842:
explanation is known. Dimensions may vary from 7.5 cm × 3.4 cm to 10 cm × 4.5 cm (3.0 in × 1.3 in to 3.9 in × 1.8 in), while weight may vary from 43.3 to 60.8 g (1.53 to 2.14 oz). Most clutches laid in captivity have been infertile, and only four successful breedings have been documented thus far. Hatchlings are about 18 in (46 cm) long and weigh around 56 g (2.0 oz). Like those of many other monitors, hatchling crocodile monitors are more colourful than adults, and feed primarily on insects and small reptiles.
678:, the gular pump of monitor lizards enables them to overcome the effect of one lung at a time being compressed by their running gait. The evolutionary development of the gular pump is functionally analogous to that of the diaphragm in mammals, which ventilates the lung independently of locomotion. Even amongst monitors, the crocodile monitor is one of the most agile species, quickly running up tree trunks and leaping from branch to branch. Investigation supports the idea that gular breathing is an evolutionary development that masks the effect of Carrier's constraint.
631:
51:
779:
95:
770:). They are known to exhibit a warning posture, in which they carry their tails rolled up behind them. According to native belief, they give a warning call if they see crocodiles. In general crocodile monitors avoid human contact, but their bite is capable of causing infection, like the Komodo dragon's. One fatality was reported from a bite in 1983 when a Papuan woman was bitten and later died from an infection.
70:
655:
865:. It faces threats from deforestation and poaching, as it is hunted and skinned by native peoples, who consider the monitor an evil spirit that "climbs trees, walks upright, breathes fire, and kills men", to make drums . In 2008, 52 individuals were maintained at 17 zoological parks in the United States, with an unknown number in private collections.
398:. Attempts at captive breeding have been mostly unsuccessful. In New Guinea, the lizard is sometimes hunted and skinned by tribesmen to make drums. It is described as an evil spirit that "climbs trees, walks upright, breathes fire, and kills men", yet the local people maintain that it gives warnings if
841:
Reproduction of crocodile monitors has only been observed in captivity, so nothing is known about its reproduction in the wild. The egg clutches, comprising four to 12 eggs, are deposited around
October to January, with the eggs showing a remarkable difference in dimensions, a phenomenon for which no
717:
Claims of crocodile monitors attaining large weights also are unverified; the species is much more slender than Komodo dragons of comparable length. The typical reported length of this species is less than 200 cm (79 in) with a matching body mass of about 20 kg (44 lb), but such a
693:
Crocodile monitors upon hatching measure about 45 cm (18 in) long and they reach maturity when about 170 cm (67 in). Reports of the maximal length vary greatly and are the subject of much dispute. It possibly attains the greatest length among extant species of lizards, although it
786:
The teeth of crocodile monitors do not resemble those of other monitor species, which typically are blunt, peglike, and face slightly rearward. Their upper teeth are long, fang-like, set vertically in the jawbone, adapted to hooking into fast-moving prey such as birds, bats, and rodents. Their lower
758:
species, the tail also may be used for defense, as captive specimens may attempt to lash keepers with their tails. This species occasionally is seen in the pet trade, but has a reputation for aggression and unpredictability. Although they are known to rest and bask in trees, they sleep on the ground
641:
A characteristic feature of the crocodile monitor is its blunt, bulbous snout, which makes it look different from every other monitor on New Guinea, and suggested the common name "tree crocodile". The body of the lizard is dark green with rings of yellow spots. The tail is banded yellow and black
722:
species. Ten specimens were found to weigh only 5 to 6.38 kg (11.0 to 14.1 lb) at a length of 1.16 to 2.25 m (3 ft 10 in to 7 ft 5 in), and a review of healthy adults in captivity found that a typical weight was up to 6 kg (13 lb). The average size of
233:
832:
This species has been observed hunting prey in a unique fashion for monitor lizards. Rather than following its prey to ambush it from behind, the crocodile monitor may stalk its prey and anticipate where it will run, meeting it headlong.
732:
1808:
1559:
645:
Its teeth are long, straight, and sharp. Its claws are prominent and strongly curved. Males reach a considerably larger size than females in both weight and length, and also tend to have a more robust head.
379:. Like all monitors, it has anatomical features that enable it to breathe more easily when running than other lizards, and is sometimes considered one of the most agile monitor species.
1812:
1563:
2787:
690:
varanid species in that the animal's tail is much longer than the snout-to-vent length in both juveniles and adults. The tail generally is 2–2.7 times the snout-to-vent length.
1679:
1202:
Vidal, N.; Marin, J.; Sassi, J.; Battistuzzi, F.U.; Donnellan, S.; Fitch, A.J.; Fry, B.G.; Vonk, F.J.; Rodriguez de la Vega, R.C.; Couloux, A.; Hedges, S.B. (2012).
2691:
2730:
723:
crocodile monitors caught in one study was 99.2 cm (39.1 in) with a weight of 2.02 kg (4.5 lb), but these must have been young specimens.
2833:
694:
is considerably less massive than the Komodo dragon. Crocodile monitors have been confirmed to reach up to 255 cm (100 in) in length (record by a
618:, so the full extent of its range is unknown. Its remote and generally inaccessible habitat is an obstacle to detailed study of this monitor in its natural
375:
to eggs, other reptiles, amphibians and carrion. Its large, backwards-curving teeth are better adapted than those of most monitors for seizing fast-moving
356:
with a dark green body marked with bands of yellowish spots. It has a characteristic blunt snout and a very long prehensile tail. It lives among the
2665:
1305:
342:, verified at up to 255 cm (100 in). Its tail is exceptionally long, with some specimens having been claimed to exceed the length of the
2848:
2704:
2843:
1465:
2868:
2853:
1772:
1747:
1524:
1492:
1425:
1371:
Owerkowicz, T.; Farmer, C. G.; Hicks, J. W.; Brainerd, E. L. (1999). "Contribution of Gular
Pumping to Lung Ventilation in Monitor Lizards".
1137:
1085:
996:
980:
940:
2873:
2858:
2179:
1858:
614:
swamps, sometimes venturing out of these areas during floods in the rainy seasons. No detailed field investigation data are available for
2863:
2053:
1683:
1666:
Hunting and
Ethnozoology Systems of Monitor Lizards (Fam. Varanidae) Utilized by Yaur Tribe at National Park of Cenderawasih Gulf
94:
674:
in the animal's throat assists lung ventilation. Unlike most lizards that cannot breathe efficiently while running, because of
1831:
1335:
754:
grip. The primary function of the tail, however, is as a counterbalance when leaping from branch to branch. As in some other
339:
2709:
598:
The largest of the seven species of monitors found on the island of New Guinea, crocodile monitors occur in both the state
935:. Ithaca, N.Y., USA: Society for the Study of Amphibians and Reptiles in cooperation with Deutsche Gesellschaft für Herp.
1011:
1456:
Auliya, M.; Koch, A. (2020). "Visual
Identification Guide to the Monitor Lizard Species of the World (Genus Varanus)".
2367:
1738:
McDade, Melissa C.; Grzimek, Bernhard; Schlager, Neil; Hutchins, Michael; Trumpey, Joseph E.; Olendorf, Donna (2004).
2161:
1204:"Molecular evidence for an Asian origin of monitor lizards followed by Tertiary dispersals to Africa and Australasia"
2279:
630:
2735:
762:
These monitors rise up on their hind legs to check their surroundings, behaviour that also has been documented in
2152:
635:
518:
857:, which requires an exportation permit for international trade. It is not listed as a threatened species on the
2362:
1942:
1851:
817:, and hunting dogs, and hauls its prey into the canopy to consume it. Captive specimens have been known to eat
1597:
Turner, Frederick B.; Jennrich, Robert I.; Weintraub, Joel D. (1969). "Home ranges and body size of lizards".
50:
2878:
2372:
675:
510:
443:
280:
2617:
2579:
2333:
2261:
2252:
2198:
2116:
1993:
1388:
1076:
Lizards: A Natural
History of Some Uncommon Creatures--Extraordinary Chameleons, Iguanas, Geckos, and More
862:
1724:
2769:
1960:
1951:
1933:
1540:
Böhme, W.; Ziegler, T. (1997). "Großwarane im Museum Koenig, mit
Bemerkungen zu Afrikas größter Echse".
1309:
586:) was also a member of the group. Monitors apparently colonized Australasia from Asia about 32 Mya; the
201:
778:
906:
2652:
2626:
2288:
2098:
2089:
2035:
1709:
1606:
1380:
1167:
426:
2470:
2216:
2062:
2017:
1844:
1393:
695:
659:
607:
59:
1429:
2026:
1622:
1284:
429:
of 48 cm (19 in), and a 114 cm (45 in) long tail. It was subordinated to the
395:
249:
89:
2774:
2696:
2544:
2756:
2722:
2639:
2427:
2297:
2270:
2107:
1768:
1743:
1520:
1488:
1406:
1276:
1235:
1133:
1081:
992:
936:
854:
740:
555:
522:
499:
482:
232:
494:), meaning "lizard". The term "monitor" is thought to have come about from confusion between
2838:
2810:
2782:
2761:
2315:
2143:
1614:
1398:
1266:
1225:
1215:
1175:
901:
667:
599:
509:, meaning "warning". The term "goanna" came about as a corruption of the name "iguana". The
31:
2631:
1704:
1426:"Research by UMass Amherst Biologist Suggests that Lizards Offer Evolutionary Freeze-Frame"
2743:
2434:
2411:
2243:
2189:
750:
lizard. It can hang onto branches with its rear legs, and occasionally use its tail as a
1610:
1384:
1171:
1155:
883:
2324:
2225:
2080:
2044:
2008:
1230:
1203:
671:
544:
434:
418:
357:
328:
293:
263:
218:
166:
1179:
394:. Little is known of its reproduction and development, as it is difficult to breed in
2827:
2717:
2420:
2342:
2125:
1514:
1271:
1254:
1074:
892:
798:
788:
699:
571:
469:
based on the color of their snout and the overall brightness of their color pattern.
422:
383:
367:
of the southeastern part of New Guinea, feeding opportunistically on everything from
343:
267:
222:
79:
74:
1288:
714:
that reportedly was 475 cm (187 in) long, but these sizes are unverified.
562:
identifies the crocodile monitor as a member of a species cluster that includes the
2644:
2537:
2516:
2488:
2234:
2170:
2134:
1969:
1343:
711:
687:
563:
1402:
718:
weight is questionable for healthy (i.e., not obese) individuals of this slender,
2800:
931:
Klipfel, Meghan; Peters, Wilhelm C. H.; Bauer, Aaron M.; Günther, Rainer (1995).
17:
2678:
2611:
2530:
2523:
2502:
2495:
2306:
1984:
1788:
763:
654:
551:
1255:"A combined evidence phylogenetic analysis of Anguimorpha (Reptilia: Squamata)"
610:. It inhabits the high and low canopies of the lowland rainforests and coastal
1912:
1867:
751:
450:
364:
335:
2602:
787:
teeth are housed in a fleshy sheath. In the wild, crocodile monitors are the
2509:
2453:
2445:
1925:
806:
603:
579:
547:
allowed the varanids to spread into what is now the
Indonesian archipelago.
540:
399:
156:
106:
1640:
1410:
1280:
1239:
1220:
2748:
702:). Some have been claimed to be much larger, including an individual from
2596:
2207:
1897:
1891:
1885:
747:
719:
703:
611:
462:
387:
350:
331:
146:
126:
1017:
731:
2670:
2551:
1626:
1112:
Best
Practice Guidelines for the Crocodile Monitor (Varanus salvadorii)
1046:
Bayless, Mark K. (June 1, 1998). "The
Artellia: Dragons of the Trees".
810:
793:
619:
535:
466:
361:
324:
136:
539:
originated in Asia about 40 million years ago (Mya). Around 15 Mya, a
2683:
2463:
2395:
1879:
826:
710:
that reportedly was 427 cm (168 in) and an individual from
707:
458:
372:
353:
116:
2573:
1618:
933:
The herpetological contributions of
Wilhelm C. H. Peters (1815-1883)
882:
Shea, G.; Allison, A.; Parker, F.; O'Shea, M.; Tallowin, O. (2016).
338:. It is the largest monitor lizard in New Guinea and is one of the
2795:
777:
730:
653:
629:
559:
430:
391:
666:
Like many monitor species, the crocodile monitor has mammal-like
2657:
1469:
1115:
858:
822:
818:
814:
376:
368:
2577:
2393:
1910:
1840:
802:
1836:
1253:
Conrad, J.L.; Ast, J.C.; Montanari, S.; Norell, M.A. (2011).
706:
that reportedly was 323 cm (127 in) long, one from
1132:. Baltimore: Johns Hopkins University Press. xiii + 296 pp.
457:, but the captive reptiles market distinguishes those from
1308:. Central Florida Zoo and Botanial Gardens. Archived from
1128:
Beolens, Bo; Watkins, Michael; Grayson, Michael (2011).
1664:
Iyai, D.A., Murwanto, A.G. & Killian, A.M. (2011)
525:, an Italian ornithologist who worked in New Guinea.
2408:
1080:. Stillwater, MN: Voyageur Press. pp. 140–141.
813:. Natives have reported that it can take down pigs,
2586:
2480:
2444:
2407:
2355:
2006:
1982:
1924:
1073:
1767:. Vol. 2. Hague: W. Junk. pp. 803–813.
907:10.2305/IUCN.UK.2016-1.RLTS.T42485775A42485784.en
578:). Morphological considerations suggest that the
987:. In Pianka, E.R.; King, D.; King, R.A. (eds.).
991:. Indiana University Press. pp. 234–244.
1852:
1451:
1449:
1447:
8:
1587:. Bloomington, IN: Indiana University Press.
1197:
1195:
503:
489:
1516:The Guinness Book of Animal Facts and Feats
974:
972:
2574:
2404:
2390:
1921:
1907:
1859:
1845:
1837:
970:
968:
966:
964:
962:
960:
958:
956:
954:
952:
425:in 1878 based on a female specimen with a
231:
68:
49:
38:
1392:
1270:
1229:
1219:
1105:
1103:
1101:
1099:
1097:
1041:
1039:
1037:
1035:
1033:
1031:
1029:
1027:
1025:
905:
791:in New Guinea, feeding on birds (such as
1705:"Varanus salvadorii (Crocodile Monitor)"
1366:
1364:
1362:
1360:
1067:
1065:
1063:
1061:
1508:
1506:
1504:
1340:Leeward Community College's Zoology 101
874:
1765:Biogeography and Ecology of New Guinea
1742:. Detroit: Thomson/Gale. p. 368.
1466:Federal Agency for Nature Conservation
1330:
1328:
1326:
1300:
1298:
1072:Netherton, J.; Badger, D. P. (2002).
382:Crocodile monitors are threatened by
7:
2180:Northern Sierra Madre forest monitor
2054:Black-spotted ridge-tailed monitor (
1682:. Wildlife World Zoo. Archived from
1583:Pianka, E., D. King, R. King. 2004.
1487:. Neptune, NJ: T.F.H. Publications.
1472:SSC Monitor Lizard Specialist Group.
926:
924:
686:Crocodile monitors are unique among
2834:IUCN Red List least concern species
1680:"Aloha to three crocodile monitors"
893:IUCN Red List of Threatened Species
634:Detail of bulbous snout — taken at
490:
1740:Grzimek's Animal Life Encyclopedia
1560:"ANIMAL BYTES - Crocodile Monitor"
746:The crocodile monitor is a highly
550:Based upon DNA sequences of three
25:
1180:10.1038/scientificamerican0399-84
1130:The Eponym Dictionary of Reptiles
853:is currently protected under the
543:connection between Australia and
449:There are no formally recognized
1272:10.1111/j.1096-0031.2010.00330.x
93:
1428:. UMass Amherst. Archived from
670:abilities; a positive pressure
590:clade then arose about 17 Mya.
288:Varanus (Papusaurus) salvadorii
2849:Reptiles of Western New Guinea
1562:. Seaworld.org. Archived from
1342:. Honolulu Zoo. Archived from
1306:"New Guinea Crocodile Monitor"
1:
2844:Monitor lizards of New Guinea
1763:Gressitt, J. Linsley (1982).
1668:. Biota Vol. 16 (2): 278−286.
1585:Varanoid Lizards of the World
1403:10.1126/science.284.5420.1661
1018:Reptarium.cz Reptile Database
989:Varanoid Lizards of the World
2869:Taxa named by Wilhelm Peters
2854:Reptiles of Papua New Guinea
2280:Short-tailed pygmy monitor (
1118:Reptile Taxon Advisory Group
2874:Taxa named by Giacomo Doria
2859:Endemic fauna of New Guinea
2895:
2864:Reptiles described in 1878
2162:Mitchell's water monitor (
1952:Rio Fuerte beaded lizard (
1943:Guatemalan beaded lizard (
1789:"Appendices I, II and III"
1483:Sprackland, R. G. (1992).
829:, chickens, and dog food.
29:
2403:
2389:
1920:
1906:
1874:
1519:. Guinness Superlatives.
801:, small mammals (such as
255:
248:
239:
230:
207:
200:
90:Scientific classification
88:
66:
57:
48:
41:
2334:Yellow-spotted monitor (
2199:Peach-throated monitor (
2153:Merten's water monitor (
2117:Kimberely rock monitor (
1994:Earless monitor lizard (
900:: e.T42485775A42485784.
809:), frogs, reptiles, and
346:, however less massive.
30:Not to be confused with
2253:Rennel Island monitor (
1961:Mexican beaded lizard (
1934:Chiapan beaded lizard (
1154:Ciofi, Claudio (1999).
782:Crocodile monitor skull
759:or submerged in water.
642:and is extremely long.
444:George Albert Boulenger
413:was first described as
390:, and are protected by
349:Crocodile monitors are
2289:Spiny-tailed monitor (
2099:Emerald tree monitor (
2036:Black-headed monitor (
1221:10.1098/rsbl.2012.0460
863:Endangered Species Act
783:
743:
663:
638:
626:Biology and morphology
504:
481:, is derived from the
2770:Paleobiology Database
2262:Rosenberg's monitor (
2217:Pygmy mulga monitor (
2063:Blue-tailed monitor (
2018:Asian water monitor (
1424:Brainerd, E. (1999).
1110:Reh, B., ed. (2021),
781:
734:
657:
633:
315:), also known as the
244:, indicated by green
2027:Black tree monitor (
1725:"Varanus Salvadorii"
1710:Animal Diversity Web
1647:. Memim Encyclopedia
676:Carrier's constraint
517:, is derived from a
427:snout-to-vent length
2471:Varanus amnhophilis
2298:Turquoise monitor (
2271:Roughneck monitor (
2090:Dumeril's monitor (
2072:Crocodile monitor (
1809:"Crocodile Monitor"
1611:1969Ecol...50.1076T
1385:1999Sci...284.1661O
1379:(5420): 1661–1663.
1336:"Crocodile Monitor"
1172:1999SciAm.280c..84C
1160:Scientific American
1156:"The Komodo Dragon"
983:Varanus salvadorii
979:Horn, H-G. (2004).
861:Red List or on the
321:Salvadori's monitor
240:Estimated range of
60:Conservation status
2632:Varanus_salvadorii
2618:Varanus salvadorii
2588:Varanus salvadorii
2368:Cretaceous lizards
2356:Related categories
2316:Savannah monitor (
2144:Mangrove monitor (
1643:Varanus salvadorii
1142:Varanus salvadorii
1013:Varanus salvadorii
886:Varanus salvadorii
851:Varanus salvadorii
784:
744:
664:
662:in the background.
660:Rainbow eucalyptus
639:
616:V. salvadorii
560:cladistic analysis
477:The generic name,
440:Varanus salvadorii
415:Monitor salvadorii
411:Varanus salvadorii
312:Varanus salvadorii
275:Varanus salvadorii
259:Monitor salvadorii
211:Varanus salvadorii
193:V. salvadorii
43:Varanus salvadorii
2821:
2820:
2757:Open Tree of Life
2580:Taxon identifiers
2569:
2568:
2565:
2564:
2561:
2560:
2385:
2384:
2381:
2380:
2318:V. exanthematicus
2244:Pilbara monitor (
2190:Peacock monitor (
1945:H. charlesbogerti
1774:978-90-6193-094-5
1749:978-0-7876-5362-0
1526:978-0-85112-235-9
1513:Wood, G. (1983).
1494:978-0-86622-634-9
1138:978-1-4214-0135-5
1087:978-0-7603-2579-7
998:978-0-253-34366-6
942:978-0-916984-35-9
855:CITES Appendix II
741:Oklahoma City Zoo
523:Tommaso Salvadori
307:crocodile monitor
303:
302:
83:
27:Species of lizard
18:Crocodile monitor
16:(Redirected from
2886:
2814:
2813:
2804:
2803:
2791:
2790:
2778:
2777:
2765:
2764:
2752:
2751:
2739:
2738:
2726:
2725:
2713:
2712:
2700:
2699:
2687:
2686:
2674:
2673:
2661:
2660:
2648:
2647:
2635:
2634:
2622:
2621:
2620:
2607:
2606:
2605:
2575:
2410:
2405:
2391:
2325:Yellow monitor (
2300:V. caerulivirens
2226:Quince monitor (
2108:Gray's monitor (
2081:Desert monitor (
2045:Bengal monitor (
1922:
1908:
1861:
1854:
1847:
1838:
1824:
1823:
1821:
1820:
1811:. Archived from
1805:
1799:
1798:
1796:
1795:
1785:
1779:
1778:
1760:
1754:
1753:
1735:
1729:
1728:
1721:
1715:
1714:
1701:
1695:
1694:
1692:
1691:
1678:Stafford, Grey.
1675:
1669:
1662:
1656:
1655:
1653:
1652:
1637:
1631:
1630:
1605:(6): 1076–1081.
1594:
1588:
1581:
1575:
1574:
1572:
1571:
1556:
1550:
1549:
1537:
1531:
1530:
1510:
1499:
1498:
1480:
1474:
1473:
1453:
1442:
1441:
1439:
1437:
1421:
1415:
1414:
1396:
1368:
1355:
1354:
1352:
1351:
1332:
1321:
1320:
1318:
1317:
1302:
1293:
1292:
1274:
1250:
1244:
1243:
1233:
1223:
1199:
1190:
1189:
1187:
1186:
1151:
1145:
1126:
1120:
1119:
1107:
1092:
1091:
1079:
1069:
1056:
1055:
1043:
1020:
1009:
1003:
1002:
976:
947:
946:
928:
919:
918:
916:
914:
909:
879:
764:Gould's monitors
600:Papua New Guinea
507:
493:
492:
297:
284:
271:
235:
213:
98:
97:
77:
72:
71:
53:
39:
32:Varanus salvator
21:
2894:
2893:
2889:
2888:
2887:
2885:
2884:
2883:
2824:
2823:
2822:
2817:
2809:
2807:
2799:
2794:
2786:
2781:
2773:
2768:
2760:
2755:
2747:
2744:Observation.org
2742:
2734:
2729:
2721:
2716:
2708:
2703:
2695:
2690:
2682:
2677:
2669:
2664:
2656:
2651:
2643:
2638:
2630:
2625:
2616:
2615:
2610:
2601:
2600:
2595:
2582:
2570:
2557:
2476:
2456:Varanus priscus
2440:
2435:Paranecrosaurus
2412:Palaeovaranidae
2399:
2377:
2363:Monitor lizards
2351:
2343:Yemen monitor (
2126:Komodo dragon (
2002:
1978:
1916:
1902:
1870:
1865:
1832:Honoluluzoo.org
1828:
1827:
1818:
1816:
1807:
1806:
1802:
1793:
1791:
1787:
1786:
1782:
1775:
1762:
1761:
1757:
1750:
1737:
1736:
1732:
1723:
1722:
1718:
1703:
1702:
1698:
1689:
1687:
1686:on July 1, 2009
1677:
1676:
1672:
1663:
1659:
1650:
1648:
1639:
1638:
1634:
1619:10.2307/1936898
1596:
1595:
1591:
1582:
1578:
1569:
1567:
1558:
1557:
1553:
1542:Tier und Museum
1539:
1538:
1534:
1527:
1512:
1511:
1502:
1495:
1482:
1481:
1477:
1455:
1454:
1445:
1435:
1433:
1432:on 30 June 2009
1423:
1422:
1418:
1370:
1369:
1358:
1349:
1347:
1334:
1333:
1324:
1315:
1313:
1312:on May 28, 2008
1304:
1303:
1296:
1252:
1251:
1247:
1208:Biology Letters
1201:
1200:
1193:
1184:
1182:
1153:
1152:
1148:
1127:
1123:
1109:
1108:
1095:
1088:
1071:
1070:
1059:
1045:
1044:
1023:
1010:
1006:
999:
978:
977:
950:
943:
930:
929:
922:
912:
910:
881:
880:
876:
871:
848:
839:
776:
768:V. gouldii
729:
684:
652:
628:
596:
531:
475:
438:under the name
408:
358:mangrove swamps
340:longest lizards
292:
290:
279:
277:
262:
261:
226:
215:
209:
196:
92:
84:
73:
69:
62:
35:
28:
23:
22:
15:
12:
11:
5:
2892:
2890:
2882:
2881:
2879:Apex predators
2876:
2871:
2866:
2861:
2856:
2851:
2846:
2841:
2836:
2826:
2825:
2819:
2818:
2816:
2815:
2805:
2792:
2779:
2766:
2753:
2740:
2727:
2714:
2701:
2688:
2675:
2662:
2649:
2636:
2623:
2608:
2592:
2590:
2584:
2583:
2578:
2567:
2566:
2563:
2562:
2559:
2558:
2556:
2555:
2548:
2541:
2534:
2527:
2520:
2513:
2506:
2499:
2492:
2484:
2482:
2478:
2477:
2475:
2474:
2467:
2460:
2450:
2448:
2442:
2441:
2439:
2438:
2431:
2424:
2416:
2414:
2401:
2400:
2394:
2387:
2386:
2383:
2382:
2379:
2378:
2376:
2375:
2373:Helodermatidae
2370:
2365:
2359:
2357:
2353:
2352:
2350:
2349:
2340:
2331:
2322:
2313:
2304:
2295:
2286:
2282:V. brevicauda
2277:
2268:
2264:V. rosenbergi
2259:
2255:V. juxtindicus
2250:
2241:
2237:V. albigularis
2235:Rock monitor (
2232:
2223:
2214:
2205:
2196:
2192:V. auffenbergi
2187:
2177:
2171:Nile monitor (
2168:
2159:
2150:
2141:
2135:Lace monitor (
2132:
2128:V. komodoensis
2123:
2114:
2105:
2096:
2087:
2078:
2069:
2060:
2051:
2047:V. bengalensis
2042:
2033:
2024:
2014:
2012:
2004:
2003:
2001:
2000:
1990:
1988:
1980:
1979:
1977:
1976:
1970:Gila monster (
1967:
1958:
1954:H. exasperatum
1949:
1940:
1930:
1928:
1926:Helodermatidae
1918:
1917:
1911:
1904:
1903:
1901:
1900:
1894:
1888:
1882:
1875:
1872:
1871:
1866:
1864:
1863:
1856:
1849:
1841:
1835:
1834:
1826:
1825:
1800:
1780:
1773:
1755:
1748:
1730:
1716:
1696:
1670:
1657:
1632:
1589:
1576:
1551:
1532:
1525:
1500:
1493:
1475:
1443:
1416:
1394:10.1.1.486.481
1356:
1322:
1294:
1265:(3): 230–277.
1245:
1214:(5): 853–855.
1191:
1146:
1121:
1114:(1 ed.),
1093:
1086:
1057:
1021:
1004:
997:
948:
941:
920:
873:
872:
870:
867:
847:
844:
838:
835:
775:
772:
735:A specimen of
728:
725:
683:
680:
658:In the trees.
651:
648:
627:
624:
595:
592:
576:V. komodoensis
545:Southeast Asia
530:
527:
474:
471:
419:Wilhelm Peters
407:
404:
329:monitor lizard
317:Papuan monitor
301:
300:
299:
298:
285:
272:
253:
252:
246:
245:
237:
236:
228:
227:
216:
205:
204:
198:
197:
190:
188:
184:
183:
176:
172:
171:
164:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
86:
85:
67:
64:
63:
58:
55:
54:
46:
45:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2891:
2880:
2877:
2875:
2872:
2870:
2867:
2865:
2862:
2860:
2857:
2855:
2852:
2850:
2847:
2845:
2842:
2840:
2837:
2835:
2832:
2831:
2829:
2812:
2806:
2802:
2797:
2793:
2789:
2784:
2780:
2776:
2771:
2767:
2763:
2758:
2754:
2750:
2745:
2741:
2737:
2732:
2728:
2724:
2719:
2715:
2711:
2706:
2702:
2698:
2693:
2689:
2685:
2680:
2676:
2672:
2667:
2663:
2659:
2654:
2650:
2646:
2641:
2637:
2633:
2628:
2624:
2619:
2613:
2609:
2604:
2598:
2594:
2593:
2591:
2589:
2585:
2581:
2576:
2572:
2554:
2553:
2549:
2547:
2546:
2542:
2540:
2539:
2535:
2533:
2532:
2528:
2526:
2525:
2521:
2519:
2518:
2514:
2512:
2511:
2507:
2505:
2504:
2500:
2498:
2497:
2493:
2491:
2490:
2486:
2485:
2483:
2479:
2473:
2472:
2468:
2466:
2465:
2461:
2459:
2457:
2452:
2451:
2449:
2447:
2443:
2437:
2436:
2432:
2430:
2429:
2425:
2423:
2422:
2421:Palaeovaranus
2418:
2417:
2415:
2413:
2406:
2402:
2397:
2392:
2388:
2374:
2371:
2369:
2366:
2364:
2361:
2360:
2358:
2354:
2348:
2346:
2345:V. yemenensis
2341:
2339:
2337:
2332:
2330:
2328:
2327:V. flavescens
2323:
2321:
2319:
2314:
2312:
2310:
2307:Sand goanna (
2305:
2303:
2301:
2296:
2294:
2292:
2291:V. acanthurus
2287:
2285:
2283:
2278:
2276:
2274:
2273:V. rudicollis
2269:
2267:
2265:
2260:
2258:
2256:
2251:
2249:
2247:
2242:
2240:
2238:
2233:
2231:
2229:
2224:
2222:
2220:
2215:
2213:
2211:
2206:
2204:
2202:
2197:
2195:
2193:
2188:
2185:
2181:
2178:
2176:
2174:
2169:
2167:
2165:
2160:
2158:
2156:
2151:
2149:
2147:
2142:
2140:
2138:
2133:
2131:
2129:
2124:
2122:
2120:
2115:
2113:
2111:
2106:
2104:
2102:
2097:
2095:
2093:
2088:
2086:
2084:
2079:
2077:
2075:
2074:V. salvadorii
2070:
2068:
2066:
2061:
2059:
2057:
2052:
2050:
2048:
2043:
2041:
2039:
2034:
2032:
2030:
2025:
2023:
2021:
2016:
2015:
2013:
2011:
2010:
2005:
1999:
1997:
1996:L. borneensis
1992:
1991:
1989:
1987:
1986:
1981:
1975:
1973:
1968:
1966:
1964:
1959:
1957:
1955:
1950:
1948:
1946:
1941:
1939:
1937:
1932:
1931:
1929:
1927:
1923:
1919:
1914:
1909:
1905:
1899:
1895:
1893:
1889:
1887:
1883:
1881:
1877:
1876:
1873:
1869:
1862:
1857:
1855:
1850:
1848:
1843:
1842:
1839:
1833:
1830:
1829:
1815:on 2007-12-14
1814:
1810:
1804:
1801:
1790:
1784:
1781:
1776:
1770:
1766:
1759:
1756:
1751:
1745:
1741:
1734:
1731:
1726:
1720:
1717:
1712:
1711:
1706:
1700:
1697:
1685:
1681:
1674:
1671:
1667:
1661:
1658:
1646:
1644:
1636:
1633:
1628:
1624:
1620:
1616:
1612:
1608:
1604:
1600:
1593:
1590:
1586:
1580:
1577:
1566:on 2007-12-14
1565:
1561:
1555:
1552:
1547:
1543:
1536:
1533:
1528:
1522:
1518:
1517:
1509:
1507:
1505:
1501:
1496:
1490:
1486:
1485:Giant lizards
1479:
1476:
1471:
1467:
1463:
1459:
1452:
1450:
1448:
1444:
1431:
1427:
1420:
1417:
1412:
1408:
1404:
1400:
1395:
1390:
1386:
1382:
1378:
1374:
1367:
1365:
1363:
1361:
1357:
1346:on 2008-10-04
1345:
1341:
1337:
1331:
1329:
1327:
1323:
1311:
1307:
1301:
1299:
1295:
1290:
1286:
1282:
1278:
1273:
1268:
1264:
1260:
1256:
1249:
1246:
1241:
1237:
1232:
1227:
1222:
1217:
1213:
1209:
1205:
1198:
1196:
1192:
1181:
1177:
1173:
1169:
1165:
1161:
1157:
1150:
1147:
1143:
1139:
1135:
1131:
1125:
1122:
1117:
1113:
1106:
1104:
1102:
1100:
1098:
1094:
1089:
1083:
1078:
1077:
1068:
1066:
1064:
1062:
1058:
1053:
1049:
1042:
1040:
1038:
1036:
1034:
1032:
1030:
1028:
1026:
1022:
1019:
1015:
1014:
1008:
1005:
1000:
994:
990:
986:
984:
975:
973:
971:
969:
967:
965:
963:
961:
959:
957:
955:
953:
949:
944:
938:
934:
927:
925:
921:
908:
903:
899:
895:
894:
889:
887:
878:
875:
868:
866:
864:
860:
856:
852:
845:
843:
836:
834:
830:
828:
824:
820:
816:
812:
808:
804:
800:
796:
795:
790:
780:
773:
771:
769:
765:
760:
757:
753:
749:
742:
738:
737:V. salvadorii
733:
726:
724:
721:
715:
713:
709:
705:
701:
700:Museum Koenig
697:
691:
689:
681:
679:
677:
673:
669:
661:
656:
649:
647:
643:
637:
632:
625:
623:
621:
617:
613:
609:
605:
601:
593:
591:
589:
585:
581:
577:
573:
572:Komodo dragon
569:
565:
561:
557:
556:mitochondrial
553:
548:
546:
542:
538:
537:
528:
526:
524:
520:
516:
512:
511:specific name
508:
506:
501:
497:
487:
484:
480:
472:
470:
468:
464:
460:
456:
455:V. salvadorii
452:
447:
445:
441:
437:
436:
432:
428:
424:
423:Giacomo Doria
420:
416:
412:
405:
403:
401:
397:
393:
389:
385:
384:deforestation
380:
378:
374:
370:
366:
363:
359:
355:
352:
347:
345:
344:Komodo dragon
341:
337:
333:
330:
326:
322:
318:
314:
313:
308:
295:
289:
286:
282:
276:
273:
269:
265:
260:
257:
256:
254:
251:
247:
243:
242:V. salvadorii
238:
234:
229:
224:
220:
214:
212:
206:
203:
202:Binomial name
199:
195:
194:
189:
186:
185:
182:
181:
177:
174:
173:
170:
169:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
96:
91:
87:
81:
76:
75:Least Concern
65:
61:
56:
52:
47:
44:
40:
37:
33:
19:
2587:
2571:
2550:
2543:
2538:Palaeosaniwa
2536:
2529:
2522:
2517:Morohasaurus
2515:
2508:
2501:
2494:
2489:Arcanosaurus
2487:
2469:
2462:
2455:
2433:
2426:
2419:
2344:
2335:
2326:
2317:
2308:
2299:
2290:
2281:
2272:
2263:
2254:
2245:
2236:
2227:
2218:
2210:V. giganteus
2209:
2201:V. jobiensis
2200:
2191:
2183:
2173:V. niloticus
2172:
2164:V. mitchelli
2163:
2154:
2145:
2136:
2127:
2118:
2110:V. olivaceus
2109:
2100:
2092:V. dumerilii
2091:
2082:
2073:
2071:
2064:
2055:
2046:
2037:
2028:
2019:
2007:
1995:
1983:
1972:H. suspectum
1971:
1962:
1953:
1944:
1935:
1817:. Retrieved
1813:the original
1803:
1792:. Retrieved
1783:
1764:
1758:
1739:
1733:
1719:
1708:
1699:
1688:. Retrieved
1684:the original
1673:
1665:
1660:
1649:. Retrieved
1642:
1635:
1602:
1598:
1592:
1584:
1579:
1568:. Retrieved
1564:the original
1554:
1545:
1541:
1535:
1515:
1484:
1478:
1461:
1458:BFN-Skripten
1457:
1434:. Retrieved
1430:the original
1419:
1376:
1372:
1348:. Retrieved
1344:the original
1339:
1314:. Retrieved
1310:the original
1262:
1258:
1248:
1211:
1207:
1183:. Retrieved
1166:(3): 84–91.
1163:
1159:
1149:
1141:
1129:
1124:
1111:
1075:
1051:
1047:
1012:
1007:
988:
982:
932:
911:. Retrieved
897:
891:
885:
877:
850:
849:
846:Conservation
840:
837:Reproduction
831:
792:
789:top predator
785:
767:
761:
755:
745:
736:
716:
712:Port Moresby
692:
685:
665:
644:
640:
615:
597:
594:Distribution
587:
583:
575:
567:
564:lace monitor
549:
534:
532:
519:Latinization
514:
502:
495:
485:
478:
476:
454:
448:
439:
433:
414:
410:
409:
402:are nearby.
381:
348:
320:
316:
311:
310:
306:
304:
287:
274:
258:
241:
210:
208:
192:
191:
179:
178:
167:
42:
36:
2679:iNaturalist
2612:Wikispecies
2531:Telmasaurus
2503:Cherminotus
2496:Aiolosaurus
2454:Megalania (
2336:V. panoptes
2219:V. gilleni
2184:V. bitatawa
2155:V. mertensi
2119:V. glauerti
2101:V. prasinus
2065:V. doreanus
2029:V. beccarii
2020:V. salvator
1985:Lanthanotus
1963:H. horridum
1936:H. alvarezi
1548:(3): 65–74.
1436:19 December
1054:(6): 32–47.
913:19 November
650:Respiration
636:Cologne Zoo
365:rainforests
2828:Categories
2788:salvadorii
2309:V. gouldii
2228:V. melinus
2208:Perentie (
2146:V. indicus
2083:V. griseus
2056:V. baritji
2038:V. tristis
1868:Varanoidea
1819:2008-12-07
1794:2008-11-08
1690:2008-08-21
1651:2015-06-09
1570:2013-04-22
1350:2008-08-21
1316:2008-08-21
1259:Cladistics
1185:2006-12-21
1144:, p. 232).
869:References
807:bandicoots
752:prehensile
672:gular pump
608:West Papua
606:region of
604:Indonesian
584:V. priscus
570:) and the
533:The genus
515:salvadorii
451:subspecies
400:crocodiles
371:and small
336:New Guinea
180:Papusaurus
175:Subgenus:
2545:Saniwides
2510:Gobiderma
2446:Varanidae
2137:V. varius
1878:Kingdom:
1389:CiteSeerX
580:megalania
568:V. varius
529:Evolution
473:Etymology
446:in 1885.
396:captivity
281:Boulenger
264:W. Peters
219:W. Peters
187:Species:
157:Varanidae
113:Kingdom:
107:Eukaryota
2796:Species+
2723:42485775
2697:10364347
2597:Wikidata
2428:Eosaniwa
2246:V. bushi
1898:Squamata
1892:Reptilia
1886:Chordata
1884:Phylum:
1880:Animalia
1411:10356394
1289:84301257
1281:34875778
1240:22809723
1048:Reptiles
748:arboreal
727:Behavior
720:arboreal
704:Konedobu
696:specimen
612:mangrove
602:and the
554:and two
541:tectonic
498:and the
463:Jayapura
406:Taxonomy
388:poaching
351:arboreal
250:Synonyms
153:Family:
147:Squamata
137:Reptilia
127:Chordata
123:Phylum:
117:Animalia
103:Domain:
80:IUCN 3.1
2839:Varanus
2671:2470727
2658:1055073
2603:Q940883
2552:Estesia
2009:Varanus
1915:species
1896:Order:
1890:Class:
1627:1936898
1607:Bibcode
1599:Ecology
1381:Bibcode
1373:Science
1231:3441001
1168:Bibcode
1016:at the
827:rodents
811:carrion
794:Cacatua
756:Varanus
739:at the
668:aerobic
620:habitat
558:genes,
552:nuclear
536:Varanus
479:Varanus
467:Merauke
435:Varanus
373:mammals
362:coastal
332:endemic
325:species
323:, is a
294:Mertens
225:, 1878)
168:Varanus
163:Genus:
143:Order:
133:Class:
78: (
2811:208865
2808:uBio:
2775:374207
2762:172898
2749:104405
2710:716538
2481:Others
2464:Saniwa
2396:Fossil
1913:Extant
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797:sp.),
708:Kikori
688:extant
588:varius
505:warnen
500:German
483:Arabic
459:Sorong
354:lizard
296:, 1962
283:, 1885
270:, 1878
266:&
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2801:10060
2736:62049
2692:IRMNG
2684:39440
2645:7FG6S
1623:JSTOR
1285:S2CID
823:frogs
496:waral
486:waral
431:genus
392:CITES
369:birds
268:Doria
223:Doria
2731:NCBI
2718:IUCN
2705:ITIS
2666:GBIF
2524:Ovoo
2398:taxa
1769:ISBN
1744:ISBN
1521:ISBN
1489:ISBN
1470:IUCN
1438:2008
1407:PMID
1277:PMID
1236:PMID
1134:ISBN
1116:EAZA
1082:ISBN
993:ISBN
937:ISBN
915:2021
898:2016
859:IUCN
819:fish
815:deer
805:and
803:rats
799:eggs
774:Diet
682:Size
465:and
421:and
386:and
377:prey
360:and
305:The
2653:EoL
2640:CoL
2627:ADW
1615:doi
1462:552
1399:doi
1377:284
1267:doi
1226:PMC
1216:doi
1176:doi
1164:280
1140:. (
902:doi
698:at
521:of
491:ورل
453:of
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