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Orinoco crocodile

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1320:, though there is little to suggest that Orinoco specimens in modern times can rival these species. Sexual maturity for Orinoco crocodiles is obtained for females at around 2.5 m (8 ft 2 in) while that of males seems to be obtained around 3 m (9 ft 10 in) length, with most adult crocodiles of the species exceeding 93 kg (205 lb). Average length of wild adult females from 1985 to 1992 was found to be 3.06 m (10 ft 0 in) while that of males is not known to have been surveyed at that time. If not culled by humans, mature males easily exceed 3.6 m (12 ft) and attain a length of as much as 4.1 to 4.8 m (13 to 16 ft) as they grow throughout life, perhaps weighing 500 to 700 kg (1,100 to 1,500 lb) in big specimens, while females may grow over the expected size of 3.25 m (10 ft 8 in) in length and may sometimes exceed 225 kg (496 lb). According to 1328:, males have been reported to reach at least 4.1 m (13 ft 5 in) in length and weigh 380 kg (840 lb), while females reach up to 3.2 m (10 ft 6 in) and weigh up to 210 kg (460 lb). The largest specimen historically reported was shot in 1800, and allegedly measured at 6.78 m (22 ft 3 in), although the source is considered reliable, unfortunately, no voucher specimen is known from this. Because of extensive hunting for their skins in the 20th century, such giants do not exist today, and modern Orinoco crocodiles have not been reported to exceed 5.1 m (16 ft 9 in) in length. Merchán listed the maximum total length attainable for a male as 5.2 m (17 ft 1 in) and the maximum length of a female as 3.6 m (11 ft 10 in). 1276: 1476:, Orinoco crocodiles retreat to burrows they excavate into the riverbanks. The adult pair mates during the drier period of the year. Adult males attract the females by roaring. The females often outnumber the males two to one and appear to select the males with the deepest roars. In a reintroduced population, only 6 of 14 potential adult males parented over 90% of eggs laid. Usually 14 weeks after mating, the female crocodile will dig a nest and lay about 40 eggs. It is a hole-nester, as are all crocodilians, and it usually makes its nest on a sand bank. The eggs incubate under a mixture of soil and rotting vegetation for around three months. The most common predator of buried eggs are 1518: 1465: 1509:, though these species are sometimes also caught and killed by the defending mother crocodile. Adults have no predators except for humans. Females have defended pods of juveniles for over three years, though closer to one year to independence is generally most common. A study of captive Orinoco crocodiles noted the aggressive behavior of adults while nesting and noted that the normally relatively docile crocodilians could not be approached while they were actively brooding. Without persecution, it is possible that Orinoco crocodiles may reach a lifespan of 70–80 years. 1382: 738: 1572:. Additionally, a large number of individuals are held at captive-breeding facilities in Venezuela. Since the early 1990s, a large number of hatchlings have been released both into private ranchlands (especially in the Llanos where nature-oriented tourism is important for the local economy) and in national parks in Venezuela. While six Venezuelan captive-breeding programs continue today, many are plagued by lack of funds or staff, as well as conflicts between private and state-owned facilities. 1141: 829: 859: 1324:, the average length of adults that they were able to examine was only 3 m (9 ft 10 in) and the largest specimen found firsthand was 4.2 m (13 ft 9 in). In captivity, at the Roberto Franco Tropical Biological Station (EBTRF), the largest male was recorded at 4.2 m (13 ft 9 in) and weighed 428 kg (944 lb), and maximum size recorded for females was 3.9 m (12 ft 10 in) long and 195 kg (430 lb) in weight. In 882: 1084: 1028: 387:; males have been reported up to 6.8 m (22 ft 4 in) in the past, weighing over 900 kg (2,000 lb), but such sizes do not exist today, 5.2 m (17 ft 1 in) being a more widely accepted maximum size. A large male today may attain 4.2 m (13 ft 9 in) in length and can weigh up to 450 kg (1,000 lb), while females are substantially smaller with the largest likely to weigh around 225 kg (496 lb). 197: 341: 1439: 155: 131: 3252: 1233: 1251:. This species generally has a pale tan hide, though at least three coloration variations are known, with some almost completely yellowish, and some a dark brownish-gray. The skin can change colour over long periods of time; this phenomenon has been recorded in other species that can gradually change the amount of 1389:
Little study has gone into the dietary biology of Orinoco crocodiles but from eye-witness accounts and partial studies from captivity and crocodile farms, the majority of the Orinoco crocodile's diet appears to consists of large fish. The species' relatively narrow snout is ideally suited to minimize
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although the tegus are sometimes caught and killed by the mother crocodile. During the night the young hatch and call to their mother; she digs them out of the nest and carries them to the water, which is considerably higher at this point. Young Orinoco crocodiles are often at risk from predation by
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Gorzula, S., & Seijas, A. E. (1986). "The Common Caiman", p. 44 in Crocodiles: Proceedings of the 7th Working Meeting of the Crocodile Specialist Group of the Species Survival Commission of the International Union for Conservation of Nature and Natural Resources, Caracas, Venezuela, 21 to 28
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that two or three adult people were killed per year by Orinoco crocodiles. A small number of better documented fatal attacks were reported in the 1900s–1930s when the species was still relatively common. The only well-documented recent attack, on a fisherman in 2009, was serious but not fatal. A
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Attacks on humans have been reported, but this is highly unlikely to be a common behavior today, given the very small population of the species and its relative isolation from large human settlements. Historically attacks were not unusual and on his trip to the region in 1800, natives told
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in their skin. These crocodiles have dark-brown markings, which present as more pronounced bands in younger specimens and as scattered markings on mature ones. One individual measuring 3.4 m (11 ft 2 in) and weighing 182 kg (401 lb) had a bite force of
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water resistance in capturing such aquatic prey. However, as an opportunistic apex predator, virtually any animal living within its range could be considered a potential meal, such as invertebrates, reptiles, birds, and mammals. Despite having a rather elongated
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The Orinoco crocodile is highly endangered due to excessive hunting for its hide. During the 1940s to the 1960s, thousands of these animals were slaughtered in the Orinoco River and the Llanos wetlands, and the species came very close to
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may approach similar dimensions, the Orinoco crocodile may be (or had been, when in healthy numbers) slightly longer. Given its possible maximum sizes, the Orinoco crocodile may rank as the third largest extant true crocodile, after the
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diet. Additionally, as they age, mature males in particular appear to manifest a broadened snout, presumably due to a shift to being able to take larger prey as is seen in several other large crocodile species. It stalks both
3640: 1530:. The Orinoco crocodile was given protected status in the 1970s, but has yet to recover. Today, it is protected both in Colombia and Venezuela, and also included on Appendix I by 1352:, but this has not been confirmed, and witnesses may have mistaken an American crocodile for the rarer species. This crocodile was once thought to have inhabited a wide range of 3676: 3367: 2513:) in Venezuela", pp. 267–285 in Crocodiles: Proceedings of the 5th Working Meeting of the Crocodile Specialist Group, IUCN–The World Conservation Union, Gland, Switzerland. 1521:
A Orinoco crocodile near its wild range in Los Llanos, Venezuela in a captive breeding program intended to bolster the severely depleted population of these crocodiles.
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Hekkala, E.; Gatesy, J.; Narechania, A.; Meredith, R.; Russello, M.; Aardema, M. L.; Jensen, E.; Montanari, S.; Brochu, C.; Norell, M.; Amato, G. (27 April 2021).
2163: 3583: 2089: 383:. Extensively hunted for their skins in the 19th and 20th centuries, it is one of the most endangered species of crocodiles. It is a very large species of 1606:"Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem" 1546:, a smaller crocodilian that can outcompete the Orinoco crocodile for fish due to its much larger population and much more accelerated breeding rates. 3114: 1916:
Erickson, G. M.; Gignac, P. M.; Steppan, S. J.; Lappin, A. K.; Vliet, K. A.; Brueggen, J. A.; Inouye, B. D.; Kledzik, D. & Webb, G. J. W. (2012).
3518: 1565: 2545: 3557: 1316:(which is closely related to it, despite its substantially different range), and additionally rank 4th amongst all extant crocodilians behind the 2529: 3701: 3105: 2408:
Lafferriere, N. A. R.; Antelo, R.; Alda, F.; Mårtensson, D.; Hailer, F.; Castroviejo-Fisher, S.; Doadrio, I.; Vilá, C. & Amato, G. (2016).
1275: 430:, challenging the general view by locals complaining about crocodiles hunting local fish to very low numbers. Reproduction takes place in the 2623: 2147: 1995: 1708: 1549:
It's unclear how many individuals remain in the wild, but estimates range between 250 and 1500. The largest subpopulation in Venezuela is in
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In spite of its somewhat narrow snout, the Orinoco crocodile is both a formidable apex predator and one of the world's largest reptiles.
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The ecology of the Orinoco crocodile is poorly documented in the wild, mostly due to its small population. It is thought to have a more
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The Orinoco crocodile can be recognised by its relatively long snout, which is narrower than that of the somewhat similar-looking
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outwards towards Southeast Asia and the Americas, although an Australia/Asia origin has also been considered.
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Promise for the Survival of the Orinoco Crocodile: a strong commitment from the private sector in Venezuela
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Balaguera-Reina, S.A.; Espinosa-Blanco, A.; Antelo, R.; Morales-Betancourt, M. & Seijas, A. (2020) .
1583:. Thanks to that program, between May 2015 and February 2016, 41 orinoco crocodiles were reintroduced at 3681: 3622: 2884: 2385: 1569: 1432: 1140: 828: 460: 365: 340: 317: 858: 881: 3505: 3456: 3013: 2986: 2977: 2818: 2522: 2484: 2425: 1929: 940: 781: 763: 733: 501: 1083: 3281: 3147: 3031: 2636: 2352:
Thorbjarnarson, J. B., & Hernández, G. (1993). "Reproductive ecology of the Orinoco crocodile (
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Does skull morphology constrain bone ornamentation? A morphometric analysis in the Crocodylia
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Thorbjarnarson, J. B.; Hernández, G. (1993). "Reproductive ecology of the Orinoco crocodile (
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is not as profound as in other crocodilian species. The coloration is light even in adults.
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Clarac, F., Souter, T., Cubo, J., De Buffrénil, V., Brochu, C., & Cornette, R. (2016).
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Crocodile Specialist Group – Status Survey and Conservation Action Plan, 2nd edition.
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Ramo, C.; Busto, B. & Utrera, A. (1992). "Breeding and rearing the Orinoco crocodile
2488: 2429: 1933: 1557:, with fewer than 500 adults remaining. A number of other smaller subpopulations exist. 3377: 3274: 3004: 2800: 2448: 2409: 1952: 1917: 1867: 1838: 1765: 1730: 1634: 1605: 1400: 1313: 1023: 505: 17: 3670: 3570: 3225: 3085: 2827: 2728: 2713: 2496: 1984: 1806: 1789: 1670: 1661: 1453: 1428: 1337: 1261: 1257: 1237: 520: 509: 435: 407: 399: 372: 164: 81: 434:
when the water level is low. It is a hole nester and digs holes in the sand for its
2931: 2916: 2851: 2842: 1823: 1456:, but any details of this attack (including when exactly it happened) are lacking. 1300: 536: 464: 3487: 1906:. IUCN/SSC Crocodile Specialist Group, IUCN, Gland, Switzerland and Cambridge, UK. 1542:
in the upper Orinoco River region. Another problem is the increased population of
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An Orinoco crocodile splashes into the water in the savanna near Mani, Colombia.
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Greer, A. E. (1974). "On the Maximum Total Length of the Salt-Water Crocodile (
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Cohen, M. M.; Gans, C. (1970). "The chromosomes of the order Crocodilia".
1579:, a program launched in 2014 to protect 10 wildlife species of Colombia's 3601: 3426: 3192: 2685: 2679: 2673: 1490: 1353: 1349: 1345: 1288: 1241: 380: 228: 96: 91: 76: 71: 61: 43: 3497: 1624: 3523: 3310: 3201: 2377: 2202: 2039: 1420: 1317: 1252: 1232: 486: 419: 238: 111: 86: 2090:"Critically Endangered Orinoco Crocodiles Coming to Gladys Porter Zoo" 2066: 2018:) in Venezuela. I. Nesting ecology and egg and clutch relationships". 1904:
Crocodiles: Status Survey and Conservation Action Plan. Second Edition
3536: 3469: 3302: 2782: 1580: 1554: 1502: 1494: 1365: 513: 423: 415: 218: 3403: 2369: 2194: 2031: 1568:, of which the largest population, 35 individuals, were kept in the 3648: 2262:"Crocodilian Species - Orinoco Crocodile (Crocodylus intermedius)" 1610: 1531: 1516: 1498: 1463: 1452:
second survivor was reported in 2011 to live in La Palmita at the
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Barrio-Amoros, C.L. (February 2011). "Orinoco Croc Encounters".
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Natural history and conservation of the Orinoco crocodile (
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in the Americas. It is arguably, on average, the largest
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diverging from its closest recent relative, the extinct
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Biología y conservación de los Crocodylia de Colombia
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prey. As a large adult, terrestrial prey may include
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on occasion. Its prey base is mostly large predatory
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Blohm, T. (1982). "Husbandry of Orinoco crocodiles (
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foothills. Today, this species is restricted to the
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IUCN - Species Survival Commission: 8. 1986:The Guinness Book of Animal Facts and Feats 1368:and associated seasonal freshwater rivers. 523:study using DNA extracted from the extinct 138:Orinoco crocodile in Los Llanos, Venezuela 3404: 3289: 3275: 3267: 3187: 3173: 3091: 2936: 2927: 2910: 2795: 2741: 2724: 2707: 2651: 2637: 2629: 2563:"Captives held in zoos registered by ISIS" 2162:: CS1 maint: location missing publisher ( 516:age) data, as revised by the 2021 Hekkala 339: 153: 129: 31: 2447: 2437: 1951: 1941: 1866: 1856: 1805: 1764: 1746: 1633: 1623: 3106:West African slender snouted crocodile ( 1604:Rio, J. P. & Mannion, P. D. (2021). 481:, around 25 million years ago, near the 1703:. Oxford University Press. p. 76. 1700:The Amazon: What Everyone Needs to Know 1596: 1394:, its base snout is wide, hinting to a 2383: 2278:. Journal of Anatomy, 229(2), 292-301. 2155: 1977: 1975: 1973: 1971: 1898: 1896: 1894: 1892: 1890: 1888: 1886: 1655: 1653: 1283:The Orinoco crocodile ranks among the 2225:Crocodile Specialist Group Newsletter 2116:Crocodile Specialist Group Newsletter 1575:The Orinoco crocodile became part of 1435:smaller individuals of its own kind. 7: 2216:Whitaker, R.; Whitaker, N. (2008). 1671:IUCN Red List of Threatened Species 1108: 1051: 1044: 1013: 988: 932: 925: 844: 814: 753: 723: 716: 708: 644: 588: 581: 574: 542: 532: 1844:Proceedings of the Royal Society B 1336:This species is restricted to the 25: 3388:Famous crocodiles and alligators 3250: 2342:, Northern Territory, Australia. 2325:, Northern Territory, Australia. 1807:10.1111/j.1558-5646.2011.01373.x 1585:El Tuparro National Natural Park 1139: 1082: 1026: 880: 857: 827: 736: 459:likely originated in Africa and 195: 2336:Orinoco crocodile, 15 June 2009 2250:. Madrid: Asociación Chelonia. 406:predatory behaviour. It is an 1: 3702:Crocodilians of South America 2960:Hall's New Guinea crocodile ( 442:and young for several years. 2497:10.1016/0006-3207(92)91160-T 2439:10.1371/journal.pone.0150245 1943:10.1371/journal.pone.0031781 3345:Madras Crocodile Bank Trust 2107:Alvaro Velasco, B. (2023). 3723: 3697:Reptiles described in 1819 3340:List of crurotarsan genera 3148:Osborn’s dwarf crocodile ( 2088:Gladys Porter Zoo (2011). 1748:10.1038/s42003-021-02017-0 1472:When water recedes in the 617:Crocodylus thorbjarnarsoni 410:and preys on a variety of 3246: 3186: 3172: 2926: 2909: 2723: 2706: 2668: 2340:Charles Darwin University 2323:Charles Darwin University 2248:Graves, 1819) in Colombia 2133:Carlos A. Lasso. (2013). 1697:Plotkin, Mark J. (2020). 1287:, as well as the largest 1131: 1113: 1106: 1074: 1056: 1049: 1042: 1018: 1011: 993: 986: 957: 937: 930: 923: 872: 849: 842: 819: 812: 776: 758: 751: 728: 721: 714: 706: 669: 649: 642: 613: 597:Crocodylus anthropophagus 593: 586: 579: 572: 547: 540: 347: 338: 323: 316: 192:Scientific classification 190: 173: 151: 142: 137: 128: 34: 3068:West African crocodile ( 2717:(Alligators and caimans) 2218:"Who's got the biggest?" 1538:, and the proposal of a 1468:Orinoco crocodile's head 1332:Distribution and habitat 438:. The females guard the 3368:Fatal alligator attacks 3117:Mecistops leptorhynchus 2885:Smooth-fronted caiman ( 2876:Cuvier's dwarf caiman ( 2477:Biological Conservation 1990:. Sterling Pub Co Inc. 1788:Oaks, Jamie R. (2011). 1577:Proyecto Vida Silvestre 1483:American black vultures 1295:in the Americas; while 1285:largest living reptiles 761:Crocodylus novaeguineae 653:Crocodylus palaeindicus 3475:crocodylus-intermedius 3462:Crocodylus_intermedius 3448:Crocodylus intermedius 3418:Crocodylus intermedius 3108:Mecistops cataphractus 3014:New Guinea crocodile ( 2987:Philippine crocodile ( 2978:Freshwater crocodile ( 2819:Broad-snouted caiman ( 2548:Crocodylus intermedius 2525:Crocodylus intermedius 2511:Crocodylus intermedius 2473:Crocodylus intermedius 2412:Crocodylus intermedius 2358:Journal of Herpetology 2354:Crocodylus intermedius 2246:Crocodylus intermedius 2183:Journal of Herpetology 2020:Journal of Herpetology 2016:Crocodylus intermedius 1858:10.1098/rspb.2018.1071 1735:Communications Biology 1664:Crocodylus intermedius 1522: 1469: 1449:Alexander von Humboldt 1443: 1386: 1360:to the streams of the 1280: 1244: 1116:Crocodylus intermedius 999:West African crocodile 961:Crocodylus falconensis 921:Africa+New World 779:Crocodylus mindorensis 361:Crocodylus intermedius 327:Crocodylus intermedius 18:Crocodylus intermedius 3692:Reptiles of Venezuela 3623:Paleobiology Database 3032:Saltwater crocodile ( 2996:Morelet's crocodile ( 1587:in eastern Colombia. 1570:Dallas World Aquarium 1520: 1467: 1441: 1384: 1278: 1236:Orinoco crocodile in 1235: 402:nature, resulting in 366:critically endangered 160:Critically Endangered 3687:Reptiles of Colombia 3297:Related articles on 2951:American crocodile ( 2756:American alligator ( 2242:Merchán, M. (2011). 1982:Wood, G. L. (1983). 1372:Behavior and ecology 1077:Crocodylus rhombifer 1059:Crocodylus moreletii 1021:Crocodylus niloticus 941:Crocodylus checchiai 875:Crocodylus palustris 852:Crocodylus siamensis 782:Philippine crocodile 764:New Guinea crocodile 734:Freshwater crocodile 731:Crocodylus johnstoni 3257:Reptiles portal 3059:Siamese crocodile ( 2969:Orinoco crocodile ( 2810:Spectacled caiman ( 2765:Chinese alligator ( 2758:A. mississippiensis 2489:1992BCons..60..101R 2430:2016PLoSO..1150245R 2288:October 1984. IUCN. 1934:2012PLoSO...731781E 1678:: e.T5661A181089024 1625:10.7717/peerj.12094 1513:Conservation status 1310:saltwater crocodile 1297:American crocodiles 1062:Morelet's crocodile 825:Saltwater crocodile 309:C. intermedius 145:Conservation status 3330:Foramen of Panizza 3041:Borneo crocodile ( 3023:Mugger crocodile ( 2597:, 25 December 2013 2594:The New York Times 2569:on 9 February 2008 2532:1 May 2006 at the 2179:Crocodylus porosus 1902:Ross, R.P. (1998) 1544:spectacled caimans 1523: 1470: 1444: 1387: 1305:American alligator 1281: 1249:American crocodile 1245: 1137:American crocodile 822:Crocodylus porosus 467:evidence supports 27:Species of reptile 3664: 3663: 3610:Open Tree of Life 3410:Taxon identifiers 3401: 3400: 3373:Crocodile attacks 3350:The Croc Festival 3325:Crocodilian armor 3264: 3263: 3242: 3241: 3238: 3237: 3168: 3167: 3164: 3163: 3160: 3159: 3139:Dwarf crocodile ( 3080: 3079: 3050:Cuban crocodile ( 2921: 2920:(True crocodiles) 2905: 2904: 2901: 2900: 2897: 2896: 2792: 2777: 2776: 2738: 2718: 2319:Orinoco crocodile 2305:Reptiles Magazine 2149:978-958-8343-87-7 2142:. . p. 113. 2067:10.1159/000130080 1997:978-0-85112-235-9 1800:(11): 3285–3297. 1710:978-0-19-066829-7 1224: 1223: 1215: 1214: 1206: 1205: 1197: 1196: 1188: 1187: 1179: 1178: 1170: 1169: 1161: 1160: 1152: 1151: 1134:Crocodylus acutus 1119:Orinoco crocodile 1095: 1094: 996:Crocodylus suchus 975: 974: 911: 910: 902: 901: 893: 892: 855:Siamese crocodile 801: 800: 792: 791: 695: 694: 686: 685: 631: 630: 389:Sexual dimorphism 356:Orinoco crocodile 352: 351: 185: 168: 35:Orinoco crocodile 16:(Redirected from 3714: 3657: 3656: 3644: 3643: 3631: 3630: 3618: 3617: 3605: 3604: 3592: 3591: 3579: 3578: 3566: 3565: 3553: 3552: 3540: 3539: 3527: 3526: 3514: 3513: 3501: 3500: 3491: 3490: 3478: 3477: 3465: 3464: 3452: 3451: 3450: 3437: 3436: 3435: 3405: 3291: 3284: 3277: 3268: 3255: 3254: 3253: 3188: 3174: 3092: 3005:Nile crocodile ( 2937: 2928: 2919: 2911: 2796: 2791: 2790: 2786: 2742: 2737: 2736: 2732: 2725: 2716: 2708: 2653: 2646: 2639: 2630: 2611: 2605: 2599: 2598: 2585: 2579: 2578: 2576: 2574: 2565:. Archived from 2559: 2553: 2543: 2537: 2520: 2514: 2507: 2501: 2500: 2468: 2462: 2461: 2451: 2441: 2405: 2396: 2395: 2389: 2381: 2349: 2343: 2332: 2326: 2315: 2309: 2308: 2300: 2289: 2285: 2279: 2272: 2266: 2265: 2258: 2252: 2251: 2239: 2233: 2232: 2222: 2213: 2207: 2206: 2174: 2168: 2167: 2161: 2153: 2141: 2130: 2124: 2123: 2113: 2104: 2098: 2097: 2085: 2079: 2078: 2050: 2044: 2043: 2011: 2002: 2001: 1989: 1979: 1966: 1965: 1955: 1945: 1913: 1907: 1900: 1881: 1880: 1870: 1860: 1834: 1828: 1827: 1809: 1785: 1779: 1778: 1768: 1750: 1726: 1715: 1714: 1694: 1688: 1687: 1685: 1683: 1657: 1648: 1647: 1637: 1627: 1601: 1425:domestic animals 1377:Hunting and diet 1358:tropical forests 1322:Guinness Records 1143: 1109: 1086: 1052: 1045: 1030: 1014: 989: 966: 946: 933: 926: 884: 878:Mugger crocodile 861: 845: 831: 815: 754: 740: 724: 717: 709: 677: 658: 645: 622: 602: 589: 582: 575: 556: 543: 533: 496:based on a 2018 343: 329: 265:Archosauriformes 252:Archosauromorpha 200: 199: 179: 162: 157: 156: 133: 123: 53: 40:Late Pleistocene 38:Temporal range: 32: 21: 3722: 3721: 3717: 3716: 3715: 3713: 3712: 3711: 3667: 3666: 3665: 3660: 3652: 3647: 3639: 3634: 3626: 3621: 3613: 3608: 3600: 3597:Observation.org 3595: 3587: 3582: 3574: 3569: 3561: 3556: 3548: 3543: 3535: 3530: 3522: 3517: 3509: 3504: 3496: 3494: 3486: 3481: 3473: 3468: 3460: 3455: 3446: 3445: 3440: 3431: 3430: 3425: 3412: 3402: 3397: 3393:Sewer alligator 3383:Crocodile tears 3360: 3354: 3313: 3295: 3265: 3260: 3251: 3249: 3234: 3226:False gharial ( 3210: 3182: 3156: 3123: 3088: 3076: 3016:C. novaeguineae 2922: 2893: 2860: 2836: 2828:Yacare caiman ( 2788: 2787: 2785: 2773: 2734: 2733: 2731: 2719: 2702: 2698:Crocodylomorpha 2664: 2657: 2620: 2615: 2614: 2606: 2602: 2587: 2586: 2582: 2572: 2570: 2561: 2560: 2556: 2544: 2540: 2534:Wayback Machine 2523:Information on 2521: 2517: 2508: 2504: 2475:in Venezuela". 2470: 2469: 2465: 2424:(3): e0150245. 2407: 2406: 2399: 2382: 2370:10.2307/1564822 2351: 2350: 2346: 2333: 2329: 2316: 2312: 2302: 2301: 2292: 2286: 2282: 2273: 2269: 2260: 2259: 2255: 2241: 2240: 2236: 2220: 2215: 2214: 2210: 2195:10.2307/1562913 2176: 2175: 2171: 2154: 2150: 2139: 2132: 2131: 2127: 2111: 2106: 2105: 2101: 2087: 2086: 2082: 2052: 2051: 2047: 2032:10.2307/1564821 2013: 2012: 2005: 1998: 1981: 1980: 1969: 1915: 1914: 1910: 1901: 1884: 1836: 1835: 1831: 1787: 1786: 1782: 1728: 1727: 1718: 1711: 1696: 1695: 1691: 1681: 1679: 1659: 1658: 1651: 1603: 1602: 1598: 1593: 1515: 1462: 1379: 1374: 1356:habitats, from 1334: 1273: 1265: 1230: 1228:Characteristics 1225: 1216: 1207: 1198: 1189: 1180: 1171: 1162: 1153: 1096: 1080:Cuban crocodile 1040:New World 976: 912: 903: 894: 802: 793: 712:Asia+Australia 696: 687: 632: 448: 334: 331: 325: 312: 194: 186: 169: 158: 154: 147: 124: 122: 121: 120: 119: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 48: 47: 36: 28: 23: 22: 15: 12: 11: 5: 3720: 3718: 3710: 3709: 3707:Apex predators 3704: 3699: 3694: 3689: 3684: 3679: 3669: 3668: 3662: 3661: 3659: 3658: 3645: 3632: 3619: 3606: 3593: 3580: 3567: 3554: 3541: 3528: 3515: 3502: 3492: 3479: 3466: 3453: 3438: 3422: 3420: 3414: 3413: 3408: 3399: 3398: 3396: 3395: 3390: 3385: 3380: 3378:Crocodile farm 3375: 3370: 3364: 3362: 3356: 3355: 3353: 3352: 3347: 3342: 3337: 3332: 3327: 3321: 3319: 3315: 3314: 3296: 3294: 3293: 3286: 3279: 3271: 3262: 3261: 3247: 3244: 3243: 3240: 3239: 3236: 3235: 3233: 3232: 3222: 3220: 3212: 3211: 3209: 3208: 3198: 3196: 3184: 3183: 3177: 3170: 3169: 3166: 3165: 3162: 3161: 3158: 3157: 3155: 3154: 3145: 3135: 3133: 3125: 3124: 3122: 3121: 3112: 3102: 3100: 3089: 3082: 3081: 3078: 3077: 3075: 3074: 3065: 3056: 3047: 3038: 3029: 3020: 3011: 3002: 2993: 2989:C. mindorensis 2984: 2975: 2971:C. intermedius 2966: 2957: 2947: 2945: 2934: 2924: 2923: 2914: 2907: 2906: 2903: 2902: 2899: 2898: 2895: 2894: 2892: 2891: 2882: 2878:P. palpebrosus 2872: 2870: 2862: 2861: 2859: 2858: 2852:Black caiman ( 2848: 2846: 2838: 2837: 2835: 2834: 2825: 2821:C. latirostris 2816: 2806: 2804: 2793: 2779: 2778: 2775: 2774: 2772: 2771: 2762: 2752: 2750: 2739: 2721: 2720: 2711: 2704: 2703: 2701: 2700: 2694: 2688: 2682: 2676: 2669: 2666: 2665: 2658: 2656: 2655: 2648: 2641: 2633: 2627: 2626: 2619: 2618:External links 2616: 2613: 2612: 2600: 2580: 2554: 2538: 2515: 2502: 2483:(2): 101–108. 2463: 2397: 2364:(4): 371–379. 2344: 2327: 2310: 2290: 2280: 2267: 2253: 2234: 2208: 2189:(4): 381–384. 2169: 2148: 2125: 2099: 2094:prnewswire.com 2080: 2045: 2026:(4): 363–370. 2003: 1996: 1967: 1908: 1882: 1829: 1780: 1716: 1709: 1689: 1649: 1595: 1594: 1592: 1589: 1564:registered by 1514: 1511: 1461: 1458: 1431:and sometimes 1429:common caimans 1378: 1375: 1373: 1370: 1366:Llanos savanna 1333: 1330: 1314:Nile crocodile 1272: 1269: 1263: 1229: 1226: 1222: 1221: 1218: 1217: 1213: 1212: 1209: 1208: 1204: 1203: 1200: 1199: 1195: 1194: 1191: 1190: 1186: 1185: 1182: 1181: 1177: 1176: 1173: 1172: 1168: 1167: 1164: 1163: 1159: 1158: 1155: 1154: 1150: 1149: 1146: 1145: 1130: 1127: 1126: 1123: 1122: 1112: 1107: 1105: 1102: 1101: 1098: 1097: 1093: 1092: 1089: 1088: 1073: 1070: 1069: 1066: 1065: 1055: 1050: 1048: 1043: 1041: 1037: 1036: 1033: 1032: 1024:Nile crocodile 1017: 1012: 1010: 1007: 1006: 1003: 1002: 992: 987: 985: 982: 981: 978: 977: 973: 972: 969: 968: 956: 953: 952: 949: 948: 936: 931: 929: 924: 922: 918: 917: 914: 913: 909: 908: 905: 904: 900: 899: 896: 895: 891: 890: 887: 886: 871: 868: 867: 864: 863: 848: 843: 841: 838: 837: 834: 833: 818: 813: 811: 808: 807: 804: 803: 799: 798: 795: 794: 790: 789: 786: 785: 775: 772: 771: 768: 767: 757: 752: 750: 747: 746: 743: 742: 727: 722: 720: 715: 713: 707: 705: 702: 701: 698: 697: 693: 692: 689: 688: 684: 683: 680: 679: 668: 665: 664: 661: 660: 648: 643: 641: 638: 637: 634: 633: 629: 628: 625: 624: 612: 609: 608: 605: 604: 592: 587: 585: 580: 578: 573: 571: 563: 562: 559: 558: 546: 541: 539: 531: 506:DNA sequencing 447: 444: 436:clutch of eggs 350: 349: 348:Range (green) 345: 344: 336: 335: 332: 321: 320: 314: 313: 306: 304: 300: 299: 292: 288: 287: 282: 278: 277: 272: 268: 267: 262: 255: 254: 249: 242: 241: 236: 232: 231: 226: 222: 221: 216: 212: 211: 206: 202: 201: 188: 187: 174: 171: 170: 152: 149: 148: 143: 140: 139: 135: 134: 126: 125: 117: 116: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 54: 37: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 3719: 3708: 3705: 3703: 3700: 3698: 3695: 3693: 3690: 3688: 3685: 3683: 3680: 3678: 3675: 3674: 3672: 3655: 3650: 3646: 3642: 3637: 3633: 3629: 3624: 3620: 3616: 3611: 3607: 3603: 3598: 3594: 3590: 3585: 3581: 3577: 3572: 3568: 3564: 3559: 3555: 3551: 3546: 3542: 3538: 3533: 3529: 3525: 3520: 3516: 3512: 3507: 3503: 3499: 3493: 3489: 3484: 3480: 3476: 3471: 3467: 3463: 3458: 3454: 3449: 3443: 3439: 3434: 3428: 3424: 3423: 3421: 3419: 3415: 3411: 3406: 3394: 3391: 3389: 3386: 3384: 3381: 3379: 3376: 3374: 3371: 3369: 3366: 3365: 3363: 3357: 3351: 3348: 3346: 3343: 3341: 3338: 3336: 3333: 3331: 3328: 3326: 3323: 3322: 3320: 3316: 3312: 3308: 3304: 3300: 3292: 3287: 3285: 3280: 3278: 3273: 3272: 3269: 3259: 3258: 3245: 3231: 3229: 3228:T. schlegelii 3224: 3223: 3221: 3219: 3218: 3213: 3207: 3205: 3204:G. gangeticus 3200: 3199: 3197: 3195: 3194: 3189: 3185: 3181: 3175: 3171: 3153: 3151: 3146: 3144: 3142: 3137: 3136: 3134: 3132: 3131: 3126: 3120: 3118: 3113: 3111: 3109: 3104: 3103: 3101: 3099: 3098: 3093: 3090: 3087: 3086:Osteolaeminae 3083: 3073: 3071: 3066: 3064: 3062: 3057: 3055: 3053: 3048: 3046: 3044: 3039: 3037: 3035: 3030: 3028: 3026: 3021: 3019: 3017: 3012: 3010: 3008: 3003: 3001: 2999: 2994: 2992: 2990: 2985: 2983: 2981: 2976: 2974: 2972: 2967: 2965: 2963: 2958: 2956: 2954: 2949: 2948: 2946: 2944: 2943: 2938: 2935: 2933: 2929: 2925: 2918: 2912: 2908: 2890: 2888: 2887:P. trigonatus 2883: 2881: 2879: 2874: 2873: 2871: 2869: 2868: 2863: 2857: 2855: 2850: 2849: 2847: 2845: 2844: 2839: 2833: 2831: 2826: 2824: 2822: 2817: 2815: 2813: 2812:C. crocodilus 2808: 2807: 2805: 2803: 2802: 2797: 2794: 2784: 2780: 2770: 2768: 2763: 2761: 2759: 2754: 2753: 2751: 2749: 2748: 2743: 2740: 2730: 2729:Alligatorinae 2726: 2722: 2715: 2714:Alligatoridae 2709: 2705: 2699: 2695: 2693: 2689: 2687: 2683: 2681: 2677: 2675: 2671: 2670: 2667: 2662: 2654: 2649: 2647: 2642: 2640: 2635: 2634: 2631: 2625: 2622: 2621: 2617: 2609: 2604: 2601: 2596: 2595: 2590: 2584: 2581: 2568: 2564: 2558: 2555: 2551: 2549: 2542: 2539: 2535: 2531: 2528: 2526: 2519: 2516: 2512: 2506: 2503: 2498: 2494: 2490: 2486: 2482: 2478: 2474: 2467: 2464: 2459: 2455: 2450: 2445: 2440: 2435: 2431: 2427: 2423: 2419: 2415: 2413: 2404: 2402: 2398: 2393: 2387: 2379: 2375: 2371: 2367: 2363: 2359: 2355: 2348: 2345: 2341: 2337: 2331: 2328: 2324: 2320: 2314: 2311: 2306: 2299: 2297: 2295: 2291: 2284: 2281: 2277: 2271: 2268: 2263: 2257: 2254: 2249: 2245: 2238: 2235: 2230: 2226: 2219: 2212: 2209: 2204: 2200: 2196: 2192: 2188: 2184: 2180: 2173: 2170: 2165: 2159: 2151: 2145: 2138: 2137: 2129: 2126: 2121: 2117: 2110: 2103: 2100: 2095: 2091: 2084: 2081: 2076: 2072: 2068: 2064: 2061:(2): 81–105. 2060: 2056: 2049: 2046: 2041: 2037: 2033: 2029: 2025: 2021: 2017: 2010: 2008: 2004: 1999: 1993: 1988: 1987: 1978: 1976: 1974: 1972: 1968: 1963: 1959: 1954: 1949: 1944: 1939: 1935: 1931: 1928:(3): e31781. 1927: 1923: 1919: 1912: 1909: 1905: 1899: 1897: 1895: 1893: 1891: 1889: 1887: 1883: 1878: 1874: 1869: 1864: 1859: 1854: 1850: 1846: 1845: 1840: 1833: 1830: 1825: 1821: 1817: 1813: 1808: 1803: 1799: 1795: 1791: 1784: 1781: 1776: 1772: 1767: 1762: 1758: 1754: 1749: 1744: 1740: 1736: 1732: 1725: 1723: 1721: 1717: 1712: 1706: 1702: 1701: 1693: 1690: 1677: 1673: 1672: 1667: 1665: 1656: 1654: 1650: 1645: 1641: 1636: 1631: 1626: 1621: 1617: 1613: 1612: 1607: 1600: 1597: 1590: 1588: 1586: 1582: 1578: 1573: 1571: 1567: 1563: 1558: 1556: 1552: 1547: 1545: 1541: 1537: 1533: 1529: 1519: 1512: 1510: 1508: 1504: 1500: 1496: 1492: 1488: 1484: 1479: 1475: 1466: 1459: 1457: 1455: 1454:Cojedes River 1450: 1440: 1436: 1434: 1433:cannibalizing 1430: 1426: 1422: 1418: 1414: 1410: 1406: 1402: 1397: 1393: 1383: 1376: 1371: 1369: 1367: 1363: 1359: 1355: 1351: 1347: 1343: 1339: 1338:Orinoco river 1331: 1329: 1327: 1323: 1319: 1315: 1311: 1306: 1302: 1301:black caimans 1298: 1294: 1290: 1286: 1277: 1270: 1268: 1266: 1259: 1254: 1250: 1243: 1239: 1238:Villavicencio 1234: 1227: 1220: 1219: 1211: 1210: 1202: 1201: 1193: 1192: 1184: 1183: 1175: 1174: 1166: 1165: 1157: 1156: 1148: 1147: 1144: 1142: 1138: 1135: 1129: 1128: 1125: 1124: 1121: 1120: 1117: 1111: 1110: 1104: 1103: 1100: 1099: 1091: 1090: 1087: 1085: 1081: 1078: 1072: 1071: 1068: 1067: 1064: 1063: 1060: 1054: 1053: 1047: 1046: 1039: 1038: 1035: 1034: 1031: 1029: 1025: 1022: 1016: 1015: 1009: 1008: 1005: 1004: 1001: 1000: 997: 991: 990: 984: 983: 980: 979: 971: 970: 967: 963: 962: 955: 954: 951: 950: 947: 943: 942: 935: 934: 928: 927: 920: 919: 916: 915: 907: 906: 898: 897: 889: 888: 885: 883: 879: 876: 870: 869: 866: 865: 862: 860: 856: 853: 847: 846: 840: 839: 836: 835: 832: 830: 826: 823: 817: 816: 810: 809: 806: 805: 797: 796: 788: 787: 784: 783: 780: 774: 773: 770: 769: 766: 765: 762: 756: 755: 749: 748: 745: 744: 741: 739: 735: 732: 726: 725: 719: 718: 711: 710: 704: 703: 700: 699: 691: 690: 682: 681: 678: 674:Tirari Desert 673: 667: 666: 663: 662: 659: 655: 654: 647: 646: 640: 639: 636: 635: 627: 626: 623: 619: 618: 611: 610: 607: 606: 603: 599: 598: 591: 590: 584: 583: 577: 576: 570: 569: 565: 564: 561: 560: 557: 553: 552: 545: 544: 538: 535: 534: 530: 528: 527: 522: 521:paleogenomics 519: 515: 511: 510:stratigraphic 507: 504:, molecular ( 503: 502:morphological 499: 495: 490: 488: 484: 480: 476: 475: 470: 466: 462: 458: 457: 453: 445: 443: 441: 437: 433: 429: 425: 421: 417: 413: 409: 408:apex predator 405: 401: 400:opportunistic 398:diet with an 397: 392: 390: 386: 382: 378: 374: 373:Orinoco river 370: 367: 363: 362: 357: 346: 342: 337: 330: 328: 322: 319: 318:Binomial name 315: 311: 310: 305: 302: 301: 298: 297: 293: 290: 289: 286: 283: 280: 279: 276: 273: 270: 269: 266: 263: 260: 257: 256: 253: 250: 247: 244: 243: 240: 237: 234: 233: 230: 227: 224: 223: 220: 217: 214: 213: 210: 207: 204: 203: 198: 193: 189: 183: 177: 172: 166: 161: 150: 146: 141: 136: 132: 127: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 52: 45: 41: 33: 30: 19: 3682:Crocodylidae 3417: 3248: 3227: 3215: 3203: 3191: 3149: 3141:O. tetraspis 3140: 3128: 3116: 3107: 3095: 3069: 3061:C. siamensis 3060: 3052:C. rhombifer 3051: 3042: 3033: 3025:C. palustris 3024: 3015: 3007:C. niloticus 3006: 2998:C. moreletii 2997: 2988: 2980:C. johnstoni 2979: 2970: 2968: 2961: 2952: 2940: 2932:Crocodylinae 2917:Crocodylidae 2886: 2877: 2865: 2853: 2843:Melanosuchus 2841: 2829: 2820: 2811: 2799: 2766: 2757: 2745: 2735:(Alligators) 2696:Superorder: 2690:(unranked): 2603: 2592: 2583: 2571:. Retrieved 2567:the original 2557: 2547: 2541: 2524: 2518: 2510: 2505: 2480: 2476: 2472: 2466: 2421: 2417: 2411: 2386:cite journal 2361: 2357: 2353: 2347: 2330: 2313: 2304: 2283: 2275: 2270: 2256: 2247: 2243: 2237: 2228: 2224: 2211: 2186: 2182: 2178: 2172: 2135: 2128: 2119: 2115: 2102: 2093: 2083: 2058: 2055:Cytogenetics 2054: 2048: 2023: 2019: 2015: 1985: 1925: 1921: 1911: 1903: 1848: 1842: 1832: 1797: 1793: 1783: 1738: 1734: 1699: 1692: 1680:. Retrieved 1675: 1669: 1663: 1615: 1609: 1599: 1576: 1574: 1559: 1548: 1524: 1505:, and other 1487:tegu lizards 1478:tegu lizards 1471: 1460:Reproduction 1445: 1388: 1335: 1282: 1260:(1,411  1246: 1133: 1132: 1118: 1115: 1114: 1076: 1075: 1058: 1057: 1020: 1019: 995: 994: 959: 958: 939: 938: 874: 873: 851: 850: 821: 820: 778: 777: 760: 759: 730: 729: 671: 670: 651: 650: 615: 614: 595: 594: 566: 549: 548: 537:Crocodylinae 524: 517: 491: 472: 468: 465:Phylogenetic 454: 449: 422:, including 393: 360: 359: 355: 353: 333:Graves, 1819 326: 324: 308: 307: 295: 285:Crocodylidae 258: 245: 29: 3641:intermedius 3532:iNaturalist 3442:Wikispecies 3361:interaction 3130:Osteolaemus 2867:Paleosuchus 2767:A. sinensis 2692:Archosauria 2661:Crocodilian 2231:(4): 26−30. 1405:terrestrial 1293:crocodilian 1256:6,276  492:Below is a 396:piscivorous 385:crocodilian 49:0.1–0  3671:Categories 3335:Gastralium 3307:crocodiles 3299:alligators 3180:Gavialidae 3150:O. osborni 3043:C. raninus 3034:C. porosus 2942:Crocodylus 2783:Caimaninae 2573:9 November 1741:(1): 505. 1682:22 January 1618:: e12094. 1591:References 1566:Species360 1528:extinction 1507:carnivores 1474:dry season 1396:generalist 672:Crocodylus 568:Crocodylus 498:tip dating 489:boundary. 479:Madagascar 469:Crocodylus 456:Crocodylus 432:dry season 404:generalist 296:Crocodylus 275:Crocodilia 178:Appendix I 3217:Tomistoma 3202:Gharial ( 3097:Mecistops 3070:C. suchus 2953:C. acutus 2830:C. yacare 2789:(Caimans) 2747:Alligator 2672:Kingdom: 2546:IUCN/SSC 2158:cite book 1794:Evolution 1757:2399-3642 1536:pollution 1491:anacondas 1342:Venezuela 1340:basin in 1326:Venezuela 494:cladogram 483:Oligocene 377:Venezuela 375:basin in 369:crocodile 303:Species: 215:Kingdom: 209:Eukaryota 3649:Species+ 3550:10368646 3427:Wikidata 3311:gharials 3193:Gavialis 2962:C. halli 2854:M. niger 2686:Reptilia 2680:Chordata 2678:Phylum: 2674:Animalia 2530:Archived 2458:26982578 2418:PLOS ONE 2307:: 48–53. 1962:22431965 1922:PLOS ONE 1877:30051855 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Index

Crocodylus intermedius
Late Pleistocene
Present
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Conservation status
Critically Endangered
IUCN 3.1
CITES
CITES
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Archosauromorpha
Archosauriformes

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