624:
3813:
643:
Mateus are a set of thoracic ribs. Two ribs are from the left side of the animal. They are T-shaped in cross section, and display plesiomorphic features, although their incomplete state makes their identification uncertain. Multiple right ribs are preserved, including both the shafts and heads. They are similar to the left ribs, which also show that they lack pneumatization. Other appendicular (non-vertebral) material includes a very incomplete and fragmentary shaft of the pubis, and over one hundred
3824:
3831:
5225:
148:
5957:
7192:
7186:
5230:
684:, with the first and second dorsals similar in anatomy to the last and second-last cervical. The dorsal vary in length from the 58 cm (23 in) of the first dorsal to the 25 cm (9.8 in) of the seventh, eight and ninth dorsals. Height in the vertebrae is also quite variable, with the shortest height being 51 cm (20 in) tall to 76 cm (30 in) tall, increasing from the first dorsal.
4687:
7181:
4679:
7175:
5949:
3819:
4699:
130:
4693:
5965:
668:. As of the original description, the thirteenth cervical was only prepared on the lateroventral portion. The length of the centrum is 71 cm (28 in), and the fourteenth cervical is quite similar overall. 63 cm (25 in) is the total measurement of the 14th cervical's centrum, which is well-preserved, complete, and concave along the bottom edge. The
1826:
663:
comes from the vertebrae, which are well represented and described. Of the cervicals, only two of the assumed fifteen are preserved. According to
Bonaparte & Mateus (1999), the cervicals would number 13 and 14. Apparently cervical 15 was lost during the excavation and removal of the holotype and
679:
A relatively complete series of dorsal vertebrae are known, which number one to seven. All of the dorsals, however, are distorted upwards due to their state of preservation. Bonaparte & Mateus (1999) noted that the position of the dorsals was not certain, and that in fact the first dorsal could
642:
The animal is not known well from non-vertebral material, currently only consisting of partial ribs and a fragment of a pelvis. One of the ribs attached to the cervicals, and is quite fragmentary. It is elongated, although that might be a feature of distortion. Also undescribed by
Bonaparte &
595:
regions, and are articulated. The two cervicals are not greatly preserved, although the twelve dorsals are articulated and in good condition. Other vertebral material includes seven centra that are fragmentary and a few neural arches, which are unattached. 12 dorsal ribs are preserved, as well as
516:
preliminarily announced ML 414 as soon as the excavations were complete. To remove the fossils from the surrounding rock, a bulldozer and tilt hammer were needed. The fossils were situated at the top of a coastal cliff, and once removed, were shipped to
LourinhĂŁ in two blocks with the help of a
2416:
1380:
possessed a whip-tail. If it did, it has been speculated that its tail could have been used like a bullwhip, with supersonic speed or, more recently, as a tactile organ to keep in touch with other members of a group. Being related to both
1669:, confirming the presence of a least two and possibly more diplodocids in the Late Jurassic of Spain and Portugal. This is unique in the variety of diplodocoids in all Europe, with the only other genera possibly non-diplodocoid (
2796:
2515:
1430:. The analysis took into account that among the hundreds of sauropods found, gastroliths are only known from a few associated specimens. Authors chose to use the three sauropods with the most associated gastroliths,
634:
was an average sized diplodocid, and had an elongated neck and tail. The main features of the genus are based on its vertebral anatomy, and multiple vertebrae from across the spine have been found. In total,
2374:
1980:
1677:. This suggests that the biogeography of primitive sauropods is incomplete, with possible primitive eusauropods and diplodocids surviving in the Late Jurassic, potentially until the
2375:"Abelisauridae (Dinosauria: Theropoda) Teeth from the Late Jurassic of Portugal and Dentition-Based Phylogeny as a Contribution for the Identification of Isolated Theropod Teeth"
1454:
had less gastroliths to body mass, an avian-style gastric mill is unlikely to have evolved in sauropods, and they instead might have used gastroliths to absorb minerals.
2002:
512:, and after the excavations which lasted from the time of discovery until 1992, the specimen was then moved into the museum, and catalogued under the number 414. Dantas
2143:
2698:
Escaso, F.; Ortega, F.; Dantas, P.; Malafaia, E.; Pimentel, N. L.; Pereda-Suberbiola, X.; Sanz, J. L.; Kullberg, J. C.; Kullberg, M. A. C.; Barriga, F. (2006).
2101:
Mocho, P.; Royo-Torres, R.; Escaso, F.; Malafaia, E.; de Miguel Chaves, C.; Narvaez, I.; PĂ©rez-GarcĂa, A.; Pimentel, N.; Silva, B.C. & Ortega, F. (2017).
2647:
607:, and only found 9 dorsals in the holotype, while also misinterpreting the pubis as a limb fragment. They also incorrectly stated that it was found in the
1442:, because of the large amount of gastroliths found in birds. When birds were typically found to have 1.05% of their body weight gastroliths, the sauropod
7332:
2886:
2915:
7362:
7357:
7281:
2417:"The large theropod fauna of the LourinhĂŁ Formation (Portugal) and its similarity to the Morrison Formation, with a description of a new species of
1931:
1768:
7352:
2842:
1903:
759:; their results are shown below. However, to either confirm or refute this hypothesis, several remains of the holotype need to be prepared.
1404:
is one of relatively few sauropods for which gastroliths were found obviously alongside the type specimen. In 2007, an experiment using
2627:
Thulborn, R.A. (1973). "Teeth of ornithischian dinosaurs from the Upper
Jurassic of Portugal, with description of a hypsilophodontid (
1661:. One sauropod, a diplodocid currently based on an unnamed specimen including vertebrae and some bones, is clearly different from
3823:
3812:
3830:
2454:
n. sp., the
Largest Terrestrial Predator from Europe, and a Proposed Terminology of the Maxilla Anatomy in Nonavian Theropods"
419:. Of the material, only the vertebrae are diagnostic, with the ribs and pubis being too fragmentary or general to distinguish
5224:
1930:
Royo-Torres, R.; Cobos, A.; Aberasturi, A.; EspĂlez, E.; Fierro, I.; González, A.; Luque, L.; Mampel, L.; Alcala, L. (2007).
7347:
1446:, which had the highest amount of gastroliths, only amassed to 0.03% body weight. This means that since the other sauropods
1322:, for the differences anatomically are quite great. Bonaparte & Mateus found that a few features present suggested that
2908:
147:
7342:
1544:
2103:"Upper Jurassic sauropod record in the Lusitanian Basin (Portugal): Geographical and lithostratigraphical distribution"
5956:
647:. The pubis displays practically no anatomical features, and the gastroliths were not described in detail by Mannion
7337:
2605:
Ruiz-Omeñaca, J.I.; Canudo, J.I. (2004). "Dinosaurios ornitĂłpodos del Cretácico inferior de la PenĂnsula IbĂ©rica".
639:
would have had an approximate length of 20–25 metres (66–82 ft) and weighed 8.8 metric tons (9.7 short tons).
623:
7191:
7185:
2323:"No gastric mill in sauropod dinosaurs: new evidence from analysis of gastrolith mass and function in ostriches"
7228:
5229:
3003:
2901:
1393:
probably possessed a squared snout. This means that it was probably a non-selective ground-feeding sauropod.
7327:
2264:"Inferences of diplodocoid (Sauropoda: Dinosauria) feeding behavior from snout shape and microwear snalyses"
4686:
1990:. Universidade de Coimbra e Centro de Estudos e Filosofia da História da Ciência Coimbra. pp. 121–126.
7249:
4698:
4678:
2633:
Contribuição para o conhecimento da Fauna do
Kimerridgiano da Mina de Lignito Guimarota (Leiria, Portugal)
1604:
1512:
915:
608:
1954:
1795:
7294:
7218:
7180:
6909:
3191:
2137:
1595:
891:
717:
550:
430:
314:
7174:
5948:
4692:
2700:"New evidence of shared dinosaur across Upper Jurassic Proto-North Atlantic: Stegosaurus from Portugal"
1703:"A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda)"
1466:
739:. However, a cladogram from 2014 found that their group was supported, but in fact more primitive than
429:, but differences were noticed and in 1999 Bonaparte and Mateus redescribed the material under the new
385:
5964:
3818:
1831:(Sauropoda: Diplodocoidea) from the Late Jurassic of Portugal, with a review of European diplodocoids"
7322:
2857:
2815:
2769:
2714:
2534:
2465:
2275:
2173:
2055:
2024:
1589:
1494:
1063:
712:
509:
291:
700:
3894:
1583:
1527:
1168:
1151:
1096:
932:
858:
545:, concluding one specimen, under the name ML 414, to be more closely related to diplodocids of the
6949:
4021:
2738:
2550:
2244:
2197:
2189:
2125:
1861:
1613:
1015:
780:
615:
by
Bonaparte and Mateus, but is now considered to be a distinct new unnamed genus of diplodocid.
546:
400:
142:
2217:"Tactile tails: a new hypothesis for the function of the elongate tails of diplodocid sauropods"
7299:
1895:
7085:
6626:
5373:
5014:
4743:
3931:
2801:, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of LourinhĂŁ, Portugal"
2730:
2680:
2493:
2397:
2352:
2303:
2236:
2083:
1946:
1909:
1899:
1853:
1787:
1734:
1657:
1651:
have been found in the Late
Jurassic of Europe. The sauropods are from around the base of the
1218:
1201:
536:
532:
377:
373:
287:
283:
17:
6956:
6801:
6741:
6383:
6275:
6181:
6072:
5874:
5854:
5793:
4912:
4765:
4612:
4329:
4283:
4263:
4207:
3510:
3287:
3267:
2865:
2823:
2777:
2722:
2670:
2662:
2585:
2542:
2483:
2473:
2389:
2342:
2334:
2293:
2283:
2228:
2181:
2117:
2073:
2063:
1887:
1845:
1724:
1714:
1674:
1574:
841:
597:
572:
404:
381:
2520:, a new turiasaurian sauropod (Dinosauria, Eusauropoda) from the Late Jurassic of Portugal"
388:. Although the precise age of the formation is not known, it can be dated around the early
7223:
7093:
6826:
6719:
6415:
6395:
6376:
6334:
6261:
6247:
6130:
5881:
5801:
5448:
4984:
4975:
4836:
4828:
4755:
4515:
4393:
4214:
4070:
3901:
3630:
3623:
3615:
3602:
3519:
3065:
3052:
3015:
2988:
2963:
2924:
1558:
1518:
1120:
221:
2861:
2819:
2773:
2718:
2648:"A new long-necked 'sauropod-mimic' stegosaur and the evolution of the plated dinosaurs"
2538:
2469:
2279:
2177:
2059:
2028:
680:
have been the last cervical or even the second dorsal. A similar numbering was found in
611:. Another pair of vertebrae, under collection number ML 418, was originally assigned to
508:
ML 414 was first uncovered in 1987 by Mr. Carlos
Anunciação. He was associated with the
7100:
7015:
6897:
6842:
6792:
6604:
6597:
6543:
6515:
6470:
6443:
6318:
6283:
6225:
6152:
6065:
5829:
5708:
5680:
5659:
5644:
5566:
5539:
5357:
5341:
5307:
4949:
4821:
4647:
4386:
4365:
4314:
4306:
4158:
4151:
4058:
3957:
3945:
3753:
3725:
3562:
3533:
3526:
3472:
3386:
3372:
3356:
3331:
3161:
2675:
2635:. Vol. 22. Serviços Geológicos de Portugal, Memória (Nova Série). pp. 89–134.
2488:
2449:
2347:
2322:
2298:
2263:
2078:
2043:
1729:
1702:
1600:
1553:
1540:
1523:
1499:
1352:
558:
541:
425:
2869:
2827:
2781:
2699:
2158:
2102:
7316:
7143:
7128:
7107:
7023:
6983:
6971:
6941:
6890:
6862:
6835:
6756:
6733:
6681:
6612:
6536:
6502:
6491:
6429:
6348:
6311:
6290:
6240:
6218:
6210:
6203:
6196:
6137:
6102:
6079:
6021:
5888:
5867:
5735:
5726:
5715:
5694:
5673:
5651:
5594:
5471:
5408:
5292:
5115:
5091:
5057:
5028:
5021:
5000:
4935:
4928:
4921:
4886:
4877:
4814:
4806:
4799:
4729:
4716:
4633:
4557:
4500:
4479:
4428:
4421:
4414:
4407:
4400:
4357:
4336:
4255:
4232:
4079:
4036:
3938:
3915:
3887:
3718:
3711:
3659:
3555:
3492:
3483:
3365:
3239:
3207:
3175:
3136:
3118:
3097:
2248:
1888:
1508:
1486:
1482:
1364:
1331:
800:
393:
250:
80:
42:
2760:
n. gen. and n. sp., an ankylosaurian dinosaur from the Upper
Jurassic of Portugal".
2742:
2554:
2201:
2129:
1865:
7151:
7121:
7114:
7051:
6992:
6918:
6850:
6808:
6777:
6763:
6748:
6726:
6710:
6644:
6633:
6584:
6565:
6558:
6436:
6422:
6408:
6362:
6355:
6326:
6297:
6233:
6174:
6116:
6109:
6058:
6051:
6014:
5996:
5983:
5911:
5895:
5836:
5822:
5786:
5779:
5765:
5743:
5637:
5629:
5622:
5601:
5573:
5521:
5507:
5493:
5478:
5429:
5350:
5322:
5315:
5134:
5123:
5100:
5082:
5007:
4993:
4900:
4857:
4850:
4792:
4785:
4548:
4463:
4271:
4221:
4193:
4186:
4165:
4137:
4123:
4043:
3987:
3966:
3652:
3540:
3444:
3400:
3393:
3345:
3317:
3278:
3104:
1619:
1532:
1503:
1490:
1478:
1427:
1418:
1358:
981:
766:
744:
720:
696:
669:
412:
263:
2232:
1566:
known. Ornithischians are well represented, with identified remains persisting to
2546:
2478:
2288:
2068:
1849:
7136:
7044:
7030:
6926:
6815:
6770:
6667:
6660:
6653:
6619:
6551:
6529:
6450:
6341:
6036:
5843:
5808:
5701:
5666:
5608:
5587:
5580:
5552:
5485:
5457:
5422:
5415:
5401:
5300:
5285:
5186:
5108:
5071:
4942:
4893:
4654:
4530:
4493:
4486:
4455:
4439:
4379:
4343:
4321:
4241:
4200:
4144:
4116:
4102:
4050:
4003:
3923:
3908:
3880:
3746:
3704:
3690:
3670:
3644:
3590:
3583:
3499:
3465:
3451:
3294:
3232:
3168:
3145:
3111:
3083:
2216:
2003:"THE SAUROPOD DINOSAUR TURIASAURUS RIODEVENSIS IN THE LATE JURASSIC OF PORTUGAL"
1644:
1609:
1579:
1568:
1549:
584:
562:
527:
482:
464:
449:
416:
362:
55:
1825:
Mannion, P.D.; Upchurch, Paul; Mateus, O.; Barnes, R.N.; Jones, M.E.H. (2012).
7037:
6964:
6933:
6695:
6522:
6369:
6304:
6268:
6254:
6189:
6160:
6123:
6086:
6029:
5615:
5559:
5514:
5464:
5436:
5394:
5330:
5278:
5260:
5247:
5171:
5164:
5157:
5143:
5050:
5043:
5036:
4956:
4868:
4843:
4573:
4523:
4507:
4471:
4372:
4248:
4172:
4086:
4029:
3980:
3732:
3548:
3437:
3301:
3260:
3225:
2954:
2726:
2393:
2185:
1678:
1631:
644:
476:
470:
453:
350:
205:
100:
65:
7272:
2240:
1950:
1932:"Riodeva sites (Teruel, Spain) shedding light to European Sauropod phylogeny"
1913:
1857:
1791:
7000:
6882:
6688:
6167:
6145:
6094:
5815:
5772:
5758:
5500:
5193:
5179:
5064:
4778:
4626:
4619:
4350:
4299:
3862:
3849:
3764:
3739:
3697:
3576:
3416:
3183:
3125:
1652:
1625:
1562:
1481:
are also from the LourinhĂŁ Formation, which contains a similar fauna to the
707:
488:
389:
237:
159:
129:
105:
49:
2734:
2684:
2666:
2497:
2401:
2356:
2338:
2307:
2087:
2044:"A Diplodocid Sauropod Survivor from the Early Cretaceous of South America"
1738:
1485:
Morrison Formation. Many theropods are known including an unnamed genus of
7266:
7008:
6674:
6484:
5687:
5528:
5387:
4640:
4130:
3973:
3683:
3458:
3430:
3338:
3253:
3246:
3153:
3090:
2972:
2948:
1474:
1470:
445:
440:, is now known to be from another Portuguese diplodocid. This means that
357:
353:
192:
179:
95:
90:
75:
70:
60:
699:
position is not certain. In 2012 during a redescription of the taxon by
7286:
6044:
4448:
4292:
4179:
3637:
2193:
110:
85:
2590:
2569:
1894:(Second ed.). Berkeley: University of California Press. pp.
1719:
6573:
5751:
3569:
2942:
2042:
Gallina, P. A.; ApesteguĂa, S. N.; Haluza, A.; Canale, J. I. (2014).
592:
580:
169:
7243:
2574:(Macronaria) and the evolutionary history of basal titanosauriforms"
1986:. In Brandao, J.M.; Callapez, P.M.; Mateus, O.; et al. (eds.).
2021:
Dinosaur Facts and Figures: The Sauropods and Other Sauropodomorphs
2893:
2448:
Hendrickx, C.; Mateus, O.V. (2014). Evans, Alistair Robert (ed.).
1981:"Paleontological Collections of the Museum of LourinhĂŁ (Portugal)"
622:
346:
2121:
1334:
features also present conclude that they branched separately and
6875:
3379:
1423:
588:
576:
408:
7247:
7207:
5981:
5245:
4714:
3847:
3050:
2935:
2897:
4537:
676:, is massive, and projects upwards towards its posterior end.
2570:"Osteology of the Late Jurassic Portuguese sauropod dinosaur
2568:
Mannion, P.D.; Upchurch, P.; Barnes, R.; Mateus, O. (2013).
2443:
2441:
517:
crane. A year before being described as a new taxon, Dantas
356:
dinosaur that is known from fossils uncovered in modern-day
2368:
2366:
1827:"New information on the anatomy and systematic position of
672:, while compressed from above compared to the cervicals of
1773:
gen. et sp. nov., from the Late Jurassic Beds of Portugal"
1762:
1760:
1758:
1756:
1754:
1752:
1750:
1748:
727:
and using a modified matrix of Whitlock (2011) found that
2428:
New Mexico Museum of Natural History and Science Bulletin
2159:"Supersonic sauropods? Tail dynamics in the diplodocids"
2023:. New Jersey: Princeton University Press. p. 256.
1531:. Sauropods are less common, with only an intermediate
695:
is not extremely well known, and as a consequence, its
1925:
1923:
1881:
1879:
1877:
1875:
423:. This material was first described as in the genus
7256:
7077:
6982:
6908:
6861:
6825:
6791:
6709:
6643:
6583:
6501:
6468:
6394:
5994:
5910:
5853:
5725:
5538:
5447:
5371:
5340:
5258:
5133:
5090:
5081:
4983:
4974:
4911:
4876:
4867:
4764:
4753:
4727:
4604:
4572:
4556:
4547:
4438:
4282:
4231:
4100:
4069:
4001:
3956:
3860:
3763:
3669:
3601:
3509:
3482:
3414:
3355:
3315:
3277:
3205:
3135:
3063:
3014:
2987:
2646:Mateus, O.; Maidment, S.; Christiansen, N. (2009).
1886:Weishampel, D.B.; Dodson, P.; OsmĂłlska, H. (2004).
603:did not recognize all the material as belonging to
1701:Tschopp, E.; Mateus, O.; Benson, R. B. J. (2015).
1696:
1694:
2509:
2507:
1820:
1818:
1816:
1814:
1812:
1810:
1808:
1780:Revista del Museo Argentino de Ciencias Naturales
1422:tested if sauropods used their gastroliths in an
2631:gen. et sp. nov.) from the Guimarota lignite".
2622:
2620:
2514:Mateus, O; Mannion, P.D.; Upchurch, P. (2014).
1988:Colecções e museus de Geologia: missão e gestão
1465:was one of many dinosaurs to have lived in the
1318:was classified within a Diplodocidae excluding
2909:
2415:Mateus, O.; Walen, A.; Antunes, M.T. (2006).
8:
2655:Proceedings of the Royal Society of London B
2262:Whitlock, J. A. (2011). Farke, A. A. (ed.).
2142:: CS1 maint: multiple names: authors list (
1350:was an intermediate diplodocoid, along with
1047:Supersaurus (=Dinheirosaurus) lourinhanensis
456:, as well as at least one other diplodocid.
7244:
7204:
7074:
6867:
6822:
6589:
6507:
6476:
6400:
6006:
6002:
5991:
5978:
5859:
5544:
5379:
5270:
5266:
5255:
5242:
5087:
4980:
4873:
4770:
4761:
4735:
4724:
4711:
4601:
4553:
4108:
4013:
4009:
3872:
3868:
3857:
3844:
3675:
3607:
3422:
3352:
3323:
3217:
3213:
3075:
3071:
3060:
3047:
2993:
2984:
2932:
2916:
2902:
2894:
2756:Galton, P.M. (1980). "Partial skeleton of
436:. Another specimen, ML 418, thought to be
128:
31:
2674:
2589:
2578:Zoological Journal of the Linnean Society
2487:
2477:
2346:
2297:
2287:
2077:
2067:
1728:
1718:
723:. A 2012 cladogram, published by Mannion
587:. The vertebrae were certainly from the
553:name. This new species was described as
1690:
492:. Among those, the closest relative to
2135:
2019:Molina-Perez & Larramendi (2020).
2157:Myhrvold, N.P.; Currie, P.J. (1997).
2008:. Journal of Vertebrate Paleontology.
1376:As a diplodocid, it is probable that
7:
2215:Baron, Matthew G. (3 October 2021).
1522:; an intermediate theropod; and the
659:The most distinguishing material of
1838:Journal of Systematic Palaeontology
1193:
1143:
1136:
1112:
1088:
1038:
1031:
1007:
987:
907:
883:
833:
826:
806:
796:
772:
762:
2841:Antunes, M.T.; Mateus, O. (2003).
2795:Mateus, O.; Antunes, M.T. (2001).
2527:Journal of Vertebrate Paleontology
2373:Hendrickx, C.; Mateus, O. (2014).
2327:Proceedings of the Royal Society B
1767:Bonaparte, J.; Mateus, O. (1999).
25:
7333:Late Jurassic dinosaurs of Europe
7209:Topics in sauropodomorph research
384:and some other material from the
7190:
7184:
7179:
7173:
5963:
5955:
5947:
5228:
5223:
4697:
4691:
4685:
4677:
3829:
3822:
3817:
3811:
2321:Wings, O.; Sander, M.P. (2007).
399:The known material includes two
360:. It may represent a species of
146:
1469:during the Late Jurassic. Many
462:is a diplodocid, a relative of
716:and together forming the most
1:
7353:Fossil taxa described in 1999
7258:Dinheirosaurus lourinhanensis
2870:10.1016/S1631-0683(03)00003-4
2828:10.1016/S0753-3969(01)88003-4
2782:10.1016/s0016-6995(80)80081-7
2233:10.1080/08912963.2020.1769092
1829:Dinheirosaurus lourinhanensis
1771:Dinheirosaurus lourinhanensis
749:Dinheirosaurus lourinhanensis
609:Camadas de Alcobaça Formation
555:Dinheirosaurus lourinhanensis
434:Dinheirosaurus lourinhanensis
369:Dinheirosaurus lourinhanensis
327:Dinheirosaurus lourinhanensis
18:Dinheirosaurus lourinhanensis
7363:Taxa named by Octávio Mateus
7358:Taxa named by José Bonaparte
2547:10.1080/02724634.2013.822875
2479:10.1371/journal.pone.0088905
2289:10.1371/journal.pone.0018304
2069:10.1371/journal.pone.0097128
1850:10.1080/14772019.2011.595432
1658:Anchispirocyclina lusitanica
1655:as based on the presence of
1545:Lourinhasaurus alenquerensis
751:was considered a species of
706:, it was recovered, in both
596:some appendicular elements.
549:, and thus warranting a new
523:Lourinhasaurus alenquerensis
333:Bonaparte & Mateus, 1999
137:3rd to 9th dorsal vertebrae
1342:. A 2004 study by Upchurch
525:, previously grouped under
7379:
2110:Palaeontologia Electronica
1504:Ceratosaurus dentisculatus
710:, to be sister species to
557:, with a full meaning of "
27:Genus of reptiles (fossil)
7214:
7203:
7171:
7073:
6870:
6592:
6510:
6479:
6403:
6009:
6005:
5990:
5977:
5944:
5862:
5547:
5382:
5273:
5269:
5254:
5241:
5221:
4773:
4738:
4723:
4710:
4674:
4600:
4111:
4016:
4012:
3875:
3871:
3856:
3843:
3809:
3678:
3610:
3425:
3326:
3220:
3216:
3078:
3074:
3059:
3046:
2996:
2983:
2931:
2727:10.1007/s00114-006-0209-8
2394:10.11646/zootaxa.3759.1.1
2186:10.1017/S0094837300019801
1338:is not the descendant of
1215:
1198:
1191:
1165:
1148:
1141:
1134:
1117:
1110:
1093:
1086:
1060:
1043:
1036:
1029:
1012:
1005:
992:
985:
929:
912:
905:
888:
881:
855:
838:
831:
824:
811:
804:
794:
777:
770:
320:
313:
143:Scientific classification
141:
136:
127:
34:
7229:List of sauropod species
2808:Annales de Paléontologie
743:, and therefore outside
731:was more primitive than
539:studied the material of
2843:"Dinosaurs of Portugal"
2850:Comptes Rendus Palevol
2758:Dracopelta zbyszewskii
2667:10.1098/rspb.2008.1909
2339:10.1098/rspb.2006.3763
1605:Dracopelta zbyszewskii
1513:Aviatyrannis jurassica
1326:was more derived than
916:Brontosaurus yahnahpin
735:and more derived than
628:
366:. The only species is
306:D. lourinhanensis
7295:Paleobiology Database
7219:Sauropod neck posture
6910:Opisthocoelicaudiinae
2572:Lusotitan atalaiensis
1801:on 28 September 2007.
1596:Stegosaurus ungulatus
892:Brontosaurus excelsus
755:in a new combination
626:
444:lived alongside many
372:, first described by
1960:on 24 September 2015
1673:), or classified in
1590:Miragaia longicollum
1569:Trimucrodon cuneatus
1495:Allosaurus europaeus
1064:Supersaurus vivianae
713:Supersaurus vivianae
504:Discovery and naming
7343:Fossils of Portugal
7078:Dubious titanosaurs
3895:Chinshakiangosaurus
2862:2003CRPal...2...77A
2820:2001AnPal..87...61M
2774:1980Geobi..13..451G
2719:2007NW.....94..367E
2707:Naturwissenschaften
2661:(1663): 1815–1821.
2539:2014JVPal..34..618M
2470:2014PLoSO...988905H
2452:Torvosaurus gurneyi
2280:2011PLoSO...618304W
2178:1997Pbio...23..393M
2060:2014PLoSO...997128G
2029:2020dffs.book.....M
2001:Mateus, O. (2009).
1979:Mateus, O. (2010).
1769:"A New Diplodocid,
1584:Dacentrurus armatus
1528:Torvosaurus gurneyi
1169:Diplodocus hallorum
1152:Diplodocus carnegii
1097:Leinkupal laticauda
933:Brontosaurus parvus
859:Apatosaurus louisae
521:assigned ML 414 to
7348:LourinhĂŁ Formation
6950:Opisthocoelicaudia
4022:Archaeodontosaurus
2799:Draconyx loureiroi
2221:Historical Biology
1626:Phyllodon henkelli
1614:Draconyx loureiroi
1467:LourinhĂŁ Formation
1016:Tornieria africana
781:Amphicoelias altus
629:
571:material included
547:Morrison Formation
411:, a fragment of a
401:cervical vertebrae
386:LourinhĂŁ Formation
7338:Jurassic Portugal
7310:
7309:
7250:Taxon identifiers
7241:
7240:
7237:
7236:
7199:
7198:
7169:
7168:
7165:
7164:
7161:
7160:
7086:Bruhathkayosaurus
7069:
7068:
7065:
7064:
7061:
7060:
6787:
6786:
6705:
6704:
6627:Panamericansaurus
6464:
6463:
6460:
6459:
5973:
5972:
5942:
5941:
5938:
5937:
5934:
5933:
5930:
5929:
5906:
5905:
5374:Titanosauriformes
5367:
5366:
5237:
5236:
5219:
5218:
5215:
5214:
5211:
5210:
5207:
5206:
5203:
5202:
5015:Brachytrachelopan
4970:
4969:
4966:
4965:
4744:Haplocanthosaurus
4706:
4705:
4672:
4671:
4668:
4667:
4664:
4663:
4605:Dubious sauropods
4596:
4595:
4592:
4591:
4588:
4587:
4096:
4095:
3997:
3996:
3932:Protognathosaurus
3839:
3838:
3807:
3806:
3803:
3802:
3799:
3798:
3795:
3794:
3791:
3790:
3787:
3786:
3783:
3782:
3410:
3409:
3311:
3310:
3201:
3200:
3042:
3041:
3038:
3037:
3034:
3033:
2629:Phyllodon henkeli
2591:10.1111/zoj.12029
2333:(1610): 635–640.
2227:(10): 2057–2066.
1905:978-0-520-94143-4
1720:10.7717/peerj.857
1477:, and especially
1311:
1310:
1302:
1301:
1293:
1292:
1284:
1283:
1275:
1274:
1266:
1265:
1257:
1256:
1248:
1247:
1239:
1238:
1230:
1229:
1219:Barosaurus lentus
1202:Kaatedocus siberi
1180:
1179:
1075:
1074:
971:
970:
962:
961:
953:
952:
944:
943:
870:
869:
757:S. lourinhanensis
664:only specimen of
510:Museu da LourinhĂŁ
338:
337:
294:
16:(Redirected from
7370:
7303:
7302:
7290:
7289:
7277:
7276:
7275:
7245:
7205:
7194:
7188:
7183:
7177:
7075:
6957:Pellegrinisaurus
6868:
6823:
6802:Bustingorrytitan
6742:Futalognkosaurus
6590:
6508:
6477:
6401:
6384:Diamantinasauria
6276:Pellegrinisaurus
6182:Jiangshanosaurus
6073:Baotianmansaurus
6007:
6003:
5992:
5979:
5968:
5967:
5960:
5959:
5952:
5951:
5875:Diamantinasaurus
5860:
5855:Diamantinasauria
5794:Jiangshanosaurus
5545:
5380:
5271:
5267:
5256:
5243:
5232:
5227:
5088:
4981:
4913:Rebbachisaurinae
4874:
4771:
4766:Rebbachisauridae
4762:
4736:
4725:
4712:
4701:
4695:
4690:
4689:
4682:
4681:
4613:Bothriospondylus
4602:
4554:
4330:Eomamenchisaurus
4284:Mamenchisauridae
4264:Lapparentosaurus
4208:Spinophorosaurus
4109:
4014:
4010:
3873:
3869:
3858:
3845:
3834:
3833:
3826:
3821:
3815:
3676:
3608:
3511:Massospondylidae
3423:
3353:
3324:
3288:Jaklapallisaurus
3268:Thecodontosaurus
3218:
3214:
3076:
3072:
3061:
3048:
2994:
2985:
2978:
2977:
2933:
2918:
2911:
2904:
2895:
2874:
2873:
2847:
2838:
2832:
2831:
2805:
2792:
2786:
2785:
2753:
2747:
2746:
2704:
2695:
2689:
2688:
2678:
2652:
2643:
2637:
2636:
2624:
2615:
2614:
2602:
2596:
2595:
2593:
2565:
2559:
2558:
2524:
2511:
2502:
2501:
2491:
2481:
2445:
2436:
2435:
2425:
2412:
2406:
2405:
2379:
2370:
2361:
2360:
2350:
2318:
2312:
2311:
2301:
2291:
2259:
2253:
2252:
2212:
2206:
2205:
2163:
2154:
2148:
2147:
2141:
2133:
2107:
2098:
2092:
2091:
2081:
2071:
2039:
2033:
2032:
2016:
2010:
2009:
2007:
1998:
1992:
1991:
1985:
1976:
1970:
1969:
1967:
1965:
1959:
1953:. Archived from
1936:
1927:
1918:
1917:
1893:
1883:
1870:
1869:
1835:
1822:
1803:
1802:
1800:
1794:. Archived from
1777:
1764:
1743:
1742:
1732:
1722:
1698:
1675:Rebbachisauridae
1575:Alocodon kuehnei
1194:
1144:
1137:
1113:
1089:
1039:
1032:
1008:
994:Unnamed species
988:
908:
884:
842:Apatosaurus ajax
834:
827:
813:Unnamed species
807:
797:
773:
763:
598:David Weishampel
405:dorsal vertebrae
329:
325:
282:
275:
262:
249:
236:
220:
151:
150:
132:
122:
52:
40:Temporal range:
32:
21:
7378:
7377:
7373:
7372:
7371:
7369:
7368:
7367:
7313:
7312:
7311:
7306:
7298:
7293:
7285:
7280:
7271:
7270:
7265:
7252:
7242:
7233:
7224:Sauropod hiatus
7210:
7195:
7157:
7094:Campylodoniscus
7057:
6978:
6904:
6857:
6827:Nemegtosauridae
6821:
6783:
6720:Argentinosaurus
6701:
6639:
6579:
6497:
6473:
6456:
6416:Atsinganosaurus
6396:Lirainosaurinae
6390:
6377:Xianshanosaurus
6262:Normanniasaurus
6248:Narambuenatitan
6131:Dongyangosaurus
5999:
5986:
5969:
5962:
5954:
5946:
5926:
5902:
5882:Sarmientosaurus
5849:
5802:Phuwiangosaurus
5721:
5534:
5449:Brachiosauridae
5443:
5376:
5363:
5336:
5263:
5250:
5233:
5199:
5129:
5077:
4985:Dicraeosauridae
4976:Flagellicaudata
4962:
4907:
4863:
4837:Nopcsaspondylus
4829:Maraapunisaurus
4758:
4756:Diplodocimorpha
4749:
4732:
4719:
4702:
4684:
4676:
4660:
4584:
4568:
4543:
4516:Tehuelchesaurus
4434:
4394:Tienshanosaurus
4278:
4227:
4215:Tehuelchesaurus
4105:
4092:
4071:Vulcanodontidae
4065:
4006:
3993:
3952:
3902:Gongxianosaurus
3865:
3852:
3835:
3828:
3779:
3759:
3665:
3624:Jingshanosaurus
3616:Chuxiongosaurus
3603:Sauropodiformes
3597:
3520:Adeopapposaurus
3505:
3478:
3419:
3406:
3351:
3320:
3307:
3273:
3210:
3197:
3131:
3068:
3066:Sauropodomorpha
3055:
3053:Sauropodomorpha
3030:
3016:Sauropodomorpha
3010:
3004:Avemetatarsalia
2989:Avemetatarsalia
2979:
2964:Avemetatarsalia
2938:
2937:
2927:
2925:Sauropodomorpha
2922:
2883:
2878:
2877:
2845:
2840:
2839:
2835:
2803:
2794:
2793:
2789:
2755:
2754:
2750:
2702:
2697:
2696:
2692:
2650:
2645:
2644:
2640:
2626:
2625:
2618:
2604:
2603:
2599:
2567:
2566:
2562:
2522:
2513:
2512:
2505:
2447:
2446:
2439:
2423:
2414:
2413:
2409:
2377:
2372:
2371:
2364:
2320:
2319:
2315:
2261:
2260:
2256:
2214:
2213:
2209:
2161:
2156:
2155:
2151:
2134:
2105:
2100:
2099:
2095:
2041:
2040:
2036:
2018:
2017:
2013:
2005:
2000:
1999:
1995:
1983:
1978:
1977:
1973:
1963:
1961:
1957:
1934:
1929:
1928:
1921:
1906:
1885:
1884:
1873:
1833:
1824:
1823:
1806:
1798:
1775:
1766:
1765:
1746:
1700:
1699:
1692:
1687:
1641:
1519:Richardoestesia
1509:coelurosaurians
1460:
1399:
1374:
1312:
1303:
1294:
1285:
1276:
1267:
1258:
1249:
1240:
1231:
1181:
1121:Galeamopus hayi
1076:
972:
963:
954:
945:
871:
690:
657:
627:Size comparison
621:
506:
334:
331:
323:
322:
309:
281:
273:
260:
247:
234:
222:Sauropodomorpha
218:
145:
123:
121:
120:
119:
118:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
47:
46:
41:
38:
28:
23:
22:
15:
12:
11:
5:
7376:
7374:
7366:
7365:
7360:
7355:
7350:
7345:
7340:
7335:
7330:
7328:Tithonian life
7325:
7315:
7314:
7308:
7307:
7305:
7304:
7291:
7278:
7262:
7260:
7254:
7253:
7248:
7239:
7238:
7235:
7234:
7232:
7231:
7226:
7221:
7215:
7212:
7211:
7208:
7201:
7200:
7197:
7196:
7172:
7170:
7167:
7166:
7163:
7162:
7159:
7158:
7156:
7155:
7148:
7140:
7133:
7125:
7118:
7111:
7104:
7101:Clasmodosaurus
7097:
7090:
7081:
7079:
7071:
7070:
7067:
7066:
7063:
7062:
7059:
7058:
7056:
7055:
7048:
7041:
7034:
7027:
7020:
7016:Maxakalisaurus
7012:
7005:
6997:
6988:
6986:
6980:
6979:
6977:
6976:
6968:
6961:
6953:
6946:
6938:
6930:
6923:
6914:
6912:
6906:
6905:
6903:
6902:
6898:Quaesitosaurus
6894:
6887:
6879:
6871:
6865:
6859:
6858:
6856:
6855:
6847:
6843:Quaesitosaurus
6839:
6831:
6829:
6820:
6819:
6812:
6805:
6797:
6795:
6793:Saltasauroidea
6789:
6788:
6785:
6784:
6782:
6781:
6774:
6767:
6760:
6753:
6745:
6738:
6730:
6723:
6715:
6713:
6707:
6706:
6703:
6702:
6700:
6699:
6692:
6685:
6678:
6671:
6664:
6657:
6649:
6647:
6641:
6640:
6638:
6637:
6630:
6623:
6616:
6609:
6605:Maxakalisaurus
6601:
6598:Adamantisaurus
6593:
6587:
6581:
6580:
6578:
6577:
6570:
6562:
6555:
6548:
6544:Epachthosaurus
6540:
6533:
6526:
6519:
6516:Antarctosaurus
6511:
6505:
6499:
6498:
6496:
6495:
6488:
6480:
6474:
6471:Eutitanosauria
6469:
6466:
6465:
6462:
6461:
6458:
6457:
6455:
6454:
6447:
6444:Mansourasaurus
6440:
6433:
6426:
6419:
6412:
6404:
6398:
6392:
6391:
6389:
6388:
6380:
6373:
6366:
6359:
6352:
6345:
6338:
6331:
6323:
6319:Tastavinsaurus
6315:
6308:
6301:
6294:
6287:
6284:Petrobrasaurus
6280:
6272:
6265:
6258:
6251:
6244:
6237:
6230:
6226:Malarguesaurus
6222:
6215:
6207:
6200:
6193:
6186:
6178:
6171:
6164:
6157:
6153:Epachthosaurus
6149:
6142:
6134:
6127:
6120:
6113:
6106:
6099:
6091:
6083:
6076:
6069:
6066:Austroposeidon
6062:
6055:
6048:
6041:
6033:
6026:
6018:
6010:
6000:
5995:
5988:
5987:
5982:
5975:
5974:
5971:
5970:
5945:
5943:
5940:
5939:
5936:
5935:
5932:
5931:
5928:
5927:
5925:
5924:
5923:
5922:
5916:
5914:
5908:
5907:
5904:
5903:
5901:
5900:
5892:
5885:
5878:
5871:
5863:
5857:
5851:
5850:
5848:
5847:
5840:
5833:
5830:Tangvayosaurus
5826:
5819:
5812:
5805:
5798:
5790:
5783:
5776:
5769:
5762:
5755:
5748:
5740:
5731:
5729:
5723:
5722:
5720:
5719:
5712:
5709:Tastavinsaurus
5705:
5698:
5691:
5684:
5681:Pukyongosaurus
5677:
5670:
5663:
5660:Liubangosaurus
5656:
5648:
5645:Liaoningotitan
5641:
5634:
5626:
5619:
5612:
5605:
5598:
5591:
5584:
5577:
5570:
5567:Astrophocaudia
5563:
5556:
5548:
5542:
5540:Somphospondyli
5536:
5535:
5533:
5532:
5525:
5518:
5511:
5504:
5497:
5490:
5482:
5475:
5468:
5461:
5453:
5451:
5445:
5444:
5442:
5441:
5433:
5426:
5419:
5412:
5405:
5398:
5391:
5383:
5377:
5372:
5369:
5368:
5365:
5364:
5362:
5361:
5358:Lourinhasaurus
5354:
5346:
5344:
5342:Camarasauridae
5338:
5337:
5335:
5334:
5327:
5319:
5312:
5308:Dashanpusaurus
5304:
5297:
5289:
5282:
5274:
5264:
5259:
5252:
5251:
5246:
5239:
5238:
5235:
5234:
5222:
5220:
5217:
5216:
5213:
5212:
5209:
5208:
5205:
5204:
5201:
5200:
5198:
5197:
5190:
5183:
5176:
5168:
5161:
5154:
5151:Dinheirosaurus
5147:
5139:
5137:
5131:
5130:
5128:
5127:
5120:
5112:
5105:
5096:
5094:
5085:
5079:
5078:
5076:
5075:
5068:
5061:
5054:
5047:
5040:
5033:
5025:
5018:
5011:
5004:
4997:
4989:
4987:
4978:
4972:
4971:
4968:
4967:
4964:
4963:
4961:
4960:
4953:
4950:Rebbachisaurus
4946:
4939:
4932:
4925:
4917:
4915:
4909:
4908:
4906:
4905:
4897:
4890:
4882:
4880:
4871:
4865:
4864:
4862:
4861:
4854:
4847:
4840:
4833:
4825:
4822:Lavocatisaurus
4818:
4811:
4803:
4796:
4789:
4782:
4774:
4768:
4759:
4754:
4751:
4750:
4748:
4747:
4739:
4733:
4728:
4721:
4720:
4715:
4708:
4707:
4704:
4703:
4675:
4673:
4670:
4669:
4666:
4665:
4662:
4661:
4659:
4658:
4651:
4648:Qinlingosaurus
4644:
4637:
4630:
4623:
4616:
4608:
4606:
4598:
4597:
4594:
4593:
4590:
4589:
4586:
4585:
4583:
4582:
4581:(see below ↓ )
4578:
4576:
4570:
4569:
4567:
4566:
4565:(see below ↓ )
4562:
4560:
4551:
4545:
4544:
4542:
4541:
4534:
4527:
4520:
4512:
4504:
4497:
4490:
4483:
4476:
4468:
4460:
4452:
4444:
4442:
4436:
4435:
4433:
4432:
4425:
4418:
4411:
4404:
4397:
4390:
4387:Rhomaleopakhus
4383:
4376:
4369:
4366:Mamenchisaurus
4362:
4354:
4347:
4340:
4333:
4326:
4318:
4315:Chuanjiesaurus
4311:
4307:Cetiosauriscus
4303:
4296:
4288:
4286:
4280:
4279:
4277:
4276:
4268:
4260:
4252:
4245:
4237:
4235:
4229:
4228:
4226:
4225:
4218:
4211:
4204:
4197:
4190:
4183:
4176:
4169:
4162:
4159:Dashanpusaurus
4155:
4152:Cetiosauriscus
4148:
4141:
4134:
4127:
4120:
4112:
4106:
4101:
4098:
4097:
4094:
4093:
4091:
4090:
4083:
4075:
4073:
4067:
4066:
4064:
4063:
4059:Zizhongosaurus
4055:
4047:
4040:
4033:
4026:
4017:
4007:
4002:
3999:
3998:
3995:
3994:
3992:
3991:
3984:
3977:
3970:
3962:
3960:
3958:Lessemsauridae
3954:
3953:
3951:
3950:
3946:Tuebingosaurus
3942:
3935:
3928:
3920:
3912:
3905:
3898:
3891:
3884:
3876:
3866:
3861:
3854:
3853:
3848:
3841:
3840:
3837:
3836:
3810:
3808:
3805:
3804:
3801:
3800:
3797:
3796:
3793:
3792:
3789:
3788:
3785:
3784:
3781:
3780:
3778:
3777:
3776:
3775:
3769:
3767:
3761:
3760:
3758:
3757:
3754:Sefapanosaurus
3750:
3743:
3736:
3729:
3726:Melanorosaurus
3722:
3715:
3708:
3701:
3694:
3687:
3679:
3673:
3667:
3666:
3664:
3663:
3656:
3649:
3641:
3634:
3627:
3620:
3611:
3605:
3599:
3598:
3596:
3595:
3587:
3580:
3573:
3566:
3563:Massospondylus
3559:
3552:
3545:
3537:
3534:Glacialisaurus
3530:
3527:Coloradisaurus
3523:
3515:
3513:
3507:
3506:
3504:
3503:
3496:
3488:
3486:
3480:
3479:
3477:
3476:
3473:Tuebingosaurus
3469:
3462:
3455:
3448:
3441:
3434:
3426:
3420:
3415:
3412:
3411:
3408:
3407:
3405:
3404:
3397:
3390:
3387:Pachysauriscus
3383:
3376:
3373:Gresslyosaurus
3369:
3361:
3359:
3357:Plateosauridae
3350:
3349:
3342:
3335:
3332:Plateosauravus
3327:
3321:
3316:
3313:
3312:
3309:
3308:
3306:
3305:
3298:
3291:
3283:
3281:
3275:
3274:
3272:
3271:
3264:
3257:
3250:
3243:
3236:
3229:
3221:
3211:
3206:
3203:
3202:
3199:
3198:
3196:
3195:
3188:
3180:
3172:
3165:
3162:Chromogisaurus
3158:
3150:
3141:
3139:
3133:
3132:
3130:
3129:
3122:
3115:
3108:
3101:
3094:
3087:
3079:
3069:
3064:
3057:
3056:
3051:
3044:
3043:
3040:
3039:
3036:
3035:
3032:
3031:
3029:
3028:
3027:
3026:
3020:
3018:
3012:
3011:
3009:
3008:
3007:
3006:
2997:
2991:
2981:
2980:
2976:
2975:
2966:
2957:
2951:
2945:
2936:
2929:
2928:
2923:
2921:
2920:
2913:
2906:
2898:
2892:
2891:
2888:Dinheirosaurus
2882:
2881:External links
2879:
2876:
2875:
2833:
2787:
2768:(3): 451–457.
2748:
2690:
2638:
2616:
2597:
2560:
2533:(3): 618–634.
2518:Zby atlanticus
2503:
2437:
2407:
2362:
2313:
2254:
2207:
2172:(4): 393–409.
2149:
2093:
2034:
2011:
1993:
1971:
1919:
1904:
1890:The Dinosauria
1871:
1844:(3): 521–551.
1804:
1744:
1689:
1688:
1686:
1683:
1671:Cetiosauriscus
1663:Dinheirosaurus
1649:Dinheirosaurus
1640:
1637:
1554:Zby atlanticus
1537:Dinheirosaurus
1483:North American
1479:ornithischians
1463:Dinheirosaurus
1459:
1456:
1448:Dinheirosaurus
1432:Dinheirosaurus
1406:Dinheirosaurus
1402:Dinheirosaurus
1398:
1395:
1391:Dinheirosaurus
1378:Dinheirosaurus
1373:
1370:
1353:Cetiosauriscus
1348:Dinheirosaurus
1336:Dinheirosaurus
1324:Dinheirosaurus
1316:Dinheirosaurus
1309:
1308:
1305:
1304:
1300:
1299:
1296:
1295:
1291:
1290:
1287:
1286:
1282:
1281:
1278:
1277:
1273:
1272:
1269:
1268:
1264:
1263:
1260:
1259:
1255:
1254:
1251:
1250:
1246:
1245:
1242:
1241:
1237:
1236:
1233:
1232:
1228:
1227:
1224:
1223:
1214:
1211:
1210:
1207:
1206:
1197:
1192:
1190:
1187:
1186:
1183:
1182:
1178:
1177:
1174:
1173:
1164:
1161:
1160:
1157:
1156:
1147:
1142:
1140:
1135:
1133:
1130:
1129:
1126:
1125:
1116:
1111:
1109:
1106:
1105:
1102:
1101:
1092:
1087:
1085:
1082:
1081:
1078:
1077:
1073:
1072:
1069:
1068:
1059:
1056:
1055:
1052:
1051:
1042:
1037:
1035:
1030:
1028:
1025:
1024:
1021:
1020:
1011:
1006:
1004:
1001:
1000:
997:
996:
991:
986:
984:
978:
977:
974:
973:
969:
968:
965:
964:
960:
959:
956:
955:
951:
950:
947:
946:
942:
941:
938:
937:
928:
925:
924:
921:
920:
911:
906:
904:
901:
900:
897:
896:
887:
882:
880:
877:
876:
873:
872:
868:
867:
864:
863:
854:
851:
850:
847:
846:
837:
832:
830:
825:
823:
820:
819:
816:
815:
810:
805:
803:
795:
793:
790:
789:
786:
785:
776:
771:
769:
761:
729:Dinheirosaurus
701:Philip Mannion
693:Dinheirosaurus
689:
688:Classification
686:
666:Dinheirosaurus
661:Dinheirosaurus
656:
653:
637:Dinheirosaurus
632:Dinheirosaurus
620:
617:
613:Dinheirosaurus
605:Dinheirosaurus
569:Dinheirosaurus
559:Porto Dinheiro
542:Lourinhasaurus
537:Octávio Mateus
533:José Bonaparte
505:
502:
494:Dinheirosaurus
460:Dinheirosaurus
442:Dinheirosaurus
438:Dinheirosaurus
426:Lourinhasaurus
421:Dinheirosaurus
378:Octávio Mateus
374:José Bonaparte
342:Dinheirosaurus
336:
335:
332:
318:
317:
311:
310:
302:
300:
296:
295:
278:Dinheirosaurus
271:
267:
266:
258:
254:
253:
245:
241:
240:
232:
225:
224:
216:
209:
208:
203:
196:
195:
190:
183:
182:
177:
173:
172:
167:
163:
162:
157:
153:
152:
139:
138:
134:
133:
125:
124:
116:
115:
114:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
53:
39:
36:Dinheirosaurus
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
7375:
7364:
7361:
7359:
7356:
7354:
7351:
7349:
7346:
7344:
7341:
7339:
7336:
7334:
7331:
7329:
7326:
7324:
7321:
7320:
7318:
7301:
7296:
7292:
7288:
7283:
7279:
7274:
7268:
7264:
7263:
7261:
7259:
7255:
7251:
7246:
7230:
7227:
7225:
7222:
7220:
7217:
7216:
7213:
7206:
7202:
7193:
7189:
7187:
7182:
7176:
7154:
7153:
7149:
7146:
7145:
7144:Mongolosaurus
7141:
7139:
7138:
7134:
7131:
7130:
7129:Macrurosaurus
7126:
7124:
7123:
7119:
7117:
7116:
7112:
7110:
7109:
7108:Hypselosaurus
7105:
7103:
7102:
7098:
7096:
7095:
7091:
7088:
7087:
7083:
7082:
7080:
7076:
7072:
7054:
7053:
7049:
7047:
7046:
7042:
7040:
7039:
7035:
7033:
7032:
7028:
7026:
7025:
7024:Neuquensaurus
7021:
7018:
7017:
7013:
7011:
7010:
7006:
7003:
7002:
6998:
6995:
6994:
6990:
6989:
6987:
6985:
6984:Saltasaurinae
6981:
6974:
6973:
6972:Zhuchengtitan
6969:
6967:
6966:
6962:
6959:
6958:
6954:
6952:
6951:
6947:
6944:
6943:
6942:Dreadnoughtus
6939:
6936:
6935:
6931:
6929:
6928:
6924:
6921:
6920:
6916:
6915:
6913:
6911:
6907:
6900:
6899:
6895:
6893:
6892:
6891:Qingxiusaurus
6888:
6885:
6884:
6880:
6878:
6877:
6873:
6872:
6869:
6866:
6864:
6863:Saltasauridae
6860:
6853:
6852:
6848:
6845:
6844:
6840:
6838:
6837:
6836:Nemegtosaurus
6833:
6832:
6830:
6828:
6824:
6818:
6817:
6813:
6811:
6810:
6806:
6804:
6803:
6799:
6798:
6796:
6794:
6790:
6780:
6779:
6775:
6773:
6772:
6768:
6766:
6765:
6761:
6759:
6758:
6757:Mendozasaurus
6754:
6751:
6750:
6746:
6744:
6743:
6739:
6736:
6735:
6734:Dzharatitanis
6731:
6729:
6728:
6724:
6722:
6721:
6717:
6716:
6714:
6712:
6708:
6698:
6697:
6693:
6691:
6690:
6686:
6684:
6683:
6682:Gondwanatitan
6679:
6677:
6676:
6672:
6670:
6669:
6665:
6663:
6662:
6658:
6656:
6655:
6651:
6650:
6648:
6646:
6642:
6636:
6635:
6631:
6629:
6628:
6624:
6622:
6621:
6617:
6615:
6614:
6613:Muyelensaurus
6610:
6607:
6606:
6602:
6600:
6599:
6595:
6594:
6591:
6588:
6586:
6582:
6576:
6575:
6571:
6568:
6567:
6563:
6561:
6560:
6556:
6554:
6553:
6549:
6546:
6545:
6541:
6539:
6538:
6537:Chucarosaurus
6534:
6532:
6531:
6527:
6525:
6524:
6520:
6518:
6517:
6513:
6512:
6509:
6506:
6504:
6503:Colossosauria
6500:
6494:
6493:
6492:Menucocelsior
6489:
6487:
6486:
6482:
6481:
6478:
6475:
6472:
6467:
6453:
6452:
6448:
6446:
6445:
6441:
6439:
6438:
6434:
6432:
6431:
6430:Lirainosaurus
6427:
6425:
6424:
6420:
6418:
6417:
6413:
6411:
6410:
6406:
6405:
6402:
6399:
6397:
6393:
6386:
6385:
6381:
6379:
6378:
6374:
6372:
6371:
6367:
6365:
6364:
6360:
6358:
6357:
6353:
6351:
6350:
6349:Trigonosaurus
6346:
6344:
6343:
6339:
6337:
6336:
6332:
6329:
6328:
6324:
6321:
6320:
6316:
6314:
6313:
6312:Ruyangosaurus
6309:
6307:
6306:
6302:
6300:
6299:
6295:
6293:
6292:
6291:Pitekunsaurus
6288:
6286:
6285:
6281:
6278:
6277:
6273:
6271:
6270:
6266:
6264:
6263:
6259:
6257:
6256:
6252:
6250:
6249:
6245:
6243:
6242:
6241:Mnyamawamtuka
6238:
6236:
6235:
6231:
6228:
6227:
6223:
6221:
6220:
6219:Magyarosaurus
6216:
6213:
6212:
6211:Ligabuesaurus
6208:
6206:
6205:
6204:Laplatasaurus
6201:
6199:
6198:
6197:Karongasaurus
6194:
6192:
6191:
6187:
6184:
6183:
6179:
6177:
6176:
6172:
6170:
6169:
6165:
6163:
6162:
6158:
6155:
6154:
6150:
6148:
6147:
6143:
6140:
6139:
6138:Dreadnoughtus
6135:
6133:
6132:
6128:
6126:
6125:
6121:
6119:
6118:
6114:
6112:
6111:
6107:
6105:
6104:
6103:Borealosaurus
6100:
6097:
6096:
6092:
6089:
6088:
6084:
6082:
6081:
6080:Barrosasaurus
6077:
6075:
6074:
6070:
6068:
6067:
6063:
6061:
6060:
6056:
6054:
6053:
6049:
6047:
6046:
6042:
6039:
6038:
6034:
6032:
6031:
6027:
6024:
6023:
6022:Aegyptosaurus
6019:
6017:
6016:
6012:
6011:
6008:
6004:
6001:
5998:
5993:
5989:
5985:
5980:
5976:
5966:
5961:
5958:
5950:
5920:
5919:
5918:
5917:
5915:
5913:
5909:
5898:
5897:
5893:
5891:
5890:
5889:Savannasaurus
5886:
5884:
5883:
5879:
5877:
5876:
5872:
5870:
5869:
5868:Australotitan
5865:
5864:
5861:
5858:
5856:
5852:
5846:
5845:
5841:
5839:
5838:
5834:
5832:
5831:
5827:
5825:
5824:
5820:
5818:
5817:
5813:
5811:
5810:
5806:
5804:
5803:
5799:
5796:
5795:
5791:
5789:
5788:
5784:
5782:
5781:
5777:
5775:
5774:
5770:
5768:
5767:
5763:
5761:
5760:
5756:
5754:
5753:
5749:
5746:
5745:
5741:
5738:
5737:
5736:Australodocus
5733:
5732:
5730:
5728:
5727:Euhelopodidae
5724:
5718:
5717:
5716:Triunfosaurus
5713:
5711:
5710:
5706:
5704:
5703:
5699:
5697:
5696:
5695:Sauroposeidon
5692:
5690:
5689:
5685:
5683:
5682:
5678:
5676:
5675:
5674:Padillasaurus
5671:
5669:
5668:
5664:
5662:
5661:
5657:
5654:
5653:
5652:Ligabuesaurus
5649:
5647:
5646:
5642:
5640:
5639:
5635:
5632:
5631:
5627:
5625:
5624:
5620:
5618:
5617:
5613:
5611:
5610:
5606:
5604:
5603:
5599:
5597:
5596:
5595:Chubutisaurus
5592:
5590:
5589:
5585:
5583:
5582:
5578:
5576:
5575:
5571:
5569:
5568:
5564:
5562:
5561:
5557:
5555:
5554:
5550:
5549:
5546:
5543:
5541:
5537:
5531:
5530:
5526:
5524:
5523:
5519:
5517:
5516:
5512:
5510:
5509:
5505:
5503:
5502:
5498:
5496:
5495:
5491:
5488:
5487:
5483:
5481:
5480:
5476:
5474:
5473:
5472:Brachiosaurus
5469:
5467:
5466:
5462:
5460:
5459:
5455:
5454:
5452:
5450:
5446:
5439:
5438:
5434:
5432:
5431:
5427:
5425:
5424:
5420:
5418:
5417:
5413:
5411:
5410:
5409:Fushanosaurus
5406:
5404:
5403:
5399:
5397:
5396:
5392:
5390:
5389:
5385:
5384:
5381:
5378:
5375:
5370:
5360:
5359:
5355:
5353:
5352:
5348:
5347:
5345:
5343:
5339:
5333:
5332:
5328:
5325:
5324:
5320:
5318:
5317:
5313:
5310:
5309:
5305:
5303:
5302:
5298:
5295:
5294:
5293:Bashunosaurus
5290:
5288:
5287:
5283:
5281:
5280:
5276:
5275:
5272:
5268:
5265:
5262:
5257:
5253:
5249:
5244:
5240:
5231:
5226:
5196:
5195:
5191:
5189:
5188:
5184:
5182:
5181:
5177:
5174:
5173:
5169:
5167:
5166:
5162:
5160:
5159:
5155:
5153:
5152:
5148:
5146:
5145:
5141:
5140:
5138:
5136:
5132:
5126:
5125:
5121:
5118:
5117:
5116:Atlantosaurus
5113:
5111:
5110:
5106:
5103:
5102:
5098:
5097:
5095:
5093:
5092:Apatosaurinae
5089:
5086:
5084:
5080:
5074:
5073:
5069:
5067:
5066:
5062:
5060:
5059:
5058:Smitanosaurus
5055:
5053:
5052:
5048:
5046:
5045:
5041:
5039:
5038:
5034:
5031:
5030:
5029:Dyslocosaurus
5026:
5024:
5023:
5022:Dicraeosaurus
5019:
5017:
5016:
5012:
5010:
5009:
5005:
5003:
5002:
5001:Amargatitanis
4998:
4996:
4995:
4991:
4990:
4988:
4986:
4982:
4979:
4977:
4973:
4959:
4958:
4954:
4952:
4951:
4947:
4945:
4944:
4940:
4938:
4937:
4936:Katepensaurus
4933:
4931:
4930:
4929:Itapeuasaurus
4926:
4924:
4923:
4922:Demandasaurus
4919:
4918:
4916:
4914:
4910:
4903:
4902:
4898:
4896:
4895:
4891:
4889:
4888:
4887:Cathartesaura
4884:
4883:
4881:
4879:
4878:Limaysaurinae
4875:
4872:
4870:
4866:
4860:
4859:
4855:
4853:
4852:
4848:
4846:
4845:
4841:
4839:
4838:
4834:
4831:
4830:
4826:
4824:
4823:
4819:
4817:
4816:
4815:Histriasaurus
4812:
4809:
4808:
4807:Dzharatitanis
4804:
4802:
4801:
4800:Comahuesaurus
4797:
4795:
4794:
4790:
4788:
4787:
4783:
4781:
4780:
4776:
4775:
4772:
4769:
4767:
4763:
4760:
4757:
4752:
4746:
4745:
4741:
4740:
4737:
4734:
4731:
4730:Diplodocoidea
4726:
4722:
4718:
4717:Diplodocoidea
4713:
4709:
4700:
4696:
4694:
4688:
4680:
4657:
4656:
4652:
4650:
4649:
4645:
4643:
4642:
4638:
4636:
4635:
4634:Gigantosaurus
4631:
4629:
4628:
4624:
4622:
4621:
4617:
4615:
4614:
4610:
4609:
4607:
4603:
4599:
4580:
4579:
4577:
4575:
4571:
4564:
4563:
4561:
4559:
4558:Diplodocoidea
4555:
4552:
4550:
4546:
4540:
4539:
4535:
4533:
4532:
4528:
4526:
4525:
4521:
4518:
4517:
4513:
4510:
4509:
4505:
4503:
4502:
4501:Narindasaurus
4498:
4496:
4495:
4491:
4489:
4488:
4484:
4482:
4481:
4480:Losillasaurus
4477:
4474:
4473:
4469:
4466:
4465:
4461:
4458:
4457:
4453:
4451:
4450:
4446:
4445:
4443:
4441:
4437:
4431:
4430:
4429:Zigongosaurus
4426:
4424:
4423:
4422:Yuanmousaurus
4419:
4417:
4416:
4415:Xinjiangtitan
4412:
4410:
4409:
4408:Wamweracaudia
4405:
4403:
4402:
4401:Tonganosaurus
4398:
4396:
4395:
4391:
4389:
4388:
4384:
4382:
4381:
4377:
4375:
4374:
4370:
4368:
4367:
4363:
4360:
4359:
4358:Klamelisaurus
4355:
4353:
4352:
4348:
4346:
4345:
4341:
4339:
4338:
4337:Huangshanlong
4334:
4332:
4331:
4327:
4324:
4323:
4319:
4317:
4316:
4312:
4309:
4308:
4304:
4302:
4301:
4297:
4295:
4294:
4290:
4289:
4287:
4285:
4281:
4274:
4273:
4269:
4266:
4265:
4261:
4258:
4257:
4256:Ferganasaurus
4253:
4251:
4250:
4246:
4244:
4243:
4239:
4238:
4236:
4234:
4233:Cetiosauridae
4230:
4224:
4223:
4219:
4217:
4216:
4212:
4210:
4209:
4205:
4203:
4202:
4198:
4196:
4195:
4191:
4189:
4188:
4184:
4182:
4181:
4177:
4175:
4174:
4170:
4168:
4167:
4163:
4161:
4160:
4156:
4154:
4153:
4149:
4147:
4146:
4142:
4140:
4139:
4135:
4133:
4132:
4128:
4126:
4125:
4121:
4119:
4118:
4114:
4113:
4110:
4107:
4104:
4099:
4089:
4088:
4084:
4082:
4081:
4080:Tazoudasaurus
4077:
4076:
4074:
4072:
4068:
4061:
4060:
4056:
4053:
4052:
4048:
4046:
4045:
4041:
4039:
4038:
4037:Ohmdenosaurus
4034:
4032:
4031:
4027:
4024:
4023:
4019:
4018:
4015:
4011:
4008:
4005:
4000:
3990:
3989:
3985:
3983:
3982:
3978:
3976:
3975:
3971:
3969:
3968:
3964:
3963:
3961:
3959:
3955:
3948:
3947:
3943:
3941:
3940:
3939:Schleitheimia
3936:
3934:
3933:
3929:
3926:
3925:
3921:
3918:
3917:
3916:Leonerasaurus
3913:
3911:
3910:
3906:
3904:
3903:
3899:
3897:
3896:
3892:
3890:
3889:
3888:Blikanasaurus
3885:
3883:
3882:
3878:
3877:
3874:
3870:
3867:
3864:
3859:
3855:
3851:
3846:
3842:
3832:
3827:
3825:
3820:
3814:
3773:
3772:
3771:
3770:
3768:
3766:
3762:
3756:
3755:
3751:
3749:
3748:
3744:
3742:
3741:
3737:
3735:
3734:
3730:
3728:
3727:
3723:
3721:
3720:
3719:Leonerasaurus
3716:
3714:
3713:
3712:Lamplughsaura
3709:
3707:
3706:
3702:
3700:
3699:
3695:
3693:
3692:
3688:
3686:
3685:
3681:
3680:
3677:
3674:
3672:
3668:
3662:
3661:
3660:Yunnanosaurus
3657:
3655:
3654:
3650:
3647:
3646:
3642:
3640:
3639:
3635:
3633:
3632:
3628:
3626:
3625:
3621:
3618:
3617:
3613:
3612:
3609:
3606:
3604:
3600:
3593:
3592:
3588:
3586:
3585:
3581:
3579:
3578:
3574:
3572:
3571:
3567:
3565:
3564:
3560:
3558:
3557:
3556:Lufengosaurus
3553:
3551:
3550:
3546:
3543:
3542:
3538:
3536:
3535:
3531:
3529:
3528:
3524:
3522:
3521:
3517:
3516:
3514:
3512:
3508:
3502:
3501:
3497:
3495:
3494:
3493:Eucnemesaurus
3490:
3489:
3487:
3485:
3484:Riojasauridae
3481:
3475:
3474:
3470:
3468:
3467:
3463:
3461:
3460:
3456:
3454:
3453:
3449:
3447:
3446:
3442:
3440:
3439:
3435:
3433:
3432:
3428:
3427:
3424:
3421:
3418:
3413:
3403:
3402:
3398:
3396:
3395:
3391:
3389:
3388:
3384:
3382:
3381:
3377:
3375:
3374:
3370:
3368:
3367:
3366:Euskelosaurus
3363:
3362:
3360:
3358:
3354:
3348:
3347:
3343:
3341:
3340:
3336:
3334:
3333:
3329:
3328:
3325:
3322:
3319:
3314:
3304:
3303:
3299:
3297:
3296:
3292:
3290:
3289:
3285:
3284:
3282:
3280:
3276:
3270:
3269:
3265:
3263:
3262:
3258:
3256:
3255:
3251:
3249:
3248:
3244:
3242:
3241:
3240:Bagualosaurus
3237:
3235:
3234:
3230:
3228:
3227:
3223:
3222:
3219:
3215:
3212:
3209:
3208:Bagualosauria
3204:
3194:
3193:
3189:
3186:
3185:
3181:
3178:
3177:
3176:Pampadromaeus
3173:
3171:
3170:
3166:
3164:
3163:
3159:
3156:
3155:
3151:
3148:
3147:
3143:
3142:
3140:
3138:
3137:Saturnaliidae
3134:
3128:
3127:
3123:
3121:
3120:
3119:Pampadromaeus
3116:
3114:
3113:
3109:
3107:
3106:
3102:
3100:
3099:
3098:Gigantoscelus
3095:
3093:
3092:
3088:
3086:
3085:
3081:
3080:
3077:
3073:
3070:
3067:
3062:
3058:
3054:
3049:
3045:
3024:
3023:
3022:
3021:
3019:
3017:
3013:
3005:
3001:
3000:
2999:
2998:
2995:
2992:
2990:
2986:
2982:
2974:
2970:
2967:
2965:
2961:
2958:
2956:
2952:
2950:
2946:
2944:
2940:
2939:
2934:
2930:
2926:
2919:
2914:
2912:
2907:
2905:
2900:
2899:
2896:
2890:
2889:
2885:
2884:
2880:
2871:
2867:
2863:
2859:
2855:
2851:
2844:
2837:
2834:
2829:
2825:
2821:
2817:
2813:
2809:
2802:
2800:
2791:
2788:
2783:
2779:
2775:
2771:
2767:
2763:
2759:
2752:
2749:
2744:
2740:
2736:
2732:
2728:
2724:
2720:
2716:
2713:(5): 367–74.
2712:
2708:
2701:
2694:
2691:
2686:
2682:
2677:
2672:
2668:
2664:
2660:
2656:
2649:
2642:
2639:
2634:
2630:
2623:
2621:
2617:
2612:
2608:
2601:
2598:
2592:
2587:
2583:
2579:
2575:
2573:
2564:
2561:
2556:
2552:
2548:
2544:
2540:
2536:
2532:
2528:
2521:
2519:
2510:
2508:
2504:
2499:
2495:
2490:
2485:
2480:
2475:
2471:
2467:
2464:(3): e88905.
2463:
2459:
2455:
2453:
2444:
2442:
2438:
2433:
2429:
2422:
2420:
2411:
2408:
2403:
2399:
2395:
2391:
2387:
2383:
2376:
2369:
2367:
2363:
2358:
2354:
2349:
2344:
2340:
2336:
2332:
2328:
2324:
2317:
2314:
2309:
2305:
2300:
2295:
2290:
2285:
2281:
2277:
2274:(4): e18304.
2273:
2269:
2265:
2258:
2255:
2250:
2246:
2242:
2238:
2234:
2230:
2226:
2222:
2218:
2211:
2208:
2203:
2199:
2195:
2191:
2187:
2183:
2179:
2175:
2171:
2167:
2160:
2153:
2150:
2145:
2139:
2131:
2127:
2123:
2119:
2115:
2111:
2104:
2097:
2094:
2089:
2085:
2080:
2075:
2070:
2065:
2061:
2057:
2054:(5): e97128.
2053:
2049:
2045:
2038:
2035:
2030:
2026:
2022:
2015:
2012:
2004:
1997:
1994:
1989:
1982:
1975:
1972:
1956:
1952:
1948:
1944:
1940:
1933:
1926:
1924:
1920:
1915:
1911:
1907:
1901:
1897:
1892:
1891:
1882:
1880:
1878:
1876:
1872:
1867:
1863:
1859:
1855:
1851:
1847:
1843:
1839:
1832:
1830:
1821:
1819:
1817:
1815:
1813:
1811:
1809:
1805:
1797:
1793:
1789:
1785:
1781:
1774:
1772:
1763:
1761:
1759:
1757:
1755:
1753:
1751:
1749:
1745:
1740:
1736:
1731:
1726:
1721:
1716:
1712:
1708:
1704:
1697:
1695:
1691:
1684:
1682:
1680:
1676:
1672:
1668:
1667:Losillasaurus
1664:
1660:
1659:
1654:
1650:
1646:
1638:
1636:
1634:
1633:
1628:
1627:
1622:
1621:
1616:
1615:
1611:
1607:
1606:
1602:
1598:
1597:
1592:
1591:
1586:
1585:
1581:
1580:stegosaurians
1577:
1576:
1571:
1570:
1565:
1564:
1560:
1559:brachiosaurid
1556:
1555:
1551:
1547:
1546:
1542:
1538:
1534:
1530:
1529:
1525:
1521:
1520:
1515:
1514:
1510:
1506:
1505:
1501:
1497:
1496:
1492:
1488:
1484:
1480:
1476:
1472:
1468:
1464:
1457:
1455:
1453:
1449:
1445:
1441:
1437:
1433:
1429:
1425:
1421:
1420:
1415:
1411:
1407:
1403:
1396:
1394:
1392:
1388:
1384:
1379:
1371:
1369:
1367:
1366:
1365:Losillasaurus
1361:
1360:
1355:
1354:
1349:
1345:
1341:
1337:
1333:
1332:plesiomorphic
1329:
1325:
1321:
1317:
1307:
1306:
1298:
1297:
1289:
1288:
1280:
1279:
1271:
1270:
1262:
1261:
1253:
1252:
1244:
1243:
1235:
1234:
1226:
1225:
1222:
1221:
1220:
1213:
1212:
1209:
1208:
1205:
1204:
1203:
1196:
1195:
1189:
1188:
1185:
1184:
1176:
1175:
1172:
1171:
1170:
1163:
1162:
1159:
1158:
1155:
1154:
1153:
1146:
1145:
1139:
1138:
1132:
1131:
1128:
1127:
1124:
1123:
1122:
1115:
1114:
1108:
1107:
1104:
1103:
1100:
1099:
1098:
1091:
1090:
1084:
1083:
1080:
1079:
1071:
1070:
1067:
1066:
1065:
1058:
1057:
1054:
1053:
1050:
1049:
1048:
1041:
1040:
1034:
1033:
1027:
1026:
1023:
1022:
1019:
1018:
1017:
1010:
1009:
1003:
1002:
999:
998:
995:
990:
989:
983:
980:
979:
976:
975:
967:
966:
958:
957:
949:
948:
940:
939:
936:
935:
934:
927:
926:
923:
922:
919:
918:
917:
910:
909:
903:
902:
899:
898:
895:
894:
893:
886:
885:
879:
878:
875:
874:
866:
865:
862:
861:
860:
853:
852:
849:
848:
845:
844:
843:
836:
835:
829:
828:
822:
821:
818:
817:
814:
809:
808:
802:
801:Apatosaurinae
799:
798:
792:
791:
788:
787:
784:
783:
782:
775:
774:
768:
765:
764:
760:
758:
754:
750:
746:
742:
738:
734:
730:
726:
722:
719:
715:
714:
709:
705:
702:
698:
694:
687:
685:
683:
677:
675:
671:
667:
662:
654:
652:
650:
646:
640:
638:
633:
625:
618:
616:
614:
610:
606:
602:
599:
594:
590:
586:
582:
578:
574:
570:
566:
564:
560:
556:
552:
548:
544:
543:
538:
534:
530:
529:
524:
520:
515:
511:
503:
501:
499:
495:
491:
490:
485:
484:
479:
478:
473:
472:
467:
466:
461:
457:
455:
451:
450:thyreophorans
448:, sauropods,
447:
443:
439:
435:
432:
428:
427:
422:
418:
414:
410:
406:
402:
397:
395:
394:Late Jurassic
391:
387:
383:
379:
375:
371:
370:
365:
364:
359:
355:
352:
348:
344:
343:
330:
328:
319:
316:
315:Binomial name
312:
308:
307:
301:
298:
297:
293:
289:
285:
280:
279:
272:
269:
268:
265:
259:
256:
255:
252:
251:Diplodocoidea
246:
244:Superfamily:
243:
242:
239:
233:
230:
227:
226:
223:
217:
214:
211:
210:
207:
204:
201:
198:
197:
194:
191:
188:
185:
184:
181:
178:
175:
174:
171:
168:
165:
164:
161:
158:
155:
154:
149:
144:
140:
135:
131:
126:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
51:
44:
43:Late Jurassic
37:
33:
30:
19:
7257:
7178:
7152:Titanosaurus
7150:
7142:
7135:
7127:
7122:Loricosaurus
7120:
7115:Iuticosaurus
7113:
7106:
7099:
7092:
7084:
7052:Yamanasaurus
7050:
7043:
7036:
7029:
7022:
7014:
7007:
6999:
6993:Abditosaurus
6991:
6970:
6963:
6955:
6948:
6940:
6932:
6925:
6919:Abditosaurus
6917:
6896:
6889:
6881:
6874:
6851:Tapuiasaurus
6849:
6841:
6834:
6814:
6809:Titanomachya
6807:
6800:
6778:Puertasaurus
6776:
6769:
6764:Notocolossus
6762:
6755:
6749:Jiangxititan
6747:
6740:
6732:
6727:Drusilasaura
6725:
6718:
6711:Lognkosauria
6694:
6687:
6680:
6673:
6666:
6659:
6652:
6645:Aeolosaurini
6634:Rinconsaurus
6632:
6625:
6618:
6611:
6603:
6596:
6585:Rinconsauria
6572:
6566:Tengrisaurus
6564:
6559:Quetecsaurus
6557:
6550:
6542:
6535:
6528:
6521:
6514:
6490:
6483:
6449:
6442:
6437:Lohuecotitan
6435:
6428:
6423:Garrigatitan
6421:
6414:
6409:Ampelosaurus
6407:
6382:
6375:
6368:
6363:Uberabatitan
6361:
6356:Sonidosaurus
6354:
6347:
6340:
6333:
6327:Tengrisaurus
6325:
6317:
6310:
6303:
6298:Rapetosaurus
6296:
6289:
6282:
6274:
6267:
6260:
6253:
6246:
6239:
6234:Malawisaurus
6232:
6224:
6217:
6209:
6202:
6195:
6188:
6180:
6175:Huabeisaurus
6173:
6166:
6159:
6151:
6144:
6136:
6129:
6122:
6117:Choconsaurus
6115:
6110:Brasilotitan
6108:
6101:
6093:
6085:
6078:
6071:
6064:
6059:Atacamatitan
6057:
6052:Argyrosaurus
6050:
6043:
6035:
6028:
6020:
6015:Abdarainurus
6013:
5997:Titanosauria
5984:Titanosauria
5953:
5912:Titanosauria
5896:Wintonotitan
5894:
5887:
5880:
5873:
5866:
5842:
5837:Yongjinglong
5835:
5828:
5823:Tambatitanis
5821:
5814:
5807:
5800:
5792:
5787:Huanghetitan
5785:
5780:Huabeisaurus
5778:
5771:
5766:Gannansaurus
5764:
5757:
5750:
5744:Chiayusaurus
5742:
5734:
5714:
5707:
5700:
5693:
5686:
5679:
5672:
5665:
5658:
5650:
5643:
5638:Jiutaisaurus
5636:
5630:Jiangxititan
5628:
5623:Garumbatitan
5621:
5614:
5607:
5602:Dongbeititan
5600:
5593:
5586:
5579:
5574:Austrosaurus
5572:
5565:
5558:
5551:
5527:
5522:Venenosaurus
5520:
5513:
5508:Sonorasaurus
5506:
5499:
5494:Giraffatitan
5492:
5484:
5479:Cedarosaurus
5477:
5470:
5463:
5456:
5435:
5430:Pelorosaurus
5428:
5421:
5414:
5407:
5400:
5393:
5386:
5356:
5351:Camarasaurus
5349:
5329:
5323:Haestasaurus
5321:
5316:Europasaurus
5314:
5306:
5299:
5291:
5284:
5277:
5192:
5185:
5178:
5170:
5163:
5156:
5150:
5149:
5142:
5135:Diplodocinae
5124:Brontosaurus
5122:
5114:
5107:
5101:Amphicoelias
5099:
5083:Diplodocidae
5070:
5063:
5056:
5049:
5042:
5035:
5027:
5020:
5013:
5008:Bajadasaurus
5006:
4999:
4994:Amargasaurus
4992:
4955:
4948:
4941:
4934:
4927:
4920:
4901:Rayososaurus
4899:
4892:
4885:
4858:Zapalasaurus
4856:
4851:Xenoposeidon
4849:
4842:
4835:
4827:
4820:
4813:
4805:
4798:
4793:Campananeyen
4791:
4786:Amazonsaurus
4784:
4777:
4742:
4683:
4653:
4646:
4639:
4632:
4625:
4618:
4611:
4549:Neosauropoda
4536:
4529:
4522:
4514:
4506:
4499:
4492:
4485:
4478:
4470:
4464:Haestasaurus
4462:
4454:
4447:
4427:
4420:
4413:
4406:
4399:
4392:
4385:
4378:
4371:
4364:
4356:
4349:
4342:
4335:
4328:
4320:
4313:
4305:
4298:
4291:
4272:Patagosaurus
4270:
4262:
4254:
4247:
4240:
4222:Volkheimeria
4220:
4213:
4206:
4199:
4194:Perijasaurus
4192:
4187:Nebulasaurus
4185:
4178:
4171:
4166:Dystrophaeus
4164:
4157:
4150:
4143:
4138:Barapasaurus
4136:
4129:
4124:Asiatosaurus
4122:
4115:
4085:
4078:
4057:
4049:
4044:Rhoetosaurus
4042:
4035:
4028:
4020:
3988:Lessemsaurus
3986:
3979:
3972:
3967:Antetonitrus
3965:
3944:
3937:
3930:
3922:
3914:
3907:
3900:
3893:
3886:
3879:
3816:
3752:
3745:
3738:
3731:
3724:
3717:
3710:
3703:
3696:
3689:
3682:
3658:
3653:Yizhousaurus
3651:
3643:
3636:
3629:
3622:
3614:
3589:
3582:
3575:
3568:
3561:
3554:
3547:
3541:Ignavusaurus
3539:
3532:
3525:
3518:
3498:
3491:
3471:
3464:
3457:
3450:
3445:Ignavusaurus
3443:
3436:
3429:
3401:Yimenosaurus
3399:
3394:Plateosaurus
3392:
3385:
3378:
3371:
3364:
3346:Xixiposaurus
3344:
3337:
3330:
3318:Plateosauria
3300:
3293:
3286:
3279:Unaysauridae
3266:
3259:
3252:
3245:
3238:
3231:
3224:
3190:
3182:
3174:
3167:
3160:
3152:
3144:
3124:
3117:
3110:
3105:Guaibasaurus
3103:
3096:
3089:
3082:
2968:
2959:
2887:
2856:(1): 77–95.
2853:
2849:
2836:
2814:(1): 61–73.
2811:
2807:
2798:
2790:
2765:
2761:
2757:
2751:
2710:
2706:
2693:
2658:
2654:
2641:
2632:
2628:
2610:
2606:
2600:
2581:
2577:
2571:
2563:
2530:
2526:
2517:
2461:
2457:
2451:
2431:
2427:
2418:
2410:
2385:
2381:
2330:
2326:
2316:
2271:
2267:
2257:
2224:
2220:
2210:
2169:
2166:Paleobiology
2165:
2152:
2138:cite journal
2122:10.26879/662
2113:
2109:
2096:
2051:
2047:
2037:
2020:
2014:
1996:
1987:
1974:
1962:. Retrieved
1955:the original
1942:
1938:
1889:
1841:
1837:
1828:
1796:the original
1786:(2): 13–29.
1783:
1779:
1770:
1710:
1706:
1670:
1666:
1662:
1656:
1648:
1647:, including
1642:
1639:Biogeography
1630:
1624:
1620:Camptosaurus
1618:
1612:
1603:
1601:ankylosaurid
1594:
1588:
1582:
1573:
1567:
1561:
1552:
1543:
1541:camarasaurid
1536:
1526:
1524:megalosaurid
1517:
1511:
1502:
1500:ceratosaurid
1493:
1462:
1461:
1458:Paleoecology
1452:Cedarosaurus
1451:
1447:
1443:
1440:Cedarosaurus
1439:
1435:
1431:
1428:gastric mill
1419:Cedarosaurus
1417:
1414:Seismosaurus
1413:
1409:
1405:
1401:
1400:
1390:
1386:
1382:
1377:
1375:
1372:Paleobiology
1363:
1359:Amphicoelias
1357:
1351:
1347:
1343:
1339:
1335:
1327:
1323:
1319:
1315:
1314:Previously,
1313:
1217:
1216:
1200:
1199:
1167:
1166:
1150:
1149:
1119:
1118:
1095:
1094:
1062:
1061:
1046:
1045:
1044:
1014:
1013:
993:
982:Diplodocinae
931:
930:
914:
913:
890:
889:
857:
856:
840:
839:
812:
779:
778:
767:Diplodocidae
756:
752:
748:
745:Diplodocinae
740:
736:
732:
728:
724:
721:diplodocines
711:
703:
697:phylogenetic
692:
691:
681:
678:
673:
670:neural spine
665:
660:
658:
648:
641:
636:
631:
630:
612:
604:
600:
568:
567:
561:lizard from
554:
540:
526:
522:
518:
513:
507:
497:
493:
487:
481:
475:
469:
463:
459:
458:
441:
437:
433:
424:
420:
398:
380:in 1999 for
368:
367:
361:
341:
340:
339:
326:
321:
305:
304:
277:
276:
264:Diplodocidae
228:
212:
199:
186:
35:
29:
7323:Diplodocids
7137:Microcoelus
7045:Saltasaurus
7031:Paralititan
6927:Alamosaurus
6816:Udelartitan
6771:Patagotitan
6668:Bravasaurus
6661:Arrudatitan
6654:Aeolosaurus
6620:Overosaurus
6552:Jainosaurus
6530:Bonitasaura
6451:Paludititan
6342:Traukutitan
6037:Angolatitan
5844:Yunmenglong
5809:Qiaowanlong
5702:Sibirotitan
5667:Oceanotitan
5609:Europatitan
5588:Brontomerus
5581:Brohisaurus
5553:Angolatitan
5486:Galvesaurus
5458:Abydosaurus
5423:Ornithopsis
5416:Fusuisaurus
5402:Eucamerotus
5301:Daanosaurus
5286:Aragosaurus
5187:Supersaurus
5109:Apatosaurus
5072:Tharosaurus
4943:Nigersaurus
4894:Limaysaurus
4655:Ultrasaurus
4531:Turiasaurus
4494:Moabosaurus
4487:Mierasaurus
4456:Bellusaurus
4440:Turiasauria
4380:Qijianglong
4344:Hudiesaurus
4322:Datousaurus
4242:Cetiosaurus
4201:Shunosaurus
4145:Bellusaurus
4117:Algoasaurus
4103:Eusauropoda
4051:Sanpasaurus
4004:Gravisauria
3924:Pulanesaura
3909:Isanosaurus
3881:Amygdalodon
3747:Pulanesaura
3705:Irisosaurus
3691:Anchisaurus
3671:Anchisauria
3645:Xingxiulong
3591:Xingxiulong
3584:Sarahsaurus
3500:Riojasaurus
3466:Sarahsaurus
3452:Kholumolumo
3295:Macrocollum
3233:Asylosaurus
3169:Nhandumirim
3146:Buriolestes
3112:Mbiresaurus
3084:Buriolestes
2613:(5): 63–65.
2388:(1): 1–74.
1945:: 183–186.
1645:eusauropods
1610:ornithopods
1535:as well as
1487:abelisaurid
1383:Apatosaurus
1346:found that
1320:Apatosaurus
753:Supersaurus
747:. In 2015,
741:Apatosaurus
737:Apatosaurus
645:gastroliths
619:Description
585:gastroliths
528:Apatosaurus
498:Supersaurus
483:Supersaurus
465:Apatosaurus
454:ornithopods
417:gastroliths
415:, and many
363:Supersaurus
7317:Categories
7038:Rocasaurus
6965:Qunkasaura
6934:Baurutitan
6696:Shingopana
6523:Baalsaurus
6370:Volgatitan
6305:Rukwatitan
6269:Nullotitan
6255:Ninjatitan
6190:Kaijutitan
6161:Gandititan
6124:Daxiatitan
6087:Baurutitan
6030:Andesaurus
5921:see below↓
5616:Fukuititan
5560:Arkharavia
5515:Soriatitan
5465:Atlasaurus
5437:Rugocaudia
5395:Duriatitan
5331:Yuzhoulong
5279:Abrosaurus
5261:Macronaria
5248:Macronaria
5172:Kaatedocus
5165:Galeamopus
5158:Diplodocus
5144:Barosaurus
5051:Pilmatueia
5044:Lingwulong
5037:Kaatedocus
4957:Tataouinea
4869:Khebbashia
4844:Sidersaura
4574:Macronaria
4524:Tendaguria
4508:Oplosaurus
4472:Janenschia
4373:Omeisaurus
4249:Chebsaurus
4173:Janenschia
4087:Vulcanodon
4030:Kotasaurus
3981:Ledumahadi
3774:see below↓
3733:Meroktenos
3549:Leyesaurus
3438:Gyposaurus
3302:Unaysaurus
3261:Pantydraco
3226:Arcusaurus
3192:Saturnalia
3025:see below↓
2973:Dinosauria
2955:Sauropsida
2584:: 98–206.
2434:: 123–129.
2419:Allosaurus
1685:References
1679:Berriasian
1632:Dryosaurus
1629:, and cf.
1557:; and the
1533:diplodocid
1516:, and cf.
1491:allosaurid
1444:Diplodocus
1436:Diplodocus
1410:Diplodocus
1387:Diplodocus
1340:Diplodocus
1328:Diplodocus
708:cladograms
682:Diplodocus
674:Diplodocus
579:, partial
477:Barosaurus
471:Diplodocus
351:diplodocid
206:Saurischia
193:Dinosauria
7001:Bonatitan
6883:Isisaurus
6689:Punatitan
6168:Hamititan
6146:Elaltitan
6095:Bonatitan
5816:Silutitan
5773:Gobititan
5759:Euhelopus
5501:Lusotitan
5194:Tornieria
5180:Leinkupal
5065:Suuwassea
4779:Agustinia
4627:Dinodocus
4620:Cardiodon
4351:Jingiella
4300:Anhuilong
3863:Sauropoda
3850:Sauropoda
3765:Sauropoda
3740:Mussaurus
3698:Camelotia
3577:Pradhania
3417:Massopoda
3184:Panphagia
3126:Panphagia
2941:Kingdom:
2607:Geo-Temas
2249:219762797
2241:0891-2963
1964:24 August
1951:0213-683X
1939:Geogaceta
1914:801843269
1858:1478-0941
1792:0524-9511
1653:Tithonian
1563:Lusotitan
1550:turiasaur
1475:sauropods
1471:theropods
1397:Digestion
655:Vertebrae
651:in 2012.
573:vertebrae
489:Tornieria
446:theropods
390:Tithonian
382:vertebrae
299:Species:
284:Bonaparte
238:Sauropoda
166:Kingdom:
160:Eukaryota
50:Tithonian
7267:Wikidata
7009:Ibirania
6675:Caieiria
6485:Inawentu
5688:Ruixinia
5529:Vouivria
5388:Astrodon
4641:Neosodon
4131:Bagualia
3974:Ingentia
3684:Aardonyx
3631:Qianlong
3459:Musankwa
3431:Gryponyx
3339:Ruehleia
3254:Nambalia
3247:Efraasia
3154:Eoraptor
3091:Eoraptor
2949:Chordata
2947:Phylum:
2943:Animalia
2743:10930309
2735:17187254
2685:19324778
2555:59387149
2498:24598585
2458:PLOS ONE
2402:24869965
2357:17254987
2308:21494685
2268:PLOS ONE
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