833:
147:
129:
115:
545:
If after 1999 a new family-group name is based on a generic name which is or ends in a Greek or Latin word or ends in a Greek or Latin suffix, but its derivation does not follow the grammatical procedures of
Articles 29.3.1 or 29.3.2, its original spelling must be maintained as the correct original
402:
Kirejtshuk, A.G.; Poschmann, M.; Prokop, J.; Garrouste, R.; Nel, A. (2014). "Evolution of the elytral venation and structural adaptations in the oldest
Palaeozoic beetles (Insecta: Coleoptera: Tshekardocoleidae)".
385:, Article 29.4, family-rank names originally published after 1999 are not subject to emendation because of incorrect spelling, so Coleopsidae would be maintained as the correct original spelling.
377:
to all other beetles. The family name was originally published in 2016 as "Coleopsidae" but a later publication claimed that this spelling was erroneous and proposed the spelling "
618:
933:
874:
373:, this interpretation has been revised and questioned, with the most recent analyses suggesting that it is best placed in its own family, and may be the
782:
700:
605:
918:
913:
923:
498:"Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea"
938:
898:
867:
465:(Insecta: Coleoptera) – morphological re-evaluation using Reflectance Transformation Imaging (RTI) and phylogenetic assessment"
928:
893:
860:
146:
908:
903:
832:
691:
558:
354:
808:
726:
644:
282:
307:
224:
141:
813:
731:
649:
623:
795:
713:
631:
370:
800:
773:
718:
636:
509:
476:
412:
211:
596:
361:, estimated to be about 297 million years old. It is currently the oldest known beetle.
844:
431:
887:
72:
676:
374:
758:
581:
539:
416:
767:
685:
590:
514:
497:
128:
47:
114:
433:Происхождение отряда жуков (Coleoptera) и значение палеонтологической летописи
332:
92:
57:
178:
158:
122:
Holotype fossil showing part (A) counterpart (C) and virtually combined (B)
97:
17:
481:
460:
752:
670:
575:
436:[Origin of the Coleoptera and significance of the fossil record]
350:
87:
82:
67:
62:
52:
787:
705:
610:
358:
346:
102:
77:
136:
Interpretive drawing of the part (A) counterpart (C) and combined (B)
840:
198:
188:
168:
552:
328:
382:
556:
848:
369:
While originally interpreted as a member of the family
345:. It is known from a single specimen from the Early
742:
660:
565:
459:Schädel, M.; Yavorskaya, M.; Beutel, R. (2022).
868:
8:
315:Coleopseidae Kirejtshuk, 2020 (Unj. Emend.)
543:29.4. Acceptance of originally formed stem.
875:
861:
553:
127:
113:
31:
513:
480:
532:
394:
341:, and is the only member of the family
335:beetles. It contains a single species,
469:Arthropod Systematics & Phylogeny
7:
829:
827:
934:Monotypic prehistoric insect genera
405:Journal of Systematic Palaeontology
25:
430:Kirejtshuk, A.G.; Nel, A (2016).
831:
145:
442:Euroasian Entomological Journal
1:
919:Cisuralian animals of Europe
914:Permian arthropods of Europe
847:. You can help Knowledge by
417:10.1080/14772019.2013.821530
924:Paleozoic insects of Europe
515:10.3390/geosciences10020073
955:
826:
242:Kirejtshuk & Nel, 2016
496:Kirejtshuk, A.G. (2020).
313:
306:
288:
281:
142:Scientific classification
140:
135:
126:
121:
112:
34:
939:Prehistoric beetle stubs
432:
365:Taxonomy and systematics
27:Extinct genus of beetles
899:Monotypic beetle genera
301:Kirejtshuk et al., 2014
261:Kirejtshuk et al., 2014
843:-related article is a
461:"The earliest beetle †
448:(Supplement 1): 66–73.
381:"; however, under the
809:Paleobiology Database
727:Paleobiology Database
645:Paleobiology Database
482:10.3897/asp.80.e86582
929:Fossil beetle genera
355:Meisenheim Formation
894:Tshekardocoleoidea
692:Coleopsis archaica
662:Coleopsis archaica
463:Coleopsis archaica
338:Coleopsis archaica
295:Coleopsis archaica
225:Tshekardocoleoidea
856:
855:
824:
823:
796:Open Tree of Life
714:Open Tree of Life
632:Open Tree of Life
559:Taxon identifiers
371:Tshekardocoleidae
320:
319:
262:
243:
16:(Redirected from
946:
877:
870:
863:
835:
828:
817:
816:
804:
803:
791:
790:
778:
777:
776:
763:
762:
761:
735:
734:
722:
721:
709:
708:
696:
695:
694:
681:
680:
679:
653:
652:
640:
639:
627:
626:
614:
613:
601:
600:
599:
586:
585:
584:
554:
547:
537:
520:
519:
517:
493:
487:
486:
484:
456:
450:
449:
439:
427:
421:
420:
399:
297:
293:
274:C. archaica
260:
253:
241:
236:
223:
210:
150:
149:
131:
117:
107:
44:
40:Temporal range:
32:
21:
954:
953:
949:
948:
947:
945:
944:
943:
909:Permian insects
904:Asselian genera
884:
883:
882:
881:
825:
820:
812:
807:
799:
794:
786:
781:
772:
771:
766:
757:
756:
751:
738:
730:
725:
717:
712:
704:
699:
690:
689:
684:
675:
674:
669:
656:
648:
643:
635:
630:
622:
617:
609:
604:
595:
594:
589:
580:
579:
574:
561:
551:
550:
538:
534:
529:
524:
523:
495:
494:
490:
458:
457:
453:
437:
434:
429:
428:
424:
401:
400:
396:
391:
367:
302:
299:
291:
290:
277:
259:
251:
240:
234:
221:
212:Protocoleoptera
208:
144:
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
42:
41:
38:
28:
23:
22:
15:
12:
11:
5:
952:
950:
942:
941:
936:
931:
926:
921:
916:
911:
906:
901:
896:
886:
885:
880:
879:
872:
865:
857:
854:
853:
836:
822:
821:
819:
818:
805:
792:
779:
764:
748:
746:
740:
739:
737:
736:
723:
710:
697:
682:
666:
664:
658:
657:
655:
654:
641:
628:
615:
602:
587:
571:
569:
563:
562:
557:
549:
548:
531:
530:
528:
525:
522:
521:
488:
451:
444:(in Russian).
422:
411:(5): 575–600.
393:
392:
390:
387:
366:
363:
327:is an extinct
318:
317:
311:
310:
304:
303:
300:
286:
285:
279:
278:
270:
268:
264:
263:
249:
245:
244:
232:
228:
227:
219:
215:
214:
206:
202:
201:
196:
192:
191:
186:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
133:
132:
124:
123:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
951:
940:
937:
935:
932:
930:
927:
925:
922:
920:
917:
915:
912:
910:
907:
905:
902:
900:
897:
895:
892:
891:
889:
878:
873:
871:
866:
864:
859:
858:
852:
850:
846:
842:
837:
834:
830:
815:
810:
806:
802:
797:
793:
789:
784:
780:
775:
769:
765:
760:
754:
750:
749:
747:
745:
741:
733:
728:
724:
720:
715:
711:
707:
702:
698:
693:
687:
683:
678:
672:
668:
667:
665:
663:
659:
651:
646:
642:
638:
633:
629:
625:
620:
616:
612:
607:
603:
598:
592:
588:
583:
577:
573:
572:
570:
568:
564:
560:
555:
544:
541:
536:
533:
526:
516:
511:
507:
503:
499:
492:
489:
483:
478:
474:
470:
466:
464:
455:
452:
447:
443:
435:
426:
423:
418:
414:
410:
406:
398:
395:
388:
386:
384:
380:
376:
372:
364:
362:
360:
356:
352:
348:
344:
340:
339:
334:
330:
326:
325:
316:
312:
309:
305:
298:
296:
287:
284:
283:Binomial name
280:
276:
275:
269:
266:
265:
258:
257:
250:
247:
246:
239:
233:
230:
229:
226:
220:
218:Superfamily:
217:
216:
213:
207:
204:
203:
200:
197:
194:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
148:
143:
139:
134:
130:
125:
120:
116:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
37:
33:
30:
19:
849:expanding it
839:This fossil
838:
743:
661:
566:
542:
535:
505:
501:
491:
472:
468:
462:
454:
445:
441:
425:
408:
404:
397:
379:Coleopseidae
378:
375:sister group
368:
342:
337:
336:
323:
322:
321:
314:
294:
289:
273:
272:
255:
254:
237:
35:
29:
774:Coleopsidae
768:Wikispecies
744:Coleopsidae
686:Wikispecies
591:Wikispecies
502:Geosciences
475:: 495–510.
357:of western
343:Coleopsidae
238:Coleopsidae
18:Coleopsidae
888:Categories
677:Q110761357
389:References
333:stem-group
205:Suborder:
199:Coleoptera
179:Arthropoda
759:Q33137206
597:Coleopsis
582:Q21215039
567:Coleopsis
540:ICZN Code
508:(2): 73.
324:Coleopsis
267:Species:
256:Coleopsis
165:Kingdom:
159:Eukaryota
36:Coleopsis
753:Wikidata
671:Wikidata
624:11913896
576:Wikidata
546:spelling
351:Asselian
308:Synonyms
231:Family:
175:Phylum:
169:Animalia
155:Domain:
43:Asselian
801:7906178
788:9295835
719:6262038
706:8488415
637:6262037
611:8583016
359:Germany
347:Permian
292:†
271:†
248:Genus:
195:Order:
189:Insecta
185:Class:
841:beetle
814:346704
732:295120
650:295119
619:IRMNG
527:Notes
438:(PDF)
329:genus
845:stub
783:GBIF
701:GBIF
606:GBIF
383:ICZN
48:PreꞒ
510:doi
477:doi
413:doi
331:of
890::
811::
798::
785::
770::
755::
729::
716::
703::
688::
673::
647::
634::
621::
608::
593::
578::
506:10
504:.
500:.
473:80
471:.
467:.
446:15
440:.
409:12
407:.
353:)
98:Pg
876:e
869:t
862:v
851:.
518:.
512::
485:.
479::
419:.
415::
349:(
252:†
235:†
222:†
209:†
103:N
93:K
88:J
83:T
78:P
73:C
68:D
63:S
58:O
53:Ꞓ
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
