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Comstock–Needham system

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Apart from the costal and the anal veins, each vein can be branched, in which case the branches are numbered from anterior to posterior. For example, the two branches of the subcostal vein will be called
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In the case where two cells are separated by a crossvein but have the same anterior longitudinal vein, they should have the same name. To avoid this, they are attributed a number. For example, the R
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Crossveins link the longitudinal veins, and are named accordingly (for example, the medio-cubital crossvein is termed
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Fraser, F. C. 1938. A note on the fallaciousness of the theory of pretracheation in the venation of Odonata.
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Comstock, J.H. & Needham, J.G. (1898) The wings of Insects. IX The Venation of the Wings of Hymenoptera.
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The Comstock and Needham system attributes different names to the veins on an insect's wing. From the
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According to the Comstock–Needham system, the cubitus forks once, producing the cubital veins Cu
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The cells are named after the vein on the anterior side; for instance, the cell between Sc
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The media may also fork twice, therefore having four branches reaching the wing margin.
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cell is divided in two by the radial cross vein: the basal cell is termed "first R
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The radius typically branches once near the base, producing anteriorly the
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Insect wing venation, showing the names after the Comstock–Needham system
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If a cell is bordered anteriorly by a forking vein, such as R
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Borror and DeLong's introduction to the study of insects
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of all insect wings. This system was based on Needham's
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North–Carolina state University course on insect wings
334:Triplehorn, Charles A.; Johnson Norman F. (2005). 235:). Some crossveins have their own name, like the 215:As there are several anal veins, they are called 35:in 1898. It was an important step in showing the 114:, in analogy with a bone in the forearm, the 8: 200:. According to some other authorities, Cu 223:, and so on. They are usually unforked. 150:, in reference to its posterior location 308: 72:(leading) edge of the wing towards the 7: 186:. The radial sector may fork twice. 16:Naming system for insect wing veins 14: 281:", and the distal cell "second R 204:may fork again, producing the Cu 43:that was later discredited by 23:is a naming system for insect 1: 317:Proc. Roy. Ent. Soc. London 243:and the sectoral crossvein 400: 384:Biological classification 328:The American Naturalist 45:Frederic Charles Fraser 21:Comstock–Needham system 65: 63: 41:pretracheation theory 179:and posteriorly the 340:Thomson Brooks/Cole 76:(rear), the major 66: 56:Longitudinal veins 80:veins are named: 391: 353: 338:(7th ed.). 320: 313: 251:Cell terminology 51:Vein terminology 399: 398: 394: 393: 392: 390: 389: 388: 369: 368: 360: 350: 333: 323: 314: 310: 306: 299: 295: 291: 284: 280: 276: 268: 262: 258: 253: 229: 211: 207: 203: 199: 195: 177: 167: 160: 58: 53: 17: 12: 11: 5: 397: 395: 387: 386: 381: 379:Insect anatomy 371: 370: 367: 366: 359: 358:External links 356: 355: 354: 348: 331: 322: 321: 307: 305: 302: 297: 293: 289: 282: 278: 274: 266: 260: 256: 252: 249: 228: 225: 209: 205: 201: 197: 193: 175: 165: 158: 152: 151: 142: 130: 118: 106: 94: 57: 54: 52: 49: 33:George Needham 15: 13: 10: 9: 6: 4: 3: 2: 396: 385: 382: 380: 377: 376: 374: 365: 362: 361: 357: 351: 349:0-03-096835-6 345: 341: 337: 332: 330:, 32:413-424. 329: 325: 324: 319:(A) 13: 60–70 318: 312: 309: 303: 301: 286: 271: 269: 250: 248: 246: 242: 238: 234: 226: 224: 222: 218: 213: 190: 187: 185: 182: 181:radial sector 178: 170: 168: 161: 149: 146: 143: 141: 137: 134: 131: 129: 125: 122: 119: 117: 113: 110: 107: 105: 104:below the rib 101: 98: 95: 93: 89: 86: 83: 82: 81: 79: 75: 71: 62: 55: 50: 48: 46: 42: 38: 34: 30: 29:John Comstock 27:, devised by 26: 22: 335: 327: 316: 311: 287: 272: 264: 254: 244: 240: 232: 230: 220: 216: 214: 191: 188: 183: 180: 173: 171: 163: 156: 153: 147: 144: 139: 135: 132: 127: 123: 120: 111: 108: 103: 99: 96: 91: 87: 84: 78:longitudinal 67: 40: 20: 18: 373:Categories 304:References 263:is called 239:crossvein 227:Crossveins 145:anal veins 138:, meaning 126:, meaning 102:, meaning 90:, meaning 25:wing veins 74:posterior 47:in 1938. 97:subcosta 70:anterior 37:homology 237:humeral 133:cubitus 346:  208:and Cu 196:and Cu 128:middle 116:radius 109:radius 292:and R 259:and R 140:elbow 121:media 85:costa 344:ISBN 233:m-cu 162:and 31:and 19:The 285:". 92:rib 375:: 342:. 300:. 270:. 265:Sc 247:. 221:A2 219:, 217:A1 212:. 210:1b 206:1a 184:Rs 169:. 164:Sc 157:Sc 136:Cu 100:Sc 352:. 298:3 294:3 290:2 283:1 279:1 275:1 267:2 261:1 257:2 245:s 241:h 202:1 198:2 194:1 176:1 174:R 166:2 159:1 148:A 124:M 112:R 88:C

Index

wing veins
John Comstock
George Needham
homology
Frederic Charles Fraser

anterior
posterior
longitudinal
radius
humeral
Thomson Brooks/Cole
ISBN
0-03-096835-6
North–Carolina state University course on insect wings
Categories
Insect anatomy
Biological classification

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