Knowledge (XXG)

Theridiidae

Source πŸ“

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are also model organisms, used for the study of sociality, because it has evolved frequently within the genus, allowing comparative studies across species, and because it contains species varying from solitary to permanently social. These spiders are also a promising model for the study of inbreeding
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Agnarsson I. 2006c. Phylogenetic placement of Echinotheridion (Araneae: Theridiidae) – do male sexual organ removal, emasculation, and sexual cannibalism in Echinotheridion and Tidarren represent evolutionary replicas? Invertebrate Systematics 20: 415–429.
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Arnedo, M.A., Coddington, J., Agnarsson, I. & Gillespie, R.G. (2004). From a comb to a tree: phylogenetic relationships of the comb-footed spiders (Araneae, Theridiidae) inferred from nuclear and mitochondrial genes.
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Arnedo MA, Agnarsson I, Gillespie RG. In Press. Molecular insights into the phylogenetic structure of the spider genus Theridion (Araneae, Theridiidae) and the origin of the Hawaiian Theridion-like fauna. Zoologica
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Benjamin, Suresh P.; Zschokke, Samuel (2002). "Untangling the tangle-web: web building behaviour of the comb-footed spider Steatoda triangulosa and comments on phylogenetic implications (Araneae: Theridiidae)".
425:) live in the webs of larger spiders and pilfer small prey caught by their host's web. They eat prey killed by the host spider, consume silk from the host web, and sometimes attack and eat the host itself. 2717:
Oxford, G.S. and Gillespie, R.G. 1996. Genetics of a colour polymorphism in Theridion grallator (Araneae: Theridiidae), the Hawaiian happy-face spider, from greater Maui. Heredity, 76: 238–248.
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Aviles, L., Maddison, W.P. and Agnarsson, I. 2006. A new independently derived social spider with explosive colony proliferation and a female size dimorphism. Biotropica, 38: 743–753.
329:. They are important to studies characterizing their venom and its clinical manifestation, but widow spiders are also used in research on spider silk and sexual biology, including 2714:
Gillespie, R.G. and Tabashnik, B.E. 1994. Foraging Behavior of the Hawaiian Happy Face Spider (Araneae, Theridiidae). Annals of the Entomological Society of America, 87: 815–822.
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trap lines leading to the soil surface. Webs remain in place for extended periods and are expanded and repaired, but no regular pattern of web replacement has been observed.
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Agnarsson I. 2004. Morphological phylogeny of cobweb spiders and their relatives (Araneae, Araneoidea, Theridiidae). Zoological Journal of the Linnean Society 141: 447–626.
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Grostal, P.; Walter, D.E. (1997). "Kleptoparasites or commensals? Effects of Argyrodes antipodianus (Araneae: Theridiidae) on nephila plumipes (Araneae: Tetragnathidae)".
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Theridiid gumfoot-webs consist of frame lines that anchor them to surroundings and of support threads, which possess viscid silk. These can either have a central retreat (
4002: 4041: 4159: 2473:"Quasistatic and continuous dynamic characterization of the mechanical properties of silk from the cobweb of the black widow spider Latrodectus hesperus" 323: 3976: 4015: 2244:"Quantum Shifts in the Genetic Control of a Colour Polymorphism in Theridion Grallator (Araneae: Theridiidae), the Hawaiian Happy-Face Spider" 2512:
Blackledge, T.A.; Zevenbergen, J.M. (2007). "Condition dependent spider web architecture in the western black widow Latrodectus hesperus".
2674: 2613:"The fossil record of spiders revisited: implications for calibrating trees and evidence for a major faunal turnover since the Mesozoic" 2201:"A revision of the New World eximius lineage of Anelosimus (Araneae, Theridiidae) and a phylogenetic analysis using worldwide exemplars" 2285:"What makes a happy face? Determinants of color pattern in the Hawaiian happy face spider Theridion grallator (Araneae, Theridiidae)" 2119:"Asymmetric female genitalia and other remarkable morphology in a new genus of cobweb spiders (Theridiidae, Araneae) from Madagascar" 2769: 438:-type). Building gum-foot lines is a unique, stereotyped behaviour, and is likely homologous for Theridiidae and its sister family 4113: 4164: 4020: 2702: 2200: 4028: 2687: 2908: 3885: 3862: 1997: 466: 241: 134: 3955: 4046: 352:
is used as a model to understand the selective forces and the genetic basis of color polymorphism within species.
112: 3968: 1624:– Asia, Africa, South America, Oceania, North America, Europe, Central America, Saint Vincent and the Grenadines 507: 3802: 2732: 512: 483: 357: 2385:
Vollrath, F. (1979). "Behavior of the Kleptoparasitic Spider Argyrodes-Elevatus (Araneae, Theridiidae)".
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Magalhaes, Ivan L. F.; Azevedo, Guilherme H. F.; Michalik, Peter; RamΓ­rez, MartΓ­n J. (February 2020).
2062:"The Habitats Humans Provide: Factors affecting the diversity and composition of arthropods in houses" 561: 500: 4154: 4126: 4118: 4056: 3924: 3849: 3207: 2589: 2429: 2163: 2073: 596: 524: 281: 221: 1976: 519: 3515: 592: 536: 476: 348: 307:, meaning that they spin sticky capture silk instead of woolly silk. They have a comb of serrated 3580: 3424: 2916: 2648: 2529: 2453: 2402: 2367: 2265: 488: 330: 129: 121: 2152:"Smaller colonies and more solitary living mark higher elevation populations of a social spider" 1974:
genera have also been placed in the family. The oldest known stem-group member of the family is
4082: 2671: 556: 4064: 3911: 3638: 3432: 3317: 3138: 2640: 2632: 2494: 2445: 2181: 2099: 1628: 715: 531: 4105: 4007: 3840: 1063:– Asia, South America, Europe, North America, New Zealand, Central America, Africa, Caribbean 4069: 3357: 3331: 3125: 2992: 2755: 2624: 2612: 2560: 2521: 2484: 2437: 2394: 2359: 2329: 2296: 2255: 2215: 2171: 2130: 2089: 2081: 1826: 400: 249: 2581: 3779: 3712: 3624: 3409: 3199: 3143: 2984: 2961: 2691: 2678: 1353: 1188: 1023: 759: 1844:– Asia, North America, Central America, Europe, South America, Africa, Oceania, Caribbean 997:– North America, Oceania, Asia, Central America, South America, Caribbean, Africa, Europe 2433: 2167: 2077: 3963: 3950: 3601: 3560: 3489: 3236: 3104: 3080: 3070: 3013: 2900: 2094: 2061: 1903: 1320: 1122: 544: 4148: 3565: 3552: 3494: 3415: 3401: 3375: 3365: 3336: 3223: 3194: 3112: 3043: 2892: 2880: 2868: 2845: 2652: 2565: 2548: 2398: 2334: 2317: 2220: 2176: 2151: 2135: 2118: 1985: 1877:– South America, Central America, Asia, North America, Caribbean, Greenland, Tanzania 1606: 1518: 1419: 1397: 1177: 638: 421: 316: 274: 206: 70: 3854: 2533: 2457: 2406: 2371: 1547:– Asia, Africa, North America, Caribbean, South America, Oceania, Europe, Costa Rica 3867: 3735: 3730: 3707: 3692: 3633: 3570: 3440: 3308: 3287: 3244: 3171: 3133: 2966: 2956: 2924: 2837: 1716: 1672: 1584: 1507: 1045: 979: 880: 847: 803: 781: 770: 581: 576: 230: 2525: 2269: 292:, and is the most common arthropod found in human dwellings throughout the world. 17: 3916: 964:– South America, North America, Oceania, Central America, Asia, Trinidad, Belgium 689:– Asia, Africa, North America, South America, Oceania, Central America, Caribbean 4033: 3989: 3834: 3702: 3646: 3610: 3539: 3523: 3472: 3463: 3454: 3295: 3265: 3161: 3021: 2804: 2225: 1947: 1727: 1661: 1639: 1573: 1540: 1474: 1452: 1364: 1298: 1232: 1144: 1133: 1034: 968: 957: 935: 891: 814: 693: 671: 627: 616: 605: 587: 430: 404: 326: 300: 176: 45: 2684: 2085: 3750: 3741: 3722: 3683: 3674: 3595: 3547: 3531: 3507: 3502: 3481: 3445: 3396: 3380: 3349: 3341: 3278: 3215: 3166: 3096: 3088: 3075: 3062: 3005: 2979: 2948: 2932: 2826: 2363: 1981: 1925: 1859: 1804: 1738: 1705: 1694: 1650: 1595: 1562: 1496: 1441: 1430: 1386: 1309: 1210: 1089: 1074:– Asia, North America, South America, Jamaica, Europe, Oceania, Africa, Panama 946: 825: 682: 439: 381: 377: 335: 90: 55: 2636: 3937: 3697: 3651: 3615: 3370: 3228: 3185: 3179: 3152: 3026: 2974: 2798: 1958: 1870: 1848: 1837: 1815: 1617: 1551: 1529: 1485: 1408: 1254: 1199: 1078: 1012: 1001: 924: 913: 836: 704: 649: 571: 453: 447: 409: 304: 166: 146: 95: 3825: 2644: 2498: 2449: 2185: 2103: 2747: 2441: 3898: 3819: 3717: 3575: 3326: 3303: 3120: 2810: 1892: 1881: 1782: 1760: 1749: 1463: 1287: 1276: 1265: 1166: 1155: 1100: 1067: 1056: 869: 858: 415: 186: 85: 80: 65: 60: 50: 2301: 2284: 2260: 2243: 3877: 3588: 3388: 2997: 1971: 1793: 1342: 1331: 1111: 990: 909:– North America, South America, Central America, Asia, Trinidad, Europe 902: 792: 748: 737: 726: 451:-type) and some that are sheet-like, which do not contain viscid silk ( 308: 285: 284:
in 1833. This diverse, globally distributed family includes over 3,000
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The family includes some model organisms for research, including the
296: 277: 196: 156: 3981: 3796: 2035: 1855:– Spain, Africa, North America, Central America, Asia, South America 3942: 1558:– North America, Asia, Africa, Jamaica, Panama, Brazil, New Zealand 4090: 2549:"Webs of theridiid spiders: construction, structure and evolution" 2318:"Webs of theridiid spiders: construction, structure and evolution" 1936: 1771: 1221: 555: 543: 530: 518: 506: 494: 482: 470: 380:, hence the common name, but Theridiidae has a large diversity of 365: 289: 1767:– South America, North America, Central America, Caribbean, Asia 1107:– South America, North America, Central America, Caribbean, Asia 898:– North America, South America, Central America, Caribbean, Asia 722:– Costa Rica, South America, Asia, Africa, Oceania, Mexico, Cuba 393: 312: 3800: 2865: 2823: 2751: 1052:– Asia, Europe, Australia, Africa, North America, South America 711:– Africa, Asia, Oceania, North America, South America, Jamaica 445:
Among webs without gumfooted lines, some contain viscid silk (
385: 1756:– Oceania, North America, Asia, Europe, South America, Africa 1305:– South America, North America, Asia, Europe, Oceania, Africa 920:– United States, South America, Seychelles, Asia, New Zealand 3929: 1514:– Asia, Oceania, Cuba, North America, Argentina, Seychelles 591:, is another large genus that includes the North American 340:
because all permanently social species are highly inbred.
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Blackledge, T.A.; Swindeman, J.E.; Hayashi, C.Y. (2005).
612:– Africa, Asia, Australia, South America, Central America 1866:– Panama, South America, Africa, Asia, Oceania, Trinidad 457:-type). However, there are many undescribed web forms. 2582:"A summary list of fossil spiders and their relatives" 1789:– United States, South America, Central America, Congo 1888:– Africa, Yemen, North America, Argentina, Costa Rica 1404:– North America, Asia, South America, Central America 303:, meaning that the females have a genital plate, and 1294:– Europe, North America, Panama, South America, Asia 565:
moving from one tree to another carrying the egg sac
3809: 3264: 3051: 3042: 2879: 656:– Central America, Caribbean, Mexico, South America 1250:– Asia, South America, Australia, Panama, Trinidad 1393:– Panama, Caribbean, South America, North America 1184:– Caribbean, Panama, North America, South America 2969:(funnel-webs or venomous funnel-web tarantulas) 953:– Ukraine, United States, Africa, Oceania, Asia 356:is known as the "happyface" spider, as certain 2547:Benjamin, Suresh P.; Zschokke, Samuel (2003). 2763: 1536:– Oceania, North America, Asia, South America 8: 3360:(wandering spiders or tropical wolf spiders) 2580:Dunlop, J.A.; Penney, D.; Jekel, D. (2015). 3797: 3270: 3055: 3048: 2885: 2876: 2862: 2820: 2770: 2756: 2748: 248: 111: 31: 2919:(atypical tarantulas or purseweb spiders) 2728:Platnick, N.I. 2006. World Spider Catalog 2564: 2553:Biological Journal of the Linnean Society 2488: 2333: 2322:Biological Journal of the Linnean Society 2300: 2283:Gillespie, R.G.; Tabashnik, B.E. (1989). 2259: 2219: 2208:Zoological Journal of the Linnean Society 2175: 2134: 2123:Biological Journal of the Linnean Society 2093: 3787:are families with more than 1000 species 1096:– Central America, South America, Mexico 2009: 1041:– North America, Asia, Papua New Guinea 2242:Oxford, G.S.; Gillespie, R.G. (1996). 2030: 2028: 1910:– North America, South America, Panama 733:– North America, Asia, Europe, Algeria 2699:Molecular Phylogenetics and Evolution 2316:Benjamin, S.P.; Zschokke, S. (2003). 2036:"Family: Theridiidae Sundevall, 1833" 1745:– Pakistan, Caribbean, Mexico, Brazil 1391:Marques, Buckup & Rodrigues, 2011 575:with over 600 species, but it is not 7: 4096:1FFE834E-0B7C-463D-E53A-594E93D0C1DD 4057:a3789e64-304e-4e95-8953-b9786ae40db8 3969:94bd443b-35d5-4428-af44-c93cb055c9cd 1635:– Europe, North America, Asia, Congo 4160:Extant Cretaceous first appearances 1448:– Asia, Seychelles, Oceania, Mexico 3218:(violin spiders, assassin spiders) 2060:Leong, Misha; et al. (2017). 25: 2733:Key to several Theridiidae genera 1833:– Tanzania, Europe, North America 1162:– South America, Panama, Trinidad 3457:(sheet weavers or money spiders) 2566:10.1046/j.1095-8312.2003.00110.x 2335:10.1046/j.1095-8312.2003.00110.x 2221:10.1111/j.1096-3642.2006.00213.x 2177:10.1111/j.1365-2656.2007.01228.x 2150:Purcell, J.; Aviles, L. (2007). 2136:10.1111/j.1095-8312.2006.00569.x 434:-type) or a peripheral retreat ( 133: 4132:urn:lsid:nmbe.ch:spiderfam:0099 2911:(Australian funnel-web spiders) 2477:Journal of Experimental Biology 775:Deeleman & Wunderlich, 2011 1008:– Panama, South America, Malta 841:Thaler & Steinberger, 1988 755:– Guinea-Bissau, Congo, Brazil 599:accepts the following genera: 376:They often build tangle space 1: 3418:(flat-bellied ground spiders) 2895:(mouse spiders and relatives) 2593:. Natural History Museum Bern 2526:10.1016/j.anbehav.2006.10.014 2042:. Natural History Museum Bern 1943:– Papua New Guinea, Australia 1822:– United States, Brazil, Asia 1580:– North America, Asia, Africa 1281:Barrion & Litsinger, 1995 1118:– Papua New Guinea, Australia 621:Barrion & Litsinger, 1995 595:. As of April 2019, the 3281:(araneomorph funnel weavers) 2399:10.1016/0003-3472(79)90186-6 1569:– Asia, United States, Spain 360:have a pattern resembling a 2935:(cork-lid trapdoor spiders) 1998:List of Theridiidae species 1886:Chamberlin & Ivie, 1934 1360:– Oceania, Asia, Seychelles 865:– Brazil, Venezuela, Guyana 467:List of Theridiidae species 368:face on their yellow body. 237: 4181: 3231:(long-legged cave spiders) 2927:(brushed trapdoor spiders) 2903:(folding trapdoor spiders) 2352:Journal of Insect Behavior 2086:10.1038/s41598-017-15584-2 1908:O. Pickard-Cambridge, 1896 1864:O. Pickard-Cambridge, 1881 1765:O. Pickard-Cambridge, 1894 1633:O. Pickard-Cambridge, 1879 1556:O. Pickard-Cambridge, 1879 1358:O. Pickard-Cambridge, 1871 1151:– Oceania, Belgium, Taiwan 918:O. Pickard-Cambridge, 1882 907:O. Pickard-Cambridge, 1882 852:O. Pickard-Cambridge, 1871 464: 388:and other ground dwelling 3775: 3273: 3249:Trogloraptor marchingtoni 3058: 2888: 2875: 2861: 2833: 2819: 2787: 2738:Tree of Life: Theridiidae 2156:Journal of Animal Ecology 2017:Sundevall, C. J. (1833). 975:– Samoa, Australia, India 247: 236: 229: 130:Scientific classification 128: 119: 110: 34: 3725:(cribellate orb weavers) 3659:(long jawed orb-weavers) 3311:(anyphaenid sac spiders) 3107:(crevice weaver spiders) 2943:(wafer trapdoor spiders) 1338:– Asia, Papua New Guinea 678:– Africa, Europe, Turkey 3526:(red and black spiders) 3466:(liocranid sac spiders) 3210:(tube-dwelling spiders) 3146:(midget ground weavers) 3008:(funnel-web tarantulas) 3000:(tree trapdoor spiders) 2951:(funnel-web tarantulas) 2364:10.1023/A:1021175507377 1921:– Mexico, South America 1699:KoΓ§ak & Kemal, 2008 1578:KoΓ§ak & Kemal, 2008 1503:– Europe, Algeria, Asia 1415:– Asia, Europe, Algeria 1272:– Chile, Turkey, Brazil 1195:– Turkey, Russia, China 786:Yoshida & Koh, 2011 273:, is a large family of 4165:Araneomorphae families 3555:(running crab spiders) 3448:(white-tailed spiders) 3290:(tangled nest spiders) 3115:(large-clawed spiders) 3091:(false violin spiders) 2199:Agnarsson, I. (2006). 2117:Agnarsson, I. (2006). 1712:– Europe, Asia, Canada 1525:– United States, Chile 843:– Europe, Angola, Asia 566: 553: 541: 540:, a black widow spider 528: 516: 504: 492: 480: 392:using elastic, sticky 4078:Paleobiology Database 3583:(nursery web spiders) 3518:(cave cobweb spiders) 3510:(spurred orb-weavers) 3427:(dwarf sheet spiders) 3383:(net-casting spiders) 3174:(palp-footed spiders) 3016:(bald-legged spiders) 2442:10.1007/s004420050273 2019:Conspectus Arachnidum 1657:– Asia, North America 569:The largest genus is 559: 547: 534: 522: 510: 498: 486: 474: 3964:Fauna Europaea (new) 3668:(tangle-web spiders) 3435:(tree trunk spiders) 3391:(intertidal spiders) 3320:(orb-weaver spiders) 2743:Theridiidae research 2590:World Spider Catalog 2040:World Spider Catalog 1019:– Asia, Sierra Leone 597:World Spider Catalog 513:Enoplognatha abrupta 282:Carl Jakob Sundevall 261:, also known as the 3641:(dwarf orb-weavers) 3128:(lampshade spiders) 3099:(woodlouse hunters) 2848:(segmented spiders) 2840:(segmented spiders) 2434:1997Oecol.111..570G 2302:10.1038/hdy.1989.50 2261:10.1038/hdy.1996.38 2168:2007JAnEc..76..590P 2078:2017NatSR...715347L 1481:– Greenland, Russia 1437:– Asia, New Zealand 593:common house spider 550:Theridion impressum 537:Latrodectus mactans 477:Chrysso pulcherrima 349:Theridion grallator 324:medically important 319:of the fourth leg. 280:first described by 271:comb-footed spiders 3675:Theridiosomatidae 3639:Symphytognathidae 3627:(huntsman spiders) 3534:(disc web spiders) 3352:(dark sac spiders) 3202:(spitting spiders) 2987:(dwarf tarantulas) 2690:2007-01-01 at the 2677:2007-09-27 at the 2617:Biological Reviews 2066:Scientific Reports 1899:– Indonesia, India 1679:– Chile, Argentina 1602:– India, Sri Lanka 1140:– Brazil, Paraguay 854:– Sri Lanka, China 567: 554: 542: 529: 517: 505: 493: 489:Coscinida japonica 481: 331:sexual cannibalism 263:tangle-web spiders 122:Enoplognatha ovata 41:Cretaceous–present 35:Tangle-web spiders 18:Comb-footed spider 4142: 4141: 4065:Open Tree of Life 3803:Taxon identifiers 3794: 3793: 3771: 3770: 3767: 3766: 3763: 3762: 3759: 3758: 3745: 3726: 3687: 3678: 3669: 3660: 3642: 3628: 3619: 3605: 3604:(jumping spiders) 3592: 3584: 3556: 3543: 3535: 3527: 3519: 3511: 3498: 3485: 3476: 3467: 3458: 3449: 3436: 3428: 3419: 3405: 3392: 3384: 3361: 3353: 3345: 3321: 3312: 3299: 3298:(termite hunters) 3291: 3282: 3260: 3259: 3252: 3245:Trogloraptoridae 3240: 3239:(armored spiders) 3232: 3219: 3211: 3203: 3189: 3175: 3156: 3147: 3139:Mecysmaucheniidae 3129: 3116: 3108: 3100: 3092: 3084: 3083:(coneweb spiders) 3066: 3065:(pelican spiders) 3038: 3037: 3030: 3029:(true tarantulas) 3017: 3009: 3001: 2988: 2985:Mecicobothriidae 2970: 2952: 2944: 2936: 2928: 2920: 2912: 2904: 2896: 2857: 2856: 2849: 2841: 2629:10.1111/brv.12559 2490:10.1242/jeb.01597 2483:(10): 1937–1949. 1964: 1953: 1942: 1931: 1920: 1909: 1898: 1887: 1876: 1865: 1854: 1843: 1832: 1821: 1810: 1799: 1788: 1777: 1766: 1755: 1744: 1733: 1722: 1711: 1700: 1689: 1678: 1667: 1656: 1645: 1634: 1623: 1612: 1601: 1590: 1579: 1568: 1557: 1546: 1535: 1524: 1513: 1502: 1491: 1480: 1469: 1458: 1447: 1436: 1425: 1414: 1403: 1392: 1381: 1370: 1359: 1348: 1337: 1326: 1315: 1304: 1293: 1282: 1271: 1260: 1249: 1238: 1227: 1216: 1205: 1194: 1183: 1172: 1161: 1150: 1139: 1128: 1117: 1106: 1095: 1084: 1073: 1062: 1051: 1040: 1029: 1018: 1007: 996: 985: 974: 963: 952: 941: 930: 919: 908: 897: 886: 875: 864: 853: 842: 831: 820: 809: 798: 787: 776: 765: 754: 743: 732: 721: 710: 699: 688: 677: 666: 655: 644: 633: 622: 611: 399:The well studied 384:forms. Many trap 256: 255: 225: 27:Family of spiders 16:(Redirected from 4172: 4135: 4134: 4122: 4121: 4109: 4108: 4099: 4098: 4086: 4085: 4073: 4072: 4060: 4059: 4050: 4049: 4037: 4036: 4034:NBNSYS0000161145 4024: 4023: 4011: 4010: 3998: 3997: 3985: 3984: 3972: 3971: 3959: 3958: 3946: 3945: 3933: 3932: 3920: 3919: 3907: 3906: 3894: 3893: 3881: 3880: 3871: 3870: 3858: 3857: 3845: 3844: 3843: 3830: 3829: 3828: 3798: 3743: 3724: 3685: 3676: 3667: 3658: 3640: 3626: 3617: 3603: 3586: 3582: 3554: 3541: 3533: 3525: 3517: 3509: 3497:(pirate spiders) 3496: 3484:(shield spiders) 3483: 3474: 3465: 3456: 3447: 3434: 3426: 3417: 3404:(velvet spiders) 3403: 3390: 3382: 3359: 3351: 3343: 3332:Cheiracanthiidae 3319: 3310: 3297: 3289: 3280: 3271: 3246: 3238: 3230: 3217: 3209: 3201: 3188:(cellar spiders) 3187: 3173: 3155:(goblin spiders) 3154: 3145: 3144:Ochyroceratidae 3127: 3114: 3106: 3098: 3090: 3082: 3064: 3056: 3049: 3028: 3015: 3007: 2999: 2993:Microstigmatidae 2986: 2968: 2950: 2942: 2941:Cyrtaucheniidae 2934: 2926: 2918: 2910: 2902: 2894: 2886: 2877: 2863: 2847: 2839: 2821: 2772: 2765: 2758: 2749: 2657: 2656: 2608: 2602: 2601: 2599: 2598: 2586: 2577: 2571: 2570: 2568: 2544: 2538: 2537: 2514:Animal Behaviour 2509: 2503: 2502: 2492: 2468: 2462: 2461: 2417: 2411: 2410: 2387:Animal Behaviour 2382: 2376: 2375: 2346: 2340: 2339: 2337: 2313: 2307: 2306: 2304: 2280: 2274: 2273: 2263: 2239: 2233: 2232: 2230: 2224:. Archived from 2223: 2205: 2196: 2190: 2189: 2179: 2147: 2141: 2140: 2138: 2114: 2108: 2107: 2097: 2072:(15347): 15347. 2057: 2051: 2050: 2048: 2047: 2032: 2023: 2022: 2014: 1962: 1951: 1940: 1929: 1919:Keyserling, 1886 1918: 1907: 1896: 1885: 1875:Keyserling, 1884 1874: 1863: 1852: 1842:Walckenaer, 1805 1841: 1830: 1819: 1809:Wunderlich, 2011 1808: 1797: 1786: 1775: 1764: 1753: 1742: 1731: 1720: 1710:Wunderlich, 1992 1709: 1698: 1687: 1676: 1668:– Africa, Europe 1666:Wunderlich, 1995 1665: 1654: 1644:Wunderlich, 2011 1643: 1632: 1621: 1611:Wunderlich, 2008 1610: 1599: 1591:– Angola, Rwanda 1588: 1577: 1566: 1555: 1544: 1533: 1522: 1511: 1500: 1489: 1479:Wunderlich, 2008 1478: 1467: 1456: 1445: 1434: 1423: 1412: 1401: 1390: 1379: 1369:Wunderlich, 2011 1368: 1357: 1346: 1335: 1325:Wunderlich, 2008 1324: 1314:Wunderlich, 1992 1313: 1303:Walckenaer, 1805 1302: 1291: 1280: 1269: 1258: 1247: 1237:Wunderlich, 1995 1236: 1225: 1214: 1204:Keyserling, 1886 1203: 1193:Wunderlich, 2008 1192: 1181: 1170: 1160:Keyserling, 1884 1159: 1148: 1137: 1127:Wunderlich, 2011 1126: 1116:Keyserling, 1890 1115: 1105:Keyserling, 1884 1104: 1094:Wunderlich, 1995 1093: 1083:Wunderlich, 1992 1082: 1071: 1061:Walckenaer, 1809 1060: 1049: 1038: 1027: 1016: 1006:Wunderlich, 1988 1005: 994: 983: 972: 961: 950: 939: 928: 917: 906: 895: 884: 873: 862: 851: 840: 830:Wunderlich, 2008 829: 818: 808:Wunderlich, 2011 807: 796: 785: 774: 763: 753:Keyserling, 1884 752: 741: 730: 719: 708: 697: 686: 676:Wunderlich, 2008 675: 664: 654:Wunderlich, 1995 653: 642: 632:Wunderlich, 2008 631: 620: 609: 562:Theridula angula 501:Dipoena martinae 252: 220: 138: 137: 115: 105: 42: 38:Temporal range: 32: 21: 4180: 4179: 4175: 4174: 4173: 4171: 4170: 4169: 4145: 4144: 4143: 4138: 4130: 4125: 4117: 4112: 4104: 4102: 4094: 4089: 4081: 4076: 4068: 4063: 4055: 4053: 4045: 4040: 4032: 4027: 4019: 4014: 4006: 4001: 3993: 3988: 3980: 3975: 3967: 3962: 3954: 3949: 3941: 3936: 3928: 3923: 3915: 3910: 3902: 3897: 3889: 3884: 3876: 3874: 3866: 3861: 3853: 3848: 3839: 3838: 3833: 3824: 3823: 3818: 3805: 3795: 3790: 3780:Spider taxonomy 3755: 3713:Trochanteriidae 3657:Tetragnathidae 3410:Gallieniellidae 3256: 3237:Tetrablemmidae 3052:Non-entelegynes 3034: 3014:Paratropididae 2962:Halonoproctidae 2901:Antrodiaetidae 2871: 2853: 2829: 2815: 2783: 2776: 2724: 2692:Wayback Machine 2679:Wayback Machine 2666: 2664:Further reading 2661: 2660: 2610: 2609: 2605: 2596: 2594: 2584: 2579: 2578: 2574: 2546: 2545: 2541: 2511: 2510: 2506: 2470: 2469: 2465: 2419: 2418: 2414: 2384: 2383: 2379: 2348: 2347: 2343: 2315: 2314: 2310: 2282: 2281: 2277: 2241: 2240: 2236: 2228: 2203: 2198: 2197: 2193: 2149: 2148: 2144: 2116: 2115: 2111: 2059: 2058: 2054: 2045: 2043: 2034: 2033: 2026: 2016: 2015: 2011: 2006: 1994: 1968: 1754:Sundevall, 1833 1189:Heterotheridion 1030:– South America 1024:Echinotheridion 742:Agnarsson, 2006 731:Sundevall, 1833 525:Epsinus nubilus 469: 463: 401:kleptoparasitic 374: 219: 132: 106: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 40: 39: 36: 28: 23: 22: 15: 12: 11: 5: 4178: 4176: 4168: 4167: 4162: 4157: 4147: 4146: 4140: 4139: 4137: 4136: 4123: 4110: 4100: 4087: 4074: 4061: 4051: 4038: 4025: 4012: 3999: 3986: 3973: 3960: 3951:Fauna Europaea 3947: 3934: 3921: 3908: 3895: 3882: 3872: 3859: 3846: 3831: 3815: 3813: 3807: 3806: 3801: 3792: 3791: 3789: 3788: 3782: 3776: 3773: 3772: 3769: 3768: 3765: 3764: 3761: 3760: 3757: 3756: 3754: 3753: 3748: 3738: 3733: 3728: 3720: 3715: 3710: 3705: 3700: 3695: 3690: 3686:(crab spiders) 3680: 3672: 3662: 3654: 3649: 3644: 3636: 3631: 3621: 3618:(bark hunters) 3613: 3608: 3598: 3593: 3578: 3573: 3568: 3563: 3561:Phrurolithidae 3558: 3553:Philodromidae 3550: 3545: 3542:(lynx spiders) 3537: 3529: 3521: 3513: 3505: 3500: 3492: 3490:Megadictynidae 3487: 3479: 3475:(wolf spiders) 3469: 3461: 3451: 3443: 3441:Homalonychidae 3438: 3430: 3422: 3412: 3407: 3399: 3394: 3386: 3378: 3373: 3368: 3363: 3355: 3347: 3339: 3334: 3329: 3324: 3314: 3306: 3301: 3293: 3285: 3274: 3268: 3262: 3261: 3258: 3257: 3255: 3254: 3242: 3234: 3226: 3221: 3213: 3205: 3197: 3192: 3182: 3177: 3169: 3164: 3159: 3149: 3141: 3136: 3131: 3123: 3118: 3113:Gradungulidae 3110: 3102: 3094: 3086: 3078: 3073: 3071:Austrochilidae 3068: 3059: 3053: 3046: 3040: 3039: 3036: 3035: 3033: 3032: 3027:Theraphosidae 3024: 3022:Porrhothelidae 3019: 3011: 3003: 2995: 2990: 2982: 2977: 2972: 2964: 2959: 2954: 2946: 2938: 2930: 2922: 2914: 2906: 2898: 2893:Actinopodidae 2889: 2883: 2873: 2872: 2866: 2859: 2858: 2855: 2854: 2852: 2851: 2846:Heptathelidae 2843: 2834: 2831: 2830: 2824: 2817: 2816: 2814: 2813: 2807: 2801: 2795: 2788: 2785: 2784: 2777: 2775: 2774: 2767: 2760: 2752: 2746: 2745: 2740: 2735: 2730: 2723: 2722:External links 2720: 2719: 2718: 2715: 2712: 2709: 2705: 2694: 2681: 2665: 2662: 2659: 2658: 2623:(1): 184–217. 2603: 2572: 2559:(3): 293–305. 2539: 2520:(5): 855–864. 2504: 2463: 2428:(4): 570–574. 2412: 2377: 2358:(6): 791–809. 2341: 2328:(3): 293–305. 2308: 2295:(3): 355–364. 2275: 2254:(3): 249–256. 2234: 2231:on 2007-09-27. 2214:(4): 453–593. 2191: 2162:(3): 590–597. 2142: 2129:(2): 211–232. 2109: 2052: 2024: 2008: 2007: 2005: 2002: 2001: 2000: 1993: 1990: 1977:Cretotheridion 1967: 1966: 1955: 1954:– Asia, Europe 1944: 1933: 1922: 1911: 1900: 1889: 1878: 1867: 1856: 1845: 1834: 1823: 1812: 1801: 1790: 1779: 1776:Saaristo, 2006 1768: 1757: 1746: 1735: 1732:Saaristo, 2006 1724: 1723:– Japan, China 1721:Yaginuma, 1957 1713: 1702: 1691: 1688:Saaristo, 2006 1680: 1669: 1658: 1647: 1636: 1625: 1614: 1603: 1592: 1581: 1570: 1559: 1548: 1545:Westwood, 1835 1537: 1526: 1515: 1504: 1493: 1482: 1471: 1470:– China, Japan 1460: 1449: 1438: 1427: 1416: 1405: 1394: 1383: 1380:Saaristo, 2006 1372: 1361: 1350: 1339: 1328: 1321:Magnopholcomma 1317: 1306: 1295: 1284: 1273: 1262: 1251: 1240: 1229: 1218: 1217:– South Africa 1207: 1196: 1185: 1174: 1163: 1152: 1141: 1130: 1123:Grancanaridion 1119: 1108: 1097: 1086: 1075: 1064: 1053: 1042: 1031: 1020: 1009: 998: 987: 976: 965: 954: 943: 932: 931:– Asia, Africa 921: 910: 899: 888: 887:– South Africa 885:Lawrence, 1947 877: 866: 855: 844: 833: 822: 811: 800: 789: 778: 767: 764:Saaristo, 2006 756: 745: 734: 723: 712: 701: 698:Saaristo, 2006 690: 679: 668: 657: 646: 645:– China, Japan 635: 624: 613: 601: 465:Main article: 462: 459: 373: 370: 364:or a grinning 267:cobweb spiders 254: 253: 245: 244: 234: 233: 227: 226: 214: 210: 209: 204: 200: 199: 194: 190: 189: 184: 180: 179: 174: 170: 169: 164: 160: 159: 154: 150: 149: 144: 140: 139: 126: 125: 117: 116: 108: 107: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 43: 37: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 4177: 4166: 4163: 4161: 4158: 4156: 4153: 4152: 4150: 4133: 4128: 4124: 4120: 4115: 4111: 4107: 4101: 4097: 4092: 4088: 4084: 4079: 4075: 4071: 4066: 4062: 4058: 4052: 4048: 4043: 4039: 4035: 4030: 4026: 4022: 4017: 4013: 4009: 4004: 4000: 3996: 3991: 3987: 3983: 3978: 3974: 3970: 3965: 3961: 3957: 3952: 3948: 3944: 3939: 3935: 3931: 3926: 3922: 3918: 3913: 3909: 3905: 3900: 3896: 3892: 3887: 3883: 3879: 3873: 3869: 3864: 3860: 3856: 3851: 3847: 3842: 3836: 3832: 3827: 3821: 3817: 3816: 3814: 3812: 3808: 3804: 3799: 3786: 3783: 3781: 3778: 3777: 3774: 3752: 3749: 3747: 3746: 3744:(ant spiders) 3739: 3737: 3734: 3732: 3729: 3727: 3721: 3719: 3716: 3714: 3711: 3709: 3706: 3704: 3701: 3699: 3696: 3694: 3691: 3689: 3688: 3681: 3679: 3677:(ray spiders) 3673: 3671: 3670: 3663: 3661: 3655: 3653: 3650: 3648: 3645: 3643: 3637: 3635: 3632: 3630: 3629: 3622: 3620: 3614: 3612: 3609: 3607: 3606: 3599: 3597: 3594: 3590: 3585: 3579: 3577: 3574: 3572: 3569: 3567: 3566:Physoglenidae 3564: 3562: 3559: 3557: 3551: 3549: 3546: 3544: 3538: 3536: 3530: 3528: 3522: 3520: 3514: 3512: 3506: 3504: 3501: 3499: 3493: 3491: 3488: 3486: 3480: 3478: 3477: 3470: 3468: 3462: 3460: 3459: 3452: 3450: 3444: 3442: 3439: 3437: 3431: 3429: 3423: 3421: 3420: 3413: 3411: 3408: 3406: 3400: 3398: 3395: 3393: 3387: 3385: 3379: 3377: 3376:Cycloctenidae 3374: 3372: 3369: 3367: 3366:Cyatholipidae 3364: 3362: 3356: 3354: 3348: 3346: 3344:(sac spiders) 3340: 3338: 3337:Cithaeronidae 3335: 3333: 3330: 3328: 3325: 3323: 3322: 3315: 3313: 3309:Anyphaenidae 3307: 3305: 3302: 3300: 3294: 3292: 3288:Amaurobiidae 3286: 3284: 3283: 3276: 3275: 3272: 3269: 3267: 3263: 3253: 3250: 3243: 3241: 3235: 3233: 3227: 3225: 3224:Stenochilidae 3222: 3220: 3214: 3212: 3208:Segestriidae 3206: 3204: 3198: 3196: 3195:Plectreuridae 3193: 3191: 3190: 3183: 3181: 3178: 3176: 3172:Palpimanidae 3170: 3168: 3165: 3163: 3160: 3158: 3157: 3150: 3148: 3142: 3140: 3137: 3135: 3132: 3130: 3126:Hypochilidae 3124: 3122: 3119: 3117: 3111: 3109: 3105:Filistatidae 3103: 3101: 3095: 3093: 3087: 3085: 3079: 3077: 3074: 3072: 3069: 3067: 3061: 3060: 3057: 3054: 3050: 3047: 3045: 3044:Araneomorphae 3041: 3031: 3025: 3023: 3020: 3018: 3012: 3010: 3004: 3002: 2996: 2994: 2991: 2989: 2983: 2981: 2980:Macrothelidae 2978: 2976: 2973: 2971: 2967:Hexathelidae 2965: 2963: 2960: 2958: 2955: 2953: 2947: 2945: 2939: 2937: 2931: 2929: 2925:Barychelidae 2923: 2921: 2915: 2913: 2907: 2905: 2899: 2897: 2891: 2890: 2887: 2884: 2882: 2881:Mygalomorphae 2878: 2874: 2870: 2869:Opisthothelae 2864: 2860: 2850: 2844: 2842: 2838:Liphistiidae 2836: 2835: 2832: 2828: 2822: 2818: 2812: 2808: 2806: 2802: 2800: 2796: 2794: 2790: 2789: 2786: 2781: 2773: 2768: 2766: 2761: 2759: 2754: 2753: 2750: 2744: 2741: 2739: 2736: 2734: 2731: 2729: 2726: 2725: 2721: 2716: 2713: 2710: 2706: 2704: 2700: 2695: 2693: 2689: 2686: 2682: 2680: 2676: 2673: 2668: 2667: 2663: 2654: 2650: 2646: 2642: 2638: 2634: 2630: 2626: 2622: 2618: 2614: 2607: 2604: 2592: 2591: 2583: 2576: 2573: 2567: 2562: 2558: 2554: 2550: 2543: 2540: 2535: 2531: 2527: 2523: 2519: 2515: 2508: 2505: 2500: 2496: 2491: 2486: 2482: 2478: 2474: 2467: 2464: 2459: 2455: 2451: 2447: 2443: 2439: 2435: 2431: 2427: 2423: 2416: 2413: 2408: 2404: 2400: 2396: 2392: 2388: 2381: 2378: 2373: 2369: 2365: 2361: 2357: 2353: 2345: 2342: 2336: 2331: 2327: 2323: 2319: 2312: 2309: 2303: 2298: 2294: 2290: 2286: 2279: 2276: 2271: 2267: 2262: 2257: 2253: 2249: 2245: 2238: 2235: 2227: 2222: 2217: 2213: 2209: 2202: 2195: 2192: 2187: 2183: 2178: 2173: 2169: 2165: 2161: 2157: 2153: 2146: 2143: 2137: 2132: 2128: 2124: 2120: 2113: 2110: 2105: 2101: 2096: 2091: 2087: 2083: 2079: 2075: 2071: 2067: 2063: 2056: 2053: 2041: 2037: 2031: 2029: 2025: 2020: 2013: 2010: 2003: 1999: 1996: 1995: 1991: 1989: 1987: 1986:Burmese amber 1983: 1979: 1978: 1973: 1961: 1960: 1956: 1952:Yoshida, 2007 1950: 1949: 1945: 1939: 1938: 1934: 1930:Yoshida, 2002 1928: 1927: 1923: 1917: 1916: 1912: 1906: 1905: 1901: 1895: 1894: 1890: 1884: 1883: 1879: 1873: 1872: 1868: 1862: 1861: 1857: 1853:Emerton, 1882 1851: 1850: 1846: 1840: 1839: 1835: 1829: 1828: 1824: 1818: 1817: 1813: 1807: 1806: 1802: 1798:Yoshida, 2001 1796: 1795: 1791: 1785: 1784: 1780: 1774: 1773: 1769: 1763: 1762: 1758: 1752: 1751: 1747: 1741: 1740: 1736: 1730: 1729: 1725: 1719: 1718: 1714: 1708: 1707: 1703: 1697: 1696: 1692: 1686: 1685: 1681: 1677:Berland, 1924 1675: 1674: 1670: 1664: 1663: 1659: 1653: 1652: 1648: 1642: 1641: 1637: 1631: 1630: 1626: 1622:L. Koch, 1872 1620: 1619: 1615: 1609: 1608: 1607:Pycnoepisinus 1604: 1598: 1597: 1593: 1587: 1586: 1582: 1576: 1575: 1571: 1565: 1564: 1560: 1554: 1553: 1549: 1543: 1542: 1538: 1534:Thorell, 1869 1532: 1531: 1527: 1521: 1520: 1519:Paratheridula 1516: 1510: 1509: 1505: 1499: 1498: 1494: 1490:Yoshida, 2009 1488: 1487: 1483: 1477: 1476: 1472: 1468:Yoshida, 2009 1466: 1465: 1461: 1457:Yoshida, 2001 1455: 1454: 1450: 1446:Yoshida, 2016 1444: 1443: 1439: 1433: 1432: 1428: 1422: 1421: 1420:Nesopholcomma 1417: 1411: 1410: 1406: 1400: 1399: 1398:Neospintharus 1395: 1389: 1388: 1384: 1378: 1377: 1373: 1367: 1366: 1362: 1356: 1355: 1351: 1347:Thorell, 1892 1345: 1344: 1340: 1334: 1333: 1329: 1323: 1322: 1318: 1312: 1311: 1307: 1301: 1300: 1296: 1290: 1289: 1285: 1283:– Philippines 1279: 1278: 1274: 1268: 1267: 1263: 1259:Yoshida, 2016 1257: 1256: 1252: 1246: 1245: 1241: 1235: 1234: 1230: 1228:– New Zealand 1226:Forster, 1955 1224: 1223: 1219: 1213: 1212: 1208: 1202: 1201: 1197: 1191: 1190: 1186: 1180: 1179: 1178:Hentziectypus 1175: 1171:Thorell, 1890 1169: 1168: 1164: 1158: 1157: 1153: 1149:Thorell, 1881 1147: 1146: 1142: 1136: 1135: 1131: 1125: 1124: 1120: 1114: 1113: 1109: 1103: 1102: 1098: 1092: 1091: 1087: 1081: 1080: 1076: 1070: 1069: 1065: 1059: 1058: 1054: 1048: 1047: 1043: 1037: 1036: 1032: 1026: 1025: 1021: 1015: 1014: 1010: 1004: 1003: 999: 995:Thorell, 1869 993: 992: 988: 984:Yoshida, 2003 982: 981: 977: 973:L. Koch, 1872 971: 970: 966: 960: 959: 955: 949: 948: 944: 938: 937: 933: 927: 926: 922: 916: 915: 911: 905: 904: 900: 894: 893: 889: 883: 882: 878: 874:Yoshida, 2009 872: 871: 867: 861: 860: 856: 850: 849: 845: 839: 838: 834: 828: 827: 823: 819:Yoshida, 2015 817: 816: 812: 806: 805: 801: 795: 794: 790: 784: 783: 779: 773: 772: 768: 762: 761: 757: 751: 750: 746: 740: 739: 735: 729: 728: 724: 720:Thorell, 1869 718: 717: 713: 707: 706: 702: 696: 695: 691: 685: 684: 680: 674: 673: 669: 665:Berland, 1927 663: 662: 658: 652: 651: 647: 641: 640: 639:Allothymoites 636: 630: 629: 625: 623:– Philippines 619: 618: 614: 608: 607: 603: 602: 600: 598: 594: 590: 589: 585:, previously 584: 583: 578: 574: 573: 564: 563: 558: 552: 551: 546: 539: 538: 533: 527: 526: 521: 515: 514: 509: 503: 502: 497: 491: 490: 485: 479: 478: 473: 468: 460: 458: 456: 455: 450: 449: 443: 441: 437: 433: 432: 426: 424: 423: 422:Neospintharus 418: 417: 412: 411: 406: 402: 397: 395: 391: 387: 383: 379: 371: 369: 367: 363: 359: 355: 351: 350: 346: 341: 338: 337: 332: 328: 327:widow spiders 325: 320: 318: 314: 310: 306: 302: 298: 293: 291: 287: 283: 279: 276: 272: 268: 264: 260: 251: 246: 243: 239: 235: 232: 228: 223: 218: 215: 212: 211: 208: 207:Araneomorphae 205: 202: 201: 198: 195: 192: 191: 188: 185: 182: 181: 178: 175: 172: 171: 168: 165: 162: 161: 158: 155: 152: 151: 148: 145: 142: 141: 136: 131: 127: 124: 123: 118: 114: 109: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 33: 30: 19: 3810: 3784: 3740: 3736:Xenoctenidae 3731:Viridasiidae 3708:Trechaleidae 3693:Titanoecidae 3682: 3666:Theridiidae 3665: 3664: 3634:Stiphidiidae 3625:Sparassidae 3623: 3616:Senoculidae 3600: 3571:Phyxelididae 3548:Penestomidae 3524:Nicodamidae 3471: 3464:Liocranidae 3455:Linyphiidae 3453: 3433:Hersiliidae 3416:Gnaphosidae 3414: 3342:Clubionidae 3316: 3296:Ammoxenidae 3277: 3248: 3184: 3180:Periegopidae 3151: 3134:Leptonetidae 2957:Euctenizidae 2701:31:225–245. 2698: 2620: 2616: 2606: 2595:. 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Retrieved 2039: 2018: 2012: 1988:of Myanmar. 1975: 1969: 1965:– St. Helena 1963:Benoit, 1977 1957: 1946: 1935: 1924: 1913: 1902: 1891: 1880: 1869: 1858: 1847: 1836: 1825: 1814: 1803: 1792: 1781: 1778:– Seychelles 1770: 1759: 1748: 1737: 1734:– Seychelles 1726: 1717:Spheropistha 1715: 1704: 1701:– Seychelles 1693: 1690:– Seychelles 1682: 1673:Selkirkiella 1671: 1660: 1655:Archer, 1950 1649: 1638: 1627: 1616: 1605: 1594: 1589:Miller, 1970 1585:Proboscidula 1583: 1572: 1567:Archer, 1950 1561: 1550: 1539: 1528: 1517: 1512:Archer, 1946 1508:Parasteatoda 1506: 1501:Archer, 1950 1495: 1484: 1473: 1462: 1451: 1440: 1435:Archer, 1950 1429: 1418: 1407: 1402:Exline, 1950 1396: 1385: 1382:– Seychelles 1374: 1363: 1352: 1341: 1330: 1319: 1308: 1297: 1286: 1275: 1270:Archer, 1950 1264: 1253: 1248:Strand, 1932 1242: 1231: 1220: 1209: 1198: 1187: 1182:Archer, 1946 1176: 1165: 1154: 1143: 1132: 1129:– Canary Is. 1121: 1110: 1099: 1088: 1085:– Canary Is. 1077: 1066: 1055: 1050:Pavesi, 1880 1046:Enoplognatha 1044: 1039:Bryant, 1945 1033: 1022: 1011: 1000: 989: 980:Deelemanella 978: 967: 962:Archer, 1946 956: 945: 934: 923: 912: 901: 890: 881:Chorizopella 879: 868: 857: 848:Cephalobares 846: 835: 824: 813: 804:Cameronidion 802: 791: 782:Brunepisinus 780: 771:Borneoridion 769: 766:– Seychelles 758: 747: 744:– Madagascar 736: 725: 714: 703: 700:– Seychelles 692: 681: 670: 659: 648: 637: 626: 615: 610:Strand, 1929 604: 586: 582:Parasteatoda 580: 577:monophyletic 570: 568: 560: 548: 535: 523: 511: 499: 487: 475: 452: 446: 444: 435: 429: 427: 420: 414: 408: 398: 375: 354:T. grallator 353: 347: 342: 334: 321: 294: 270: 266: 262: 258: 257: 242:3028 species 216: 203:Infraorder: 120: 29: 4155:Theridiidae 3990:iNaturalist 3868:Theridiidae 3855:Theridiidae 3841:Theridiidae 3835:Wikispecies 3811:Theridiidae 3742:Zodariidae 3723:Uloboridae 3703:Trachelidae 3684:Thomisidae 3652:Synotaxidae 3647:Synaphridae 3611:Selenopidae 3602:Salticidae 3587:(including 3581:Pisauridae 3532:Oecobiidae 3516:Nesticidae 3508:Mysmenidae 3482:Malkaridae 3446:Lamponidae 3381:Deinopidae 3350:Corinnidae 3279:Agelenidae 3266:Entelegynae 3216:Sicariidae 3200:Scytodidae 3162:Orsolobidae 3121:Huttoniidae 3097:Dysderidae 3089:Drymusidae 3081:Diguetidae 3063:Archaeidae 3006:Nemesiidae 2949:Dipluridae 2933:Ctenizidae 2805:Chelicerata 2803:Subphylum: 2393:: 515–521. 1948:Yunohamella 1941:Baert, 1984 1897:Simon, 1895 1831:Simon, 1881 1787:Simon, 1894 1743:Hentz, 1850 1728:Spinembolia 1662:Sardinidion 1640:Ruborridion 1600:Simon, 1894 1574:Platnickina 1541:Phoroncidia 1475:Ohlertidion 1453:Nipponidion 1413:Menge, 1868 1365:Montanidion 1336:Simon, 1894 1327:– Australia 1299:Latrodectus 1292:Simon, 1881 1233:Jamaitidion 1215:Simon, 1895 1173:– Indonesia 1145:Hadrotarsus 1138:Baert, 1984 1134:Guaraniella 1072:Menge, 1868 1035:Emertonella 1017:Simon, 1909 986:– Indonesia 969:Cyllognatha 958:Cryptachaea 951:Menge, 1868 940:Simon, 1894 936:Craspedisia 929:Simon, 1895 896:Simon, 1894 892:Chrosiothes 863:Simon, 1894 815:Campanicola 799:– Venezuela 788:– Indonesia 777:– Indonesia 709:Simon, 1864 694:Argyrodella 687:Simon, 1891 672:Anatolidion 667:– Australia 628:Achaeridion 617:Achaearyopa 606:Achaearanea 588:Achaearanea 436:Latrodectus 431:Achaearanea 405:Argyrodinae 403:members of 362:smiley face 305:ecribellate 275:araneomorph 259:Theridiidae 217:Theridiidae 177:Chelicerata 173:Subphylum: 4149:Categories 3751:Zoropsidae 3596:Psechridae 3540:Oxyopidae 3503:Miturgidae 3495:Mimetidae 3473:Lycosidae 3425:Hahniidae 3397:Dictynidae 3318:Araneidae 3229:Telemidae 3186:Pholcidae 3167:Pacullidae 3153:Oonopidae 3076:Caponiidae 2909:Atracidae 2827:Mesothelae 2799:Arthropoda 2597:2015-11-06 2046:2019-04-25 2004:References 1982:Cenomanian 1926:Yaginumena 1860:Thwaitesia 1820:Levi, 1957 1811:– Malaysia 1805:Tamanidion 1739:Spintharus 1706:Simitidion 1695:Seycellesa 1651:Rugathodes 1596:Propostira 1563:Phylloneta 1523:Levi, 1957 1497:Paidiscura 1442:Nihonhimea 1431:Nesticodes 1387:Neopisinus 1371:– Malaysia 1310:Macaridion 1211:Histagonia 1090:Exalbidion 1028:Levi, 1963 947:Crustulina 826:Canalidion 810:– Malaysia 797:Levi, 1964 683:Anelosimus 440:Nesticidae 382:spider web 336:Anelosimus 301:entelegyne 295:Theridiid 238:124 genera 167:Arthropoda 3698:Toxopidae 3402:Eresidae 3371:Cybaeidae 3358:Ctenidae 2975:Idiopidae 2917:Atypidae 2867:Suborder 2825:Suborder 2811:Arachnida 2791:Kingdom: 2653:207937170 2637:1464-7931 2422:Oecologia 1980:from the 1970:About 35 1959:Zercidium 1871:Thymoites 1849:Theridula 1838:Theridion 1816:Tekellina 1618:Rhomphaea 1552:Phycosoma 1530:Pholcomma 1486:Okumaella 1424:Ono, 2010 1409:Neottiura 1255:Keijiella 1239:– Jamaica 1200:Hetschkia 1079:Eurypoena 1013:Dipoenura 1002:Dipoenata 925:Coscinida 914:Coleosoma 837:Carniella 705:Argyrodes 650:Ameridion 643:Ono, 2007 572:Theridion 454:Coleosoma 448:Theridion 410:Argyrodes 315:) on the 299:are both 231:Diversity 222:Sundevall 187:Arachnida 153:Kingdom: 147:Eukaryota 3899:BugGuide 3875:BioLib: 3820:Wikidata 3718:Udubidae 3576:Pimoidae 3389:Desidae 3327:Arkyidae 3304:Anapidae 2998:Migidae 2797:Phylum: 2793:Animalia 2782:families 2708:Scripta. 2688:Archived 2675:Archived 2645:31713947 2534:53182880 2499:15879074 2458:11217593 2450:28308120 2407:53177663 2372:26850827 2289:Heredity 2248:Heredity 2186:17439475 2104:29127355 1992:See also 1893:Tomoxena 1882:Tidarren 1783:Styposis 1761:Stemmops 1750:Steatoda 1629:Robertus 1464:Nojimaia 1316:– Europe 1288:Lasaeola 1277:Landoppo 1266:Kochiura 1206:– Brazil 1167:Helvidia 1156:Helvibis 1101:Faiditus 1068:Euryopis 1057:Episinus 942:– Brazil 870:Chikunia 859:Cerocida 832:– Russia 716:Ariamnes 634:– Turkey 416:Faiditus 345:Hawaiian 309:bristles 213:Family: 163:Phylum: 157:Animalia 143:Domain: 4106:7087238 3589:Halidae 2809:Class: 2780:Araneae 2778:Extant 2430:Bibcode 2164:Bibcode 2095:5681556 2074:Bibcode 1972:extinct 1827:Theonoe 1794:Takayus 1646:– India 1613:– Kenya 1492:– Japan 1459:– Japan 1426:– Japan 1343:Molione 1332:Meotipa 1112:Gmogala 991:Dipoena 903:Chrysso 793:Cabello 760:Bardala 749:Audifia 738:Asygyna 727:Asagena 390:insects 297:spiders 288:in 124 286:species 278:spiders 197:Araneae 193:Order: 183:Class: 4119:231930 4103:uBio: 4070:917915 4054:NZOR: 4008:101228 3943:1THERF 3826:Q12019 2651:  2643:  2635:  2532:  2497:  2456:  2448:  2405:  2370:  2270:824014 2268:  2184:  2102:  2092:  1932:– Asia 1915:Wirada 1800:– Asia 1684:Sesato 1376:Nanume 1354:Moneta 1349:– Asia 1261:– Asia 1244:Janula 876:– Asia 821:– Asia 661:Anatea 461:Genera 419:, and 358:morphs 317:tarsus 290:genera 224:, 1833 4114:WoRMS 4091:Plazi 4083:57477 4047:34643 4021:82740 4003:IRMNG 3995:47371 3956:10655 2649:S2CID 2585:(PDF) 2530:S2CID 2454:S2CID 2403:S2CID 2368:S2CID 2266:S2CID 2229:(PDF) 2204:(PDF) 1984:aged 1937:Yoroa 1904:Wamba 1772:Stoda 1222:Icona 366:clown 313:setae 4042:NCBI 4016:ITIS 3982:9583 3977:GBIF 3938:EPPO 3904:1960 3891:1249 3886:BOLD 3785:Bold 2641:PMID 2633:ISSN 2495:PMID 2446:PMID 2182:PMID 2100:PMID 394:silk 386:ants 378:webs 372:Webs 343:The 269:and 46:Preκž’ 4127:WSC 4029:NBN 3930:169 3925:EoL 3917:H6N 3912:CoL 3878:968 3863:AFD 3850:ADW 2703:PDF 2685:PDF 2672:PDF 2625:doi 2561:doi 2522:doi 2485:doi 2481:208 2438:doi 2426:111 2395:doi 2360:doi 2330:doi 2297:doi 2256:doi 2216:doi 2212:146 2172:doi 2131:doi 2090:PMC 2082:doi 4151:: 4129:: 4116:: 4093:: 4080:: 4067:: 4044:: 4031:: 4018:: 4005:: 3992:: 3979:: 3966:: 3953:: 3940:: 3927:: 3914:: 3901:: 3888:: 3865:: 3852:: 3837:: 3822:: 2647:. 2639:. 2631:. 2621:95 2619:. 2615:. 2587:. 2557:78 2555:. 2551:. 2528:. 2518:73 2516:. 2493:. 2479:. 2475:. 2452:. 2444:. 2436:. 2424:. 2401:. 2391:27 2389:. 2366:. 2356:15 2354:. 2326:78 2324:. 2320:. 2293:62 2291:. 2287:. 2264:. 2252:76 2250:. 2246:. 2210:. 2206:. 2180:. 2170:. 2160:76 2158:. 2154:. 2127:87 2125:. 2121:. 2098:. 2088:. 2080:. 2068:. 2064:. 2038:. 2027:^ 579:. 442:. 413:, 333:. 265:, 240:, 96:Pg 3591:) 3251:) 3247:( 2771:e 2764:t 2757:v 2655:. 2627:: 2600:. 2569:. 2563:: 2536:. 2524:: 2501:. 2487:: 2460:. 2440:: 2432:: 2409:. 2397:: 2374:. 2362:: 2338:. 2332:: 2305:. 2299:: 2272:. 2258:: 2218:: 2188:. 2174:: 2166:: 2139:. 2133:: 2106:. 2084:: 2076:: 2070:7 2049:. 2021:. 407:( 311:( 101:N 91:K 86:J 81:T 76:P 71:C 66:D 61:S 56:O 51:κž’ 20:)

Index

Comb-footed spider
Preκž’
κž’
O
S
D
C
P
T
J
K
Pg
N

Enoplognatha ovata
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Chelicerata
Arachnida
Araneae
Araneomorphae
Theridiidae
Sundevall
Diversity
124 genera
3028 species

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