:9 3292:. 3262:: 3254:: 3248:6 3197:. 3172:. 3108:. 3077:. 3043:. 3015:. 3011:: 3003:: 2980:. 2968:: 2960:: 2937:. 2902:. 2888:: 2860:" 2832:" 2812:. 2790:: 2784:2 2763:. 2739:: 2731:: 2701:. 2671:. 2647:: 2639:: 2607:. 2585:. 2571:: 2565:9 2515:. 2503:: 2495:: 2472:. 2460:: 2434:. 2407:. 2384:. 2372:: 2364:: 2322:. 2276:. 2264:: 2256:: 2233:. 2227:: 2219:: 2192:. 2168:: 2141:. 2119:: 2092:. 2074:" 2058:. 2044:: 1982:. 1952:. 1940:: 1932:: 412:. 342:) 338:( 292:4 288:1 285:+ 283:3 281:– 278:4 274:3 271:+ 269:2 264:2 260:1 257:+ 255:6 52:( 20:)

Index

Clown fish

Ocellaris clownfish
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Actinopterygii
Percomorpha
Ovalentaria
Pomacentridae
Amphiprioninae
Allen
Amphiprion
Bloch
Schneider
Premnas
Cuvier
fishes
Pomacentridae
species
Premnas
Amphiprion
symbiotic
mutualisms
sea anemones
Red Sea
Great Barrier Reef
reefs

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