Knowledge (XXG)

Aublysodon

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was a mystery for a long time since no further skeletal elements were found that could be assigned with certainty to the teeth. In the early twentieth century it was assumed by some workers that it represented a member of the
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Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B.
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was for a time also used in definitions of higher-level taxa. Holtz proposed a node clade definition of the Tyrannosauridae in 2001 as "all descendants of the most recent common ancestor of
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in 1977/1978. The skull, specimen LACM 28741 at forty-five centimetres the length of an average human arm, bore pointed teeth attached to a long narrow snout. First thought to be a juvenile
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as an anchor taxon for the Tyrannosauridae, although his definition was problematic for other reasons. These concepts have now been redefined without the dubious name.
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was by Paul in 1988 thought to belong to a unique subfamily of tyrannosaurids called the Aublysodontinae, a name already coined, together with an Aublysodontidae, by
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over the years several other species have been named. These are now all considered either dubious or identical to other species or as having no close connection to
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Leidy, J., 1856, "Notices of the remains of extinct reptiles and fishes, discovered by Dr. F.V. Hayden in the badlands of the Judith River, Nebraska Territory",
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further limited its type by choosing a single small nonserrated premaxillary tooth with a D-shaped cross-section, specimen ANSP 9535, as the lectotype of
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Carpenter, K., 1982, "Baby dinosaurs from the Late Cretaceous Lance and Hell Creek formations and a description of a new species of theropod",
1058:. Marsh however, in 1892 was misled by the small size of the teeth, their D-shaped cross-section and their lack of serrations into considering 5913: 481:
alleged to distinguish the Aublysodontinae, especially lack of serrations on premaxillary teeth could have been caused by tooth wear in life,
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The Fossil Record: A Symposium with Documentation, jointly sponsored by the Geological Society of London and the Palaeontological Association
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exceptionally large for the Cretaceous. By the early twentieth century it was again generally understood that
477:, which was present in contemporary formations, and which matches specific details of the original tooth. The 5920: 1602:, gen. et sp. nov., a new tyrannosauroid from New Mexico and the origin of deep snouts in Tyrannosauroidea", 6070: 1751: 629: 1722:
Holtz, TR, Jr. (2001), "The phylogeny and taxonomy of the Tyrannosauridae" in: K Carpenter & D Tanke ,
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Currie, P.J., J.H. Hurum, and K. Sabath, 2003, "Skull structure and evolution in tyrannosaurid dinosaurs",
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Rozhdestvensky, A.K., 1965, "Возрастная изменчивость и некоторые вопросы систематики динозавров Азии",
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Olshevsky, G.; Ford, T.L. & Yamamoto, S., 1995, "The origin and evolution of the tyrannosaurids",
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Jacobsen, A.R. 2001. "Tooth-marked small theropod bone: An extremely rare trace". p. 58-63 In:
1682:"Diversity of late Maastrichtian Tyrannosauridae (Dinosauria: Theropoda) from western North America" 4330: 4117: 1043: 895: 593: 4599: 1127: 658:. Figures 37-40 show ANSP 9533, and figures 33 and 34 specimen ANSP 9534, the later lectotypes of 5785: 4306: 4166: 4022: 3962: 3397: 3307: 3190: 2054: 1744: 1407:
Cope, E.D., 1876, "Descriptions of some vertebrate remains from the Fort Union Beds of Montana",
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Cope, E.D., 1876, "Descriptions of some vertebrate remains from the Fort Union Beds of Montana",
879: 844:, based on specimen AMNH 3956, a tooth of a juvenile tyrannosaur which has been synonymized with 523: 372: 152: 858:, respectively based on teeth YPM 296 and YPM 297; the latter has also been placed in the genus 616:
or explanation of the intended meaning. It is possible the genus name is derived from Greek αὖ,
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Currie, P. (2005). "Theropods, including birds." Pp. 367-397 in Currie and Koppelhus (eds).,
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Molnar, R.E., 1978, "A new theropod dinosaur from the Upper Cretaceous of central Montana",
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Cope, E.D., 1870, "Synopsis of the extinct Batrachia, Reptilia and Aves of North America",
1341:*Lambe, L.M., 1902, "New genera and species from the Belly River Series (mid-Cretaceous)", 5834: 5572: 5535: 5467: 5322: 5315: 5143: 5085: 5058: 5048: 5008: 4843: 4702: 4686: 4504: 4476: 4454: 4341: 4186: 3893: 3667: 3550: 3528: 3449: 3278: 3255: 3248: 3226: 3212: 3205: 3166: 3017: 2864: 2791: 2780: 2716: 2604: 2491: 2068: 1977: 1831: 1806: 1075: 982: 964: 870: 787: 464: 257: 244: 41: 908:, but most later researchers have considered this a synonym of the Campanian tyrannosaur 451:(front) tooth. Although this specimen is now lost, similar teeth have been found in many 5799: 5762: 5743: 5705: 5694: 5491: 5204: 5170: 5152: 5132: 5029: 4743: 4488: 4350: 4177: 4159: 3987: 3954: 3940: 3901: 3675: 3536: 3493: 3486: 3408: 3384: 3337: 3296: 3233: 3219: 2947: 2932: 2759: 2593: 2457: 2425: 2414: 2391: 2353: 2214: 2184: 2108: 2047: 1946: 1019: 1013: 977:, then interpreted as a large dromaeosaurid, this "Jordan theropod" was given the name 774: 744: 478: 473: 444: 1252:
Cope, E.D., 1866, "Discovery of a gigantic dinosaur in the Cretaceous of New Jersey",
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Lehman, T.M., Carpenter, K., 1990, "A partial skeleton of the tyrannosaurid dinosaur
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by George Olshevsky in 1995, but was later, in 2004, reinterpreted to be a juvenile
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in 1868. The meaning of the generic name is uncertain because Leidy himself gave no
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Sahni, A. (1972). "The vertebrate fauna of the Judith River Formation, Montana."
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several other species have been named within the genus. In 1876 Cope created an
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no longer considered the name to represent a real biological taxon, but to be a
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Hay, O.P., 1899, "On the nomenclature of certain American fossil vertebrates",
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Charig, A.J., 1967, "Archosauria", pp 708-718 in: Walter Brian Harland (ed.)
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period (about 75 million years ago). The only currently recognized species,
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Second Bibliography and Catalogue of the Fossil Vertebrata of North America
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Today it is known that similar teeth are found in juvenile specimens of
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Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte
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in 1928. This was a concept which afterwards enjoyed some popularity:
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was a theropod reptile; later it would be typically assigned to the
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Geological Survey of Canada Contributions to Canadian Palaeontology
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Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed
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by Thomas Carr and Tom Williamson. Another partial skeleton from
650:: the small tooth shown in figures 41-45 is ANSP 9535, the later 463:. These teeth almost certainly belong to juvenile tyrannosaurine 5884: 4511: 460: 5944: 5382: 4628: 3822: 3137: 2317: 1893: 1778: 1740: 1409:
Proceedings of the Academy of Natural Sciences of Philadelphia
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Proceedings of the Academy of Natural Sciences of Philadelphia
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Proceedings of the Academy of Natural Sciences of Philadelphia
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came from that genus. In October 2000, the type specimen of
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Marsh, O.C., 1892, "Notes on Mesozoic vertebrate fossils",
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In 1856 Joseph Leidy had named fourteen teeth collected by
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Illustrations of the teeth by Leidy and O. C. Marsh, 4:
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Bakker, R.T., Williams, M. & Currie, P.J., 1988, "
942:; the last represents either a juvenile individual of 492:. Most other "aublysodontine"-type teeth may be from 1663: 1661: 955:
The first skeletal material referred to an original
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when it was not yet known this group was toothless.
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revealed the presence of large predatory dinosaurs.
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Monographien zur Geologie und Palaeontologie. 596:chose three nonserrated teeth from the original 1326:Contributions to Geology, University of Wyoming 620:, "again", "backwards", "contrariwise", βλύζω, 1238:Proceedings of the Academy of Natural Sciences 669:was based on the same type, it was at first a 522:In the mid-to-late 19th century many dinosaur 1752: 566:turning up in many localities in the Western 8: 1648:Nopcsa, F., 1928, "The genera of reptiles", 1423: 1421: 1205: 1203: 1096:definition of the Aublysodontinae in 2001, " 1545: 1543: 1541: 1303: 1301: 1299: 792:Academy of Natural Sciences of Philadelphia 6061:Late Cretaceous dinosaurs of North America 5941: 5672: 5589: 5564: 5448: 5444: 5403: 5392: 5379: 5290: 5258: 5090: 5054: 5014: 4883: 4879: 4764: 4760: 4749: 4649: 4638: 4625: 4416: 4406: 4322: 4234: 4122: 4069: 3979: 3932: 3928: 3878: 3843: 3832: 3819: 3750: 3659: 3567: 3520: 3389: 3364: 3360: 3288: 3284: 3273: 3158: 3147: 3134: 3062: 2881: 2872: 2701: 2516: 2467: 2463: 2376: 2372: 2338: 2327: 2314: 2113: 1996: 1914: 1903: 1890: 1836: 1827: 1775: 1759: 1745: 1737: 1582:from the Upper Cretaceous of New Mexico", 778:, and it is likely that teeth referred to 127: 31: 1697: 1686:Zoological Journal of the Linnean Society 1680:Carr, T.D.; and Williamson, T.E. (2004). 1553:. Simon & Schuster, New York, pp 464 1213:. Bloomington: Indiana University Press. 1145:based on material probably belonging to 898:classified a fragmentary skeleton named 850:. In 1892 Marsh named two more species: 1377: 1375: 1373: 1371: 1185: 930:; these had originally been species of 1726:. Indiana Univ. Press, pp. 64–83. 1486:, Geological Society of London, pp 827 1361:. Carnegie Institution of Washington. 1197:"Aublysodon." Dodson, et al. Page 128. 888:. This probably represents a tooth of 1598:Carr, T.D., Williamson, T.E., 2010, " 608:. Leidy named these same three teeth 7: 6038:0385493E-FFFD-FFA4-25DE-FECE384885F9 526:were named for isolated teeth; such 439:in 1868. It is sometimes considered 1667:"Results," in Holtz (2001). Pg. 66. 1444:Hatcher, J.B., 1903, "Osteology of 1227:"Results," in Holtz (2001). Pg. 67. 421:, which has been dated to the late 1604:Journal of Vertebrate Paleontology 632:means "wonderful" or "strange" in 624:, "to spout", "to flow" and ὀδών, 25: 6091:Campanian genus first appearances 735:. The names were thus separated. 723:became an independent genus when 711:is a junior objective synonym of 402:(“backwards-flowing tooth") is a 5925: 5919: 5912: 5364: 5357: 5350: 5344: 4610: 4603: 4598: 4591: 4585: 3804: 3797: 3790: 3119: 3112: 3105: 3098: 2300: 2294: 1699:10.1111/j.1096-3642.2004.00130.x 1551:Predatory Dinosaurs of the World 1175:Timeline of tyrannosaur research 1159: 786:went missing when being sent by 156: 1025:Shanshanosaurus huoyanshanensis 993:. It was made a separate genus 796:Field Museum of Natural History 556:started revealing the bones of 1450:Memoirs of the Carnegie Museum 751:in 1902 referred the tooth to 435:, was named by paleontologist 1: 6076:Fossil taxa described in 1868 1620:Acta Palaeontologica Polonica 798:. Because of the presence of 681:was preoccupied by the snake 447:consists only of an isolated 1130:. Paul Sereno has also used 1100:and all taxa sharing a more 707:pointed out Cope's mistake; 646:of the original syntypes of 6096:Campanian genus extinctions 1381:Carr and Williamson (2004). 1309:American Journal of Science 1074:, a group today called the 1031:. It probably represents a 719:had been. However, in 1892 6112: 6081:Taxa named by Joseph Leidy 1029:Aublysodon huoyanshanensis 866:as they were found in the 765:, forgetting in that case 600:series of fourteen as the 577:in 1854 and 1855 from the 575:Ferdinand Vandeveer Hayden 27:Extinct genus of dinosaurs 5909: 5675: 5592: 5567: 5451: 5447: 5406: 5391: 5378: 5342: 5289: 5257: 5093: 5017: 4886: 4882: 4767: 4763: 4652: 4637: 4624: 4583: 4419: 4405: 4325: 4237: 4125: 3982: 3935: 3931: 3881: 3846: 3831: 3818: 3787: 3749: 3662: 3570: 3523: 3392: 3367: 3363: 3291: 3287: 3161: 3146: 3133: 3095: 3061: 2884: 2704: 2519: 2470: 2466: 2379: 2375: 2341: 2326: 2313: 2291: 2116: 1999: 1917: 1902: 1889: 1839: 1826: 1774: 1497:Палеонтологический журнал 967:in 1966 and described by 500:of other tyrannosaurids. 378: 371: 346: 341: 322: 315: 153:Scientific classification 151: 135: 126: 34: 1731:Mesozoic Vertebrate Life 1724:Mesozoic Vertebrate Life 1392:Paleontological Bulletin 671:junior objective synonym 6086:Paleontology in Montana 1584:Journal of Paleontology 1530:Journal of Paleontology 989:, applying the correct 703:a valid genus. In 1899 388:(Leidy, 1868) Hay, 1930 1600:Bistahieversor sealeyi 1141:is today considered a 1102:recent common ancestor 959:species was a partial 757:; Hay in 1930 renamed 738:The tooth-based taxon 662: 415:Judith River Formation 6020:Paleobiology Database 4318:Carcharodontosaurinae 4230:Carcharodontosauridae 1465:Huene, F. von, 1932, 924:Aublysodon novojilovi 920:Aublysodon lancinator 918:named three species: 886:Aublysodon explanatus 769:would have priority. 763:Ornithomimus mirandus 749:Lawrence Morris Lambe 725:Othniel Charles Marsh 642: 5294:Dubious coelurosaurs 4007:Metriacanthosaurinae 3975:Metriacanthosauridae 2257:Dubious neotheropods 1709:The Age of Dinosaurs 1564:Kyoryugaku Saizensen 1429:Bulletin of the AMNH 946:or a separate genus 928:Aublysodon lancensis 904:by Marsh in 1890 as 901:Ornithomimus grandis 856:Aublysodon cristatus 841:Aublysodon lateralis 413:known only from the 4331:Carcharodontosaurus 4118:Carcharodontosauria 979:Aublysodon molnaris 896:Friedrich von Huene 832:Aublysodon mirandus 806:, such as those of 729:Aublysodon mirandus 717:Aublysodon mirandus 709:Aublysodon horridus 693:Aublysodon horridus 667:Aublysodon mirandus 656:Aublysodon mirandus 610:Aublysodon mirandus 594:Edward Drinker Cope 508:Aublysodon mirandus 469:Aublysodon mirandus 432:Aublysodon mirandus 328:Aublysodon mirandus 5786:Archaeornithomimus 4307:Veterupristisaurus 4167:Veterupristisaurus 4023:Metriacanthosaurus 3963:Piatnitzkysauridae 3398:Eustreptospondylus 3308:Piatnitzkysauridae 3191:Chuandongocoelurus 2055:Notatesseraeraptor 1549:Paul, G.S., 1988, 1288:American Geologist 1104:with it than with 1017:and ultimately to 987:Aublysodon molnari 906:Aublysodon grandis 884:Cope 1876 into an 663: 548:. Even before the 18:Deinodon cristatus 6048: 6047: 6007:Open Tree of Life 5947:Taxon identifiers 5938: 5937: 5934: 5933: 5907: 5906: 5903: 5902: 5899: 5898: 5895: 5894: 5862: 5861: 5858: 5857: 5668:Macrocheiriformes 5585:Ornithomimosauria 5559:Maniraptoriformes 5552: 5551: 5398:Maniraptoromorpha 5385:Maniraptoromorpha 5374: 5373: 5340: 5339: 5336: 5335: 5332: 5331: 5285: 5284: 5281: 5280: 5263:Maniraptoromorpha 5253: 5252: 5249: 5248: 5245: 5244: 5241: 5240: 5237: 5236: 5002: 5001: 4867: 4866: 4821:Proceratosauridae 4773:Chingkankousaurus 4620: 4619: 4581: 4580: 4577: 4576: 4573: 4572: 4549: 4548: 4401: 4400: 4397: 4396: 4393: 4392: 4389: 4388: 4385: 4384: 4381: 4380: 4057: 4056: 4053: 4052: 3814: 3813: 3785: 3784: 3781: 3780: 3777: 3776: 3773: 3772: 3745: 3744: 3741: 3740: 3737: 3736: 3733: 3732: 3649: 3648: 3605:Vallibonavenatrix 3508: 3507: 3504: 3503: 3348: 3347: 3129: 3128: 3093: 3092: 3089: 3088: 3085: 3084: 3057: 3056: 3053: 3052: 3049: 3048: 3045: 3044: 3041: 3040: 2689: 2688: 2685: 2684: 2451: 2450: 2309: 2308: 2289: 2288: 2285: 2284: 2281: 2280: 2232: 2231: 1885: 1884: 1881: 1880: 1877: 1876: 1446:Haplocanthosaurus 1357:Hay, O.P., 1930, 1000:Tyrannosaurus rex 911:Deinodon horridus 891:Saurornitholestes 875:John Bell Hatcher 852:Aublysodon amplus 847:Deinodon horridus 733:Deinodon horridus 713:Deinodon horridus 689:Deinodon horridus 675:Deinodon horridus 665:Because the name 660:Deinodon horridus 606:Deinodon horridus 443:now, because the 395: 394: 389: 362: 311: 285: 16:(Redirected from 6103: 6041: 6040: 6028: 6027: 6015: 6014: 6002: 6001: 5989: 5988: 5987: 5974: 5973: 5972: 5942: 5929: 5923: 5916: 5772:Aepyornithomimus 5673: 5590: 5565: 5529:Sinocalliopteryx 5449: 5445: 5439:Neocoelurosauria 5404: 5393: 5380: 5368: 5361: 5354: 5348: 5291: 5259: 5228:Zhuchengtyrannus 5160:Thanatotheristes 5091: 5055: 5023:Appalachiosaurus 5015: 4884: 4880: 4874:Pantyrannosauria 4765: 4761: 4755:Tyrannosauroidea 4750: 4725:Xinjiangovenator 4650: 4639: 4626: 4614: 4607: 4602: 4595: 4589: 4417: 4407: 4323: 4243:Acrocanthosaurus 4235: 4123: 4070: 3996:Yangchuanosaurus 3980: 3933: 3929: 3879: 3860:Lourinhanosaurus 3844: 3833: 3820: 3808: 3801: 3794: 3751: 3717:Sigilmassasaurus 3660: 3617:Ceratosuchopsini 3568: 3521: 3466:Dubreuillosaurus 3390: 3373:Streptospondylus 3365: 3361: 3331:Piatnitzkysaurus 3289: 3285: 3274: 3159: 3148: 3135: 3123: 3116: 3109: 3102: 3063: 2898:Ekrixinatosaurus 2882: 2873: 2702: 2517: 2468: 2464: 2377: 2373: 2339: 2328: 2315: 2304: 2298: 2208:Procompsognathus 2114: 1997: 1915: 1904: 1891: 1837: 1828: 1821: 1820: 1776: 1761: 1754: 1747: 1738: 1703: 1701: 1668: 1665: 1656: 1646: 1640: 1629: 1623: 1616: 1610: 1596: 1590: 1576: 1570: 1560: 1554: 1547: 1536: 1526: 1520: 1509: 1503: 1493: 1487: 1480: 1474: 1463: 1457: 1442: 1436: 1425: 1416: 1405: 1399: 1388: 1382: 1379: 1366: 1355: 1349: 1339: 1333: 1322: 1316: 1305: 1294: 1284: 1278: 1267: 1261: 1250: 1244: 1234: 1228: 1225: 1214: 1207: 1198: 1195: 1169: 1167:Dinosaurs portal 1164: 1163: 1162: 1092:proposed a stem 916:Alan Jack Charig 826:Referred species 705:Oliver Perry Hay 455:states, western 425:age of the late 387: 357: 351: 325: 302: 295: 274: 269: 256: 243: 161: 160: 131: 121: 51: 40:Temporal range: 32: 21: 6111: 6110: 6106: 6105: 6104: 6102: 6101: 6100: 6051: 6050: 6049: 6044: 6036: 6031: 6023: 6018: 6010: 6005: 5997: 5992: 5983: 5982: 5977: 5968: 5967: 5962: 5949: 5939: 5930: 5917: 5891: 5854: 5835:Sinornithomimus 5757: 5700: 5662: 5579: 5573:Compsognathidae 5561: 5548: 5536:Sinosauropteryx 5468:Compsognathidae 5462: 5441: 5432: 5400: 5387: 5370: 5369: 5355: 5328: 5323:Shanyangosaurus 5316:Phaedrolosaurus 5277: 5233: 5199: 5165: 5144:Daspletosaurini 5138: 5111: 5086:Tyrannosaurinae 5080: 5059:Albertosaurinae 5051: 5049:Tyrannosauridae 5042: 5011: 5009:Eutyrannosauria 4998: 4876: 4863: 4844:Proceratosaurus 4815: 4786: 4757: 4746: 4737: 4703:Richardoestesia 4687:Chilantaisaurus 4646: 4633: 4616: 4615: 4596: 4569: 4545: 4505:Australovenator 4483: 4477:Chilantaisaurus 4455:Phuwiangvenator 4377: 4342:Giganotosaurini 4336: 4312: 4224: 4187:Chilantaisaurus 4172: 4112: 4066: 4049: 4001: 3969: 3925: 3916: 3894:Monolophosaurus 3875: 3866: 3840: 3827: 3810: 3809: 3795: 3769: 3729: 3702: 3668:Camarillasaurus 3645: 3611: 3557: 3551:Ostafrikasaurus 3529:Camarillasaurus 3517: 3500: 3450:Afrovenatorinae 3444: 3403: 3379: 3357: 3344: 3302: 3281: 3279:Megalosauroidea 3270: 3261: 3256:Vectaerovenator 3249:Szechuanosaurus 3227:Monolophosaurus 3213:Kaijiangosaurus 3206:Cryolophosaurus 3167:Calamospondylus 3155: 3142: 3125: 3124: 3117: 3110: 3103: 3081: 3037: 3018:Pycnonemosaurus 2960: 2871: 2865:Majungasaurinae 2850: 2792:Majungasaurinae 2786: 2781:Xenotarsosaurus 2717:Dryptosauroides 2698: 2681: 2633: 2605:Elaphrosaurinae 2599: 2506: 2492:Ornithomimoides 2460: 2447: 2420: 2369: 2360: 2335: 2322: 2305: 2299: 2277: 2252: 2228: 2156: 2103: 2069:Shuangbaisaurus 1993: 1984: 1911: 1898: 1873: 1853: 1847:Avemetatarsalia 1832:Avemetatarsalia 1822: 1807:Avemetatarsalia 1781: 1780: 1770: 1765: 1679: 1676: 1671: 1666: 1659: 1650:Palaeobiologica 1647: 1643: 1630: 1626: 1617: 1613: 1597: 1593: 1577: 1573: 1561: 1557: 1548: 1539: 1527: 1523: 1510: 1506: 1494: 1490: 1481: 1477: 1473:(1-2), pp 1-361 1464: 1460: 1443: 1439: 1426: 1419: 1406: 1402: 1389: 1385: 1380: 1369: 1365:(II), pp 1-1074 1356: 1352: 1340: 1336: 1323: 1319: 1306: 1297: 1285: 1281: 1268: 1264: 1251: 1247: 1235: 1231: 1226: 1217: 1208: 1201: 1196: 1187: 1183: 1165: 1160: 1158: 1155: 1076:Tyrannosauridae 1041: 1027:Dong 1977 into 983:Gregory S. Paul 965:Jordan, Montana 871:Lance Formation 828: 788:registered mail 628:, "tooth". The 585:as the species 520: 503:Apart from the 337: 331: 307: 301: 293: 279: 273: 271:Aublysodontinae 267: 258:Tyrannosauridae 254: 245:Eutyrannosauria 241: 155: 122: 120: 119: 118: 117: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 46: 45: 42:Late Cretaceous 38: 28: 23: 22: 15: 12: 11: 5: 6109: 6107: 6099: 6098: 6093: 6088: 6083: 6078: 6073: 6071:Tyrannosaurids 6068: 6063: 6053: 6052: 6046: 6045: 6043: 6042: 6029: 6016: 6003: 5990: 5975: 5959: 5957: 5951: 5950: 5945: 5936: 5935: 5932: 5931: 5911: 5910: 5908: 5905: 5904: 5901: 5900: 5897: 5896: 5893: 5892: 5890: 5889: 5888: 5887: 5881: 5872: 5870: 5864: 5863: 5860: 5859: 5856: 5855: 5853: 5852: 5845: 5838: 5831: 5824: 5817: 5810: 5803: 5800:Dromiceiomimus 5796: 5789: 5782: 5775: 5767: 5765: 5763:Ornithomimidae 5759: 5758: 5756: 5755: 5748: 5744:Paraxenisaurus 5740: 5732: 5725: 5718: 5710: 5708: 5706:Deinocheiridae 5702: 5701: 5699: 5698: 5695:Shenzhousaurus 5691: 5684: 5676: 5670: 5664: 5663: 5661: 5660: 5653: 5645: 5638: 5631: 5624: 5617: 5609: 5601: 5593: 5587: 5581: 5580: 5578: 5577: 5568: 5562: 5557: 5554: 5553: 5550: 5549: 5547: 5546: 5539: 5532: 5525: 5517: 5510: 5503: 5496: 5492:Beipiaognathus 5488: 5481: 5472: 5470: 5464: 5463: 5461: 5460: 5452: 5442: 5437: 5434: 5433: 5431: 5430: 5423: 5416: 5407: 5401: 5396: 5389: 5388: 5383: 5376: 5375: 5372: 5371: 5363: 5349: 5343: 5341: 5338: 5337: 5334: 5333: 5330: 5329: 5327: 5326: 5319: 5312: 5305: 5297: 5295: 5287: 5286: 5283: 5282: 5279: 5278: 5276: 5275: 5274: 5273: 5267: 5265: 5255: 5254: 5251: 5250: 5247: 5246: 5243: 5242: 5239: 5238: 5235: 5234: 5232: 5231: 5224: 5217: 5209: 5207: 5205:Tyrannosaurini 5201: 5200: 5198: 5197: 5190: 5183: 5175: 5173: 5171:Teratophoneini 5167: 5166: 5164: 5163: 5156: 5153:Daspletosaurus 5148: 5146: 5140: 5139: 5137: 5136: 5133:Qianzhousaurus 5129: 5121: 5119: 5113: 5112: 5110: 5109: 5102: 5094: 5088: 5082: 5081: 5079: 5078: 5071: 5063: 5061: 5052: 5047: 5044: 5043: 5041: 5040: 5033: 5030:Bistahieversor 5026: 5018: 5012: 5007: 5004: 5003: 5000: 4999: 4997: 4996: 4988: 4981: 4974: 4966: 4959: 4952: 4945: 4938: 4931: 4924: 4917: 4910: 4902: 4895: 4887: 4877: 4872: 4869: 4868: 4865: 4864: 4862: 4861: 4854: 4847: 4840: 4833: 4825: 4823: 4817: 4816: 4814: 4813: 4805: 4797: 4795: 4788: 4787: 4785: 4784: 4776: 4768: 4758: 4753: 4747: 4744:Tyrannoraptora 4742: 4739: 4738: 4736: 4735: 4728: 4721: 4714: 4706: 4699: 4691: 4683: 4676: 4668: 4661: 4653: 4647: 4642: 4635: 4634: 4629: 4622: 4621: 4618: 4617: 4609: 4590: 4584: 4582: 4579: 4578: 4575: 4574: 4571: 4570: 4568: 4567: 4566: 4565: 4559: 4557: 4551: 4550: 4547: 4546: 4544: 4543: 4536: 4529: 4522: 4515: 4508: 4501: 4493: 4491: 4489:Megaraptoridae 4485: 4484: 4482: 4481: 4473: 4465: 4458: 4451: 4444: 4436: 4428: 4420: 4414: 4403: 4402: 4399: 4398: 4395: 4394: 4391: 4390: 4387: 4386: 4383: 4382: 4379: 4378: 4376: 4375: 4368: 4361: 4354: 4351:Giganotosaurus 4346: 4344: 4338: 4337: 4335: 4334: 4326: 4320: 4314: 4313: 4311: 4310: 4303: 4296: 4289: 4282: 4275: 4268: 4261: 4254: 4246: 4238: 4232: 4226: 4225: 4223: 4222: 4214: 4206: 4199: 4191: 4182: 4180: 4178:Neovenatoridae 4174: 4173: 4171: 4170: 4163: 4160:Ulughbegsaurus 4156: 4148: 4141: 4134: 4126: 4120: 4114: 4113: 4111: 4110: 4102: 4094: 4086: 4078: 4076: 4067: 4062: 4059: 4058: 4055: 4054: 4051: 4050: 4048: 4047: 4040: 4033: 4026: 4019: 4011: 4009: 4003: 4002: 4000: 3999: 3992: 3988:Xuanhanosaurus 3983: 3977: 3971: 3970: 3968: 3967: 3959: 3955:Xuanhanosaurus 3951: 3944: 3941:Asfaltovenator 3936: 3926: 3921: 3918: 3917: 3915: 3914: 3906: 3902:Megalosauridae 3898: 3890: 3882: 3876: 3871: 3868: 3867: 3865: 3864: 3856: 3847: 3841: 3836: 3829: 3828: 3823: 3816: 3815: 3812: 3811: 3803: 3789: 3788: 3786: 3783: 3782: 3779: 3778: 3775: 3774: 3771: 3770: 3768: 3767: 3766: 3765: 3759: 3757: 3747: 3746: 3743: 3742: 3739: 3738: 3735: 3734: 3731: 3730: 3728: 3727: 3720: 3712: 3710: 3704: 3703: 3701: 3700: 3693: 3686: 3679: 3676:Ichthyovenator 3672: 3663: 3657: 3651: 3650: 3647: 3646: 3644: 3643: 3636: 3629: 3621: 3619: 3613: 3612: 3610: 3609: 3601: 3593: 3586: 3579: 3571: 3565: 3559: 3558: 3556: 3555: 3547: 3540: 3537:Cristatusaurus 3533: 3524: 3518: 3513: 3510: 3509: 3506: 3505: 3502: 3501: 3499: 3498: 3494:Poekilopleuron 3490: 3487:Piveteausaurus 3483: 3476: 3469: 3462: 3454: 3452: 3446: 3445: 3443: 3442: 3435: 3428: 3421: 3413: 3411: 3409:Megalosaurinae 3405: 3404: 3402: 3401: 3393: 3387: 3385:Megalosauridae 3381: 3380: 3378: 3377: 3368: 3358: 3353: 3350: 3349: 3346: 3345: 3343: 3342: 3338:Xuanhanosaurus 3334: 3327: 3320: 3312: 3310: 3304: 3303: 3301: 3300: 3297:Yunyangosaurus 3292: 3282: 3277: 3271: 3266: 3263: 3262: 3260: 3259: 3252: 3245: 3237: 3234:Pandoravenator 3230: 3223: 3220:Kayentavenator 3216: 3209: 3202: 3194: 3187: 3179: 3171: 3162: 3156: 3151: 3144: 3143: 3138: 3131: 3130: 3127: 3126: 3118: 3111: 3104: 3097: 3096: 3094: 3091: 3090: 3087: 3086: 3083: 3082: 3080: 3079: 3078: 3077: 3071: 3069: 3059: 3058: 3055: 3054: 3051: 3050: 3047: 3046: 3043: 3042: 3039: 3038: 3036: 3035: 3028: 3021: 3014: 3007: 2999: 2992: 2985: 2978: 2970: 2968: 2962: 2961: 2959: 2958: 2951: 2948:Skorpiovenator 2944: 2936: 2929: 2922: 2915: 2908: 2901: 2894: 2885: 2879: 2870: 2869: 2860: 2858: 2852: 2851: 2849: 2848: 2841: 2833: 2826: 2819: 2812: 2804: 2796: 2794: 2788: 2787: 2785: 2784: 2777: 2770: 2763: 2760:Spectrovenator 2756: 2749: 2742: 2735: 2728: 2721: 2713: 2705: 2699: 2694: 2691: 2690: 2687: 2686: 2683: 2682: 2680: 2679: 2672: 2665: 2658: 2651: 2643: 2641: 2635: 2634: 2632: 2631: 2624: 2617: 2609: 2607: 2601: 2600: 2598: 2597: 2594:Spinostropheus 2590: 2583: 2576: 2568: 2560: 2552: 2544: 2537: 2529: 2520: 2514: 2508: 2507: 2505: 2504: 2496: 2488: 2480: 2471: 2461: 2458:Abelisauroidea 2456: 2453: 2452: 2449: 2448: 2446: 2445: 2438: 2430: 2428: 2426:Ceratosauridae 2422: 2421: 2419: 2418: 2415:Saltriovenator 2411: 2403: 2396: 2392:Fosterovenator 2388: 2380: 2370: 2365: 2362: 2361: 2359: 2358: 2354:Lophostropheus 2350: 2342: 2336: 2331: 2324: 2323: 2318: 2311: 2310: 2307: 2306: 2293: 2292: 2290: 2287: 2286: 2283: 2282: 2279: 2278: 2276: 2275: 2268: 2260: 2258: 2254: 2253: 2251: 2250: 2249: 2248: 2242: 2240: 2234: 2233: 2230: 2229: 2227: 2226: 2219: 2215:Pterospondylus 2211: 2204: 2196: 2188: 2185:Lucianovenator 2181: 2174: 2166: 2164: 2158: 2157: 2155: 2154: 2147: 2140: 2133: 2126: 2117: 2111: 2109:Coelophysoidea 2105: 2104: 2102: 2101: 2094: 2086: 2079: 2072: 2065: 2058: 2051: 2048:Lophostropheus 2044: 2037: 2030: 2023: 2016: 2008: 2000: 1994: 1989: 1986: 1985: 1983: 1982: 1974: 1966: 1958: 1950: 1947:Erythrovenator 1943: 1935: 1927: 1918: 1912: 1907: 1900: 1899: 1894: 1887: 1886: 1883: 1882: 1879: 1878: 1875: 1874: 1872: 1871: 1870: 1869: 1863: 1861: 1855: 1854: 1852: 1851: 1850: 1849: 1840: 1834: 1824: 1823: 1819: 1818: 1809: 1800: 1794: 1788: 1779: 1772: 1771: 1766: 1764: 1763: 1756: 1749: 1741: 1735: 1734: 1727: 1720: 1704: 1692:(4): 479–523. 1675: 1672: 1670: 1669: 1657: 1641: 1624: 1611: 1591: 1571: 1569:: 92-119/75-99 1555: 1537: 1521: 1504: 1488: 1475: 1458: 1437: 1417: 1400: 1383: 1367: 1350: 1334: 1317: 1295: 1279: 1262: 1245: 1229: 1215: 1199: 1184: 1182: 1179: 1178: 1177: 1171: 1170: 1154: 1151: 1147:Daspletosaurus 1040: 1039:Classification 1037: 1020:Bistahieversor 1014:Daspletosaurus 827: 824: 804:Daspletosaurus 775:Daspletosaurus 745:Ornithomimidae 519: 516: 479:synapomorphies 474:Daspletosaurus 465:tyrannosaurids 393: 392: 391: 390: 376: 375: 369: 368: 364: 363: 344: 343: 339: 338: 332: 320: 319: 313: 312: 291: 287: 286: 265: 261: 260: 252: 248: 247: 239: 232: 231: 226: 219: 218: 213: 206: 205: 200: 193: 192: 187: 183: 182: 177: 173: 172: 167: 163: 162: 149: 148: 133: 132: 124: 123: 115: 114: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 52: 39: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 6108: 6097: 6094: 6092: 6089: 6087: 6084: 6082: 6079: 6077: 6074: 6072: 6069: 6067: 6064: 6062: 6059: 6058: 6056: 6039: 6034: 6030: 6026: 6021: 6017: 6013: 6008: 6004: 6000: 5995: 5991: 5986: 5980: 5976: 5971: 5965: 5961: 5960: 5958: 5956: 5952: 5948: 5943: 5928: 5924: 5922: 5915: 5886: 5882: 5880: 5876: 5875: 5874: 5873: 5871: 5869: 5865: 5851: 5850: 5846: 5844: 5843: 5842:Struthiomimus 5839: 5837: 5836: 5832: 5830: 5829: 5825: 5823: 5822: 5818: 5816: 5815: 5811: 5809: 5808: 5804: 5802: 5801: 5797: 5795: 5794: 5793:"Coelosaurus" 5790: 5788: 5787: 5783: 5781: 5780: 5776: 5774: 5773: 5769: 5768: 5766: 5764: 5760: 5754: 5753: 5749: 5746: 5745: 5741: 5738: 5737: 5733: 5731: 5730: 5726: 5724: 5723: 5719: 5717: 5716: 5712: 5711: 5709: 5707: 5703: 5697: 5696: 5692: 5690: 5689: 5688:Pelecanimimus 5685: 5683: 5682: 5678: 5677: 5674: 5671: 5669: 5665: 5659: 5658: 5654: 5651: 5650: 5649:Thecocoelurus 5646: 5644: 5643: 5639: 5637: 5636: 5632: 5630: 5629: 5628:Kinnareemimus 5625: 5623: 5622: 5618: 5615: 5614: 5610: 5607: 5606: 5602: 5600: 5599: 5595: 5594: 5591: 5588: 5586: 5582: 5575: 5574: 5570: 5569: 5566: 5563: 5560: 5555: 5545: 5544: 5540: 5538: 5537: 5533: 5531: 5530: 5526: 5523: 5522: 5518: 5516: 5515: 5511: 5509: 5508: 5507:Huaxiagnathus 5504: 5502: 5501: 5500:Compsognathus 5497: 5494: 5493: 5489: 5487: 5486: 5482: 5479: 5478: 5474: 5473: 5471: 5469: 5465: 5459: 5458: 5454: 5453: 5450: 5446: 5443: 5440: 5435: 5429: 5428: 5427:Ornitholestes 5424: 5422: 5421: 5417: 5414: 5413: 5409: 5408: 5405: 5402: 5399: 5394: 5390: 5386: 5381: 5377: 5367: 5362: 5360: 5353: 5347: 5325: 5324: 5320: 5318: 5317: 5313: 5311: 5310: 5306: 5304: 5303: 5299: 5298: 5296: 5292: 5288: 5271: 5270: 5269: 5268: 5266: 5264: 5260: 5256: 5230: 5229: 5225: 5223: 5222: 5221:Tyrannosaurus 5218: 5216: 5215: 5211: 5210: 5208: 5206: 5202: 5196: 5195: 5194:Teratophoneus 5191: 5189: 5188: 5184: 5182: 5181: 5177: 5176: 5174: 5172: 5168: 5162: 5161: 5157: 5155: 5154: 5150: 5149: 5147: 5145: 5141: 5135: 5134: 5130: 5128: 5127: 5123: 5122: 5120: 5118: 5114: 5108: 5107: 5103: 5101: 5100: 5096: 5095: 5092: 5089: 5087: 5083: 5077: 5076: 5072: 5070: 5069: 5068:Albertosaurus 5065: 5064: 5062: 5060: 5056: 5053: 5050: 5045: 5039: 5038: 5034: 5032: 5031: 5027: 5025: 5024: 5020: 5019: 5016: 5013: 5010: 5005: 4994: 4993: 4989: 4987: 4986: 4985:Xiongguanlong 4982: 4980: 4979: 4975: 4972: 4971: 4967: 4965: 4964: 4963:Suskityrannus 4960: 4958: 4957: 4956:Stokesosaurus 4953: 4951: 4950: 4949:Santanaraptor 4946: 4944: 4943: 4939: 4937: 4936: 4932: 4930: 4929: 4925: 4923: 4922: 4918: 4916: 4915: 4911: 4908: 4907: 4903: 4901: 4900: 4896: 4894: 4893: 4892:Alectrosaurus 4889: 4888: 4885: 4881: 4878: 4875: 4870: 4860: 4859: 4855: 4853: 4852: 4848: 4846: 4845: 4841: 4839: 4838: 4834: 4832: 4831: 4827: 4826: 4824: 4822: 4818: 4811: 4810: 4809:Tanycolagreus 4806: 4804: 4803: 4799: 4798: 4796: 4793: 4789: 4782: 4781: 4777: 4775: 4774: 4770: 4769: 4766: 4762: 4759: 4756: 4751: 4748: 4745: 4740: 4734: 4733: 4729: 4727: 4726: 4722: 4720: 4719: 4715: 4712: 4711: 4707: 4705: 4704: 4700: 4697: 4696: 4692: 4689: 4688: 4684: 4682: 4681: 4677: 4674: 4673: 4672:Bahariasaurus 4669: 4667: 4666: 4662: 4660: 4659: 4655: 4654: 4651: 4648: 4645: 4644:Coelurosauria 4640: 4636: 4632: 4631:Coelurosauria 4627: 4623: 4613: 4608: 4606: 4601: 4594: 4588: 4563: 4562: 4561: 4560: 4558: 4556: 4555:Coelurosauria 4552: 4542: 4541: 4537: 4535: 4534: 4530: 4528: 4527: 4523: 4521: 4520: 4516: 4514: 4513: 4509: 4507: 4506: 4502: 4500: 4499: 4495: 4494: 4492: 4490: 4486: 4480: 4478: 4474: 4472: 4470: 4466: 4464: 4463: 4459: 4457: 4456: 4452: 4450: 4449: 4445: 4442: 4441: 4437: 4434: 4433: 4432:Bahariasaurus 4429: 4427: 4426: 4422: 4421: 4418: 4415: 4412: 4408: 4404: 4374: 4373: 4369: 4367: 4366: 4362: 4360: 4359: 4355: 4353: 4352: 4348: 4347: 4345: 4343: 4339: 4333: 4332: 4328: 4327: 4324: 4321: 4319: 4315: 4309: 4308: 4304: 4302: 4301: 4297: 4295: 4294: 4290: 4288: 4287: 4283: 4281: 4280: 4276: 4274: 4273: 4272:Kelmayisaurus 4269: 4267: 4266: 4262: 4260: 4259: 4255: 4252: 4251: 4247: 4245: 4244: 4240: 4239: 4236: 4233: 4231: 4227: 4220: 4219: 4215: 4212: 4211: 4207: 4205: 4204: 4200: 4197: 4196: 4192: 4189: 4188: 4184: 4183: 4181: 4179: 4175: 4169: 4168: 4164: 4162: 4161: 4157: 4154: 4153: 4149: 4147: 4146: 4142: 4140: 4139: 4135: 4133: 4132: 4128: 4127: 4124: 4121: 4119: 4115: 4108: 4107: 4106:Saurophaganax 4103: 4100: 4099: 4095: 4092: 4091: 4087: 4085: 4084: 4080: 4079: 4077: 4075: 4071: 4068: 4065: 4060: 4046: 4045: 4041: 4039: 4038: 4037:Siamotyrannus 4034: 4032: 4031: 4027: 4025: 4024: 4020: 4018: 4017: 4013: 4012: 4010: 4008: 4004: 3998: 3997: 3993: 3990: 3989: 3985: 3984: 3981: 3978: 3976: 3972: 3965: 3964: 3960: 3957: 3956: 3952: 3950: 3949: 3945: 3943: 3942: 3938: 3937: 3934: 3930: 3927: 3924: 3923:Allosauroidea 3919: 3912: 3911: 3910:Spinosauridae 3907: 3904: 3903: 3899: 3896: 3895: 3891: 3889: 3888: 3884: 3883: 3880: 3877: 3874: 3869: 3862: 3861: 3857: 3854: 3853: 3849: 3848: 3845: 3842: 3839: 3834: 3830: 3826: 3821: 3817: 3807: 3802: 3800: 3793: 3763: 3762: 3761: 3760: 3758: 3756: 3752: 3748: 3726: 3725: 3721: 3719: 3718: 3714: 3713: 3711: 3709: 3705: 3699: 3698: 3694: 3692: 3691: 3687: 3685: 3684: 3680: 3678: 3677: 3673: 3670: 3669: 3665: 3664: 3661: 3658: 3656: 3655:Spinosaurinae 3652: 3642: 3641: 3637: 3635: 3634: 3633:Riparovenator 3630: 3628: 3627: 3626:Ceratosuchops 3623: 3622: 3620: 3618: 3614: 3607: 3606: 3602: 3599: 3598: 3594: 3592: 3591: 3590:Riojavenatrix 3587: 3585: 3584: 3580: 3578: 3577: 3573: 3572: 3569: 3566: 3564: 3563:Baryonychinae 3560: 3553: 3552: 3548: 3546: 3545: 3541: 3539: 3538: 3534: 3531: 3530: 3526: 3525: 3522: 3519: 3516: 3515:Spinosauridae 3511: 3496: 3495: 3491: 3489: 3488: 3484: 3482: 3481: 3477: 3475: 3474: 3470: 3468: 3467: 3463: 3461: 3460: 3456: 3455: 3453: 3451: 3447: 3441: 3440: 3439:Wiehenvenator 3436: 3434: 3433: 3429: 3427: 3426: 3422: 3420: 3419: 3415: 3414: 3412: 3410: 3406: 3400: 3399: 3395: 3394: 3391: 3388: 3386: 3382: 3375: 3374: 3370: 3369: 3366: 3362: 3359: 3356: 3351: 3340: 3339: 3335: 3333: 3332: 3328: 3326: 3325: 3321: 3319: 3318: 3314: 3313: 3311: 3309: 3305: 3299: 3298: 3294: 3293: 3290: 3286: 3283: 3280: 3275: 3272: 3269: 3264: 3258: 3257: 3253: 3251: 3250: 3246: 3243: 3242: 3238: 3236: 3235: 3231: 3229: 3228: 3224: 3222: 3221: 3217: 3215: 3214: 3210: 3208: 3207: 3203: 3200: 3199: 3195: 3193: 3192: 3188: 3185: 3184: 3180: 3177: 3176: 3175:Chienkosaurus 3172: 3169: 3168: 3164: 3163: 3160: 3157: 3154: 3149: 3145: 3141: 3136: 3132: 3122: 3115: 3108: 3101: 3075: 3074: 3073: 3072: 3070: 3068: 3064: 3060: 3034: 3033: 3029: 3027: 3026: 3022: 3020: 3019: 3015: 3013: 3012: 3008: 3005: 3004: 3000: 2998: 2997: 2993: 2991: 2990: 2986: 2984: 2983: 2979: 2977: 2976: 2972: 2971: 2969: 2967: 2966:Furileusauria 2963: 2957: 2956: 2952: 2950: 2949: 2945: 2942: 2941: 2940:Rahiolisaurus 2937: 2935: 2934: 2930: 2928: 2927: 2923: 2921: 2920: 2916: 2914: 2913: 2909: 2907: 2906: 2902: 2900: 2899: 2895: 2892: 2891: 2887: 2886: 2883: 2880: 2878: 2874: 2867: 2866: 2862: 2861: 2859: 2857: 2856:Carnotaurinae 2853: 2847: 2846: 2842: 2839: 2838: 2837:Rahiolisaurus 2834: 2832: 2831: 2830:Majungasaurus 2827: 2825: 2824: 2820: 2818: 2817: 2813: 2810: 2809: 2805: 2803: 2802: 2798: 2797: 2795: 2793: 2789: 2783: 2782: 2778: 2776: 2775: 2771: 2769: 2768: 2767:Tarascosaurus 2764: 2762: 2761: 2757: 2755: 2754: 2750: 2748: 2747: 2743: 2741: 2740: 2736: 2734: 2733: 2729: 2727: 2726: 2725:Eoabelisaurus 2722: 2719: 2718: 2714: 2712: 2711: 2710:Chenanisaurus 2707: 2706: 2703: 2700: 2697: 2696:Abelisauridae 2692: 2678: 2677: 2673: 2671: 2670: 2666: 2664: 2663: 2659: 2657: 2656: 2655:Masiakasaurus 2652: 2650: 2649: 2645: 2644: 2642: 2640: 2636: 2630: 2629: 2625: 2623: 2622: 2621:Huinculsaurus 2618: 2616: 2615: 2614:Elaphrosaurus 2611: 2610: 2608: 2606: 2602: 2596: 2595: 2591: 2589: 2588: 2584: 2582: 2581: 2577: 2574: 2573: 2569: 2566: 2565: 2561: 2558: 2557: 2553: 2550: 2549: 2545: 2543: 2542: 2538: 2535: 2534: 2533:Austrocheirus 2530: 2527: 2526: 2522: 2521: 2518: 2515: 2513: 2509: 2502: 2501: 2497: 2494: 2493: 2489: 2486: 2485: 2481: 2478: 2477: 2473: 2472: 2469: 2465: 2462: 2459: 2454: 2444: 2443: 2439: 2437: 2436: 2432: 2431: 2429: 2427: 2423: 2417: 2416: 2412: 2409: 2408: 2404: 2402: 2401: 2397: 2394: 2393: 2389: 2387: 2386: 2385:Berberosaurus 2382: 2381: 2378: 2374: 2371: 2368: 2363: 2356: 2355: 2351: 2349: 2348: 2344: 2343: 2340: 2337: 2334: 2329: 2325: 2321: 2316: 2312: 2303: 2297: 2274: 2273: 2272:Halticosaurus 2269: 2267: 2266: 2265:Teinurosaurus 2262: 2261: 2259: 2255: 2246: 2245: 2244: 2243: 2241: 2239: 2235: 2225: 2224: 2220: 2217: 2216: 2212: 2210: 2209: 2205: 2202: 2201: 2197: 2194: 2193: 2192:Megapnosaurus 2189: 2187: 2186: 2182: 2180: 2179: 2175: 2173: 2172: 2168: 2167: 2165: 2163: 2162:Coelophysidae 2159: 2153: 2152: 2151:Powellvenator 2148: 2146: 2145: 2141: 2139: 2138: 2134: 2132: 2131: 2127: 2124: 2123: 2119: 2118: 2115: 2112: 2110: 2106: 2100: 2099: 2095: 2092: 2091: 2087: 2085: 2084: 2080: 2078: 2077: 2073: 2071: 2070: 2066: 2064: 2063: 2059: 2057: 2056: 2052: 2050: 2049: 2045: 2043: 2042: 2041:Liliensternus 2038: 2036: 2035: 2031: 2029: 2028: 2024: 2022: 2021: 2017: 2014: 2013: 2012:Dolichosuchus 2009: 2007: 2006: 2005:Dilophosaurus 2002: 2001: 1998: 1995: 1992: 1987: 1980: 1979: 1978:Herrerasauria 1975: 1972: 1971: 1967: 1964: 1963: 1959: 1956: 1955: 1951: 1949: 1948: 1944: 1941: 1940: 1936: 1933: 1932: 1931:Daemonosaurus 1928: 1925: 1924: 1920: 1919: 1916: 1913: 1910: 1905: 1901: 1897: 1892: 1888: 1867: 1866: 1865: 1864: 1862: 1860: 1856: 1848: 1844: 1843: 1842: 1841: 1838: 1835: 1833: 1829: 1825: 1817: 1813: 1810: 1808: 1804: 1801: 1799: 1795: 1793: 1789: 1787: 1783: 1782: 1777: 1773: 1769: 1762: 1757: 1755: 1750: 1748: 1743: 1742: 1739: 1732: 1728: 1725: 1721: 1718: 1717:0-7853-0443-6 1714: 1710: 1705: 1700: 1695: 1691: 1687: 1683: 1678: 1677: 1673: 1664: 1662: 1658: 1654: 1651: 1645: 1642: 1638: 1635:, New Series 1634: 1628: 1625: 1621: 1615: 1612: 1608: 1605: 1601: 1595: 1592: 1588: 1585: 1581: 1575: 1572: 1568: 1565: 1559: 1556: 1552: 1546: 1544: 1542: 1538: 1534: 1531: 1525: 1522: 1518: 1514: 1508: 1505: 1501: 1498: 1492: 1489: 1485: 1479: 1476: 1472: 1468: 1462: 1459: 1455: 1451: 1447: 1441: 1438: 1434: 1430: 1424: 1422: 1418: 1414: 1410: 1404: 1401: 1397: 1393: 1387: 1384: 1378: 1376: 1374: 1372: 1368: 1364: 1360: 1354: 1351: 1347: 1344: 1338: 1335: 1331: 1327: 1321: 1318: 1314: 1310: 1304: 1302: 1300: 1296: 1292: 1289: 1283: 1280: 1276: 1272: 1266: 1263: 1259: 1255: 1249: 1246: 1242: 1239: 1233: 1230: 1224: 1222: 1220: 1216: 1212: 1206: 1204: 1200: 1194: 1192: 1190: 1186: 1180: 1176: 1173: 1172: 1168: 1157: 1152: 1150: 1148: 1144: 1140: 1135: 1133: 1129: 1125: 1121: 1117: 1116:Tyrannosaurus 1113: 1109: 1107: 1106:Tyrannosaurus 1103: 1099: 1095: 1091: 1087: 1083: 1079: 1077: 1073: 1072:Deinodontidae 1069: 1065: 1061: 1057: 1053: 1049: 1045: 1038: 1036: 1034: 1030: 1026: 1022: 1021: 1016: 1015: 1010: 1006: 1002: 1001: 996: 992: 988: 984: 980: 976: 975: 974:Tyrannosaurus 970: 966: 963:unearthed in 962: 958: 953: 951: 950: 945: 944:Tyrannosaurus 941: 940: 935: 934: 929: 925: 921: 917: 913: 912: 907: 903: 902: 897: 893: 892: 887: 883: 881: 876: 872: 869: 868:Maastrichtian 865: 861: 857: 853: 849: 848: 843: 842: 837: 833: 825: 823: 821: 820: 815: 811: 810: 809:Tyrannosaurus 805: 801: 797: 793: 789: 785: 781: 777: 776: 770: 768: 764: 760: 756: 755: 754:Struthiomimus 750: 746: 741: 736: 734: 730: 726: 722: 718: 714: 710: 706: 702: 698: 694: 690: 686: 685: 680: 676: 672: 668: 661: 657: 653: 649: 645: 641: 637: 635: 631: 630:specific name 627: 623: 619: 615: 611: 607: 603: 599: 595: 591: 589: 584: 580: 576: 571: 569: 568:United States 565: 561: 560: 559:Tyrannosaurus 555: 554:North America 551: 547: 546: 541: 540: 539:Palaeoscincus 535: 534: 529: 525: 517: 515: 513: 509: 506: 501: 499: 498:sexual morphs 495: 491: 487: 484: 480: 476: 475: 470: 466: 462: 458: 454: 450: 446: 445:type specimen 442: 438: 434: 433: 428: 424: 420: 416: 412: 409: 405: 401: 400: 386: 384: 380: 379: 377: 374: 370: 367: 366:and see text 361: 356: 355: 348: 347: 345: 340: 335: 330: 329: 321: 318: 314: 310: 305: 300: 299: 292: 289: 288: 284: 283: 277: 272: 266: 263: 262: 259: 253: 250: 249: 246: 240: 237: 234: 233: 230: 227: 224: 221: 220: 217: 214: 211: 208: 207: 204: 201: 198: 195: 194: 191: 188: 185: 184: 181: 178: 175: 174: 171: 168: 165: 164: 159: 154: 150: 147: 143: 139: 134: 130: 125: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 50: 43: 37: 33: 30: 19: 6066:Nomina dubia 5954: 5918: 5847: 5840: 5833: 5826: 5819: 5814:Ornithomimus 5812: 5805: 5798: 5791: 5784: 5777: 5770: 5752:Tyrannomimus 5750: 5742: 5734: 5727: 5722:Deinocheirus 5720: 5713: 5693: 5686: 5679: 5655: 5647: 5642:Nqwebasaurus 5640: 5635:Nedcolbertia 5633: 5626: 5619: 5613:Calamosaurus 5611: 5605:Aviatyrannis 5603: 5596: 5571: 5541: 5534: 5527: 5519: 5512: 5505: 5498: 5490: 5485:Aristosuchus 5483: 5477:Aniksosaurus 5475: 5457:Aniksosaurus 5455: 5425: 5418: 5410: 5356: 5321: 5314: 5307: 5300: 5226: 5219: 5212: 5192: 5185: 5180:Dynamoterror 5178: 5158: 5151: 5131: 5124: 5104: 5099:Asiatyrannus 5097: 5073: 5066: 5037:Dryptosaurus 5035: 5028: 5021: 4990: 4983: 4976: 4968: 4961: 4954: 4947: 4940: 4933: 4928:Jinbeisaurus 4926: 4919: 4912: 4904: 4899:Aviatyrannis 4897: 4890: 4856: 4851:Sinotyrannus 4849: 4842: 4835: 4828: 4807: 4800: 4778: 4771: 4730: 4723: 4716: 4708: 4701: 4693: 4685: 4680:Bicentenaria 4678: 4670: 4665:Asiamericana 4663: 4656: 4597: 4538: 4531: 4524: 4517: 4510: 4503: 4496: 4475: 4467: 4460: 4453: 4446: 4440:Deltadromeus 4438: 4430: 4423: 4372:Tyrannotitan 4370: 4363: 4356: 4349: 4329: 4305: 4300:Taurovenator 4298: 4291: 4284: 4279:Lajasvenator 4277: 4270: 4263: 4258:Concavenator 4256: 4248: 4241: 4216: 4208: 4201: 4193: 4185: 4165: 4158: 4150: 4143: 4136: 4129: 4104: 4096: 4088: 4081: 4074:Allosauridae 4042: 4035: 4030:Shidaisaurus 4028: 4021: 4014: 3994: 3986: 3961: 3953: 3946: 3939: 3908: 3900: 3892: 3885: 3858: 3850: 3838:Avetheropoda 3825:Avetheropoda 3796: 3755:Avetheropoda 3722: 3715: 3708:Spinosaurini 3695: 3688: 3681: 3674: 3666: 3638: 3631: 3624: 3603: 3595: 3588: 3583:Protathlitis 3581: 3574: 3549: 3542: 3535: 3527: 3492: 3485: 3478: 3473:Leshansaurus 3471: 3464: 3457: 3437: 3430: 3425:Megalosaurus 3423: 3418:Duriavenator 3416: 3396: 3371: 3355:Megalosauria 3336: 3329: 3324:Marshosaurus 3322: 3317:Condorraptor 3315: 3295: 3254: 3247: 3239: 3232: 3225: 3218: 3211: 3204: 3198:Cruxicheiros 3196: 3189: 3181: 3173: 3165: 3030: 3025:Quilmesaurus 3023: 3016: 3009: 3001: 2994: 2987: 2980: 2973: 2953: 2946: 2938: 2931: 2924: 2917: 2910: 2903: 2896: 2888: 2877:Brachyrostra 2863: 2843: 2835: 2828: 2821: 2814: 2806: 2799: 2779: 2774:Tralkasaurus 2772: 2765: 2758: 2751: 2744: 2737: 2730: 2723: 2715: 2708: 2676:Vespersaurus 2674: 2669:Velocisaurus 2667: 2660: 2653: 2646: 2626: 2619: 2612: 2592: 2585: 2578: 2570: 2564:Deltadromeus 2562: 2554: 2548:Compsosuchus 2546: 2539: 2531: 2523: 2498: 2490: 2484:Lametasaurus 2482: 2474: 2440: 2435:Ceratosaurus 2433: 2413: 2405: 2398: 2390: 2383: 2367:Ceratosauria 2352: 2345: 2270: 2263: 2221: 2213: 2206: 2198: 2190: 2183: 2176: 2169: 2149: 2144:Podokesaurus 2142: 2135: 2128: 2120: 2096: 2088: 2081: 2074: 2067: 2060: 2053: 2046: 2039: 2032: 2027:Gojirasaurus 2025: 2020:Dracovenator 2018: 2010: 2003: 1991:Neotheropoda 1976: 1968: 1960: 1954:Guaibasaurus 1952: 1945: 1937: 1929: 1923:Chindesaurus 1921: 1811: 1802: 1730: 1723: 1708: 1689: 1685: 1652: 1649: 1644: 1636: 1632: 1627: 1619: 1614: 1606: 1603: 1599: 1594: 1586: 1583: 1579: 1574: 1566: 1563: 1558: 1550: 1532: 1529: 1524: 1516: 1513:Nanotyrannus 1512: 1507: 1499: 1496: 1491: 1483: 1478: 1470: 1466: 1461: 1453: 1449: 1445: 1440: 1432: 1428: 1412: 1408: 1403: 1395: 1391: 1386: 1362: 1358: 1353: 1345: 1342: 1337: 1332:(2): 123-134 1329: 1325: 1320: 1312: 1308: 1290: 1287: 1282: 1274: 1270: 1265: 1257: 1253: 1248: 1240: 1237: 1232: 1210: 1146: 1143:nomen dubium 1142: 1138: 1136: 1131: 1128:anchor taxon 1123: 1119: 1115: 1111: 1110: 1105: 1097: 1090:Thomas Holtz 1086:Franz Nopcsa 1081: 1080: 1067: 1059: 1047: 1042: 1032: 1028: 1024: 1018: 1012: 1008: 998: 995:Stygivenator 994: 986: 978: 972: 969:Ralph Molnar 956: 954: 949:Nanotyrannus 947: 943: 937: 931: 927: 923: 919: 909: 905: 899: 889: 885: 878: 863: 859: 855: 851: 845: 839: 835: 831: 829: 819:nomen dubium 817: 807: 803: 799: 783: 779: 773: 771: 766: 762: 758: 752: 739: 737: 732: 728: 720: 716: 712: 708: 700: 696: 692: 688: 687:and renamed 682: 678: 674: 666: 664: 659: 655: 647: 625: 621: 617: 609: 605: 586: 581:Badlands of 579:Judith River 572: 557: 543: 537: 531: 521: 511: 507: 505:type species 502: 472: 468: 449:premaxillary 437:Joseph Leidy 431: 430: 398: 397: 396: 383:Ornithomimus 381: 365: 360:nomen dubium 359: 358:Leidy, 1868 353: 352: 327: 326: 317:Type species 309:nomen dubium 308: 297: 296: 282:nomen dubium 280: 270: 235: 222: 209: 196: 145: 141: 137: 35: 29: 5979:Wikispecies 5879:Maniraptora 5868:Maniraptora 5849:Tototlmimus 5779:Anserimimus 5729:Garudimimus 5715:Beishanlong 5657:Valdoraptor 5598:Arkansaurus 5543:Xunmenglong 5420:Juravenator 5214:Tarbosaurus 5106:Nanuqsaurus 5075:Gorgosaurus 4992:Megaraptora 4978:Timurlengia 4710:Sciurumimus 4658:Aratasaurus 4526:Murusraptor 4448:Fukuiraptor 4411:Megaraptora 4293:Shaochilong 4265:Eocarcharia 4218:Megaraptora 4138:Lusovenator 4016:Alpkarakush 3873:Carnosauria 3724:Spinosaurus 3697:Siamosaurus 3597:Suchosaurus 3544:Iberospinus 3480:Magnosaurus 3459:Afrovenator 3432:Torvosaurus 3183:Chilesaurus 2996:Carnotaurus 2989:Caletodraco 2975:Abelisaurus 2801:Arcovenator 2648:Laevisuchus 2639:Noasaurinae 2541:Berthasaura 2512:Noasauridae 2442:Genyodectes 2407:Lukousaurus 2400:Jubbulpuria 2200:Panguraptor 2178:Coelophysis 2171:Camposaurus 2130:Panguraptor 2122:Dracoraptor 2098:Zupaysaurus 2083:Tachiraptor 2062:Sarcosaurus 1962:Nhandumirim 1622:48: 227–234 1589:: 1026-1032 1502:(3): 95-109 1033:Tarbosaurus 939:Tarbosaurus 933:Gorgosaurus 836:A. horridus 830:Apart from 814:Thomas Carr 759:A. mirandus 644:Lithography 512:A. mirandus 494:ontogenetic 408:carnivorous 354:A. mirandus 264:Subfamily: 138:A. mirandus 6055:Categories 5985:Aublysodon 5955:Aublysodon 5807:Gallimimus 5736:Harpymimus 5681:Harpymimus 5302:Iliosuchus 5272:see below↓ 5117:Alioramini 4935:Juratyrant 4921:Eotyrannus 4906:Bagaraatan 4858:Yutyrannus 4792:Coeluridae 4718:Vayuraptor 4564:see below↓ 4540:Tratayenia 4533:Orkoraptor 4519:Megaraptor 4425:Aoniraptor 4358:Mapusaurus 4286:Sauroniops 4250:Altispinax 4203:Neovenator 4145:Siamraptor 4131:Datanglong 4098:Epanterias 4090:Antrodemus 4083:Allosaurus 4064:Allosauria 3887:Altispinax 3852:Gasosaurus 3764:see below↓ 3640:Suchomimus 3241:Sinosaurus 3076:see below↓ 3032:Viavenator 2982:Aucasaurus 2919:Ilokelesia 2890:Dahalokely 2845:Rajasaurus 2823:Indosaurus 2816:Genusaurus 2808:Dahalokely 2732:Indosuchus 2628:Limusaurus 2587:Ligabueino 2580:Kiyacursor 2572:Genusaurus 2556:Dahalokely 2476:Betasuchus 2347:Dornraptor 2247:see below↓ 2223:Segisaurus 2076:Sinosaurus 1939:Eodromaeus 1868:see below↓ 1816:Dinosauria 1798:Sauropsida 1674:References 1639:(1): 1-252 1580:Aublysodon 1535:(1): 73-82 1519:1(5): 1-30 1348:(2): 25-81 1139:Aublysodon 1132:Aublysodon 1124:Aublysodon 1120:Aublysodon 1112:Aublysodon 1098:Aublysodon 1082:Aublysodon 1068:Aublysodon 1060:Aublysodon 1048:Aublysodon 1035:specimen. 1009:Aublysodon 1005:New Mexico 957:Aublysodon 914:. In 1967 894:. In 1932 882:explanatus 873:. In 1903 800:Aublysodon 784:Aublysodon 780:Aublysodon 767:Aublysodon 740:Aublysodon 721:Aublysodon 715:, just as 701:Aublysodon 602:lectotypes 592:. In 1866 496:stages or 483:postmortem 427:Cretaceous 399:Aublysodon 298:Aublysodon 216:Saurischia 203:Dinosauria 36:Aublysodon 5883:includes 5828:Rativates 5821:Qiupalong 5521:Scipionyx 5514:Mirischia 5187:Lythronax 5126:Alioramus 4780:Labocania 4498:Aerosteon 4044:Sinraptor 3948:Erectopus 3683:Irritator 3268:Orionides 3153:Tetanurae 3140:Tetanurae 3067:Tetanurae 3003:Elemgasem 2926:Llukalkan 2905:Elemgasem 2662:Noasaurus 2525:Afromimus 2333:Averostra 2320:Averostra 2238:Averostra 2090:Velocipes 1909:Theropoda 1896:Theropoda 1859:Theropoda 1784:Kingdom: 1768:Theropoda 1655:: 163-188 1609:(1): 1-16 1456:(1): 1-72 1415:: 248-261 1315:: 170-176 1293:: 345-349 1277:: 233-242 1260:: 275-279 1181:Footnotes 1122:", using 1056:Theropoda 1052:Goniopoda 1046:assigned 790:from the 652:lectotype 614:etymology 533:Trachodon 490:digestion 423:Campanian 411:dinosaurs 229:Theropoda 176:Kingdom: 170:Eukaryota 146:cristatus 5964:Wikidata 5412:Coelurus 4837:Kileskus 4830:Guanlong 4802:Coelurus 4695:Gualicho 4195:Gualicho 3576:Baryonyx 2912:Guemesia 2739:Kryptops 2500:Ozraptor 2137:Pendraig 1792:Chordata 1790:Phylum: 1786:Animalia 1517:Hunteria 1153:See also 1137:Because 991:genitive 877:renamed 860:Deinodon 697:Deinodon 691:into an 679:Deinodon 648:Deinodon 590:horridus 588:Deinodon 550:badlands 530:include 486:abrasion 385:mirandus 373:Synonyms 342:Species 251:Family: 190:Chordata 186:Phylum: 180:Animalia 166:Domain: 144:6: "A." 140:5: "A." 47:75  6012:4126365 5999:4821751 5970:Q132625 4970:Timimus 4732:Zuolong 4462:Rapator 4365:Meraxes 3690:Oxalaia 3011:Koleken 2034:Lepidus 1796:Class: 1243:(2): 72 1050:to the 880:Laelaps 794:to the 684:Dinodon 598:syntype 583:Montana 545:Troodon 518:History 441:dubious 419:Montana 306:, 1868 290:Genus: 278:, 1928 116:↓ 5621:Hexing 5309:Kakuru 4914:Dilong 2955:Thanos 2933:Niebla 2753:Rugops 2746:Kurupi 1715:  1398:: 1-14 1126:as an 1064:mammal 864:T. rex 542:, and 528:genera 459:, and 457:Canada 336:, 1868 276:Nopcsa 142:amplus 6033:Plazi 6025:38614 5885:birds 4942:Moros 4469:Siats 4210:Siats 4152:Siats 1812:Clade 1803:Clade 1094:clade 961:skull 761:into 695:. If 634:Latin 622:blyzo 564:teeth 488:, or 404:genus 334:Leidy 304:Leidy 236:Clade 223:Clade 210:Clade 197:Clade 5994:GBIF 5877:see 4512:Maip 1970:Tawa 1845:see 1713:ISBN 1567:9/10 1500:1965 1435:(6). 1118:and 1044:Cope 926:and 854:and 834:and 626:odon 524:taxa 461:Asia 55:PreꞒ 1694:doi 1690:142 1433:147 1363:390 1108:". 981:by 673:of 654:of 604:of 552:of 417:in 406:of 6057:: 6035:: 6022:: 6009:: 5996:: 5981:: 5966:: 1814:: 1805:: 1688:. 1684:. 1660:^ 1637:14 1607:30 1587:64 1540:^ 1533:52 1452:, 1431:, 1420:^ 1413:28 1411:, 1396:22 1394:, 1370:^ 1330:20 1328:, 1313:44 1311:, 1298:^ 1291:24 1275:20 1273:, 1258:18 1256:, 1218:^ 1202:^ 1188:^ 1078:. 1062:a 952:. 922:, 822:. 636:. 618:au 562:, 536:, 514:. 453:US 238:: 225:: 212:: 199:: 105:Pg 49:Ma 44:, 5747:? 5739:? 5652:? 5616:? 5608:? 5576:? 5524:? 5495:? 5480:? 5415:? 4995:? 4973:? 4909:? 4812:? 4794:? 4783:? 4713:? 4698:? 4690:? 4675:? 4479:? 4471:? 4443:? 4435:? 4413:? 4253:? 4221:? 4213:? 4198:? 4190:? 4155:? 4109:? 4101:? 4093:? 3991:? 3966:? 3958:? 3913:? 3905:? 3897:? 3863:? 3855:? 3671:? 3608:? 3600:? 3554:? 3532:? 3497:? 3376:? 3341:? 3244:? 3201:? 3186:? 3178:? 3170:? 3006:? 2943:? 2893:? 2868:? 2840:? 2811:? 2720:? 2575:? 2567:? 2559:? 2551:? 2536:? 2528:? 2503:? 2495:? 2487:? 2479:? 2410:? 2395:? 2357:? 2218:? 2203:? 2195:? 2125:? 2093:? 2015:? 1981:? 1973:? 1965:? 1957:? 1942:? 1934:? 1926:? 1760:e 1753:t 1746:v 1719:. 1702:. 1696:: 1653:1 1471:4 1454:2 1346:3 1241:8 350:† 324:† 294:† 268:† 255:† 242:† 110:N 100:K 95:J 90:T 85:P 80:C 75:D 70:S 65:O 60:Ꞓ 20:)

Index

Deinodon cristatus
Late Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Saurischia
Theropoda
Eutyrannosauria
Tyrannosauridae
Aublysodontinae
Nopcsa
nomen dubium
Aublysodon

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