342:
536:
671:
294:
49:
431:
386:
74:
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585:
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716:
205:, occurring in Africa and Madagascar. These two scientific families of lizards, known as Cordyliformes or Cordyloidea, are sometimes combined into a larger concept of Cordylidae. Recent molecular analyses confirm the clade made up of Cordylidae and Gerrhosauridae (Cordyloidea) and place it in a larger clade including
225:. They are terrestrial, mostly inhabiting crevices in rocky terrain, although at least one species digs burrows and another lives under exfoliating bark on trees. They have flattened heads and bodies, and are distinguished by a heavy armour of
229:
and large, rectangular, scales, arranged in regular rows around the body and tail. Many species have rings of spines on the tail, that aid in wedging the animal into sheltering crevices, and also in dissuading predators.
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780:
2431:
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770:"Cordylidae." Bill Branch. 1998. Field Guide to Snakes and other reptiles of Southern Africa. Struik Publishers, Cape Town.
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1984:
558:
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73:
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1975:
1903:
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548:
541:
528:
2426:
2130:
2122:
2020:
1870:
1861:
1696:
1660:
1474:
1388:
404:
2089:
1804:
1543:
670:
56:
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1822:
1762:
1729:
1552:
1516:
1456:
597:
590:
503:
443:
436:
293:
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are almost entirely limbless, with only tiny spikes in place of the hind limbs. The family includes both
2383:
2029:
2002:
1993:
1966:
1786:
1720:
1687:
1597:
1579:
398:
391:
2271:
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2011:
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1213:
933:
728:
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554:
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1744:
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1358:
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1223:
1198:
1125:
1050:
423:
378:
162:
2284:
182:
is a family of small- to medium-sized lizards that occur in southern and eastern
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1135:
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1000:
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31:
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85:
17:
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1073:
1068:
996:
889:
877:
858:
334:
125:
105:
1150:
1040:
883:
115:
781:
The molecular evolutionary tree of lizards, snakes, and amphisbaenians
2323:
1418:
1308:
871:
183:
95:
2310:
2200:
827:
1353:
981:
915:
2204:
1377:
831:
2297:
795:
Bauer, Aaron M. (1998). Cogger, H.G.; Zweifel, R.G. (eds.).
233:
Most species have four limbs, but those in the genus
2213:
2111:
2042:
1883:
1859:
1835:
1775:
1742:
1709:
1505:
1445:
1279:
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1232:
1149:
1096:
1049:
1021:
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962:
914:
898:
737:Stanley, Bauer, Jackman, Branch & Mouton, 2011
610:Stanley, Bauer, Jackman, Branch & Mouton, 2011
565:Stanley, Bauer, Jackman, Branch & Mouton, 2011
516:Stanley, Bauer, Jackman, Branch & Mouton, 2011
799:. San Diego: Academic Press. pp. 160–161.
1389:
843:
8:
761:"Cordylidae." Encyclopædia Britannica. 2007.
201:Cordylidae is closely related to the family
2201:
1396:
1382:
1374:
1238:
1005:
850:
836:
828:
254:
47:
38:
797:Encyclopedia of Reptiles and Amphibians
754:
790:
788:
7:
1679:East African spiny-tailed lizard (
25:
714:
669:
628:
583:
534:
489:
429:
384:
340:
292:
221:Girdled lizards are diurnal and
72:
2432:Taxa named by Leopold Fitzinger
779:Nicolas Vidal; S. Blair Hedges
1607:Western dwarf girdled lizard (
1:
1985:Orange-throated flat lizard (
1847:Blue-spotted girdled lizard (
1763:African spiny-tailed lizard (
1571:Coastal spiny-tailed lizard (
186:. They are commonly known as
2139:Zoutpansberg girdled lizard
1895:Attenborough's flat lizard (
1652:N’Dolondolo girdled lizard (
1589:McLachlan's girdled lizard (
742:
739:
736:
726:
713:
701:
698:
695:
681:
668:
656:
653:
650:
640:
627:
615:
612:
609:
595:
582:
570:
568:Blue-spotted girdled lizard
567:
564:
546:
533:
521:
518:
515:
501:
488:
476:
473:
451:
441:
428:
416:
413:
410:
396:
383:
371:
368:
362:
352:
339:
327:
324:
318:
304:
291:
1796:Lawrence's girdled lizard (
1787:Campbell's girdled lizard (
1643:Oelofsen's girdled lizard (
213:Vidal & Hedges, 2009).
170:
2448:
2155:Mozambique girdled lizard
2021:Soutpansberg flat lizard (
1871:Armadillo girdled lizard (
1697:Transvaal girdled lizard (
1670:Ethiopian girdled lizard (
1661:Rhodesian girdled lizard (
1616:Kaokoveld girdled lizard (
1562:Machado's girdled lizard (
1475:Large-scale grass lizard (
29:
2179:Van Dam's girdled lizard
2131:Waterberg girdled lizard
2123:Barberton girdled lizard
2090:Drakensberg crag lizard (
1754:Jordan's girdled lizard (
1544:Rooiberg girdled lizard (
1413:
1008:
866:
613:Armadillo girdled lizard
168:
161:
69:Scientific classification
67:
55:
46:
41:
2187:Warren's girdled lizard
1976:Mitchell's flat lizard (
1904:Broadley's flat lizard (
1814:Peers’s girdled lizard (
1805:Namaqua girdled lizard (
1580:Marungu girdled lizard (
1553:Limpopo girdled lizard (
1517:Angolan girdled lizard (
1457:Transvaal grass lizard (
519:Namaqua girdled lizards
217:Description and behavior
30:Not to be confused with
2003:Sekukhune flat lizard (
1994:Ocellated flat lizard (
1967:Waterberg flat lizard (
1823:Herero girdled lizard (
1688:Ukinga girdled lizard (
1598:Mecula girdled lizard (
1526:Maasai girdled lizard (
62:Cordylus tropidosternum
57:Tropical girdled lizard
2099:Northern crag lizard (
2063:Highveld crag lizard (
1634:Nyika girdled lizard (
1625:Black girdled lizard (
1484:Zambian grass lizard (
2384:Paleobiology Database
2163:Regal girdled lizard
2147:Giant girdled lizard
2030:Striped flat lizard (
1958:Spotted flat lizard (
1949:Lebombo flat lizard (
1931:Emperor flat lizard (
1535:Cape girdled lizard (
2171:Swazi dragon lizard
2054:Lang's crag lizard (
2012:Pungwe flat lizard (
1940:Common flat lizard (
1849:N. coeruleopunctatus
1730:Dwarf cliff lizard (
549:N. coeruleopunctatus
542:N. coeruleopunctatus
2125:(S. barbertonensis)
2081:Spiny crag lizard (
1922:Dwarf flat lizard (
1721:Cape cliff lizard (
1493:Cape snake lizard (
1466:Cape grass lizard (
196:girdle-tail lizards
2072:Cape crag lizard (
1913:Cape flat lizard (
260:
2409:
2408:
2371:Open Tree of Life
2207:Taxon identifiers
2198:
2197:
2101:P. transvaalensis
2074:P. microlepidotus
1681:C. tropidosternum
1371:
1370:
1367:
1366:
1092:
1091:
1023:Gymnophthalmoidea
954:Sphaerodactylidae
747:
746:
684:P. microlepidotus
258:
192:spinytail lizards
177:
176:
157:
27:Family of lizards
16:(Redirected from
2439:
2402:
2401:
2392:
2391:
2379:
2378:
2366:
2365:
2353:
2352:
2340:
2339:
2327:
2326:
2314:
2313:
2301:
2300:
2288:
2287:
2275:
2274:
2262:
2261:
2249:
2248:
2247:
2234:
2233:
2232:
2202:
2157:(S. mossambicus)
1897:P. attenboroughi
1398:
1391:
1384:
1375:
1261:Leptotyphlopidae
1239:
1036:Gymnophthalmidae
1006:
944:Phyllodactylidae
929:Carphodactylidae
852:
845:
838:
829:
811:
810:
792:
783:
777:
771:
768:
762:
759:
740:Girdled lizards
734:
718:
693:
673:
648:
632:
607:
587:
562:
538:
513:
493:
449:
433:
408:
388:
369:Girdled lizards
360:
359:(Linnaeus, 1758)
344:
316:
296:
255:
152:
77:
76:
51:
39:
21:
2447:
2446:
2442:
2441:
2440:
2438:
2437:
2436:
2427:Lizard families
2412:
2411:
2410:
2405:
2397:
2395:
2387:
2382:
2374:
2369:
2361:
2356:
2348:
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2322:
2317:
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2304:
2296:
2291:
2283:
2278:
2270:
2265:
2257:
2252:
2243:
2242:
2237:
2228:
2227:
2222:
2209:
2199:
2194:
2107:
2038:
1951:P. lebomboensis
1879:
1873:O. cataphractus
1855:
1831:
1771:
1738:
1705:
1582:C. marunguensis
1501:
1441:
1409:
1402:
1372:
1363:
1324:Cylindrophiidae
1304:Atractaspididae
1275:
1271:Xenotyphlopidae
1228:
1214:Phrynosomatidae
1145:
1088:
1045:
1017:
991:
958:
934:Diplodactylidae
910:
894:
862:
856:
819:
814:
807:
794:
793:
786:
778:
774:
769:
765:
760:
756:
752:
732:
719:
687:
674:
646:
633:
601:
598:O. cataphractus
591:O. cataphractus
588:
552:
539:
507:
494:
474:Karusa lizards
447:
434:
402:
389:
358:
345:
310:
297:
253:
219:
188:girdled lizards
169:10 genera (see
151:
71:
60:
35:
28:
23:
22:
15:
12:
11:
5:
2445:
2443:
2435:
2434:
2429:
2424:
2414:
2413:
2407:
2406:
2404:
2403:
2393:
2380:
2367:
2354:
2341:
2328:
2315:
2302:
2289:
2276:
2263:
2250:
2235:
2219:
2217:
2211:
2210:
2205:
2196:
2195:
2193:
2192:
2184:
2176:
2168:
2160:
2152:
2149:(S. giganteus)
2144:
2141:(S. depressus)
2136:
2128:
2119:
2117:
2109:
2108:
2106:
2105:
2096:
2087:
2078:
2069:
2060:
2050:
2048:
2045:Pseudocordylus
2040:
2039:
2037:
2036:
2027:
2018:
2014:P. pungweensis
2009:
2000:
1991:
1982:
1973:
1964:
1955:
1946:
1942:P. intermedius
1937:
1928:
1919:
1910:
1901:
1891:
1889:
1881:
1880:
1878:
1877:
1867:
1865:
1857:
1856:
1854:
1853:
1843:
1841:
1833:
1832:
1830:
1829:
1820:
1811:
1807:N. namaquensis
1802:
1793:
1783:
1781:
1773:
1772:
1770:
1769:
1760:
1750:
1748:
1740:
1739:
1737:
1736:
1727:
1717:
1715:
1707:
1706:
1704:
1703:
1694:
1685:
1676:
1667:
1663:C. rhodesianus
1658:
1654:C. phonolithos
1649:
1640:
1631:
1622:
1613:
1604:
1595:
1586:
1577:
1573:C. macropholis
1568:
1559:
1550:
1541:
1532:
1523:
1513:
1511:
1503:
1502:
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1499:
1490:
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1403:
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1400:
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1386:
1378:
1369:
1368:
1365:
1364:
1362:
1361:
1356:
1351:
1346:
1344:Tropidophiidae
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1336:
1331:
1326:
1321:
1316:
1311:
1306:
1301:
1296:
1291:
1285:
1283:
1281:Alethinophidia
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1276:
1274:
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1268:
1263:
1258:
1253:
1251:Anomalepididae
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1245:
1236:
1230:
1229:
1227:
1226:
1221:
1216:
1211:
1206:
1201:
1196:
1194:Leiocephalidae
1191:
1186:
1181:
1176:
1171:
1169:Corytophanidae
1166:
1164:Chamaeleonidae
1161:
1155:
1153:
1147:
1146:
1144:
1143:
1141:Diploglossidae
1138:
1133:
1128:
1123:
1121:Helodermatidae
1118:
1113:
1108:
1102:
1100:
1094:
1093:
1090:
1089:
1087:
1086:
1081:
1076:
1071:
1066:
1061:
1059:Amphisbaenidae
1055:
1053:
1047:
1046:
1044:
1043:
1038:
1033:
1031:Alopoglossidae
1027:
1025:
1019:
1018:
1016:
1015:
1009:
1003:
993:
992:
990:
989:
984:
979:
977:Gerrhosauridae
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817:External links
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664:Pseudocordylus
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414:Cliff lizards
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382:
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325:Grass lizards
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275:
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262:
261:
259:List of genera
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249:
218:
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203:Gerrhosauridae
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2240:
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2216:
2212:
2208:
2203:
2191:
2190:
2185:
2183:
2182:
2177:
2175:
2174:
2173:(S. swazicus)
2169:
2167:
2166:
2161:
2159:
2158:
2153:
2151:
2150:
2145:
2143:
2142:
2137:
2135:
2134:
2129:
2127:
2126:
2121:
2120:
2118:
2116:
2115:
2110:
2104:
2102:
2097:
2095:
2093:
2092:P. subviridis
2088:
2086:
2084:
2079:
2077:
2075:
2070:
2068:
2066:
2061:
2059:
2057:
2052:
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2049:
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2046:
2041:
2035:
2033:
2028:
2026:
2024:
2019:
2017:
2015:
2010:
2008:
2006:
2005:P. orientalis
2001:
1999:
1997:
1992:
1990:
1988:
1987:P. monotropis
1983:
1981:
1979:
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1863:
1858:
1852:
1850:
1845:
1844:
1842:
1840:
1839:
1834:
1828:
1826:
1825:N. pustulatus
1821:
1819:
1817:
1812:
1810:
1808:
1803:
1801:
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1794:
1792:
1790:
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1693:
1691:
1690:C. ukingensis
1686:
1684:
1682:
1677:
1675:
1673:
1668:
1666:
1664:
1659:
1657:
1655:
1650:
1648:
1646:
1641:
1639:
1637:
1632:
1630:
1628:
1623:
1621:
1619:
1618:C. namakuiyus
1614:
1612:
1610:
1605:
1603:
1601:
1596:
1594:
1592:
1591:C. mclachlani
1587:
1585:
1583:
1578:
1576:
1574:
1569:
1567:
1565:
1560:
1558:
1556:
1551:
1549:
1547:
1542:
1540:
1538:
1533:
1531:
1529:
1528:C. beraduccii
1524:
1522:
1520:
1519:C. angolensis
1515:
1514:
1512:
1510:
1509:
1504:
1498:
1496:
1491:
1489:
1487:
1482:
1480:
1478:
1477:C. macrolepis
1473:
1471:
1469:
1464:
1462:
1460:
1455:
1454:
1452:
1450:
1449:
1444:
1438:
1434:
1432:
1428:
1426:
1422:
1420:
1416:
1415:
1412:
1407:
1399:
1394:
1392:
1387:
1385:
1380:
1379:
1376:
1360:
1357:
1355:
1352:
1350:
1347:
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1340:
1337:
1335:
1332:
1330:
1327:
1325:
1322:
1320:
1317:
1315:
1312:
1310:
1307:
1305:
1302:
1300:
1299:Anomochilidae
1297:
1295:
1292:
1290:
1289:Acrochordidae
1287:
1286:
1284:
1282:
1278:
1272:
1269:
1267:
1264:
1262:
1259:
1257:
1256:Gerrhopilidae
1254:
1252:
1249:
1248:
1246:
1244:
1243:Scolecophidia
1240:
1237:
1235:
1231:
1225:
1222:
1220:
1219:Polychrotidae
1217:
1215:
1212:
1210:
1207:
1205:
1202:
1200:
1197:
1195:
1192:
1190:
1187:
1185:
1184:Hoplocercidae
1182:
1180:
1177:
1175:
1174:Crotaphytidae
1172:
1170:
1167:
1165:
1162:
1160:
1157:
1156:
1154:
1152:
1148:
1142:
1139:
1137:
1134:
1132:
1129:
1127:
1124:
1122:
1119:
1117:
1114:
1112:
1111:Lanthanotidae
1109:
1107:
1106:Shinisauridae
1104:
1103:
1101:
1099:
1095:
1085:
1084:Trogonophidae
1082:
1080:
1077:
1075:
1072:
1070:
1067:
1065:
1062:
1060:
1057:
1056:
1054:
1052:
1048:
1042:
1039:
1037:
1034:
1032:
1029:
1028:
1026:
1024:
1020:
1014:
1011:
1010:
1007:
1004:
1002:
998:
994:
988:
985:
983:
980:
978:
975:
973:
970:
969:
967:
965:
961:
955:
952:
950:
947:
945:
942:
940:
939:Eublepharidae
937:
935:
932:
930:
927:
925:
922:
921:
919:
917:
913:
907:
904:
903:
901:
897:
891:
887:
885:
881:
879:
875:
873:
869:
868:
865:
860:
853:
848:
846:
841:
839:
834:
833:
830:
824:
821:
820:
816:
808:
806:0-12-178560-2
802:
798:
791:
789:
785:
782:
776:
773:
767:
764:
758:
755:
749:
733:(Smith, 1844)
731:
730:
724:
723:
717:
711:
710:
706:
705:
699:Crag lizards
691:
686:
685:
679:
678:
677:P. subviridis
672:
666:
665:
661:
660:
654:Flat lizards
645:
644:
638:
637:
631:
625:
624:
620:
619:
605:
600:
599:
593:
592:
586:
580:
579:
575:
574:
560:
556:
551:
550:
544:
543:
537:
531:
530:
526:
525:
511:
506:
505:
504:N. pustulatus
499:
498:
492:
486:
485:
481:
480:
470:
466:
462:
458:
454:
448:(Smith, 1838)
446:
445:
439:
438:
432:
426:
425:
421:
420:
406:
401:
400:
394:
393:
387:
381:
380:
376:
375:
365:
357:
356:
350:
349:
343:
337:
336:
332:
331:
321:
314:
309:
308:
302:
301:
295:
289:
288:
284:
283:
279:
276:
274:Taxon author
273:
271:Type species
270:
267:
264:
263:
257:
256:
250:
248:
246:
245:ovoviviparous
242:
238:
237:
231:
228:
224:
223:insectivorous
216:
214:
212:
211:Cordylomorpha
208:
204:
199:
197:
193:
189:
185:
181:
172:
167:
164:
160:
155:
150:
147:
144:
143:
140:
139:Cordyliformes
137:
134:
131:
130:
127:
124:
121:
120:
117:
114:
111:
110:
107:
104:
101:
100:
97:
94:
91:
90:
87:
84:
81:
80:
75:
70:
66:
63:
58:
54:
50:
45:
40:
37:
33:
19:
2214:
2189:(S. warreni)
2188:
2181:(S. vandami)
2180:
2172:
2164:
2156:
2148:
2140:
2133:(S. breyeri)
2132:
2124:
2112:
2100:
2091:
2082:
2073:
2065:P. melanotus
2064:
2055:
2043:
2032:P. torquatus
2031:
2022:
2013:
2004:
1996:P. ocellatus
1995:
1986:
1978:P. mitchelli
1977:
1968:
1960:P. maculatus
1959:
1950:
1941:
1933:P. imperator
1932:
1923:
1914:
1906:P. broadleyi
1905:
1896:
1884:
1872:
1860:
1848:
1836:
1824:
1815:
1806:
1797:
1789:N. campbelli
1788:
1776:
1765:K. polyzonus
1764:
1755:
1745:Karusasaurus
1743:
1732:H. nebulosus
1731:
1722:
1712:Hemicordylus
1710:
1698:
1689:
1680:
1671:
1662:
1653:
1645:C. oelofseni
1644:
1635:
1626:
1617:
1608:
1599:
1590:
1581:
1572:
1563:
1554:
1545:
1536:
1527:
1518:
1506:
1494:
1486:C. miopropus
1485:
1476:
1467:
1458:
1446:
1405:
1359:Xenopeltidae
1224:Tropiduridae
1199:Leiosauridae
1126:Xenosauridae
1051:Amphisbaenia
971:
796:
775:
766:
757:
729:S. giganteus
727:
722:S. giganteus
720:
707:
696:Smith, 1838
682:
675:
662:
651:Smith, 1844
641:
634:
621:
596:
589:
576:
547:
540:
527:
502:
495:
482:
444:K. polyzonus
442:
437:K. polyzonus
435:
424:Karusasaurus
422:
411:Smith, 1838
397:
390:
379:Hemicordylus
377:
353:
346:
333:
305:
298:
285:
277:Common name
234:
232:
220:
200:
195:
191:
187:
179:
178:
148:
132:
61:
36:
2319:iNaturalist
2239:Wikispecies
2165:(S. regius)
2083:P. spinosus
2023:P. relictus
1924:P. guttatus
1915:P. capensis
1886:Platysaurus
1798:N. lawrenci
1778:Namazonurus
1723:H. capensis
1699:C. vittifer
1564:C. machadoi
1537:C. cordylus
1448:Chamaesaura
1349:Uropeltidae
1334:Loxocemidae
1314:Bolyeriidae
1266:Typhlopidae
1204:Liolaemidae
1179:Dactyloidae
1136:Anniellidae
1098:Anguimorpha
1079:Rhineuridae
1001:Lacertoidea
987:Xantusiidae
949:Pygopodidae
647:Smith, 1844
643:P. capensis
636:P. relictus
623:Platysaurus
484:Namazonurus
399:H. capensis
392:H. capensis
355:C. cordylus
287:Chamaesaura
236:Chamaesaura
207:Xantusiidae
42:Cordylidae
32:Corydalidae
18:Cordyloidea
2422:Cordylidae
2416:Categories
2259:Cordylidae
2245:Cordylidae
2215:Cordylidae
1756:K. jordani
1600:C. meculae
1555:C. jonesii
1495:C. tenuior
1468:C. anguina
1406:Cordylidae
1339:Pythonidae
1319:Colubridae
1013:Lacertidae
972:Cordylidae
964:Scincoidea
924:Gekkonidae
823:Cordylidae
750:References
307:C. anguina
300:C. anguina
241:egg-laying
227:osteoderms
180:Cordylidae
149:Cordylidae
1862:Ouroboros
1816:N. peersi
1636:C. nyikae
1546:C. imkeae
1417:Kingdom:
1354:Viperidae
1294:Aniliidae
1234:Serpentes
1209:Opluridae
1189:Iguanidae
1116:Varanidae
1064:Bipedidae
982:Scincidae
906:Dibamidae
870:Kingdom:
578:Ouroborus
510:W. Peters
497:N. peersi
320:Schneider
247:species.
163:Diversity
154:Fitzinger
92:Kingdom:
86:Eukaryota
2224:Wikidata
2056:P. langi
1969:P. minor
1672:C. rivae
1627:C. niger
1609:C. minor
1508:Cordylus
1459:C. aenea
1437:Squamata
1431:Reptilia
1425:Chordata
1423:Phylum:
1419:Animalia
1329:Elapidae
1159:Agamidae
1131:Anguidae
1074:Cadeidae
1069:Blanidae
997:Laterata
890:Squamata
884:Reptilia
878:Chordata
876:Phylum:
872:Animalia
861:families
859:Squamata
364:Laurenti
348:C. niger
335:Cordylus
313:Linnaeus
280:Species
145:Family:
126:Squamata
116:Reptilia
106:Chordata
102:Phylum:
96:Animalia
82:Domain:
2230:Q840980
1838:Ninurta
1435:Order:
1429:Class:
1408:species
1404:Extant
1151:Iguania
1041:Teiidae
916:Gekkota
899:Dibamia
888:Order:
882:Class:
692:, 1829)
606:, 1828)
604:F. Boie
561:, 1913)
555:Methuen
529:Ninurta
512:, 1862)
471:, 2011
461:Jackman
453:Stanley
407:, 1838)
366:, 1768
322:, 1801
315:, 1758)
122:Order:
112:Class:
2399:215815
2396:uBio:
2376:269281
2350:209057
2337:114227
1309:Boidae
803:
690:Cuvier
559:Hewitt
557:&
469:Mouton
467:&
465:Branch
268:Image
265:Genus
251:Genera
184:Africa
156:, 1826
2389:37965
2363:52173
2332:IRMNG
2324:33023
2114:Smaug
709:Smaug
457:Bauer
405:Smith
194:, or
133:Clade
2358:NCBI
2345:ITIS
2311:6173
2306:GBIF
2298:1721
2272:1100
2267:BOLD
801:ISBN
243:and
171:text
2293:EoL
2285:8K5
2280:CoL
2254:ADW
657:16
372:21
2418::
2386::
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2334::
2321::
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2295::
2282::
2269::
2256::
2241::
2226::
999:/
787:^
743:9
702:6
616:1
571:1
522:5
477:2
463:,
459:,
455:,
417:2
328:5
198:.
190:,
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135::
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1397:e
1390:t
1383:v
851:e
844:t
837:v
809:.
688:(
602:(
553:(
508:(
403:(
311:(
209:(
34:.
20:)
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