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long with expanded tips. The stems which did not bear sporangia had coiled (circinnate) tips. Fertile stems bore disc-shaped sporangia laterally on stalks up to 5 mm long, forming open spikes. The sporangia, which were about 3.7 millimetres (0.15 in) in diameter, had spines like the stems, and split (dehisced) along their margin to release the trilete
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consisted of creeping stems (axes) up to 5 millimetres (0.20 in) in diameter with many K- or H-shaped branches as well as upturned or trailing stems, slightly smaller in diameter, which also branched. All stems had irregularly arranged multicellular spines up to 2.5 millimetres (0.098 in)
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The early Devonian Posongchong flora of Yunnan: a contribution to an understanding of the evolution and early diversification of vascular plants
977: 842: 756: 718: 676: 256:(spore-forming organs). The genus was first described by Hao in 1989 based on fossil specimens from the Posongchong Formation, 967: 646:. The Gosslingiales are considered to be zosterophylls with indeterminate growth and fertile branches generally showing 737:
Boyce, C.K. (2005), "The evolutionary history of roots and leaves", in Holbrook, N.M. & Zwieniecki, M.A. (eds.),
783: 116: 972: 881: 301:, but differs in branching pattern and in the arrangement of the sporangia, including their long stalks. 647: 674:
Hao, Shou-Gang (1989), "A new zosterophyll from the Lower Denovian (Siegenian) of Yunnan, China",
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Crane, P.R.; Herendeen, P. & Friis, E.M. (2004), "Fossils and plant phylogeny",
317:(club-mosses and allies) has been much discussed. Hao considered that the spines of 309:
The evolutionary relationship between spines in early plants and the small leaves ('
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could be precursors of lycophylls. Boyce regarded the occurrence of the spines of
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on sporangia as evidence that they were not related to lycophylls in this way.
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List of Latin and Greek words commonly used in systematic names
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This article is about the genus. For other taxonomic uses, see
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Paleobotany, The Biology and Evolution of Fossil Plants
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A cladogram published in 2004 by Crane et al. places
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Taylor, T.N.; Taylor, E.L. & Krings, M. (2009),
888: 870: 630:Hao and Xue in 2013 used the absence of terminal 743:, Burlington: Academic Press, pp. 479–499, 364:        347:(living and extinct clubmosses and relatives). 8: 831:Hao, Shougang & Xue, Jinzhuang (2013), 776: 774: 876: 837:, Beijing: Science Press, pp. 52–54, 669: 667: 665: 663: 27: 796: 659: 246:). The name is derived from the Greek 677:Review of Palaeobotany and Palynology 436:Z. llanoveranum, Z. fertile 16:Extinct genus of spore-bearing plants 7: 732: 730: 700: 698: 289:having annular, spiral thickenings. 818: 709:(2nd ed.), Amsterdam; Boston: 554: 386: 360: 350: 14: 871:Crane, Herendeen & Friis 2004 558: †basal groups  390: †basal groups  749:10.1016/B978-012088457-5/50025-3 115: 285:was present in the stems, with 252:, referring to the disc-shaped 1: 978:Prehistoric lycophyte genera 740:Vascular Transport in Plants 690:10.1016/0034-6667(89)90018-3 591:extant and extinct members 239:or Siegenian stage, around 994: 784:American Journal of Botany 634:to place the genus in the 248: 18: 589: 560: 552: 453: 392: 384: 366: 358: 203: 198: 112:Scientific classification 110: 30: 650:(initially curled up). 798:10.3732/ajb.91.10.1683 968:Early Devonian plants 504:B. citrulliforme 447: †'core'  339:of broadly defined " 242:411 to 408 648:circinate vernation 295:somewhat resembles 821:, pp. 169–170 432: myretonianum 955: 954: 940:Open Tree of Life 882:Taxon identifiers 844:978-7-03-036616-0 758:978-0-12-088457-5 720:978-0-12-373972-8 627: 626: 618: 617: 609: 608: 600: 599: 492:Protobarinophyton 260:, Yunnan, China. 244:million years ago 216: 215: 211: 194: 985: 948: 947: 935: 934: 922: 921: 909: 908: 907: 877: 854: 853: 852: 851: 828: 822: 816: 810: 809: 800: 778: 769: 767: 766: 765: 734: 725: 723: 702: 693: 692: 684:(3–4): 155–171, 671: 642:, in the family 555: 387: 361: 351: 343:", basal to the 258:Wenshan district 251: 250: 245: 209: 192: 185: 174: 169: 120: 119: 105: 42: 36:Temporal range: 28: 993: 992: 988: 987: 986: 984: 983: 982: 958: 957: 956: 951: 943: 938: 930: 925: 917: 912: 903: 902: 897: 884: 863: 858: 857: 849: 847: 845: 830: 829: 825: 817: 813: 791:(10): 1683–99, 780: 779: 772: 763: 761: 759: 736: 735: 728: 721: 704: 703: 696: 673: 672: 661: 656: 628: 619: 610: 601: 307: 283:Vascular tissue 266: 240: 229:vascular plants 191: 183: 172: 167: 114: 106: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 38: 37: 34: 24: 17: 12: 11: 5: 991: 989: 981: 980: 975: 970: 960: 959: 953: 952: 950: 949: 936: 923: 910: 894: 892: 886: 885: 880: 874: 873: 862: 861:External links 859: 856: 855: 843: 823: 811: 770: 757: 726: 719: 711:Academic Press 694: 658: 657: 655: 652: 625: 624: 621: 620: 616: 615: 612: 611: 607: 606: 603: 602: 598: 597: 594: 593: 588: 580: 579: 576: 575: 572: deciduum 570:Zosterophyllum 559: 553: 551: 548: 547: 544: 543: 500: obscurum 457:Zosterophyllum 452: 444: 443: 440: 439: 430:Zosterophyllum 408:Distichophytum 391: 385: 383: 380: 379: 376: 375: 365: 359: 357: 349: 306: 303: 268:The leafless 265: 262: 233:Early Devonian 214: 213: 210:S.G.Hao (1989) 201: 200: 196: 195: 193:S.G.Hao (1989) 181: 177: 176: 165: 158: 157: 152: 145: 144: 139: 132: 131: 126: 122: 121: 108: 107: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 43: 40:Early Devonian 35: 15: 13: 10: 9: 6: 4: 3: 2: 990: 979: 976: 974: 973:Zosterophylls 971: 969: 966: 965: 963: 946: 941: 937: 933: 928: 924: 920: 915: 911: 906: 900: 896: 895: 893: 891: 887: 883: 878: 872: 868: 865: 864: 860: 846: 840: 836: 835: 827: 824: 820: 815: 812: 808: 804: 799: 794: 790: 786: 785: 777: 775: 771: 760: 754: 750: 746: 742: 741: 733: 731: 727: 722: 716: 712: 708: 701: 699: 695: 691: 687: 683: 679: 678: 670: 668: 666: 664: 660: 653: 651: 649: 645: 641: 640:Gosslingiales 637: 633: 623: 622: 614: 613: 605: 604: 596: 595: 592: 586: 582: 581: 578: 577: 574: 573: 571: 566: 565: 557: 556: 550: 549: 546: 545: 542: 541: 540: 539:Crenaticaulis 535: 534: 533:Serrulacaulis 529: 528: 523: 522: 517: 516: 511: 510: 505: 501: 499: 494: 493: 488: 487: 486:Thrinkophyton 482: 478: 477: 472: 471: 466: 465: 460: 458: 450: 449:zosterophylls 446: 445: 442: 441: 438: 437: 433: 431: 426: 425: 420: 419: 414: 410: 409: 404: 403: 398: 397: 396:Adoketophyton 389: 388: 382: 381: 378: 377: 374: 373: 372: 363: 362: 356: 353: 352: 348: 346: 342: 341:zosterophylls 338: 335: 331: 326: 324: 320: 316: 312: 304: 302: 300: 299: 294: 293:D. longistipa 290: 288: 284: 280: 275: 274:D. longistipa 271: 263: 261: 259: 255: 243: 238: 234: 230: 226: 222: 221: 212: 208: 207:D. longistipa 202: 197: 190: 189: 182: 179: 178: 175: 171:Zosterophylls 166: 163: 160: 159: 156: 153: 150: 147: 146: 143: 142:Tracheophytes 140: 137: 134: 133: 130: 127: 124: 123: 118: 113: 109: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 41: 33: 29: 26: 22: 889: 848:, retrieved 833: 826: 814: 788: 782: 762:, retrieved 739: 706: 681: 675: 644:Discaliaceae 636:paraphyletic 629: 590: 568: 562: 561: 537: 531: 525: 519: 515:Deheubarthia 513: 507: 503: 498:Barinophyton 496: 490: 484: 480: 474: 468: 462: 455: 454: 435: 428: 422: 416: 412: 406: 401: 400: 394: 393: 369: 367: 334:paraphyletic 329: 327: 322: 318: 313:') of later 308: 296: 292: 291: 273: 267: 219: 218: 217: 206: 204: 187: 186: 161: 148: 135: 31: 25: 527:Anisophyton 459:divaricatum 264:Description 227:of extinct 962:Categories 850:2019-10-25 764:2011-02-06 654:References 476:Gosslingia 371:Hicklingia 355:lycophytes 337:stem group 315:lycoposids 311:lycophylls 270:sporophyte 155:Lycophytes 867:Cladogram 632:sporangia 585:lycopsids 345:lycopsids 305:Phylogeny 287:tracheids 254:sporangia 173: (?) 125:Kingdom: 905:Q5281411 899:Wikidata 890:Discalis 819:Hao 1989 807:21652317 768:, p. 491 724:, p. 254 587:  521:Konioria 509:Sawdonia 470:Oricilla 451:  413:Rebuchia 402:Discalis 330:Discalis 323:Discalis 319:Discalis 298:Sawdonia 220:Discalis 199:Species 188:Discalis 32:Discalis 945:4738295 932:1313422 919:4893993 583:  464:Tarella 368:†  237:Pragian 231:of the 180:Genus: 162:Plesion 129:Plantae 841:  805:  755:  717:  638:order 564:Nothia 418:Gumuia 279:spores 249:δίσκος 927:IRMNG 869:from 332:in a 225:genus 223:is a 149:Clade 136:Clade 914:GBIF 839:ISBN 803:PMID 753:ISBN 715:ISBN 481:Hsua 424:Huia 46:PreꞒ 793:doi 745:doi 686:doi 415:), 272:of 964:: 942:: 929:: 916:: 901:: 801:, 789:91 787:, 773:^ 751:, 729:^ 713:, 697:^ 682:57 680:, 662:^ 567:, 536:, 530:, 524:, 518:, 512:, 506:, 502:, 495:, 489:, 483:, 479:, 473:, 467:, 461:, 434:, 427:, 421:, 411:(= 405:, 399:, 164:: 151:: 138:: 96:Pg 795:: 747:: 688:: 235:( 205:† 184:† 168:† 101:N 91:K 86:J 81:T 76:P 71:C 66:D 61:S 56:O 51:Ꞓ 23:.

Index

List of Latin and Greek words commonly used in systematic names
Early Devonian
PreꞒ

O
S
D
C
P
T
J
K
Pg
N
Scientific classification
Edit this classification
Plantae
Tracheophytes
Lycophytes
Zosterophylls (?)
Discalis
genus
vascular plants
Early Devonian
Pragian
411 to 408
sporangia
Wenshan district
sporophyte
spores

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