6061:
480:
matrix, that are recurved with a broader base and have a length of three millimetres. The authors identified lower jaws with a preserved length of fifteen millimetres. The cervical vertebrae are short and broad. Six dorsal vertebrae have been preserved, nine ribs and six belly ribs at the left side. The tail has a preserved length of five centimetres, but part of this is accounted for by a section that might have been added to enhance the value of the fossil. The tail vertebrae at the base, the authenticity of which is certain, are short.
5901:
4668:
118:
6049:
3470:
147:
3476:
2853:
4684:
5908:
2858:
2312:
5914:
2320:
2327:
4676:
869:
prey they were chasing. Its large eyes that were forward facing and the small claws on its appendages had researchers assuming that the species would match closely to tree bark or dark night colors to blend in with its surroundings and ambush insects for nocturnal hunting, just like what is seem with modern-day nightjars or bats.
868:
would have had great eyesight. They could use these features to quickly spot insects and be able to follow their motion while following behind them. With their smaller wingspan and skeletal body, along with very flexible joints, these traits gave them great movability to catch the small insects and
479:
had a wingspan of about forty centimetres, making it one of the smallest known pterosaurs. Of the type specimen, most parts of the skull have become detached so that its shape is difficult to determine, but it was generally short and broad. Eleven teeth have been preserved scattered throughout the
423:
who first prepared it. It consists of a near-complete skeleton of a subadult individual and is crushed. Most elements are present, exceptions include the sternum, the tail end, sacrals and the fourth phalanx of the wing finger. This specimen was originally thought to be from the
Jianshangou Bed of
491:
is 35.5 millimetres long. The metacarpals are short with seven millimetres length for the first three, 9.3 millimetres for the fourth wing-bearing metacarpal. The first three fingers are well developed with the first having an elongated first phalanx. They bear short but sharp claws. The first
449:
In 2010 a second specimen, of a juvenile, was announced, that proved that a more elongated tail was present after all, albeit not so long as the faked tail of the holotype: about 85% of femur length. This specimen eventually was designated as the holotype of a new species,
454:, by Hone and Lü in 2012. The specimen was originally stored in the Guilin Geological Museum and designated GLGMV 0002; later it was moved to the Jinzhou Paleontological Museum and designated JZMP-04-07-3. In 2020, Hone recognized that
492:
phalanx of the fourth, wing, finger has a length of 44.5, the second of 35.6 millimetres. The size of the third cannot be established because of damage. A short and slender pteroid, 5.9 millimetres long, points towards the elbow. The
520:, mainly in view of the general long bone proportions. It was in 2000 identified as an anurognathid, and it was confirmed that the fossil had been doctored prior to its description.
504:
is reduced, reaching about half-way downwards along the tibia shaft. The foot is long with the metatarsals having a length of 12.1 millimetres. The fifth toe is elongated.
5993:
353:
6096:
5980:
6116:
1313:
1127:"On the systematic and stratigraphic significance of pterosaurs from the Lower Cretaceous Yixian Formation (Jehol Group) of Liaoning, China"
553:, and proposed Batrachognathinae as a replacement name. An analysis by Lü and Qiang Ji in 2006 resolved the relations even further, finding
6121:
440:
was actually found in the Middle
Jurassic deposits; the authors note that all other Chinese anurognathids are Jurassic in age, and that
5900:
1211:
Lü, J.; Ji, Q. (2006). "Preliminary results of a phylogenetic analysis of the pterosaurs from western
Liaoning and surrounding areas".
428:, dated to about 124.6 million years old. However, the specimen was obtained by amateur collectors and altered to contain a piece of a
6111:
883:
146:
4667:
3475:
786:
6039:
3469:
745:
589:
member of this subfamily in Wei and colleagues' study. Below is a cladogram representing their phylogenetic analysis:
378:
446:
was also initially thought to be a
Cretaceous taxon until subsequent studies established it to be from the Jurassic.
992:
Lü, J.; Hone, D.W.E. (2012). "A New
Chinese Anurognathid Pterosaur and the Evolution of Pterosaurian Tail Lengths".
134:
6106:
2852:
1306:
1410:
878:
721:
549:. He later admitted Asiaticognathidae was an inappropriate name, as the clade's definition did not include an
2311:
6101:
6091:
6086:
2888:
383:
2857:
2319:
6081:
5933:
5808:
3966:
1220:
661:
610:
5971:
5907:
6019:
5913:
566:
245:
462:
as other species of anurognathids were from each other, and elevated it to a new genus and combination
4683:
4061:
2500:
2410:
2326:
1299:
949:
324:
4675:
1225:
6053:
5565:
5254:
4941:
4240:
4167:
1935:
1655:
769:
345:
117:
1164:(Pterosauria, Anurognathidae, Batrachognathinae) and the structure of the pterosaur wing membrane"
1158:
Kellner, A.W.A.; Wang, X.; Tischlinger, H.; de
Almeida Campos, D.; Hone, D.W.E.; Meng, X. (2010).
5094:
3568:
3411:
2554:
2357:
2344:
2258:
2100:
1731:
1706:
1051:
1009:
967:
644:
270:
141:
6024:
6006:
5998:
5520:
4335:
4264:
3506:
3493:
3436:
3199:
3097:
2999:
2917:
2672:
2568:
2109:
2025:
1977:
1951:
1664:
1569:
1277:
1193:
630:
546:
528:
517:
349:
6011:
5749:
5632:
5624:
5416:
5345:
5270:
4812:
4715:
4702:
4634:
4514:
4503:
3753:
3610:
3547:
3322:
3304:
3142:
2966:
2782:
2768:
2726:
2232:
2218:
2150:
1829:
1379:
1267:
1257:
1183:
1175:
1138:
1106:(Pterosauria: Anurognathidae) with a Long Tail and the Evolution of the Pterosaurian Tail".
1082:
1043:
1001:
957:
425:
376:, "snout" in reference to it being assumed a tree-dweller and presumed a close relative of
5838:
5681:
5530:
5315:
5219:
5194:
5187:
5174:
5107:
4993:
4891:
4857:
4753:
4733:
4587:
4445:
4386:
4373:
4344:
4220:
4131:
3990:
3863:
3818:
3690:
3619:
3598:
3424:
3085:
3065:
3037:
3013:
2949:
2941:
2934:
2926:
2681:
2661:
2491:
2467:
2417:
2225:
2208:
2198:
2175:
2131:
2123:
2091:
2080:
2059:
2013:
2005:
1944:
1860:
1800:
1719:
1688:
1578:
1557:
1510:
1395:
1370:
1032:"A review of the taxonomy and palaeoecology of the Anurognathidae (Reptilia, Pterosauria)"
541:
321:
30:
953:
5815:
5788:
5766:
5590:
5504:
5489:
5441:
5380:
5365:
5331:
5156:
5140:
5128:
5114:
5067:
5024:
4984:
4898:
4844:
4572:
4565:
4558:
4530:
4437:
4358:
4285:
4196:
3952:
3844:
3779:
3720:
3660:
3626:
3346:
3273:
3245:
3164:
3114:
3053:
3022:
2622:
2598:
2576:
2522:
2480:
2424:
2392:
2073:
1990:
1967:
1846:
1772:
1740:
1593:
1542:
1490:
1272:
1241:
1188:
1159:
1031:
683:
596:
574:
513:
433:
420:
340:
280:
204:
1071:"Evolutionary radiation of the Jehol Biota: chronological and ecological perspectives"
910:
Ji, S.-A.; Ji, Q. (1998). "A new fossil pterosaur (Rhamphorhynchoidea) from
Liaoning,
6075:
5852:
5823:
5799:
5780:
5738:
5723:
5672:
5661:
5554:
5546:
5496:
5433:
5404:
5388:
5372:
5358:
5299:
5277:
5228:
5121:
4882:
4825:
4744:
4579:
4551:
4523:
4394:
4277:
4068:
3998:
3870:
3853:
3709:
3646:
3590:
3355:
3311:
3192:
3185:
3178:
3171:
3156:
3106:
3030:
2987:
2980:
2906:
2753:
2746:
2739:
2588:
2530:
2515:
2508:
2456:
2239:
1908:
1868:
1818:
1787:
1748:
1643:
1635:
1613:
1517:
1501:
1055:
1013:
429:
68:
1246:, a new anurognathid pterosaur from the Jurassic of China and comments on the group"
6065:
5845:
5773:
5644:
5539:
5473:
5425:
5235:
5163:
5041:
5009:
4777:
4544:
4472:
4413:
4308:
3945:
3930:
3914:
3905:
3894:
3734:
3697:
3583:
3533:
3376:
3289:
3282:
3262:
3234:
3227:
3213:
3206:
3121:
3078:
3045:
2761:
2703:
2646:
2631:
2611:
2436:
2167:
2159:
2139:
2066:
2045:
2038:
1893:
1681:
1627:
1586:
971:
442:
361:
306:
577:, which is unlike the former studies where it was recovered as closely related to
5965:
5731:
5691:
5610:
5574:
5308:
5083:
5016:
4959:
4905:
4762:
4430:
4301:
4293:
4253:
4212:
4204:
4079:
4046:
4014:
4006:
3977:
3938:
3922:
3881:
3810:
3802:
3795:
3741:
3682:
3674:
3335:
3297:
3220:
2875:
2808:
2775:
2711:
2052:
1921:
1876:
1853:
1528:
43:
5956:
512:
Because of the presumed long tail, the authors rejected a placement within the
5699:
5654:
5582:
5457:
5449:
5284:
5263:
5208:
5076:
5051:
5001:
4966:
4912:
4800:
4792:
4785:
4537:
4460:
4452:
4351:
4315:
4153:
4054:
4033:
4022:
3959:
3576:
3369:
3362:
2797:
2688:
1695:
1352:
88:
53:
1143:
1126:
5759:
5201:
5149:
4871:
4864:
4832:
4770:
4496:
4489:
4481:
4401:
4179:
4145:
4095:
4088:
3825:
3763:
3667:
3255:
2654:
2371:
1998:
1838:
1673:
1550:
1472:
1459:
1422:
1361:
1322:
1240:
Wei, X.; Pêgas, R. V.; Shen, C.; Guo, Y.; Ma, W.; Sun, D.; Zhou, X. (2021).
1047:
1005:
586:
524:
464:
396:
357:
309:
191:
158:
93:
37:
1281:
1197:
1179:
5950:
5831:
5715:
5707:
5481:
5465:
5396:
4973:
4323:
4228:
4160:
3832:
3787:
3560:
2789:
2449:
1346:
409:
313:
178:
83:
78:
63:
58:
48:
1262:
5985:
484:
419:
originally discovered around 1995 and obtained by science from illegal
98:
73:
496:
has a length of 26.7 millimetres and is about a third longer than the
4422:
4189:
3771:
2379:
2246:
1340:
501:
487:
is robustly built but elongated with a length of 27 millimetres. The
416:
168:
5927:
1087:
1070:
864:
as an insectivore. With very large eye sockets and a rounded head,
3520:
2116:
962:
937:
536:
497:
493:
387:
332:
328:
302:
488:
5931:
4700:
3491:
2873:
2342:
1457:
1333:
1295:
1291:
860:
The describers postulated a tree-dwelling lifestyle for
364:. It was therefore renamed in 1999. The type species is
16:
Genus of anurognathid pterosaur from the Middle
Jurassic
436:
in 2012 considered it more likely that the holotype of
6037:
5940:
5798:
5748:
5680:
5671:
5643:
5608:
5564:
5529:
5519:
5415:
5344:
5329:
5298:
5253:
5218:
5173:
5139:
5093:
5066:
5039:
4983:
4939:
4881:
4843:
4811:
4752:
4743:
4713:
4633:
4513:
4471:
4412:
4372:
4334:
4263:
4239:
4178:
4129:
4078:
4032:
3976:
3904:
3880:
3852:
3843:
3752:
3718:
3707:
3644:
3609:
3546:
3531:
3504:
3435:
3410:
3345:
3321:
3272:
3244:
3140:
3096:
3064:
3012:
2997:
2964:
2916:
2886:
2807:
2725:
2671:
2621:
2597:
2586:
2552:
2490:
2466:
2435:
2390:
2355:
2257:
2207:
2197:
2149:
2099:
2090:
2024:
1976:
1965:
1934:
1919:
1891:
1828:
1798:
1770:
1730:
1705:
1663:
1654:
1611:
1568:
1527:
1500:
1470:
1421:
1394:
569:conducted by Xuefang Wei and colleagues recovered
394:, was described in 2012, and moved to a new genus
1102:Hone, D.W.E.; Lü, J. (2010). "A New Specimen of
938:"Biostratigraphy of new pterosaurs from China"
1307:
1125:Unwin, D.M.; Lü, J.; Bakhurina, N.N. (2000).
8:
516:and classified it instead as a long-tailed
5928:
5677:
5616:
5526:
5350:
5341:
5337:
5295:
5291:
5179:
5099:
5063:
5059:
4951:
4947:
4849:
4817:
4749:
4725:
4721:
4710:
4697:
4630:
4468:
4409:
4378:
4269:
4245:
4137:
4038:
3982:
3901:
3886:
3849:
3726:
3715:
3652:
3552:
3543:
3539:
3512:
3501:
3488:
3416:
3407:
3327:
3148:
3070:
3009:
3005:
2972:
2898:
2894:
2883:
2870:
2731:
2603:
2594:
2560:
2472:
2441:
2402:
2398:
2363:
2352:
2339:
2204:
2096:
2030:
1982:
1973:
1931:
1927:
1900:
1810:
1806:
1779:
1711:
1660:
1619:
1534:
1482:
1478:
1467:
1454:
1400:
1391:
1330:
1314:
1300:
1292:
1213:Journal of the Geological Society of Korea
1025:
1023:
557:to be the sister taxon of clade formed by
432:tail before being studied. David Hone and
116:
20:
1271:
1261:
1224:
1187:
1142:
1086:
961:
987:
985:
983:
981:
931:
929:
905:
903:
901:
899:
6044:
1036:Acta Geologica Sinica - English Edition
895:
368:. The genus name is derived from Greek
936:Ji, S.-A.; Ji, Q.; Padian, K. (1999).
7:
535:being found to form an anurognathid
483:The wings are relatively short. The
761:
737:
713:
689:
636:
626:
602:
592:
6097:Middle Jurassic pterosaurs of Asia
1168:Proceedings of the Royal Society B
14:
6059:
6047:
5912:
5906:
5899:
4682:
4674:
4666:
3474:
3468:
2856:
2851:
2325:
2318:
2310:
452:Dendrorhynchoides mutoudengensis
145:
698:Dendrorhynchoides curvidentatus
366:Dendrorhynchoides curvidentatus
258:Dendrorhynchoides curvidentatus
884:Timeline of pterosaur research
531:in 2003 had the same outcome,
1:
6117:Fossil taxa described in 1999
787:Vesperopterylus lamadongensis
352:, but that name proved to be
912:Dendrorhynchus curvidentatus
746:Jeholopterus ningchengensis
585:was found to have been the
468:, named after the late Lü.
360:protozoan named in 1920 by
6138:
6122:Taxa named by Kevin Padian
386:means "curved-toothed" in
338:The genus was first named
135:Geological Museum of China
5896:
5619:
5353:
5340:
5294:
5182:
5102:
5062:
4954:
4950:
4852:
4820:
4728:
4724:
4709:
4696:
4663:
4629:
4381:
4272:
4248:
4140:
4041:
3985:
3889:
3729:
3655:
3555:
3542:
3515:
3500:
3487:
3465:
3419:
3406:
3330:
3151:
3073:
3008:
2975:
2901:
2897:
2882:
2869:
2849:
2734:
2606:
2563:
2475:
2444:
2405:
2401:
2366:
2351:
2338:
2307:
2033:
1985:
1930:
1903:
1813:
1809:
1782:
1714:
1622:
1537:
1485:
1481:
1466:
1453:
1403:
1390:
1329:
783:
766:
759:
742:
735:
718:
711:
694:
687:
658:
641:
634:
624:
607:
600:
276:
269:
251:
244:
224:Ji, Ji & Padian, 1999
142:Scientific classification
140:
124:
115:
23:
1144:10.1002/mmng.20000030109
1108:Acta Geoscientica Sinica
879:List of pterosaur genera
722:Luopterus mutoudengensis
611:"Dimorphodon" weintraubi
465:Luopterus mutoudengensis
2889:Archaeopterodactyloidea
1048:10.1111/1755-6724.14585
1006:10.1111/1755-6724.12002
320:that is known from the
1180:10.1098/rspb.2009.0846
662:Batrachognathus volans
408:The genus is based on
327:of Qinglong, northern
6112:Paleontology in Hebei
6020:Paleobiology Database
1030:Hone, D.W.E. (2020).
994:Acta Geologica Sinica
573:within the subfamily
567:phylogenetic analysis
458:was as distinct from
237:D. curvidentatus
2501:Archaeoistiodactylus
2411:Archaeoistiodactylus
1160:"The soft tissue of
914:, gen. et sp. nov".
404:Discovery and naming
390:. A second species,
325:Tiaojishan Formation
312:containing only the
5566:Targaryendraconidae
5255:Ornithocheiriformes
4942:Ornithocheiromorpha
4241:Neopterodactyloidea
4168:Dsungaripteromorpha
1936:Campylognathoididae
1656:Eudimorphodontoidea
1263:10.7717/peerj.11161
954:1999Natur.398..573J
770:Anurognathus ammoni
264:(Ji & Ji, 1998)
5095:Istiodactyliformes
3569:Lonchognathosaurus
3412:Eupterodactyloidea
2555:Pterodactyliformes
2358:Pterodactylomorpha
2345:Pterodactylomorpha
2259:Pterodactylomorpha
2101:Angustinaripterini
1732:Eudimorphodontinae
1707:Eudimorphodontidae
1244:Sinomacrops bondei
1075:Geological Journal
645:Sinomacrops bondei
6035:
6034:
6007:Open Tree of Life
5972:Dendrorhynchoides
5942:Dendrorhynchoides
5934:Taxon identifiers
5925:
5924:
5921:
5920:
5894:
5893:
5890:
5889:
5886:
5885:
5882:
5881:
5878:
5877:
5874:
5873:
5870:
5869:
5866:
5865:
5862:
5861:
5604:
5603:
5600:
5599:
5521:Targaryendraconia
5515:
5514:
5325:
5324:
5249:
5248:
5245:
5244:
5035:
5034:
4935:
4934:
4931:
4930:
4927:
4926:
4923:
4922:
4692:
4691:
4661:
4660:
4657:
4656:
4653:
4652:
4625:
4624:
4621:
4620:
4617:
4616:
4613:
4612:
4609:
4608:
4605:
4604:
4601:
4600:
4597:
4596:
4368:
4367:
4336:Chaoyangopterinae
4265:Chaoyangopteridae
4125:
4124:
4121:
4120:
4117:
4116:
4113:
4112:
4109:
4108:
4105:
4104:
3640:
3639:
3636:
3635:
3507:Ornithocheiroidea
3494:Ornithocheiroidea
3483:
3482:
3463:
3462:
3459:
3458:
3455:
3454:
3437:Ornithocheiroidea
3402:
3401:
3398:
3397:
3394:
3393:
3390:
3389:
3386:
3385:
3200:Gladocephaloideus
3136:
3135:
3132:
3131:
3098:Aurorazhdarchidae
3000:Ctenochasmatoidea
2960:
2959:
2918:Germanodactylidae
2865:
2864:
2847:
2846:
2843:
2842:
2839:
2838:
2835:
2834:
2831:
2830:
2827:
2826:
2721:
2720:
2696:Dendrorhynchoides
2673:Batrachognathinae
2639:Dendrorhynchoides
2569:Changchengopterus
2548:
2547:
2544:
2543:
2540:
2539:
2334:
2333:
2305:
2304:
2301:
2300:
2297:
2296:
2293:
2292:
2289:
2288:
2285:
2284:
2281:
2280:
2277:
2276:
2193:
2192:
2189:
2188:
2185:
2184:
2110:Angustinaripterus
2026:Rhamphorhynchinae
1978:Rhamphorhynchidae
1961:
1960:
1952:Campylognathoides
1887:
1886:
1766:
1765:
1762:
1761:
1758:
1757:
1665:Raeticodactylidae
1607:
1606:
1603:
1602:
1570:Austriadraconidae
1449:
1448:
1445:
1444:
1441:
1440:
1174:(1679): 321–329.
1114:(Supp. 1): 29–30.
1104:Dendrorhynchoides
1069:Zhou, Z. (2006).
948:(6728): 573–574.
866:Dendrorhynchoides
862:Dendrorhynchoides
852:
851:
843:
842:
834:
833:
825:
824:
816:
815:
807:
806:
798:
797:
673:
672:
631:Batrachognathinae
583:Dendrorhynchoides
571:Dendrorhynchoides
555:Dendrorhynchoides
547:Asiaticognathidae
533:Dendrorhynchoides
529:Alexander Kellner
477:Dendrorhynchoides
456:D. mutoudengensis
392:D. mutoudengensis
298:Dendrorhynchoides
294:
293:
288:
287:Ji & Ji, 1998
225:
219:Dendrorhynchoides
25:Dendrorhynchoides
6129:
6107:Fossils of China
6064:
6063:
6062:
6052:
6051:
6050:
6043:
6028:
6027:
6015:
6014:
6002:
6001:
5989:
5988:
5976:
5975:
5974:
5961:
5960:
5959:
5929:
5916:
5910:
5904:
5903:
5750:Coloborhynchinae
5678:
5633:Ornithocheiridae
5625:Brasileodactylus
5617:
5527:
5417:Ornithocheirinae
5351:
5346:Ornithocheiridae
5342:
5338:
5296:
5292:
5271:Brasileodactylus
5180:
5100:
5068:Lonchodraconidae
5064:
5060:
4952:
4948:
4850:
4818:
4813:Nyctosauromorpha
4750:
4726:
4722:
4716:Pteranodontoidea
4711:
4703:Pteranodontoidea
4698:
4687:
4686:
4679:
4678:
4671:
4670:
4635:Pteranodontoidea
4631:
4515:Quetzalcoatlinae
4504:Mistralazhdarcho
4469:
4410:
4379:
4270:
4246:
4138:
4039:
3983:
3902:
3887:
3850:
3754:Thalassodromidae
3727:
3716:
3653:
3611:Dsungaripterinae
3553:
3548:Dsungaripteridae
3544:
3540:
3513:
3502:
3489:
3478:
3473:
3472:
3417:
3408:
3328:
3323:Ctenochasmatinae
3305:Plataleorhynchus
3149:
3143:Ctenochasmatidae
3071:
3010:
3006:
2973:
2967:Euctenochasmatia
2899:
2895:
2884:
2871:
2860:
2855:
2783:Propterodactylus
2769:Ningchengopterus
2732:
2727:Pterodactyloidea
2604:
2595:
2561:
2473:
2442:
2403:
2399:
2364:
2353:
2340:
2329:
2323:
2322:
2315:
2314:
2233:Jianchangopterus
2219:Fenghuangopterus
2205:
2151:Rhamphorhynchini
2097:
2031:
1983:
1974:
1932:
1928:
1901:
1830:Dimorphodontidae
1811:
1807:
1780:
1712:
1661:
1620:
1535:
1483:
1479:
1468:
1455:
1401:
1392:
1385:
1384:
1380:Pterosauromorpha
1331:
1316:
1309:
1302:
1293:
1286:
1285:
1275:
1265:
1237:
1231:
1230:
1228:
1208:
1202:
1201:
1191:
1155:
1149:
1148:
1146:
1122:
1116:
1115:
1099:
1093:
1092:
1090:
1081:(3–4): 377–393.
1066:
1060:
1059:
1042:(5): 1676–1692.
1027:
1018:
1017:
1000:(6): 1317–1325.
989:
976:
975:
965:
933:
924:
923:
907:
762:
738:
714:
690:
637:
627:
603:
593:
545:, that he named
460:D. curvidentatus
438:D. curvidentatus
426:Yixian Formation
318:D. curvidentatus
286:
260:
256:
223:
216:
203:
190:
150:
149:
120:
110:
40:
29:Temporal range:
21:
6137:
6136:
6132:
6131:
6130:
6128:
6127:
6126:
6072:
6071:
6070:
6060:
6058:
6048:
6046:
6038:
6036:
6031:
6023:
6018:
6010:
6005:
5997:
5992:
5984:
5979:
5970:
5969:
5964:
5955:
5954:
5949:
5936:
5926:
5917:
5898:
5858:
5839:Liaoningopterus
5794:
5744:
5682:Tropeognathinae
5667:
5639:
5613:
5596:
5560:
5531:Cimoliopteridae
5511:
5411:
5334:
5321:
5316:Zhenyuanopterus
5290:
5241:
5220:Istiodactylinae
5214:
5195:Longchengpterus
5188:Lingyuanopterus
5175:Istiodactylidae
5169:
5135:
5108:Hongshanopterus
5089:
5058:
5044:
5031:
4994:Hongshanopterus
4979:
4944:
4919:
4892:Barbaridactylus
4877:
4839:
4807:
4754:Pteranodontidae
4739:
4734:Santanadactylus
4718:
4705:
4688:
4681:
4673:
4665:
4649:
4593:
4588:Zhejiangopterus
4509:
4467:
4446:Palaeocursornis
4408:
4387:Montanazhdarcho
4374:Azhdarchiformes
4364:
4345:Chaoyangopterus
4330:
4259:
4235:
4221:Montanazhdarcho
4174:
4134:
4132:Azhdarchomorpha
4101:
4074:
4028:
3991:Aymberedactylus
3972:
3900:
3876:
3864:Aymberedactylus
3839:
3819:Thalassodromeus
3748:
3723:
3712:
3703:
3691:Montanazhdarcho
3649:
3632:
3620:Domeykodactylus
3605:
3599:Tendaguripterus
3536:
3527:
3509:
3496:
3479:
3467:
3451:
3431:
3425:Altmuehlopterus
3382:
3341:
3317:
3268:
3240:
3145:
3128:
3092:
3086:Gallodactylidae
3066:Aurorazhdarchia
3060:
3038:Normannognathus
3014:Gallodactylidae
3002:
2993:
2969:
2956:
2950:Tendaguripterus
2942:Normannognathus
2935:Germanodactylus
2927:Altmuehlopterus
2912:
2891:
2878:
2861:
2823:
2803:
2717:
2682:Batrachognathus
2667:
2662:Vesperopterylus
2617:
2591:
2582:
2557:
2536:
2492:Wukongopterinae
2486:
2468:Wukongopteridae
2462:
2431:
2418:Normannognathus
2395:
2386:
2360:
2347:
2330:
2317:
2309:
2273:
2253:
2226:Jianchangnathus
2209:Scaphognathidae
2199:Digibrevisauria
2181:
2176:Rhamphorhynchus
2145:
2132:Qinglongopterus
2124:Harpactognathus
2092:Rhamphorhynchae
2086:
2081:Qinglongopterus
2060:Harpactognathus
2020:
2014:Scaphognathinae
2006:Parapsicephalus
1970:
1957:
1945:Bergamodactylus
1924:
1915:
1897:
1883:
1861:Parapsicephalus
1824:
1803:
1801:Macronychoptera
1794:
1776:
1754:
1726:
1720:Arcticodactylus
1701:
1689:Raeticodactylus
1650:
1616:
1599:
1579:Arcticodactylus
1564:
1558:Raeticodactylus
1523:
1511:Austriadactylus
1496:
1475:
1462:
1437:
1417:
1411:Avemetatarsalia
1396:Avemetatarsalia
1386:
1371:Avemetatarsalia
1336:
1335:
1325:
1320:
1290:
1289:
1239:
1238:
1234:
1226:10.1.1.694.4816
1210:
1209:
1205:
1157:
1156:
1152:
1124:
1123:
1119:
1101:
1100:
1096:
1088:10.1002/gj.1045
1068:
1067:
1063:
1029:
1028:
1021:
991:
990:
979:
935:
934:
927:
916:Jiangsu Geology
909:
908:
897:
892:
875:
858:
853:
844:
835:
826:
817:
808:
799:
674:
579:Batrachognathus
559:Batrachognathus
551:Asiaticognathus
542:Batrachognathus
518:rhamphorhynchid
510:
474:
406:
379:Rhamphorhynchus
322:Middle Jurassic
285:
265:
262:
254:
253:
240:
222:
214:
201:
188:
144:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
31:Middle Jurassic
27:
17:
12:
11:
5:
6135:
6133:
6125:
6124:
6119:
6114:
6109:
6104:
6102:Jurassic China
6099:
6094:
6092:Oxfordian life
6089:
6087:Callovian life
6084:
6074:
6073:
6069:
6068:
6056:
6033:
6032:
6030:
6029:
6016:
6003:
5990:
5977:
5962:
5946:
5944:
5938:
5937:
5932:
5923:
5922:
5919:
5918:
5897:
5895:
5892:
5891:
5888:
5887:
5884:
5883:
5880:
5879:
5876:
5875:
5872:
5871:
5868:
5867:
5864:
5863:
5860:
5859:
5857:
5856:
5849:
5842:
5835:
5828:
5820:
5816:Caulkicephalus
5812:
5804:
5802:
5796:
5795:
5793:
5792:
5789:Uktenadactylus
5785:
5777:
5770:
5767:Coloborhynchus
5763:
5755:
5753:
5746:
5745:
5743:
5742:
5735:
5728:
5720:
5712:
5704:
5696:
5687:
5685:
5675:
5669:
5668:
5666:
5665:
5658:
5650:
5648:
5641:
5640:
5638:
5637:
5629:
5620:
5614:
5609:
5606:
5605:
5602:
5601:
5598:
5597:
5595:
5594:
5591:Targaryendraco
5587:
5579:
5570:
5568:
5562:
5561:
5559:
5558:
5551:
5543:
5535:
5533:
5524:
5517:
5516:
5513:
5512:
5510:
5509:
5505:Uktenadactylus
5501:
5493:
5490:Ornithocheirus
5486:
5478:
5470:
5462:
5454:
5446:
5442:Coloborhynchus
5438:
5430:
5421:
5419:
5413:
5412:
5410:
5409:
5401:
5393:
5385:
5381:Caulkicephalus
5377:
5369:
5366:Arthurdactylus
5362:
5354:
5348:
5335:
5332:Ornithocheirae
5330:
5327:
5326:
5323:
5322:
5320:
5319:
5312:
5304:
5302:
5289:
5288:
5281:
5274:
5267:
5259:
5257:
5251:
5250:
5247:
5246:
5243:
5242:
5240:
5239:
5232:
5224:
5222:
5216:
5215:
5213:
5212:
5205:
5198:
5191:
5183:
5177:
5171:
5170:
5168:
5167:
5160:
5157:Linlongopterus
5153:
5145:
5143:
5141:Mimodactylidae
5137:
5136:
5134:
5133:
5129:Lonchodectidae
5125:
5118:
5115:Linlongopterus
5111:
5103:
5097:
5091:
5090:
5088:
5087:
5080:
5072:
5070:
5057:
5056:
5047:
5045:
5040:
5037:
5036:
5033:
5032:
5030:
5029:
5025:Targaryendraco
5021:
5013:
5006:
4998:
4989:
4987:
4985:Lonchodectidae
4981:
4980:
4978:
4977:
4970:
4963:
4955:
4945:
4940:
4937:
4936:
4933:
4932:
4929:
4928:
4925:
4924:
4921:
4920:
4918:
4917:
4909:
4902:
4899:Muzquizopteryx
4895:
4887:
4885:
4879:
4878:
4876:
4875:
4868:
4861:
4853:
4847:
4845:Aponyctosauria
4841:
4840:
4838:
4837:
4829:
4821:
4815:
4809:
4808:
4806:
4805:
4797:
4789:
4782:
4774:
4767:
4758:
4756:
4747:
4741:
4740:
4738:
4737:
4729:
4719:
4714:
4707:
4706:
4701:
4694:
4693:
4690:
4689:
4664:
4662:
4659:
4658:
4655:
4654:
4651:
4650:
4648:
4647:
4646:
4645:
4639:
4637:
4627:
4626:
4623:
4622:
4619:
4618:
4615:
4614:
4611:
4610:
4607:
4606:
4603:
4602:
4599:
4598:
4595:
4594:
4592:
4591:
4584:
4576:
4573:Thanatosdrakon
4569:
4566:Quetzalcoatlus
4562:
4559:Phosphatodraco
4555:
4548:
4541:
4534:
4531:Arambourgiania
4527:
4519:
4517:
4511:
4510:
4508:
4507:
4500:
4493:
4486:
4477:
4475:
4466:
4465:
4457:
4449:
4442:
4438:Navajodactylus
4434:
4427:
4418:
4416:
4407:
4406:
4398:
4391:
4382:
4376:
4370:
4369:
4366:
4365:
4363:
4362:
4359:Shenzhoupterus
4355:
4348:
4340:
4338:
4332:
4331:
4329:
4328:
4320:
4312:
4305:
4298:
4290:
4286:Argentinadraco
4282:
4273:
4267:
4261:
4260:
4258:
4257:
4249:
4243:
4237:
4236:
4234:
4233:
4225:
4217:
4209:
4201:
4197:Argentinadraco
4193:
4185:
4183:
4176:
4175:
4173:
4172:
4164:
4157:
4150:
4141:
4135:
4130:
4127:
4126:
4123:
4122:
4119:
4118:
4115:
4114:
4111:
4110:
4107:
4106:
4103:
4102:
4100:
4099:
4092:
4084:
4082:
4076:
4075:
4073:
4072:
4065:
4058:
4051:
4042:
4036:
4030:
4029:
4027:
4026:
4019:
4011:
4003:
3995:
3986:
3980:
3974:
3973:
3971:
3970:
3963:
3956:
3953:Nemicolopterus
3949:
3942:
3935:
3927:
3919:
3910:
3908:
3899:
3898:
3890:
3884:
3878:
3877:
3875:
3874:
3867:
3859:
3857:
3847:
3845:Tapejariformes
3841:
3840:
3838:
3837:
3829:
3822:
3815:
3807:
3799:
3792:
3784:
3780:Argentinadraco
3776:
3768:
3759:
3757:
3750:
3749:
3747:
3746:
3738:
3730:
3724:
3721:Tapejaromorpha
3719:
3713:
3708:
3705:
3704:
3702:
3701:
3694:
3687:
3679:
3671:
3664:
3661:Argentinadraco
3656:
3650:
3645:
3642:
3641:
3638:
3637:
3634:
3633:
3631:
3630:
3627:Dsungaripterus
3623:
3615:
3613:
3607:
3606:
3604:
3603:
3595:
3587:
3580:
3573:
3565:
3556:
3550:
3537:
3532:
3529:
3528:
3526:
3525:
3516:
3510:
3505:
3498:
3497:
3492:
3485:
3484:
3481:
3480:
3466:
3464:
3461:
3460:
3457:
3456:
3453:
3452:
3450:
3449:
3448:
3447:
3441:
3439:
3433:
3432:
3430:
3429:
3420:
3414:
3404:
3403:
3400:
3399:
3396:
3395:
3392:
3391:
3388:
3387:
3384:
3383:
3381:
3380:
3373:
3366:
3359:
3351:
3349:
3347:Pterodaustrini
3343:
3342:
3340:
3339:
3331:
3325:
3319:
3318:
3316:
3315:
3308:
3301:
3294:
3286:
3278:
3276:
3274:Gnathosaurinae
3270:
3269:
3267:
3266:
3259:
3251:
3249:
3246:Moganopterinae
3242:
3241:
3239:
3238:
3231:
3224:
3217:
3210:
3203:
3196:
3189:
3182:
3175:
3168:
3165:Balaenognathus
3161:
3152:
3146:
3141:
3138:
3137:
3134:
3133:
3130:
3129:
3127:
3126:
3118:
3115:Aurorazhdarcho
3111:
3102:
3100:
3094:
3093:
3091:
3090:
3082:
3074:
3068:
3062:
3061:
3059:
3058:
3054:Moganopterinae
3050:
3042:
3034:
3027:
3023:Aurorazhdarcho
3018:
3016:
3003:
2998:
2995:
2994:
2992:
2991:
2984:
2976:
2970:
2965:
2962:
2961:
2958:
2957:
2955:
2954:
2946:
2938:
2931:
2922:
2920:
2914:
2913:
2911:
2910:
2902:
2892:
2887:
2880:
2879:
2874:
2867:
2866:
2863:
2862:
2850:
2848:
2845:
2844:
2841:
2840:
2837:
2836:
2833:
2832:
2829:
2828:
2825:
2824:
2822:
2821:
2820:
2819:
2813:
2811:
2805:
2804:
2802:
2801:
2794:
2786:
2779:
2772:
2765:
2758:
2750:
2743:
2735:
2729:
2723:
2722:
2719:
2718:
2716:
2715:
2708:
2700:
2692:
2685:
2677:
2675:
2669:
2668:
2666:
2665:
2658:
2651:
2643:
2635:
2627:
2625:
2623:Anurognathinae
2619:
2618:
2616:
2615:
2607:
2601:
2599:Anurognathidae
2592:
2587:
2584:
2583:
2581:
2580:
2577:Douzhanopterus
2573:
2564:
2558:
2553:
2550:
2549:
2546:
2545:
2542:
2541:
2538:
2537:
2535:
2534:
2527:
2523:Douzhanopterus
2519:
2512:
2505:
2496:
2494:
2488:
2487:
2485:
2484:
2481:Kunpengopterus
2476:
2470:
2464:
2463:
2461:
2460:
2453:
2445:
2439:
2433:
2432:
2430:
2429:
2425:Anurognathidae
2421:
2414:
2406:
2396:
2393:Monofenestrata
2391:
2388:
2387:
2385:
2384:
2376:
2367:
2361:
2356:
2349:
2348:
2343:
2336:
2335:
2332:
2331:
2308:
2306:
2303:
2302:
2299:
2298:
2295:
2294:
2291:
2290:
2287:
2286:
2283:
2282:
2279:
2278:
2275:
2274:
2272:
2271:
2270:
2269:
2263:
2261:
2255:
2254:
2252:
2251:
2243:
2236:
2229:
2222:
2214:
2212:
2202:
2195:
2194:
2191:
2190:
2187:
2186:
2183:
2182:
2180:
2179:
2172:
2164:
2155:
2153:
2147:
2146:
2144:
2143:
2136:
2128:
2120:
2113:
2105:
2103:
2094:
2088:
2087:
2085:
2084:
2077:
2074:Orientognathus
2070:
2063:
2056:
2049:
2042:
2034:
2028:
2022:
2021:
2019:
2018:
2010:
2002:
1995:
1991:Dolicorhamphus
1986:
1980:
1971:
1968:Breviquartossa
1966:
1963:
1962:
1959:
1958:
1956:
1955:
1948:
1940:
1938:
1925:
1920:
1917:
1916:
1914:
1913:
1904:
1898:
1892:
1889:
1888:
1885:
1884:
1882:
1881:
1873:
1865:
1857:
1850:
1847:Caelestiventus
1843:
1834:
1832:
1826:
1825:
1823:
1822:
1814:
1804:
1799:
1796:
1795:
1793:
1792:
1783:
1777:
1773:Zambellisauria
1771:
1768:
1767:
1764:
1763:
1760:
1759:
1756:
1755:
1753:
1752:
1745:
1741:Carniadactylus
1736:
1734:
1728:
1727:
1725:
1724:
1715:
1709:
1703:
1702:
1700:
1699:
1692:
1685:
1678:
1669:
1667:
1658:
1652:
1651:
1649:
1648:
1640:
1632:
1623:
1617:
1612:
1609:
1608:
1605:
1604:
1601:
1600:
1598:
1597:
1594:Seazzadactylus
1590:
1583:
1574:
1572:
1566:
1565:
1563:
1562:
1554:
1547:
1543:Carniadactylus
1538:
1532:
1525:
1524:
1522:
1521:
1514:
1506:
1504:
1498:
1497:
1495:
1494:
1491:Daohugoupterus
1486:
1476:
1471:
1464:
1463:
1458:
1451:
1450:
1447:
1446:
1443:
1442:
1439:
1438:
1436:
1435:
1434:
1433:
1427:
1425:
1419:
1418:
1416:
1415:
1414:
1413:
1404:
1398:
1388:
1387:
1383:
1382:
1373:
1364:
1355:
1349:
1343:
1334:
1327:
1326:
1321:
1319:
1318:
1311:
1304:
1296:
1288:
1287:
1232:
1219:(1): 239–261.
1203:
1150:
1117:
1094:
1061:
1019:
977:
925:
894:
893:
891:
888:
887:
886:
881:
874:
871:
857:
854:
850:
849:
846:
845:
841:
840:
837:
836:
832:
831:
828:
827:
823:
822:
819:
818:
814:
813:
810:
809:
805:
804:
801:
800:
796:
795:
792:
791:
782:
779:
778:
775:
774:
765:
760:
758:
755:
754:
751:
750:
741:
736:
734:
731:
730:
727:
726:
717:
712:
710:
707:
706:
703:
702:
693:
688:
686:
684:Anurognathinae
680:
679:
676:
675:
671:
670:
667:
666:
657:
654:
653:
650:
649:
640:
635:
633:
625:
623:
620:
619:
616:
615:
606:
601:
599:
597:Anurognathidae
591:
575:Anurognathinae
514:Anurognathidae
509:
508:Classification
506:
473:
470:
421:fossil dealers
405:
402:
341:Dendrorhynchus
292:
291:
290:
289:
281:Dendrorhynchus
274:
273:
267:
266:
263:
249:
248:
242:
241:
233:
231:
227:
226:
212:
208:
207:
205:Anurognathidae
199:
195:
194:
186:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
6134:
6123:
6120:
6118:
6115:
6113:
6110:
6108:
6105:
6103:
6100:
6098:
6095:
6093:
6090:
6088:
6085:
6083:
6082:Anurognathids
6080:
6079:
6077:
6067:
6057:
6055:
6045:
6041:
6026:
6021:
6017:
6013:
6008:
6004:
6000:
5995:
5991:
5987:
5982:
5978:
5973:
5967:
5963:
5958:
5952:
5948:
5947:
5945:
5943:
5939:
5935:
5930:
5915:
5911:
5909:
5902:
5855:
5854:
5853:Maaradactylus
5850:
5848:
5847:
5843:
5841:
5840:
5836:
5834:
5833:
5829:
5826:
5825:
5824:Cearadactylus
5821:
5818:
5817:
5813:
5811:
5810:
5806:
5805:
5803:
5801:
5800:Anhanguerinae
5797:
5791:
5790:
5786:
5783:
5782:
5781:Siroccopteryx
5778:
5776:
5775:
5771:
5769:
5768:
5764:
5762:
5761:
5757:
5756:
5754:
5751:
5747:
5741:
5740:
5739:Tropeognathus
5736:
5734:
5733:
5729:
5726:
5725:
5724:Siroccopteryx
5721:
5718:
5717:
5713:
5710:
5709:
5705:
5702:
5701:
5697:
5694:
5693:
5689:
5688:
5686:
5683:
5679:
5676:
5674:
5673:Anhangueridae
5670:
5664:
5663:
5662:Iberodactylus
5659:
5657:
5656:
5652:
5651:
5649:
5646:
5642:
5635:
5634:
5630:
5627:
5626:
5622:
5621:
5618:
5615:
5612:
5607:
5593:
5592:
5588:
5585:
5584:
5580:
5577:
5576:
5572:
5571:
5569:
5567:
5563:
5557:
5556:
5555:Cimoliopterus
5552:
5549:
5548:
5547:Camposipterus
5544:
5542:
5541:
5537:
5536:
5534:
5532:
5528:
5525:
5522:
5518:
5507:
5506:
5502:
5499:
5498:
5497:Siroccopteryx
5494:
5492:
5491:
5487:
5484:
5483:
5479:
5476:
5475:
5471:
5468:
5467:
5463:
5460:
5459:
5455:
5452:
5451:
5447:
5444:
5443:
5439:
5436:
5435:
5434:Camposipterus
5431:
5428:
5427:
5423:
5422:
5420:
5418:
5414:
5407:
5406:
5405:Tropeognathus
5402:
5399:
5398:
5394:
5391:
5390:
5389:Cimoliopterus
5386:
5383:
5382:
5378:
5375:
5374:
5373:Camposipterus
5370:
5368:
5367:
5363:
5361:
5360:
5359:Araripesaurus
5356:
5355:
5352:
5349:
5347:
5343:
5339:
5336:
5333:
5328:
5318:
5317:
5313:
5311:
5310:
5306:
5305:
5303:
5301:
5300:Boreopteridae
5297:
5293:
5287:
5286:
5282:
5280:
5279:
5278:Cearadactylus
5275:
5273:
5272:
5268:
5266:
5265:
5261:
5260:
5258:
5256:
5252:
5238:
5237:
5233:
5231:
5230:
5229:Istiodactylus
5226:
5225:
5223:
5221:
5217:
5211:
5210:
5206:
5204:
5203:
5199:
5197:
5196:
5192:
5190:
5189:
5185:
5184:
5181:
5178:
5176:
5172:
5166:
5165:
5161:
5159:
5158:
5154:
5152:
5151:
5147:
5146:
5144:
5142:
5138:
5131:
5130:
5126:
5124:
5123:
5122:Yixianopterus
5119:
5117:
5116:
5112:
5110:
5109:
5105:
5104:
5101:
5098:
5096:
5092:
5086:
5085:
5081:
5079:
5078:
5074:
5073:
5071:
5069:
5065:
5061:
5054:
5053:
5049:
5048:
5046:
5043:
5038:
5027:
5026:
5022:
5019:
5018:
5014:
5012:
5011:
5007:
5004:
5003:
4999:
4996:
4995:
4991:
4990:
4988:
4986:
4982:
4976:
4975:
4971:
4969:
4968:
4964:
4962:
4961:
4957:
4956:
4953:
4949:
4946:
4943:
4938:
4915:
4914:
4910:
4908:
4907:
4903:
4901:
4900:
4896:
4894:
4893:
4889:
4888:
4886:
4884:
4883:Nyctosauridae
4880:
4874:
4873:
4869:
4867:
4866:
4862:
4860:
4859:
4855:
4854:
4851:
4848:
4846:
4842:
4835:
4834:
4830:
4828:
4827:
4826:Alamodactylus
4823:
4822:
4819:
4816:
4814:
4810:
4803:
4802:
4798:
4795:
4794:
4790:
4788:
4787:
4783:
4780:
4779:
4775:
4773:
4772:
4768:
4765:
4764:
4760:
4759:
4757:
4755:
4751:
4748:
4746:
4745:Pteranodontia
4742:
4736:
4735:
4731:
4730:
4727:
4723:
4720:
4717:
4712:
4708:
4704:
4699:
4695:
4685:
4680:
4677:
4669:
4643:
4642:
4641:
4640:
4638:
4636:
4632:
4628:
4590:
4589:
4585:
4582:
4581:
4580:Wellnhopterus
4577:
4575:
4574:
4570:
4568:
4567:
4563:
4561:
4560:
4556:
4554:
4553:
4552:Hatzegopteryx
4549:
4547:
4546:
4542:
4540:
4539:
4535:
4533:
4532:
4528:
4526:
4525:
4524:Aralazhdarcho
4521:
4520:
4518:
4516:
4512:
4506:
4505:
4501:
4499:
4498:
4494:
4492:
4491:
4487:
4484:
4483:
4479:
4478:
4476:
4474:
4470:
4463:
4462:
4458:
4455:
4454:
4450:
4448:
4447:
4443:
4440:
4439:
4435:
4433:
4432:
4428:
4425:
4424:
4420:
4419:
4417:
4415:
4411:
4404:
4403:
4399:
4397:
4396:
4395:Radiodactylus
4392:
4389:
4388:
4384:
4383:
4380:
4377:
4375:
4371:
4361:
4360:
4356:
4354:
4353:
4349:
4347:
4346:
4342:
4341:
4339:
4337:
4333:
4326:
4325:
4321:
4318:
4317:
4313:
4311:
4310:
4306:
4304:
4303:
4299:
4296:
4295:
4291:
4288:
4287:
4283:
4280:
4279:
4278:Apatorhamphus
4275:
4274:
4271:
4268:
4266:
4262:
4256:
4255:
4251:
4250:
4247:
4244:
4242:
4238:
4231:
4230:
4226:
4223:
4222:
4218:
4215:
4214:
4210:
4207:
4206:
4202:
4199:
4198:
4194:
4192:
4191:
4187:
4186:
4184:
4181:
4177:
4170:
4169:
4165:
4163:
4162:
4158:
4156:
4155:
4151:
4148:
4147:
4143:
4142:
4139:
4136:
4133:
4128:
4098:
4097:
4093:
4091:
4090:
4086:
4085:
4083:
4081:
4077:
4071:
4070:
4069:Tupandactylus
4066:
4064:
4063:
4059:
4057:
4056:
4052:
4049:
4048:
4044:
4043:
4040:
4037:
4035:
4031:
4025:
4024:
4020:
4017:
4016:
4012:
4009:
4008:
4004:
4001:
4000:
3999:Caupedactylus
3996:
3993:
3992:
3988:
3987:
3984:
3981:
3979:
3975:
3969:
3968:
3964:
3962:
3961:
3957:
3955:
3954:
3950:
3948:
3947:
3943:
3941:
3940:
3936:
3933:
3932:
3928:
3925:
3924:
3920:
3917:
3916:
3912:
3911:
3909:
3907:
3903:
3897:
3896:
3892:
3891:
3888:
3885:
3883:
3879:
3873:
3872:
3871:Caupedactylus
3868:
3866:
3865:
3861:
3860:
3858:
3855:
3854:Caupedactylia
3851:
3848:
3846:
3842:
3835:
3834:
3830:
3828:
3827:
3823:
3821:
3820:
3816:
3813:
3812:
3808:
3805:
3804:
3800:
3798:
3797:
3793:
3790:
3789:
3785:
3782:
3781:
3777:
3774:
3773:
3769:
3766:
3765:
3761:
3760:
3758:
3755:
3751:
3744:
3743:
3739:
3737:
3736:
3732:
3731:
3728:
3725:
3722:
3717:
3714:
3711:
3710:Neoazhdarchia
3706:
3700:
3699:
3695:
3693:
3692:
3688:
3685:
3684:
3680:
3677:
3676:
3672:
3670:
3669:
3665:
3663:
3662:
3658:
3657:
3654:
3651:
3648:
3647:Azhdarchoidea
3643:
3629:
3628:
3624:
3622:
3621:
3617:
3616:
3614:
3612:
3608:
3601:
3600:
3596:
3593:
3592:
3591:Puntanipterus
3588:
3586:
3585:
3581:
3579:
3578:
3574:
3571:
3570:
3566:
3563:
3562:
3558:
3557:
3554:
3551:
3549:
3545:
3541:
3538:
3535:
3530:
3523:
3522:
3518:
3517:
3514:
3511:
3508:
3503:
3499:
3495:
3490:
3486:
3477:
3471:
3445:
3444:
3443:
3442:
3440:
3438:
3434:
3427:
3426:
3422:
3421:
3418:
3415:
3413:
3409:
3405:
3379:
3378:
3374:
3372:
3371:
3367:
3365:
3364:
3360:
3358:
3357:
3356:Beipiaopterus
3353:
3352:
3350:
3348:
3344:
3338:
3337:
3333:
3332:
3329:
3326:
3324:
3320:
3314:
3313:
3312:Tacuadactylus
3309:
3307:
3306:
3302:
3300:
3299:
3295:
3292:
3291:
3287:
3285:
3284:
3280:
3279:
3277:
3275:
3271:
3265:
3264:
3260:
3258:
3257:
3253:
3252:
3250:
3247:
3243:
3237:
3236:
3232:
3230:
3229:
3225:
3223:
3222:
3218:
3216:
3215:
3211:
3209:
3208:
3204:
3202:
3201:
3197:
3195:
3194:
3193:Forfexopterus
3190:
3188:
3187:
3186:Elanodactylus
3183:
3181:
3180:
3179:Cratonopterus
3176:
3174:
3173:
3172:Cathayopterus
3169:
3167:
3166:
3162:
3159:
3158:
3157:Ardeadactylus
3154:
3153:
3150:
3147:
3144:
3139:
3124:
3123:
3119:
3117:
3116:
3112:
3109:
3108:
3107:Ardeadactylus
3104:
3103:
3101:
3099:
3095:
3088:
3087:
3083:
3081:
3080:
3076:
3075:
3072:
3069:
3067:
3063:
3056:
3055:
3051:
3048:
3047:
3043:
3040:
3039:
3035:
3033:
3032:
3031:Cycnorhamphus
3028:
3025:
3024:
3020:
3019:
3017:
3015:
3011:
3007:
3004:
3001:
2996:
2990:
2989:
2988:Pterodactylus
2985:
2983:
2982:
2981:Diopecephalus
2978:
2977:
2974:
2971:
2968:
2963:
2952:
2951:
2947:
2944:
2943:
2939:
2937:
2936:
2932:
2929:
2928:
2924:
2923:
2921:
2919:
2915:
2909:
2908:
2907:Prejanopterus
2904:
2903:
2900:
2896:
2893:
2890:
2885:
2881:
2877:
2872:
2868:
2859:
2854:
2817:
2816:
2815:
2814:
2812:
2810:
2806:
2800:
2799:
2795:
2792:
2791:
2787:
2785:
2784:
2780:
2778:
2777:
2773:
2771:
2770:
2766:
2764:
2763:
2759:
2756:
2755:
2754:Herbstosaurus
2751:
2749:
2748:
2747:Eurolimnornis
2744:
2742:
2741:
2740:Dermodactylus
2737:
2736:
2733:
2730:
2728:
2724:
2714:
2713:
2709:
2706:
2705:
2701:
2698:
2697:
2693:
2691:
2690:
2686:
2684:
2683:
2679:
2678:
2676:
2674:
2670:
2664:
2663:
2659:
2657:
2656:
2652:
2649:
2648:
2644:
2641:
2640:
2636:
2634:
2633:
2629:
2628:
2626:
2624:
2620:
2614:
2613:
2609:
2608:
2605:
2602:
2600:
2596:
2593:
2590:
2589:Caelidracones
2585:
2579:
2578:
2574:
2571:
2570:
2566:
2565:
2562:
2559:
2556:
2551:
2533:
2532:
2531:Wukongopterus
2528:
2525:
2524:
2520:
2518:
2517:
2516:Darwinopterus
2513:
2511:
2510:
2509:Cuspicephalus
2506:
2503:
2502:
2498:
2497:
2495:
2493:
2489:
2483:
2482:
2478:
2477:
2474:
2471:
2469:
2465:
2459:
2458:
2457:Pterorhynchus
2454:
2452:
2451:
2447:
2446:
2443:
2440:
2438:
2434:
2427:
2426:
2422:
2420:
2419:
2415:
2413:
2412:
2408:
2407:
2404:
2400:
2397:
2394:
2389:
2382:
2381:
2377:
2374:
2373:
2369:
2368:
2365:
2362:
2359:
2354:
2350:
2346:
2341:
2337:
2328:
2324:
2321:
2313:
2267:
2266:
2265:
2264:
2262:
2260:
2256:
2249:
2248:
2244:
2242:
2241:
2240:Scaphognathus
2237:
2235:
2234:
2230:
2228:
2227:
2223:
2221:
2220:
2216:
2215:
2213:
2210:
2206:
2203:
2200:
2196:
2178:
2177:
2173:
2170:
2169:
2165:
2162:
2161:
2157:
2156:
2154:
2152:
2148:
2142:
2141:
2137:
2134:
2133:
2129:
2126:
2125:
2121:
2119:
2118:
2114:
2112:
2111:
2107:
2106:
2104:
2102:
2098:
2095:
2093:
2089:
2083:
2082:
2078:
2076:
2075:
2071:
2069:
2068:
2064:
2062:
2061:
2057:
2055:
2054:
2050:
2048:
2047:
2043:
2041:
2040:
2036:
2035:
2032:
2029:
2027:
2023:
2016:
2015:
2011:
2008:
2007:
2003:
2001:
2000:
1996:
1993:
1992:
1988:
1987:
1984:
1981:
1979:
1975:
1972:
1969:
1964:
1954:
1953:
1949:
1947:
1946:
1942:
1941:
1939:
1937:
1933:
1929:
1926:
1923:
1918:
1911:
1910:
1909:Eudimorphodon
1906:
1905:
1902:
1899:
1895:
1890:
1879:
1878:
1874:
1871:
1870:
1869:Peteinosaurus
1866:
1863:
1862:
1858:
1856:
1855:
1851:
1849:
1848:
1844:
1841:
1840:
1836:
1835:
1833:
1831:
1827:
1821:
1820:
1819:Herbstosaurus
1816:
1815:
1812:
1808:
1805:
1802:
1797:
1790:
1789:
1788:Peteinosaurus
1785:
1784:
1781:
1778:
1774:
1769:
1751:
1750:
1749:Eudimorphodon
1746:
1743:
1742:
1738:
1737:
1735:
1733:
1729:
1722:
1721:
1717:
1716:
1713:
1710:
1708:
1704:
1698:
1697:
1693:
1691:
1690:
1686:
1684:
1683:
1679:
1676:
1675:
1671:
1670:
1668:
1666:
1662:
1659:
1657:
1653:
1646:
1645:
1644:Preondactylia
1641:
1638:
1637:
1636:Peteinosaurus
1633:
1630:
1629:
1625:
1624:
1621:
1618:
1615:
1614:Eopterosauria
1610:
1596:
1595:
1591:
1589:
1588:
1584:
1581:
1580:
1576:
1575:
1573:
1571:
1567:
1560:
1559:
1555:
1553:
1552:
1548:
1545:
1544:
1540:
1539:
1536:
1533:
1530:
1526:
1520:
1519:
1518:Preondactylus
1515:
1513:
1512:
1508:
1507:
1505:
1503:
1502:Preondactylia
1499:
1493:
1492:
1488:
1487:
1484:
1480:
1477:
1474:
1469:
1465:
1461:
1456:
1452:
1431:
1430:
1429:
1428:
1426:
1424:
1420:
1412:
1408:
1407:
1406:
1405:
1402:
1399:
1397:
1393:
1389:
1381:
1377:
1374:
1372:
1368:
1365:
1363:
1359:
1356:
1354:
1350:
1348:
1344:
1342:
1338:
1337:
1332:
1328:
1324:
1317:
1312:
1310:
1305:
1303:
1298:
1297:
1294:
1283:
1279:
1274:
1269:
1264:
1259:
1255:
1251:
1247:
1245:
1236:
1233:
1227:
1222:
1218:
1214:
1207:
1204:
1199:
1195:
1190:
1185:
1181:
1177:
1173:
1169:
1165:
1163:
1154:
1151:
1145:
1140:
1136:
1132:
1131:Fossil Record
1128:
1121:
1118:
1113:
1109:
1105:
1098:
1095:
1089:
1084:
1080:
1076:
1072:
1065:
1062:
1057:
1053:
1049:
1045:
1041:
1037:
1033:
1026:
1024:
1020:
1015:
1011:
1007:
1003:
999:
995:
988:
986:
984:
982:
978:
973:
969:
964:
963:10.1038/19221
959:
955:
951:
947:
943:
939:
932:
930:
926:
922:(4): 199–206.
921:
917:
913:
906:
904:
902:
900:
896:
889:
885:
882:
880:
877:
876:
872:
870:
867:
863:
856:Palaeobiology
855:
848:
847:
839:
838:
830:
829:
821:
820:
812:
811:
803:
802:
794:
793:
790:
789:
788:
781:
780:
777:
776:
773:
772:
771:
764:
763:
757:
756:
753:
752:
749:
748:
747:
740:
739:
733:
732:
729:
728:
725:
724:
723:
716:
715:
709:
708:
705:
704:
701:
700:
699:
692:
691:
685:
682:
681:
678:
677:
669:
668:
665:
664:
663:
656:
655:
652:
651:
648:
647:
646:
639:
638:
632:
629:
628:
622:
621:
618:
617:
614:
613:
612:
605:
604:
598:
595:
594:
590:
588:
584:
580:
576:
572:
568:
565:. In 2021, a
564:
560:
556:
552:
548:
544:
543:
538:
534:
530:
526:
521:
519:
515:
507:
505:
503:
499:
495:
490:
486:
481:
478:
471:
469:
467:
466:
461:
457:
453:
447:
445:
444:
439:
435:
431:
430:dromaeosaurid
427:
422:
418:
414:
411:
403:
401:
399:
398:
393:
389:
385:
384:specific name
381:
380:
375:
372:, "tree" and
371:
367:
363:
359:
355:
351:
347:
343:
342:
336:
334:
330:
326:
323:
319:
315:
311:
308:
304:
300:
299:
284:
283:curvidentatus
282:
278:
277:
275:
272:
268:
261:
259:
250:
247:
246:Binomial name
243:
239:
238:
232:
229:
228:
221:
220:
213:
210:
209:
206:
200:
197:
196:
193:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
148:
143:
139:
136:
132:
131:curvidentatus
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
36:164–159
32:
26:
22:
19:
6054:Paleontology
5941:
5905:
5851:
5846:Ludodactylus
5844:
5837:
5830:
5822:
5814:
5807:
5787:
5779:
5774:Nicorhynchus
5772:
5765:
5758:
5737:
5730:
5722:
5714:
5706:
5698:
5690:
5660:
5653:
5645:Hamipteridae
5631:
5623:
5589:
5581:
5573:
5553:
5545:
5540:Aetodactylus
5538:
5503:
5495:
5488:
5480:
5474:Ludodactylus
5472:
5464:
5456:
5448:
5440:
5432:
5426:Aetodactylus
5424:
5403:
5395:
5387:
5379:
5371:
5364:
5357:
5314:
5307:
5283:
5276:
5269:
5262:
5236:Liaoxipterus
5234:
5227:
5207:
5200:
5193:
5186:
5164:Mimodactylus
5162:
5155:
5148:
5127:
5120:
5113:
5106:
5082:
5075:
5050:
5042:Lanceodontia
5023:
5015:
5010:Lonchodectes
5008:
5000:
4992:
4972:
4965:
4958:
4911:
4904:
4897:
4890:
4870:
4863:
4856:
4831:
4824:
4799:
4791:
4784:
4778:Ornithostoma
4776:
4769:
4761:
4732:
4672:
4586:
4578:
4571:
4564:
4557:
4550:
4545:Eurazhdarcho
4543:
4536:
4529:
4522:
4502:
4495:
4488:
4480:
4473:Azhdarchinae
4459:
4451:
4444:
4436:
4429:
4421:
4414:Azhdarchidae
4400:
4393:
4385:
4357:
4350:
4343:
4322:
4314:
4309:Meilifeilong
4307:
4300:
4292:
4284:
4276:
4252:
4227:
4219:
4211:
4203:
4195:
4188:
4166:
4159:
4152:
4144:
4094:
4087:
4067:
4060:
4053:
4045:
4021:
4013:
4005:
3997:
3989:
3965:
3958:
3951:
3946:Huaxiapterus
3944:
3937:
3931:Eopteranodon
3929:
3921:
3915:Afrotapejara
3913:
3906:Sinopterinae
3895:Afrotapejara
3893:
3869:
3862:
3831:
3824:
3817:
3809:
3801:
3794:
3786:
3778:
3770:
3762:
3740:
3735:Bennettazhia
3733:
3698:Ornithostoma
3696:
3689:
3681:
3673:
3666:
3659:
3625:
3618:
3597:
3589:
3584:Ordosipterus
3582:
3575:
3567:
3559:
3534:Tapejaroidea
3519:
3423:
3377:Pterodaustro
3375:
3368:
3361:
3354:
3334:
3310:
3303:
3296:
3290:Huanhepterus
3288:
3283:Gnathosaurus
3281:
3263:Moganopterus
3261:
3254:
3235:Pterofiltrus
3233:
3228:Petrodactyle
3226:
3219:
3214:Liaodactylus
3212:
3207:Kepodactylus
3205:
3198:
3191:
3184:
3177:
3170:
3163:
3155:
3122:Huanhepterus
3120:
3113:
3105:
3084:
3079:Aerodactylus
3077:
3052:
3046:Petrodactyle
3044:
3036:
3029:
3021:
2986:
2979:
2948:
2940:
2933:
2925:
2905:
2796:
2788:
2781:
2774:
2767:
2762:Kryptodrakon
2760:
2752:
2745:
2738:
2710:
2704:Jeholopterus
2702:
2695:
2694:
2687:
2680:
2660:
2653:
2647:Jeholopterus
2645:
2638:
2637:
2632:Anurognathus
2630:
2612:Mesadactylus
2610:
2575:
2567:
2529:
2521:
2514:
2507:
2499:
2479:
2455:
2448:
2437:Darwinoptera
2423:
2416:
2409:
2378:
2370:
2316:
2245:
2238:
2231:
2224:
2217:
2174:
2168:Nesodactylus
2166:
2160:Cacibupteryx
2158:
2140:Sericipterus
2138:
2130:
2122:
2115:
2108:
2079:
2072:
2067:Nesodactylus
2065:
2058:
2051:
2046:Cacibupteryx
2044:
2039:Bellubrunnus
2037:
2012:
2004:
1997:
1989:
1950:
1943:
1907:
1894:Lonchognatha
1875:
1867:
1859:
1852:
1845:
1837:
1817:
1786:
1747:
1739:
1718:
1694:
1687:
1682:Pachagnathus
1680:
1672:
1642:
1634:
1628:Austriadraco
1626:
1592:
1587:Austriadraco
1585:
1577:
1556:
1549:
1541:
1516:
1509:
1489:
1375:
1366:
1357:
1253:
1249:
1243:
1235:
1216:
1212:
1206:
1171:
1167:
1162:Jeholopterus
1161:
1153:
1134:
1130:
1120:
1111:
1107:
1103:
1097:
1078:
1074:
1064:
1039:
1035:
997:
993:
945:
941:
919:
915:
911:
865:
861:
859:
785:
784:
768:
767:
744:
743:
720:
719:
697:
696:
695:
660:
659:
643:
642:
609:
608:
582:
578:
570:
563:Jeholopterus
562:
558:
554:
550:
540:
532:
527:analysis by
522:
511:
482:
476:
475:
463:
459:
455:
451:
448:
443:Jeholopterus
441:
437:
412:
407:
395:
391:
377:
373:
369:
365:
362:David Keilin
339:
337:
317:
307:anurognathid
297:
296:
295:
279:
257:
252:
236:
235:
218:
217:
130:
126:
125:Holotype of
24:
18:
5966:Wikispecies
5732:Thapunngaka
5692:Amblydectes
5611:Anhangueria
5575:Aussiedraco
5309:Boreopterus
5084:Lonchodraco
5017:Lonchodraco
4960:Aussiedraco
4906:Nyctosaurus
4763:Bogolubovia
4431:Bogolubovia
4302:Lacusovagus
4294:Eoazhdarcho
4254:Eoazhdarcho
4213:Leptostomia
4205:Keresdrakon
4080:Caiuajarina
4047:Bakonydraco
4015:Lacusovagus
4007:Keresdrakon
3978:Tapejarinae
3939:Huaxiadraco
3923:Bakonydraco
3882:Tapejaridae
3811:Leptostomia
3803:Lacusovagus
3796:Kariridraco
3742:Keresdrakon
3683:Leptostomia
3675:Keresdrakon
3336:Ctenochasma
3298:Lusognathus
3221:Otogopterus
2876:Lophocratia
2809:Lophocratia
2776:Pangupterus
2712:Sinomacrops
2053:Dorygnathus
1922:Novialoidea
1877:Rhamphinion
1854:Dimorphodon
1529:Caviramidae
1473:Pterosauria
1460:Pterosauria
1423:Pterosauria
1362:Archosauria
1323:Pterosauria
1137:: 181–206.
472:Description
434:Lü Junchang
354:preoccupied
344:in 1998 by
192:Pterosauria
6076:Categories
5809:Anhanguera
5700:Ferrodraco
5655:Hamipterus
5583:Barbosania
5458:Ferrodraco
5450:Draigwenia
5285:Hamipterus
5264:Barbosania
5209:Nurhachius
5077:Ikrandraco
5052:Draigwenia
5002:Ikrandraco
4967:Serradraco
4913:Volgadraco
4801:Volgadraco
4793:Tethydraco
4786:Pteranodon
4644:see below↓
4538:Cryodrakon
4461:Volgadraco
4453:Tethydraco
4352:Jidapterus
4316:Microtuban
4154:Microtuban
4055:Europejara
4034:Tapejarini
4023:Vectidraco
3960:Sinopterus
3577:Noripterus
3446:see below↓
3370:Gegepterus
3363:Eosipterus
2818:see below↓
2798:Wenupteryx
2689:Cascocauda
2268:see below↓
1696:Yelaphomte
1432:see below↓
1353:Sauropsida
1256:: e11161.
890:References
331:Province,
5760:Aerodraco
5202:Luchibang
5150:Haopterus
4872:Simurghia
4865:Epapatelo
4833:Cretornis
4771:Dawndraco
4497:Azhdarcho
4490:Albadraco
4482:Aerotitan
4402:Alanqidae
4180:Alanqidae
4146:Cretornis
4096:Torukjara
4089:Caiuajara
3826:Tupuxuara
3764:Aerotitan
3668:Inabtanin
3256:Feilongus
2655:Luopterus
2372:Allkaruen
1999:Klobiodon
1839:Allkaruen
1674:Caviramus
1551:Caviramus
1339:Kingdom:
1221:CiteSeerX
1056:225169094
1014:140741278
587:basalmost
525:cladistic
400:in 2020.
397:Luopterus
358:parasitic
346:Ji Shu'an
310:pterosaur
230:Species:
165:Kingdom:
159:Eukaryota
5957:Q2855046
5951:Wikidata
5832:Guidraco
5716:Mythunga
5708:Haliskia
5482:Mythunga
5466:Guidraco
5397:Haliskia
4974:Unwindia
4324:Xericeps
4229:Xericeps
4161:Xericeps
4062:Tapejara
3833:Xericeps
3788:Banguela
3561:Banguela
2790:Samrukia
2450:Ceoptera
1347:Chordata
1345:Phylum:
1341:Animalia
1282:33850665
1198:19656798
873:See also
410:holotype
374:rhynkhos
350:Ji Qiang
316:species
314:holotype
271:Synonyms
198:Family:
179:Chordata
175:Phylum:
169:Animalia
155:Domain:
6040:Portals
6012:4947106
5999:1139047
5986:4818451
4858:Alcione
3967:Wightia
1351:Class:
1273:8019321
1189:2842671
972:4316586
950:Bibcode
485:humerus
413:GMV2128
370:dendron
255:†
234:†
211:Genus:
185:Order:
105:↓
4423:Alanqa
4190:Alanqa
3772:Alanqa
2380:Sordes
2247:Sordes
1280:
1270:
1223:
1196:
1186:
1054:
1012:
970:
942:Nature
502:fibula
500:. The
417:fossil
382:. The
301:was a
6066:China
6025:68167
5994:IRMNG
3521:Piksi
2117:Dearc
1376:Clade
1367:Clade
1358:Clade
1250:PeerJ
1052:S2CID
1010:S2CID
968:S2CID
539:with
537:clade
498:femur
494:tibia
388:Latin
356:by a
333:China
329:Hebei
303:genus
5981:GBIF
1409:see
1278:PMID
1194:PMID
561:and
489:ulna
424:the
415:, a
348:and
44:PreꞒ
1268:PMC
1258:doi
1184:PMC
1176:doi
1172:277
1139:doi
1083:doi
1044:doi
1002:doi
958:doi
946:398
305:of
6078::
6022::
6009::
5996::
5983::
5968::
5953::
1378::
1369::
1360::
1276:.
1266:.
1252:.
1248:.
1217:22
1215:.
1192:.
1182:.
1170:.
1166:.
1133:.
1129:.
1112:31
1110:.
1079:41
1077:.
1073:.
1050:.
1040:94
1038:.
1034:.
1022:^
1008:.
998:86
996:.
980:^
966:.
956:.
944:.
940:.
928:^
920:22
918:.
898:^
581:.
523:A
335:.
133:,
129:.
94:Pg
38:Ma
33:,
6042::
5827:?
5819:?
5784:?
5752:?
5727:?
5719:?
5711:?
5703:?
5695:?
5684:?
5647:?
5636:?
5628:?
5586:?
5578:?
5550:?
5523:?
5508:?
5500:?
5485:?
5477:?
5469:?
5461:?
5453:?
5445:?
5437:?
5429:?
5408:?
5400:?
5392:?
5384:?
5376:?
5132:?
5055:?
5028:?
5020:?
5005:?
4997:?
4916:?
4836:?
4804:?
4796:?
4781:?
4766:?
4583:?
4485:?
4464:?
4456:?
4441:?
4426:?
4405:?
4390:?
4327:?
4319:?
4297:?
4289:?
4281:?
4232:?
4224:?
4216:?
4208:?
4200:?
4182:?
4171:?
4149:?
4050:?
4018:?
4010:?
4002:?
3994:?
3934:?
3926:?
3918:?
3856:?
3836:?
3814:?
3806:?
3791:?
3783:?
3775:?
3767:?
3756:?
3745:?
3686:?
3678:?
3602:?
3594:?
3572:?
3564:?
3524:?
3428:?
3293:?
3248:?
3160:?
3125:?
3110:?
3089:?
3057:?
3049:?
3041:?
3026:?
2953:?
2945:?
2930:?
2793:?
2757:?
2707:?
2699:?
2650:?
2642:?
2572:?
2526:?
2504:?
2428:?
2383:?
2375:?
2250:?
2211:?
2201:?
2171:?
2163:?
2135:?
2127:?
2017:?
2009:?
1994:?
1912:?
1896:?
1880:?
1872:?
1864:?
1842:?
1791:?
1775:?
1744:?
1723:?
1677:?
1647:?
1639:?
1631:?
1582:?
1561:?
1546:?
1531:?
1315:e
1308:t
1301:v
1284:.
1260::
1254:9
1242:"
1229:.
1200:.
1178::
1147:.
1141::
1135:3
1091:.
1085::
1058:.
1046::
1016:.
1004::
974:.
960::
952::
215:†
202:†
189:†
127:D
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
