Knowledge (XXG)

Dwarf pufferfish

Source πŸ“

45: 1241: 1660:, or on the substrate hidden within plants. A female will scatter approximately 1–5 eggs, 1.43 mm (0.056 in) in mean diameter. The eggs are adhesive and appear transparent and round, with a mass of small oil globules. After laying, the eggs are then fertilized externally by the male. Spawning has been observed in the evening, with the female resting on the spawning site and the male slowly approaching. After spawning, both fish will leave the site. The male will then quickly return to guard the eggs. Sneaking ejaculation by other males has also been observed. The pair may spawn multiple times in 1–4 day intervals. 1648: 1577: 1386: 89: 479: 64: 2401: 1655:
The mid-dorsal and mid-ventral skin ridges of males become brown in colour during the spawning season. The belly of the female will swell, and the courting male will frequently chase the female and nibble at its belly. The female will then search for a suitable location to spawn while the male chases
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surface. Males can also have a dark stripe down the centre of their pale belly and iridescent, blue "eye wrinkle" patterns that females do not have. Females are more rounded, tend to be larger than males, and may have more small spots between their larger dark markings. Their abdomens are white, and
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Pagnussat, NatΓ‘lia; Piato, Angelo L.; Schaefer, Isabel C.; Blank, Martina; Tamborski, AngΓ©lica R.; Guerim, Laura D.; Bonan, Carla D.; Vianna, MΓ΄nica R.M.; Lara, Diogo R. (September 2013). "One for All and All for One: The Importance of Shoaling on Behavioral and Stress Responses in Zebrafish".
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The maximum documented size is 3.5 cm (1.4 in) total length (TL), with individuals typically reaching less than 2.5 cm (0.98 in) TL, making dwarf pufferfish one of the smallest pufferfish in the world. Both sexes are primarily greenish-yellow, with dark green to brown-black
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is larger than the other fins and truncate, in that it terminates in a more-or-less vertical edge. Their bodies are round and oblong. As with other members of the genus, sexual dimorphism is apparent in mature fish, with males being more brightly coloured than females and having a yellow
1677:(a genus of rotifers), frozen bloodworms, and brine shrimp when they are a week old. There is little information on what the larvae eat in the wild. Hatched larvae are a mean of 3.15 mm (0.124 in) total length, with eyes incompletely developed and the body a red-brown. The 1696:
When first described in 1941, K. Nair noted that dwarf pufferfish were a favourite of children, who would catch and use the fish as playthings. They were regarded as "frog tadpoles" by local fishermen and otherwise given little consideration. They are of no interest to
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Although closely related to marine pufferfish, they are not found in salt water, and reports to the contrary are based on misidentification. Dwarf pufferfish are one of only 27 known species of Tetraodontidae adapted to freshwater. They are a
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period. Within the Pamba River, the minimum size at which half the population becomes sexually mature is approximately 1.83 cm (0.72 in). Environmental and dietary conditions may influence the maturation rate of individuals.
467:. A polyphyletic genus is one in which members share common characteristics, but do not necessarily share an immediate common ancestor, and their placement is based on these shared traits rather than confirmed, empirical evidence. " 1363:
was mistaken as the same species, and the two could be found together in aquaria and were widely available in the international aquarium trade, both sold as dwarf pufferfish. Dwarf pufferfish can be distinguished from their
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for the aquarium trade. In 2010, some researchers estimated that the population would decline by 30–40% from 2005 to 2015, while others estimated it had already declined by the same amount between 2005 and 2010.
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have smaller, faint blotches compared to dwarf pufferfish, as well as greatly reduced body spination. Male dwarf pufferfish have a darker yellow colouration and an iridescent "eye wrinkle" not found in males of
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Raghavan, R.; Prasad, G.; Ali, A.P.H.; Pereira, B. (2008). "Fish fauna of Chalakudy River, part of Western Ghats biodiversity hotspot, Kerala, India: patterns of distribution, threats and conservation needs".
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Raghavan, R.; Prasad, G.; Ali, A.P.H.; Pereira, B. (2008). "Fish fauna of Chalakudy River, part of Western Ghats biodiversity hotspot, Kerala, India: patterns of distribution, threats and conservation needs".
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small animals. Unlike other species of pufferfish, they are found in large groups in the wild. They breed throughout most of the year, with spawning pairs producing 1–5 eggs in 1–4 day intervals.
1713:", and relative ease of maintenance. The dwarf pufferfish is also one of the few aquarium fish to regularly eat small, live snails and thus can be helpful in controlling snail populations. 1355:
is a species of related pufferfish closely resembling dwarf pufferfish. They are of similar size, shape, patterns, and colouration. Both species are found within the same region and may be
1335:. The pufferfish's ability to inflate makes it difficult to swallow or bite, and the tetrodotoxin makes it unpalatable or even deadly for other fish. Biologists believe these adaptations 3976: 2790:
Doi, Hiroyuki; Sakai, Harumi; Yamanoue, Yusuke; Sonoyama, Takayuki; Ishibashi, Toshiaki (March 2015). "Spawning of eight Southeast Asian brackish and freshwater puffers of the genera
1623:, occasionally consisting of hundreds of individuals. They are found mainly during the summer months of January to May and rarely during the rainy season. Shoaling fish experience 3046: 2473: 437:, which other members of the genus have, but which had not been confirmed in dwarf pufferfish until one year later in aquarium literature. Specifically, dwarf pufferfish possess 471:" is represented in quotation marks to signify its polyphyletic status. Freshwater species are denoted with "FW". The cladogram is incomplete due to the absence of species like 3251: 2910: 3843: 3882: 3991: 3971: 3981: 3961: 3433: 44: 1639:
have been found during all months of the year but December and January, with the peak spawning period extending from May to August, concurring with the
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All pufferfish can inflate, or "puff up", their bodies by quickly ingesting large amounts of water (or air when necessary) into their highly elastic
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Sathyajith, C.; Yamanoue, Y.; Yokobori, S.; Thampy, S.; Vattiringal Jayadradhan, R.K. (December 2019). "Mitogenome analysis of dwarf pufferfish (
3655: 3986: 3956: 2846: 2233: 2179: 1929: 1615:(bottom-dwelling) species. Unlike many pufferfish species, which are primarily solitary and potentially aggressive or territorial between 1573:. Sand and detritus, presumably ingested by mistake when feeding on small, bottom-dwelling animals, have also been found in their gut. 1533:
as alternate feed when the availability of preferable prey decreases. Their diet in the wild mainly consists of small animals such as
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larvae and other small animals they would predate in the wild. Dwarf pufferfish are commonly associated with plants in the genus
3200:"Waterbird-Mediated Chance-Dispersal of Fishes: A Natural Process of Affecting Range of Distribution and Biogeography of Fishes" 1240: 3951: 1681:
is consumed in four days, and the larvae commence swimming after six days, at which point their eyes are completely developed.
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Santini, F.; Nguyen, M. T. T.; Sorenson, L.; Waltzek, T. B.; Lynch Alfaro, J. W.; Eastman, J. M.; Alfaro, M. E. (May 2013).
2915:(Hora & Nair, 1941) from the Neyyar wildlife sanctuary with a note on its feeding habit and length-weight relationship" 2669:(Hora & Nair, 1941) from the Neyyar wildlife sanctuary with a note on its feeding habit and length-weight relationship" 88: 3062:"On the Fish Collections from Kerala, Deposited in Southern Regional Station, Zoological Survey of India by NRM Stockholm" 2843:
Global Register of Migratory Species: From Global to Regional Scales : Final Report of the R&D-Projekt 808 05 081
2610:"On the Fish Collections from Kerala, Deposited in Southern Regional Station, Zoological Survey of India by NRM Stockholm" 1299:. Inflating tires pufferfish and can put them at greater risk of predation. Like many other pufferfish, in the absence of 2937: 2664: 3757: 3401:
Renjithkumar, C.R.; Roshni, K.; Ranjeet, K. (24 September 2020). "Feeding ecology of the endemic freshwater puffer fish
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of 3.5 cm, dwarf pufferfish are one of the smallest pufferfish in the world. They closely resemble the related
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in aquaria. Dwarf pufferfish have become popular as aquarium fish thanks to their attractive colours, small size, "
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away other males. In the aquarium, dwarf pufferfish are often plant-spawners, laying eggs in plants, including
1351: 1332: 805: 441: 397: 291: 20: 2055:"Do habitat shifts drive diversification in teleost fishes? An example from the pufferfishes (Tetraodontidae)" 1307:
on most of their body. These spines become erect when the fish is inflated. Dwarf pufferfish, like many other
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Easa, P.S.; Shaji, C.P. (1997). "Freshwater fish diversity in Kera a part of the Nilgiri Biosphere Reserve".
2271: 1576: 1060: 598: 380: 339: 2967:"A Survey on the Habitat and Distribution of Stream Fishes in the Kerala part of Nilgiri Biosphere Reserve" 2638:"A Survey on the Habitat and Distribution of Stream Fishes in the Kerala part of Nilgiri Biosphere Reserve" 2303:
Shono, T.; Thiery, A.P.; Cooper, R. L.; Kurokawa, D.; Britz, R.; Okabe, M.; Fraser, G.J. (September 2019).
3705: 3476: 2365:"Analysis of Poisoning Cases, Monitoring and Risk Warning for Marine Toxins (TTX, PSP and CTXs) in Taiwan" 2209: 1620: 1616: 968: 3770: 3061: 3040: 2609: 2467: 1640: 951: 927: 903: 183: 3294:"Freshwater Fishes of Southern Kerala with Notes on the Distribution of Endemic and Endangered Species" 3913: 1385: 3946: 3791: 3694: 3577: 3333: 3199: 2807: 2516: 2316: 1765: 1398:
species, meaning they migrate within freshwater river systems, and are endemic to rivers, lakes, and
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are situated opposite each other toward the posterior of the fish, both short and round, while the
674: 519: 53: 3601: 3349: 3274: 3222: 3096:"Fishes of River Bharathapuzha, Kerala, India: diversity, distribution, threats and conservation" 2823: 2704:"Fishes of River Bharathapuzha, Kerala, India: diversity, distribution, threats and conservation" 2532: 2082: 2030: 1482: 1478: 1365: 657: 401:, a superficially similar and closely related species previously mistaken for dwarf pufferfish. " 209: 83: 68: 3848: 1608:, and the presence of these plants has been shown to reduce mortality among captive specimens. 3895: 3778: 3664: 3634: 3593: 3515: 3376: 3218: 3028: 2929: 2850: 2755: 2683: 2455: 2342: 2175: 2147: 2115: 2074: 2022: 1973: 1893: 1585: 1308: 1093: 449: 367: 140: 3252:"Distribution, endemism and threat status of freshwater fishes in the Western Ghats of India" 2054: 1465:. Their presence in small, isolated water bodies is likely due to unintentional dispersal by 3900: 3874: 3585: 3507: 3414: 3341: 3266: 3214: 3107: 3020: 2875: 2815: 2715: 2589: 2524: 2447: 2376: 2332: 2324: 2066: 2014: 1760: 1624: 1407: 1258: 1142: 543: 385: 286: 3094:
Bijukumar, A.; Philip, Siby; Ali, Anvar; Sushama, S.; Raghavan, Rajeev (26 November 2013).
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Bijukumar, A.; Philip, Siby; Ali, Anvar; Sushama, S.; Raghavan, Rajeev (26 November 2013).
1798:"Notes on Fishes in the Indian Museum: XLI. New Records of Freshwater Fish from Travancore" 26:"Pygmy pufferfish" redirects here. For the pygmy pufferfish native to Central America, see 1706: 1304: 1254: 478: 335: 200: 160: 3581: 3337: 2811: 2520: 2320: 1710: 2337: 2304: 2005:) endemic to southwest India and its implications in the phylogeny of Tetraodontidae". 1742: 414: 319: 315: 150: 130: 3293: 3161: 3128: 2545: 1831: 1481:, though with no rationale provided. The dwarf pufferfish is officially classified as 3930: 3869: 3605: 3561: 3367:
Kurup, B.M.; Manojkumar, T.G. (2010). "Central Marine Fisheries Research Institute".
3270: 3226: 2871: 2827: 2034: 1957: 1925: 1877: 1751: 1664: 1612: 1494: 1486: 1439: 1431: 1403: 1395: 1262: 1175: 73: 3278: 2086: 1422:
with silt and sand, the species is reported from 13 rivers in Kerala, lakes such as
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There is no information available on the lifespans of these fish in the wild, but
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Dwarf pufferfish have dark, iridescent patches on their flanks and dorsal surfaces
2966: 2637: 2575:"Checklist of freshwater fishes collected from Ernakulam District, Kerala, India" 1414:
of 7.5–8.3 and temperatures of 22–28 Β°C (72–82 Β°F). Inhabiting heavily
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Anupama, K.M.; Hari Sankar, H.S.; Rithin Raj, M.; Harikrishnan, M. (July 2019).
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for which there is no antidote. The neurotoxin accumulates in the glands of the
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in that they consume a wide variety of animals. Dwarf pufferfish favour insect
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surface. Patterns and colouration vary considerably between individuals. Their
3589: 3345: 2879: 2819: 2593: 2528: 2018: 1673: 1518: 1498: 1455: 1316: 1300: 1278: 1270: 1266: 1250: 347: 267: 264: 3728: 3663:. Lucknow, India: National Bureau of Fish Genetic Resources. pp. 13–32. 3597: 3380: 3032: 2933: 2687: 2459: 1977: 1897: 1584:
In captivity, dwarf pufferfish benefit from a varied diet and will eat small
2380: 1702: 1698: 1668: 1657: 1597: 1554: 1466: 1451: 1447: 1419: 1415: 1399: 1356: 1320: 1274: 456: 438: 307: 303: 278: 100: 3519: 3418: 3129:"Freshwater fish diversity in Kera a part of the Nilgiri Biosphere Reserve" 2346: 2078: 2026: 19:"Malabar pufferfish" redirects here. For the dwarf Malabar pufferfish, see 3511: 3804: 3722: 1938: 1685: 1678: 1601: 1558: 1530: 1427: 1423: 1296: 355: 282: 120: 3809: 2174:. Lucknow: National Bureau of Fish Genetic Resources. pp. 227–228. 1956:
Britz, R.; Ali, A.; Philip, S.; Kumar, K.; Raghavan, R. (October 2012).
1882:, a new freshwater pufferfish from India (Teleostei: Tetraodontiformes)" 3822: 3160:
Biju, C.R., C.R.; Thomas, R.K.; Ajithkumar, C.R.; George, M.J. (1999).
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Biju, C.R., C.R.; Thomas, R.K.; Ajithkumar, C.R.; George, M.J. (1999).
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Eggs hatch after five days at 27 Β°C (81 Β°F), with larvae and
1605: 1589: 1566: 1550: 1542: 1538: 1490: 1336: 1324: 1283: 270: 3024: 2451: 2305:"Evolution and Developmental Diversity of Skin Spines in Pufferfishes" 2170:
Anna Mercy, T.V.; Gopalakrishnan, A.; Kapoor, D.; Lakra, W.S. (2007).
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Dahanukar, Neelesh; Raut, Rupesh; Bhat, Anuradha (22 December 2003).
3166:(Hora and Nair) in the Chalakudy, Periyar and Kechery rivers, Kerala" 2872:
Some observations on fish migration, with emphasis on tropical waters
2550:(Hora and Nair) in the Chalakudy, Periyar and Kechery rivers, Kerala" 1636: 1593: 1522: 274: 110: 3699: 444:
and a reduced number of total vertebrae similar to other members of
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in Peninsular India (Teleostei: Tetraodontiformes: Tetraodontidae)"
1619:(members of the same species), dwarf pufferfish are found in large 1646: 1575: 1562: 1458: 1384: 1328: 1239: 477: 406: 375: 1688:
report specimens live for approximately five years in captivity.
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in 1993 and K.K.P. Lim and M. Kottelat in 1995. The first use of
3375:(1). India: Central Marine Fisheries Research Institute: 81–85. 434: 3796: 3703: 1736: 1734: 1732: 1730: 1728: 1726: 1600:. They will also readily eat live foods such as red and white 395:
was in 1999 by R. Britz and M. Kottelat when first describing
1892:(1). Sri Lanka: Wildlife Heritage Trust of Sri Lanka: 39–47. 1627:
or weight loss when kept in solitude or in groups too small.
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Pufferfish Enthusiasts Worldwide Dwarf Pufferfish Care Sheet
2752:
Brackish-water Fishes: All about Species, Care and Breeding
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Beevi, K.S. Jameela; Ramachandran, A. (26 September 2009).
1411: 3405:(Hora & Nair, 1941) in Western Ghats hotspot, India". 2298: 2296: 1580:
Dwarf pufferfish shoaling in the wild, seen from above
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was not recognised as a different species until 1999.
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for their bright colours and small size. At a maximum
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Coral fish stress out if separated from 'shoal-mates'
2358: 2356: 2228: 2226: 1269:", that is to say, slightly indented at the tip. The 3538:. Manila Bulletin Publishing Corp. 22 September 2016 3396: 3394: 3392: 3390: 2909:
Prasad, G.; Sabu, K.; Prathibhakumari, P.V. (2012).
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Prasad, G.; Sabu, K.; Prathibhakumari, P.V. (2012).
1461:, and artificial tanks or abandoned water bodies in 459:
is based on molecular evidence and demonstrates the
3712: 3060:Devi, K.R.; Indra, T.J.; Emiliyamma, K. J. (1996). 2608:Devi, K.R.; Indra, T.J.; Emiliyamma, K. J. (1996). 1830:Devi, K.R.; Indra, T.J.; Raghunathan, M.B. (2000). 1359:, possibly inhabiting the same waters. Until 1999, 1339:because of the slow swimming speeds of pufferfish. 2874:(Report). Federal Agency for Nature Conservation. 2165: 2163: 1265:are short, fan-shaped, and described as "slightly 452:, also found amongst other members of this genus. 448:, in addition to a small size and the presence of 374:until 1978, when J.C. Tyler treated it as a valid 3245: 3243: 3045:: CS1 maint: DOI inactive as of September 2024 ( 2472:: CS1 maint: DOI inactive as of September 2024 ( 1996: 1994: 2483:"New records of freshwater fish from Travancore" 2272:"Puffed-Up Blowfish Aren't Holding Their Breath" 1766:10.2305/IUCN.UK.2011-1.RLTS.T166591A174788004.en 1477:Several researchers have considered the species 475:, which have not undergone mitogenome analyses. 302:Dwarf pufferfish dwell at the bottom of heavily 3555: 3553: 3292:Thomas, K.; George, M.; Biju, C. (April 2002). 3013:International Journal of Environmental Sciences 2998: 2996: 2994: 2440:International Journal of Environmental Sciences 2265: 2263: 2195: 2193: 2191: 2172:Ornamental fishes of the Western Ghats of India 2105: 2103: 322:threaten wild populations of dwarf pufferfish. 295:, and the two can be difficult to distinguish. 3977:Species endangered by agricultural development 2202:"The Dwarf Puffer: A Pleasant Little Surprise" 1287:they may have a yellow patch on their throat. 3624: 3622: 3298:Journal of the Bombay Natural History Society 3170:Journal of the Bombay Natural History Society 2554:Journal of the Bombay Natural History Society 2112:Inland fishes of India and adjacent countries 1840:Journal of the Bombay Natural History Society 1565:, with some amounts of plant matter, largely 8: 3562:"Reproductive Biology of Malabar Pufferfish 3193: 3191: 3155: 3153: 3151: 3149: 2960: 2958: 2904: 2902: 2900: 2898: 2896: 2870:Gopalakrishnan, Omkar; Riede, Klaus (2004). 1951: 1949: 1871: 1869: 1867: 1865: 1863: 1861: 330:The dwarf pufferfish was first described as 3005:"Improved survival of Malabar puffer fish, 2785: 2783: 2781: 2779: 2777: 2775: 2773: 2771: 2432:"Improved survival of Malabar puffer fish, 2363:Lin, W.-F.; Hwang, D.-F. (1 January 2012). 1825: 1823: 1791: 1789: 1787: 1785: 1783: 425:" (teeth). The justification for moving to 3700: 3654:Ponniah, A.G.; Gopalakrishnan, A. (2000). 2048: 2046: 2044: 1920: 1918: 1916: 1914: 62: 43: 34: 3207:Records of the Zoological Survey of India 3111: 3009:(Hora and Nair, 1941) in planted aquaria" 2911:"The first report of the Malabar puffer, 2719: 2665:"The first report of the Malabar puffer, 2436:(Hora and Nair, 1941) in planted aquaria" 2336: 1966:Ichthyological Exploration of Freshwaters 1764: 433:evidence and not on the presence of skin 3407:International Journal of Aquatic Biology 3003:Anupama, K.M.; Harikrishnan, M. (2015). 2430:Anupama, K.M.; Harikrishnan, M. (2015). 2270:Geggel, Laura Geggel (4 December 2014). 2137: 2135: 2133: 2131: 1489:due to declining populations because of 1402:in Kerala and southern Karnataka in the 281:in Southwest India. They are popular in 3657:Endemic fish diversity of Western Ghats 2144:The Puffers of Fresh and Brackish Water 2114:. New Delhi: Oxford & IBH Pub. Co. 1722: 1418:waters with beds of gravel and rock or 3444:. Aquarium Glaser GmbH. Archived from 3198:Gopi, K.C.; Radhakrishnan, C. (2007). 3038: 2847:Federal Agency for Nature Conservation 2465: 2402:"Tetrodotoxin: Biotoxin | NIOSH | CDC" 2110:Talwar, P. K.; Jhingran, A.G. (1991). 1886:Journal of South Asian Natural History 1836:, from south Kanara, Karnataka, India" 1611:Dwarf pufferfish are a slow-swimming, 1549:, and aquatic larvae such as those of 3069:Records of Zoological Survey of India 2617:Records of Zoological Survey of India 1438:. They have been reported as rare in 1410:. They can be found in waters with a 7: 1876:Britz, R.; Kottelat, M. (May 1999). 350:". The use of parentheses indicates 332:Tetraodon (Monotretus) travancoricus 3992:Species endangered by deforestation 3972:Species endangered by the pet trade 3432:SchΓ€fer, Frank (20 November 2020). 1834:Tetradon (Monotretus) travancoricus 1752:IUCN Red List of Threatened Species 1167: 1134: 1127: 1085: 1052: 1045: 1038: 943: 919: 895: 888: 855: 848: 780: 773: 740: 733: 726: 649: 590: 566: 559: 535: 511: 504: 497: 489: 384:, which was an opinion followed by 3982:Species endangered by urbanization 3962:Endemic fauna of the Western Ghats 3469:"In Search of the Peaceful Puffer" 2234:"Pufferfish | National Geographic" 14: 3631:Practical Fishkeeping: Pufferfish 3219:10.26515/rzsi/v107/i1/2007/159160 2922:Journal on New Biological Reports 2676:Journal on New Biological Reports 1368:(a member of the same genus), as 3937:IUCN Red List vulnerable species 3271:10.1046/j.0305-0270.2003.01016.x 3127:Easa, P.S.; Shaji, C.P. (1997). 2965:Easa, P.S.; Basha, C.C. (1995). 2636:Easa, P.S.; Basha, C.C. (1995). 2426:Based on the following sources: 1497:for agriculture, pollution from 1446:. The species was also found in 1331:of pufferfish. These traits are 87: 2481:Hora, S.L.; Nair, K.K. (1941). 2244:. 12 March 2010. Archived from 2146:. Aqualog. pp. 19, 46–49. 2059:Journal of Evolutionary Biology 1958:"First record from the wild of 1796:Hora, S.L.; Nair, K.K. (1941). 1592:, as well as foods like frozen 749:"Carinotetraodon" travancoricus 393:"Carinotetraodon travancoricus" 3467:Monks, Neale (December 2007). 3027:(inactive 12 September 2024). 2454:(inactive 12 September 2024). 1: 3987:Species endangered by damming 3957:Taxa named by Sunder Lal Hora 3758:carinotetraodon-travancoricus 3744:Carinotetraodon travancoricus 3714:Carinotetraodon travancoricus 3564:Carinotetraodon travancoricus 3436:Carinotetraodon travancoricus 3326:Biodiversity and Conservation 3007:Carinotetraodon travancoricus 2974:Kerala Forest Research Report 2913:Carinotetraodon travancoricus 2667:Carinotetraodon travancoricus 2645:Kerala Forest Research Report 2509:Biodiversity and Conservation 2434:Carinotetraodon travancoricus 2003:Carinotetraodon travancoricus 1932:Carinotetraodon travancoricus 1745:Carinotetraodon travancoricus 1635:In the wild, males with ripe 1454:, as well as ditches, ponds, 1303:, dwarf pufferfish have skin 248:Carinotetraodon travancoricus 193:Carinotetraodon travancoricus 3403:Carinotetradon travancoricus 2487:Records of the Indian Museum 2200:Hellweg, Mike (March 2007). 1802:Records of the Indian Museum 232:(Hora & K.K. Nair, 1941) 3369:Indian Journal of Fisheries 3113:10.11609/JoTT.o3640.4979-93 2721:10.11609/JoTT.o3640.4979-93 1667:in captivity initially fed 789:Dichotomyctere nigroviridis 616:"Carinotetraodon" irrubesco 575:"Carinotetraodon" salivator 484:Dichotomyctere nigroviridis 413:" (keel-shaped, shell) and 4008: 3100:Journal of Threatened Taxa 2708:Journal of Threatened Taxa 2582:Journal of Threatened Taxa 2329:10.1016/j.isci.2019.06.003 223:Hora & K.K. Nair, 1941 28:Sphoeroides trichocephalus 25: 18: 3633:. Ringpress. p. 61. 3590:10.1134/S0032945219040027 3346:10.1007/s10531-007-9293-0 2880:10.13140/RG.2.1.5099.0969 2820:10.1007/s12562-014-0842-7 2594:10.11609/JoTT.o1559.493-4 2529:10.1007/s10531-007-9293-0 2019:10.1007/s12041-019-1151-9 1705:, and are only valued as 1444:Nilgiri Biosphere Reserve 1436:Neyyar Wildlife Sanctuary 1333:anti-predator adaptations 1323:(surface layer of skin), 1172: 1165: 1139: 1132: 1125: 1090: 1083: 1057: 1050: 1043: 1036: 965: 948: 941: 924: 917: 900: 893: 886: 860: 853: 846: 802: 785: 778: 771: 745: 738: 731: 724: 671: 654: 647: 612: 599:"Carinotetraodon" lorteti 595: 588: 571: 564: 557: 540: 533: 516: 509: 502: 495: 215: 208: 189: 182: 84:Scientific classification 82: 60: 51: 42: 37: 3967:Freshwater fish of India 2238:www.nationalgeograph.orc 1960:Carinotetraodon imitator 1928:; Pauly, Daniel (eds.). 1880:Carinotetraodon imitator 1381:Distribution and habitat 1352:Carinotetraodon imitator 1345:Carinotetraodon imitator 806:Dichotomyctere ocellatus 398:Carinotetraodon imitator 338:and K.K. Nair, with the 292:Carinotetraodon imitator 228:Monotretus travancoricus 21:Carinotetraodon imitator 3259:Journal of Biogeography 3164:Tetraodon travancoricus 2750:SchΓ€fer, Frank (2005). 2548:Tetraodon travancoricus 2381:10.6227/jfda.2012200404 1759:: e.T166591A174788004. 1692:Association with humans 1061:Chelonodon pleurospilus 381:Carinotetraodon lorteti 219:Tetraodon travancoricus 3952:Fish described in 1941 3570:Journal of Ichthyology 3419:10.22034/ijab.v8i5.922 1741:Dahanukar, N. (2011). 1652: 1581: 1390: 1245: 969:Canthigaster valentini 486: 405:" is derived from the 203:& K.K. Nair, 1941) 3629:Ralph, Chris (2003). 3512:10.1089/zeb.2013.0867 3442:www.aquariumglaser.de 2754:. A.C.S. p. 34. 2142:Klaus, Ebert (2001). 1650: 1596:meat or other frozen 1579: 1514:Dwarf pufferfish are 1388: 1243: 952:Canthigaster jactator 928:Canthigaster rivulata 904:Canthigaster coronata 481: 429:was based largely on 251:), also known as the 175:C. travancoricus 2841:Reid, Klaus (2004). 1942:. June 2023 version. 864:Pelagocephalus marki 482:The closely related 463:nature of the genus 3582:2019JIch...59..545A 3338:2008BiCon..17.3119R 2812:2015FisSc..81..291D 2521:2008BiCon..17.3119R 2321:2019iSci...19.1248S 2242:National Geographic 2206:www.tfhmagazine.com 2007:Journal of Genetics 1473:Conservation status 1389:Global distribution 675:Pao cochinchinensis 520:Auriglobus modestus 54:Conservation status 3448:on 14 October 2023 1653: 1641:South-West monsoon 1582: 1510:Diet and behaviour 1501:, and, primarily, 1391: 1315:, a highly lethal 1246: 658:Pao palembangensis 487: 253:Malabar pufferfish 3924: 3923: 3896:Open Tree of Life 3706:Taxon identifiers 3566:(Tetraodontidae)" 3332:(13): 3119–3131. 3106:(15): 4979–4993. 3025:10.6088/ijes.6016 2800:Fisheries Science 2714:(15): 4979–4993. 2515:(13): 3119–3131. 2452:10.6088/ijes.6016 2181:978-81-902951-8-5 2071:10.1111/jeb.12112 1586:freshwater snails 1525:but will rely on 1493:, indiscriminate 1309:Tetraodontiformes 1232: 1231: 1223: 1222: 1214: 1213: 1205: 1204: 1196: 1195: 1187: 1186: 1154: 1153: 1114: 1113: 1105: 1104: 1094:Chelonodon patoca 1072: 1071: 1025: 1024: 1016: 1015: 1007: 1006: 998: 997: 989: 988: 980: 979: 875: 874: 835: 834: 826: 825: 817: 816: 760: 759: 713: 712: 704: 703: 695: 694: 686: 685: 636: 635: 627: 626: 450:sexual dimorphism 378:in a paper about 366:was considered a 239: 238: 233: 224: 141:Tetraodontiformes 77: 38:Dwarf pufferfish 3999: 3917: 3916: 3904: 3903: 3891: 3890: 3878: 3877: 3865: 3864: 3852: 3851: 3839: 3838: 3826: 3825: 3813: 3812: 3800: 3799: 3787: 3786: 3774: 3773: 3761: 3760: 3748: 3747: 3746: 3733: 3732: 3731: 3701: 3682: 3681: 3679: 3677: 3662: 3651: 3645: 3644: 3626: 3617: 3616: 3614: 3612: 3557: 3548: 3547: 3545: 3543: 3530: 3524: 3523: 3494: 3488: 3487: 3485: 3483: 3464: 3458: 3457: 3455: 3453: 3429: 3423: 3422: 3398: 3385: 3384: 3364: 3358: 3357: 3320: 3314: 3313: 3311: 3309: 3289: 3283: 3282: 3256: 3247: 3238: 3237: 3235: 3233: 3204: 3195: 3186: 3185: 3183: 3181: 3157: 3144: 3143: 3133: 3124: 3118: 3117: 3115: 3091: 3085: 3084: 3082: 3080: 3066: 3057: 3051: 3050: 3044: 3036: 3000: 2989: 2988: 2986: 2984: 2971: 2962: 2953: 2952: 2950: 2948: 2942: 2936:. Archived from 2919: 2906: 2891: 2890: 2888: 2886: 2867: 2861: 2860: 2838: 2832: 2831: 2787: 2766: 2765: 2747: 2741: 2738: 2725: 2723: 2698: 2696: 2694: 2673: 2659: 2657: 2655: 2642: 2632: 2630: 2628: 2614: 2604: 2602: 2600: 2579: 2569: 2567: 2565: 2540: 2502: 2500: 2498: 2477: 2471: 2463: 2424: 2418: 2417: 2415: 2413: 2398: 2392: 2391: 2389: 2387: 2360: 2351: 2350: 2340: 2300: 2291: 2290: 2288: 2286: 2267: 2258: 2257: 2255: 2253: 2248:on 29 March 2021 2230: 2221: 2220: 2218: 2216: 2197: 2186: 2185: 2167: 2158: 2157: 2139: 2126: 2125: 2107: 2098: 2097: 2095: 2093: 2065:(5): 1003–1018. 2050: 2039: 2038: 1998: 1989: 1988: 1986: 1984: 1953: 1944: 1943: 1922: 1909: 1908: 1906: 1904: 1873: 1856: 1855: 1853: 1851: 1832:"On a report of 1827: 1818: 1817: 1815: 1813: 1793: 1778: 1777: 1775: 1773: 1768: 1738: 1408:Peninsular India 1291:Natural defenses 1168: 1143:Tetraodon miurus 1135: 1128: 1086: 1053: 1046: 1039: 944: 920: 896: 889: 856: 849: 781: 774: 741: 734: 727: 650: 591: 567: 560: 544:Leiodon cutcutia 536: 512: 505: 498: 490: 257:pygmy pufferfish 243:dwarf pufferfish 231: 222: 195: 92: 91: 71: 66: 65: 47: 35: 4007: 4006: 4002: 4001: 4000: 3998: 3997: 3996: 3942:Carinotetraodon 3927: 3926: 3925: 3920: 3912: 3907: 3899: 3894: 3886: 3881: 3873: 3868: 3860: 3855: 3847: 3842: 3834: 3829: 3821: 3816: 3808: 3803: 3795: 3790: 3782: 3777: 3769: 3764: 3756: 3751: 3742: 3741: 3736: 3727: 3726: 3721: 3708: 3691: 3686: 3685: 3675: 3673: 3671: 3660: 3653: 3652: 3648: 3641: 3628: 3627: 3620: 3610: 3608: 3559: 3558: 3551: 3541: 3539: 3532: 3531: 3527: 3496: 3495: 3491: 3481: 3479: 3473:tfhmagazine.com 3466: 3465: 3461: 3451: 3449: 3431: 3430: 3426: 3400: 3399: 3388: 3366: 3365: 3361: 3322: 3321: 3317: 3307: 3305: 3291: 3290: 3286: 3254: 3249: 3248: 3241: 3231: 3229: 3202: 3197: 3196: 3189: 3179: 3177: 3162:"Occurrence of 3159: 3158: 3147: 3136:Current Science 3131: 3126: 3125: 3121: 3093: 3092: 3088: 3078: 3076: 3064: 3059: 3058: 3054: 3037: 3002: 3001: 2992: 2982: 2980: 2969: 2964: 2963: 2956: 2946: 2944: 2943:on 13 July 2020 2940: 2917: 2908: 2907: 2894: 2884: 2882: 2869: 2868: 2864: 2857: 2849:. p. 308. 2840: 2839: 2835: 2798:in captivity". 2796:Carinotetraodon 2789: 2788: 2769: 2762: 2749: 2748: 2744: 2731:Current Science 2728: 2701: 2692: 2690: 2671: 2662: 2653: 2651: 2640: 2635: 2626: 2624: 2612: 2607: 2598: 2596: 2577: 2572: 2563: 2561: 2546:"Occurrence of 2543: 2505: 2496: 2494: 2480: 2464: 2429: 2425: 2421: 2411: 2409: 2400: 2399: 2395: 2385: 2383: 2362: 2361: 2354: 2302: 2301: 2294: 2284: 2282: 2276:livescience.com 2269: 2268: 2261: 2251: 2249: 2232: 2231: 2224: 2214: 2212: 2199: 2198: 2189: 2182: 2169: 2168: 2161: 2154: 2141: 2140: 2129: 2122: 2109: 2108: 2101: 2091: 2089: 2052: 2051: 2042: 2000: 1999: 1992: 1982: 1980: 1955: 1954: 1947: 1924: 1923: 1912: 1902: 1900: 1875: 1874: 1859: 1849: 1847: 1829: 1828: 1821: 1811: 1809: 1795: 1794: 1781: 1771: 1769: 1740: 1739: 1724: 1719: 1707:ornamental fish 1694: 1633: 1512: 1475: 1383: 1348: 1343:Resemblance to 1293: 1253:patches on the 1238: 1233: 1224: 1215: 1206: 1197: 1188: 1155: 1115: 1106: 1073: 1026: 1017: 1008: 999: 990: 981: 876: 836: 827: 818: 761: 714: 705: 696: 687: 637: 628: 493:Tetraodontidae 469:Carinotetraodon 465:Carinotetraodon 446:Carinotetraodon 427:Carinotetraodon 403:Carinotetraodon 364:Carinotetraodon 328: 230: 221: 204: 197: 191: 178: 162:Carinotetraodon 86: 78: 67: 63: 56: 31: 24: 17: 16:Species of fish 12: 11: 5: 4005: 4003: 3995: 3994: 3989: 3984: 3979: 3974: 3969: 3964: 3959: 3954: 3949: 3944: 3939: 3929: 3928: 3922: 3921: 3919: 3918: 3905: 3892: 3879: 3866: 3853: 3840: 3827: 3814: 3801: 3788: 3775: 3762: 3749: 3734: 3718: 3716: 3710: 3709: 3704: 3698: 3697: 3690: 3689:External links 3687: 3684: 3683: 3669: 3646: 3639: 3618: 3576:(4): 545–554. 3549: 3525: 3506:(3): 338–342. 3489: 3459: 3424: 3413:(5): 300–310. 3386: 3359: 3315: 3284: 3239: 3187: 3145: 3119: 3086: 3052: 3019:(1): 138–144. 2990: 2954: 2892: 2862: 2855: 2833: 2806:(2): 291–299. 2767: 2760: 2742: 2740: 2739: 2726: 2699: 2660: 2633: 2605: 2588:(9): 493–494. 2570: 2541: 2503: 2478: 2446:(1): 138–144. 2419: 2393: 2375:(4): 763–771. 2352: 2292: 2259: 2222: 2187: 2180: 2159: 2152: 2127: 2120: 2099: 2040: 1990: 1972:(2): 105–109. 1945: 1926:Froese, Rainer 1910: 1857: 1819: 1779: 1721: 1720: 1718: 1715: 1711:puppy dog eyes 1693: 1690: 1632: 1629: 1511: 1508: 1474: 1471: 1450:brickyards in 1434:, part of the 1430:harbours, and 1382: 1379: 1347: 1341: 1292: 1289: 1237: 1234: 1230: 1229: 1226: 1225: 1221: 1220: 1217: 1216: 1212: 1211: 1208: 1207: 1203: 1202: 1199: 1198: 1194: 1193: 1190: 1189: 1185: 1184: 1181: 1180: 1171: 1166: 1164: 1161: 1160: 1157: 1156: 1152: 1151: 1148: 1147: 1138: 1133: 1131: 1126: 1124: 1121: 1120: 1117: 1116: 1112: 1111: 1108: 1107: 1103: 1102: 1099: 1098: 1089: 1084: 1082: 1079: 1078: 1075: 1074: 1070: 1069: 1066: 1065: 1056: 1051: 1049: 1044: 1042: 1037: 1035: 1032: 1031: 1028: 1027: 1023: 1022: 1019: 1018: 1014: 1013: 1010: 1009: 1005: 1004: 1001: 1000: 996: 995: 992: 991: 987: 986: 983: 982: 978: 977: 974: 973: 964: 961: 960: 957: 956: 947: 942: 940: 937: 936: 933: 932: 923: 918: 916: 913: 912: 909: 908: 899: 894: 892: 887: 885: 882: 881: 878: 877: 873: 872: 869: 868: 859: 854: 852: 847: 845: 842: 841: 838: 837: 833: 832: 829: 828: 824: 823: 820: 819: 815: 814: 811: 810: 801: 798: 797: 794: 793: 784: 779: 777: 772: 770: 767: 766: 763: 762: 758: 757: 754: 753: 744: 739: 737: 732: 730: 725: 723: 720: 719: 716: 715: 711: 710: 707: 706: 702: 701: 698: 697: 693: 692: 689: 688: 684: 683: 680: 679: 670: 667: 666: 663: 662: 653: 648: 646: 643: 642: 639: 638: 634: 633: 630: 629: 625: 624: 621: 620: 611: 608: 607: 604: 603: 594: 589: 587: 584: 583: 580: 579: 570: 565: 563: 558: 556: 553: 552: 549: 548: 539: 534: 532: 529: 528: 525: 524: 515: 510: 508: 503: 501: 496: 494: 488: 455:The following 421:" (four) and " 327: 324: 320:aquarium trade 316:overharvesting 261:pea pufferfish 237: 236: 235: 234: 225: 213: 212: 206: 205: 198: 187: 186: 180: 179: 172: 170: 166: 165: 158: 154: 153: 151:Tetraodontidae 148: 144: 143: 138: 134: 133: 131:Actinopterygii 128: 124: 123: 118: 114: 113: 108: 104: 103: 98: 94: 93: 80: 79: 61: 58: 57: 52: 49: 48: 40: 39: 15: 13: 10: 9: 6: 4: 3: 2: 4004: 3993: 3990: 3988: 3985: 3983: 3980: 3978: 3975: 3973: 3970: 3968: 3965: 3963: 3960: 3958: 3955: 3953: 3950: 3948: 3945: 3943: 3940: 3938: 3935: 3934: 3932: 3915: 3910: 3906: 3902: 3897: 3893: 3889: 3884: 3880: 3876: 3871: 3867: 3863: 3858: 3854: 3850: 3845: 3841: 3837: 3832: 3828: 3824: 3819: 3815: 3811: 3806: 3802: 3798: 3793: 3789: 3785: 3780: 3776: 3772: 3767: 3763: 3759: 3754: 3750: 3745: 3739: 3735: 3730: 3724: 3720: 3719: 3717: 3715: 3711: 3707: 3702: 3696: 3693: 3692: 3688: 3672: 3670:81-901014-2-0 3666: 3659: 3658: 3650: 3647: 3642: 3640:1-86054-233-6 3636: 3632: 3625: 3623: 3619: 3607: 3603: 3599: 3595: 3591: 3587: 3583: 3579: 3575: 3571: 3567: 3565: 3556: 3554: 3550: 3537: 3536: 3529: 3526: 3521: 3517: 3513: 3509: 3505: 3501: 3493: 3490: 3478: 3474: 3470: 3463: 3460: 3447: 3443: 3439: 3437: 3428: 3425: 3420: 3416: 3412: 3408: 3404: 3397: 3395: 3393: 3391: 3387: 3382: 3378: 3374: 3370: 3363: 3360: 3355: 3351: 3347: 3343: 3339: 3335: 3331: 3327: 3319: 3316: 3303: 3299: 3295: 3288: 3285: 3280: 3276: 3272: 3268: 3264: 3260: 3253: 3246: 3244: 3240: 3228: 3224: 3220: 3216: 3212: 3208: 3201: 3194: 3192: 3188: 3175: 3171: 3167: 3165: 3156: 3154: 3152: 3150: 3146: 3142:(2): 180–182. 3141: 3137: 3130: 3123: 3120: 3114: 3109: 3105: 3101: 3097: 3090: 3087: 3074: 3070: 3063: 3056: 3053: 3048: 3042: 3034: 3030: 3026: 3022: 3018: 3014: 3010: 3008: 2999: 2997: 2995: 2991: 2979: 2975: 2968: 2961: 2959: 2955: 2939: 2935: 2931: 2927: 2923: 2916: 2914: 2905: 2903: 2901: 2899: 2897: 2893: 2881: 2877: 2873: 2866: 2863: 2858: 2856:9783784338453 2852: 2848: 2844: 2837: 2834: 2829: 2825: 2821: 2817: 2813: 2809: 2805: 2801: 2797: 2793: 2786: 2784: 2782: 2780: 2778: 2776: 2774: 2772: 2768: 2763: 2761:9783936027822 2757: 2753: 2746: 2743: 2737:(2): 180–182. 2736: 2732: 2727: 2722: 2717: 2713: 2709: 2705: 2700: 2689: 2685: 2681: 2677: 2670: 2668: 2661: 2650: 2646: 2639: 2634: 2622: 2618: 2611: 2606: 2595: 2591: 2587: 2583: 2576: 2571: 2559: 2555: 2551: 2549: 2542: 2538: 2534: 2530: 2526: 2522: 2518: 2514: 2510: 2504: 2492: 2488: 2484: 2479: 2475: 2469: 2461: 2457: 2453: 2449: 2445: 2441: 2437: 2435: 2428: 2427: 2423: 2420: 2408:. 25 May 2023 2407: 2403: 2397: 2394: 2382: 2378: 2374: 2370: 2366: 2359: 2357: 2353: 2348: 2344: 2339: 2334: 2330: 2326: 2322: 2318: 2315:: 1248–1259. 2314: 2310: 2306: 2299: 2297: 2293: 2281: 2277: 2273: 2266: 2264: 2260: 2247: 2243: 2239: 2235: 2229: 2227: 2223: 2211: 2207: 2203: 2196: 2194: 2192: 2188: 2183: 2177: 2173: 2166: 2164: 2160: 2155: 2153:3-931702-60-X 2149: 2145: 2138: 2136: 2134: 2132: 2128: 2123: 2121:9788120406391 2117: 2113: 2106: 2104: 2100: 2088: 2084: 2080: 2076: 2072: 2068: 2064: 2060: 2056: 2049: 2047: 2045: 2041: 2036: 2032: 2028: 2024: 2020: 2016: 2013:(5): 98–105. 2012: 2008: 2004: 1997: 1995: 1991: 1979: 1975: 1971: 1967: 1963: 1961: 1952: 1950: 1946: 1941: 1940: 1935: 1933: 1927: 1921: 1919: 1917: 1915: 1911: 1899: 1895: 1891: 1887: 1883: 1881: 1872: 1870: 1868: 1866: 1864: 1862: 1858: 1845: 1841: 1837: 1835: 1826: 1824: 1820: 1807: 1803: 1799: 1792: 1790: 1788: 1786: 1784: 1780: 1767: 1762: 1758: 1754: 1753: 1748: 1746: 1737: 1735: 1733: 1731: 1729: 1727: 1723: 1716: 1714: 1712: 1708: 1704: 1700: 1691: 1689: 1687: 1682: 1680: 1676: 1675: 1670: 1666: 1661: 1659: 1649: 1645: 1642: 1638: 1630: 1628: 1626: 1622: 1618: 1614: 1609: 1607: 1603: 1599: 1595: 1591: 1587: 1578: 1574: 1572: 1568: 1564: 1560: 1556: 1552: 1548: 1544: 1540: 1536: 1532: 1528: 1524: 1520: 1517: 1509: 1507: 1504: 1500: 1496: 1495:deforestation 1492: 1488: 1487:IUCN Red List 1484: 1480: 1472: 1470: 1468: 1464: 1460: 1457: 1453: 1449: 1445: 1441: 1440:Bharathapuzha 1437: 1433: 1432:Kallar Stream 1429: 1425: 1421: 1417: 1413: 1409: 1405: 1404:Western Ghats 1401: 1397: 1396:potamodromous 1387: 1380: 1378: 1376: 1371: 1367: 1362: 1358: 1354: 1353: 1346: 1342: 1340: 1338: 1334: 1330: 1326: 1322: 1318: 1314: 1310: 1306: 1302: 1298: 1290: 1288: 1285: 1280: 1276: 1272: 1268: 1264: 1263:pectoral fins 1260: 1256: 1252: 1242: 1235: 1228: 1227: 1219: 1218: 1210: 1209: 1201: 1200: 1192: 1191: 1183: 1182: 1179: 1178: 1177: 1176:Tetraodon mbu 1170: 1169: 1163: 1162: 1159: 1158: 1150: 1149: 1146: 1145: 1144: 1137: 1136: 1130: 1129: 1123: 1122: 1119: 1118: 1110: 1109: 1101: 1100: 1097: 1096: 1095: 1088: 1087: 1081: 1080: 1077: 1076: 1068: 1067: 1064: 1063: 1062: 1055: 1054: 1048: 1047: 1041: 1040: 1034: 1033: 1030: 1029: 1021: 1020: 1012: 1011: 1003: 1002: 994: 993: 985: 984: 976: 975: 972: 971: 970: 963: 962: 959: 958: 955: 954: 953: 946: 945: 939: 938: 935: 934: 931: 930: 929: 922: 921: 915: 914: 911: 910: 907: 906: 905: 898: 897: 891: 890: 884: 883: 880: 879: 871: 870: 867: 866: 865: 858: 857: 851: 850: 844: 843: 840: 839: 831: 830: 822: 821: 813: 812: 809: 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Index

Carinotetraodon imitator
Sphoeroides trichocephalus
A side profile of an adult dwarf pufferfish against a white background, facing left and slightly upward.
Conservation status
Vulnerable
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Actinopterygii
Tetraodontiformes
Tetraodontidae
Carinotetraodon
Binomial name
Hora
Synonyms
freshwater
pufferfish
endemic
Kerala
Karnataka
aquaria
total length
Carinotetraodon imitator
vegetated
predating
Habitat loss
overharvesting

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