427:
439:
44:
463:
31:
451:
717:
395:
400:
731:
242:
undergoes a six-stage life cycle from egg to adult, with three larval instars and a single pupal stage. The time from egg to pupal eclosion is estimated to take approximately two years. The pupal stage lasts about 30 days. Adults emerge from the pupal chamber between the months of August through
211:
Adult body sizes of both sexes vary from 3.5 to 6.0 cm (1.4–2.4 in) (apex of elytra to apex of thoracic horn), with captive-reared individuals reaching up to 8.0 cm (3.1 in) Coloration of the
341:
are very similar and it is possible for the two species to produce hybrids. Some scientists believe that the two beetles are actually subspecies given the similarities of the male genitalia.
243:
October and typically live between two and four months in the wild. In captivity, the adults may live up to nine months. After mating the female oviposits over 100 eggs.
871:
910:
216:
varies from white to grayish-white, often with irregular black spots of various size and number. At high humidity levels, the elytra may appear black.
845:
884:
614:
961:
956:
889:
806:
275:. This species typically inhabits highland forest habitats, at or above 1,600 m (5,200 ft) above sea level.
735:
689:
915:
43:
438:
426:
333:. Despite differences in geographic origin and morphology, namely horn length and coloration of the elytra,
749:
811:
138:
787:
721:
291:. The beetles strip the bark from small branches and feed on the sap produced by the exposed cambium.
819:
462:
353:, with differentiation between species often only possible through examination of geographic origin.
374:
The significant protrusion forward of the bottom tooth at the base of the thoracic horn in males of
951:
450:
902:
671:
536:
38:
876:
923:
663:
620:
610:
528:
223:
180:
928:
655:
582:
302:
belongs to the White
Hercules beetle lineage which includes five allopatrically distributed
480:
411:
382:
376:
366:
349:
329:
322:
312:
307:
639:
945:
318:
303:
268:
675:
30:
105:
394:
897:
858:
781:
716:
659:
390:
geographic origin is generally the strongest factor in correct identification.
399:
264:
188:
772:
667:
624:
570:
532:
832:
586:
75:
55:
798:
730:
766:
370:
is typically yellowish to brownish where it is white to grayish-white in
288:
235:
115:
571:"The species versus subspecies conundrum in the Hercules beetles (genus
850:
540:
512:
284:
260:
213:
200:
184:
176:
863:
604:
272:
196:
95:
85:
65:
743:
837:
690:"Hercules Beetles of Kentucky - University of Kentucky Entomology"
226:, with the females lacking the characteristic horns of the male.
824:
256:
192:
747:
380:
is also a distinguishing factor between the two species. While
203:, large size, and characteristic horn of the adult males.
575:) and its biogeographic and conservation implications"
638:
Krell, Frank-Thorsten; Krell, Victoria H. I. (2015).
756:
606:沉醉兜鍬 = For the love of rhinoceros and stag beetles
199:. This species is known for its grayish-white
195:in the United States and in parts of northern
506:
504:
502:
500:
498:
496:
8:
644:Horn (Coleoptera: Scarabaeidae: Dynastinae)"
640:"Longevity of the Western Hercules Beetle,
511:Menke, Arnold S.; Parker, Frank D. (1988).
283:Adults have been documented feeding on the
744:
386:can be distinguished morphologically from
29:
20:
579:2016 International Congress of Entomology
564:
562:
560:
558:
556:
554:
552:
550:
492:
422:
7:
598:
596:
581:. Entomological Society of America.
513:"Adult feeding and distribution of
347:is very similar morphologically to
14:
729:
715:
517:Horn (Coleoptera: Scarabaeidae)"
461:
449:
437:
425:
398:
393:
42:
1:
405:A comparison between a male
978:
660:10.1649/0010-065X-69.4.760
648:The Coleopterists Bulletin
521:The Coleopterists Bulletin
962:Beetles described in 1870
255:may be found in southern
144:
137:
39:Scientific classification
37:
28:
23:
957:Beetles of North America
271:, and parts of Northern
247:Distribution and habitat
24:Western Hercules beetle
587:10.1603/ice.2016.109568
569:Huang, Jen-Pan (2016).
364:The body coloration of
171:, often misspelled as "
163:western Hercules beetle
740:at Wikimedia Commons
603:(動物學), 賴廷奇 (2008).
409:(left) and a male
224:sexually dimorphic
939:
938:
924:Open Tree of Life
750:Taxon identifiers
734:Media related to
616:978-957-41-5817-1
419:Additional images
388:Dynastes grantii,
372:Dynastes grantii.
357:Distinction from
181:rhinoceros beetle
159:
158:
16:Species of beetle
969:
932:
931:
919:
918:
906:
905:
893:
892:
880:
879:
867:
866:
854:
853:
841:
840:
828:
827:
815:
814:
802:
801:
792:
791:
790:
788:Dynastes grantii
777:
776:
775:
758:Dynastes grantii
745:
737:Dynastes grantii
733:
723:Dynastes grantii
720:Data related to
719:
704:
703:
701:
700:
686:
680:
679:
642:Dynastes grantii
635:
629:
628:
600:
591:
590:
566:
545:
544:
508:
465:
453:
441:
429:
407:Dynastes grantii
402:
397:
345:Dynastes grantii
339:Dynastes grantii
310:taxa, including
308:Central American
300:Dynastes grantii
253:Dynastes grantii
240:Dynastes grantii
220:Dynastes grantii
168:Dynastes grantii
150:
148:Dynastes grantii
47:
46:
33:
21:
977:
976:
972:
971:
970:
968:
967:
966:
942:
941:
940:
935:
927:
922:
914:
909:
901:
896:
888:
883:
875:
870:
862:
857:
849:
844:
836:
831:
823:
818:
810:
805:
797:
795:
786:
785:
780:
771:
770:
765:
752:
712:
707:
698:
696:
688:
687:
683:
637:
636:
632:
617:
609:. Lai ting qi.
602:
601:
594:
568:
567:
548:
515:Dynastes granti
510:
509:
494:
490:
481:Dynastes tityus
476:
469:
466:
457:
454:
445:
442:
433:
430:
421:
412:Dynastes hyllus
383:Dynastes hyllus
377:Dynastes hyllus
367:Dynastes hyllus
362:
350:Dynastes hyllus
335:Dynastes tityus
330:Dynastes tityus
323:Dynastes moroni
313:Dynastes hyllus
297:
295:Similar species
281:
251:Populations of
249:
236:Hercules beetle
232:
209:
155:
152:
146:
133:
130:D. grantii
41:
17:
12:
11:
5:
975:
973:
965:
964:
959:
954:
944:
943:
937:
936:
934:
933:
920:
907:
894:
881:
868:
855:
842:
829:
816:
803:
793:
778:
762:
760:
754:
753:
748:
742:
741:
727:
726:at Wikispecies
711:
710:External links
708:
706:
705:
681:
630:
615:
592:
546:
527:(2): 161–164.
491:
489:
486:
485:
484:
475:
472:
471:
470:
467:
460:
458:
455:
448:
446:
443:
436:
434:
431:
424:
420:
417:
361:
355:
304:North American
296:
293:
280:
279:Feeding habits
277:
248:
245:
231:
228:
208:
205:
183:that lives in
157:
156:
153:
142:
141:
135:
134:
127:
125:
121:
120:
113:
109:
108:
103:
99:
98:
93:
89:
88:
83:
79:
78:
73:
69:
68:
63:
59:
58:
53:
49:
48:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
974:
963:
960:
958:
955:
953:
950:
949:
947:
930:
925:
921:
917:
912:
908:
904:
899:
895:
891:
886:
882:
878:
873:
869:
865:
860:
856:
852:
847:
843:
839:
834:
830:
826:
821:
817:
813:
808:
804:
800:
794:
789:
783:
779:
774:
768:
764:
763:
761:
759:
755:
751:
746:
739:
738:
732:
728:
725:
724:
718:
714:
713:
709:
695:
691:
685:
682:
677:
673:
669:
665:
661:
657:
653:
649:
645:
643:
634:
631:
626:
622:
618:
612:
608:
607:
599:
597:
593:
588:
584:
580:
576:
574:
565:
563:
561:
559:
557:
555:
553:
551:
547:
542:
538:
534:
530:
526:
522:
518:
516:
507:
505:
503:
501:
499:
497:
493:
487:
483:
482:
478:
477:
473:
464:
459:
452:
447:
440:
435:
428:
423:
418:
416:
414:
413:
408:
403:
401:
396:
391:
389:
385:
384:
379:
378:
373:
369:
368:
360:
356:
354:
352:
351:
346:
342:
340:
336:
332:
331:
326:
324:
320:
319:Dynastes maya
315:
314:
309:
305:
301:
294:
292:
290:
286:
278:
276:
274:
270:
269:United States
266:
262:
258:
254:
246:
244:
241:
237:
229:
227:
225:
221:
217:
215:
206:
204:
202:
198:
194:
190:
186:
182:
178:
174:
170:
169:
164:
151:
149:
143:
140:
139:Binomial name
136:
132:
131:
126:
123:
122:
119:
118:
114:
111:
110:
107:
104:
101:
100:
97:
94:
91:
90:
87:
84:
81:
80:
77:
74:
71:
70:
67:
64:
61:
60:
57:
54:
51:
50:
45:
40:
36:
32:
27:
22:
19:
757:
736:
722:
697:. Retrieved
693:
684:
651:
647:
641:
633:
605:
578:
572:
524:
520:
514:
479:
468:Adult female
444:Adult female
410:
406:
404:
392:
387:
381:
375:
371:
365:
363:
358:
348:
344:
343:
338:
334:
328:
317:
311:
299:
298:
282:
252:
250:
239:
233:
219:
218:
210:
172:
167:
166:
162:
160:
147:
145:
129:
128:
116:
106:Scarabaeidae
18:
898:NatureServe
859:iNaturalist
782:Wikispecies
694:www.uky.edu
234:Like other
207:Description
952:Dynastinae
946:Categories
699:2021-03-16
654:(4): 760.
488:References
456:Adult male
432:Adult male
265:New Mexico
263:, western
230:Life cycle
189:New Mexico
154:Horn, 1870
96:Coleoptera
76:Arthropoda
668:0010-065X
625:862438489
533:0010-065X
359:D. hyllus
289:ash trees
238:species,
124:Species:
62:Kingdom:
56:Eukaryota
903:2.114732
877:10380128
796:BioLib:
767:Wikidata
676:85764498
573:Dynastes
474:See also
415:(right)
175:") is a
117:Dynastes
102:Family:
72:Phylum:
66:Animalia
52:Domain:
929:7769068
851:1074535
825:1026726
773:Q644777
541:4008541
285:cambium
267:of the
261:Arizona
185:Arizona
177:species
112:Genus:
92:Order:
86:Insecta
82:Class:
890:929192
864:424045
838:DYNSGR
812:381200
799:336519
674:
666:
623:
613:
539:
531:
273:Mexico
214:elytra
201:elytra
197:Mexico
173:granti
916:60843
872:IRMNG
672:S2CID
537:JSTOR
911:NCBI
885:ITIS
846:GBIF
833:EPPO
807:BOLD
664:ISSN
621:OCLC
611:ISBN
529:ISSN
337:and
327:and
306:and
257:Utah
193:Utah
191:and
161:The
820:EoL
656:doi
583:doi
287:of
222:is
179:of
948::
926::
913::
900::
887::
874::
861::
848::
835::
822::
809::
784::
769::
692:.
670:.
662:.
652:69
650:.
646:.
619:.
595:^
577:.
549:^
535:.
525:42
523:.
519:.
495:^
321:,
316:,
259:,
187:,
702:.
678:.
658::
627:.
589:.
585::
543:.
325:,
165:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
