983:
350:". Adults are about 33 cm in length (including its tail, which is about 13 cm long), and weigh between 150 and 300 grams. Their appearance varies according to the season. In the summer, they are greyish or almost greenish-brown on their backs, and pale orange on the chest and belly, while legs and feet appear brown. In the winter, the coat is browner and the underside is grayer; also, the ears appear even tuftier than they do in summer. Like many squirrels, Douglas squirrels have a white eye-ring.
513:
354:
232:
82:
38:
993:
499:
57:
988:
423:
Douglas squirrels are territorial; in winter, each squirrel occupies a territory of about 10 000 square metres, but during the breeding season a mated pair will defend a single territory together. Douglas squirrels are active by day, throughout the year, often chattering noisily at intruders. On
495:; although they quickly acclimatize to human presence, humans can be a threat to them, through robbing of their cone caches to find seeds for tree cultivation and through the destruction of old-growth forest. However, the squirrels' numbers appear to be unaffected by commercial thinning of forests.
424:
summer nights, they sleep in ball-shaped nests that they make in the trees, but in the winter they use holes in trees as nests. Groups of squirrels seen together during the summer are likely to be juveniles from a single litter.
472:. As the squirrel peels the scales of cones to get at the seeds, the discarded scales accumulate into piles that can grow to several meters across as the same site is used by generations of squirrels.
377:) are weaned at about eight weeks of age. There may be up to six kits in a litter, though four is more usual. In the southern and lower parts of their range, they produce two litters each year.
1721:
1731:
1569:
1660:
534:
1701:
1726:
862:
1543:
709:
Hope, Andrew G.; Malaney, Jason L.; Bell, Kayce C.; Salazar-Miralles, Fernando; Chavez, Andreas S.; Barber, Brian R.; Cook, Joseph A. (2016-07-01).
1595:
533:
1711:
324:
1741:
1736:
693:
660:
711:"Revision of widespread red squirrels (genus: Tamiasciurus) highlights the complexity of speciation within North American forests"
420:, which inhabits the coniferous forests of the rest of North America. The two species have very minimal overlap in the territory.
1621:
357:
982:
945:
855:
522:
1639:
1600:
624:
840:
830:
1379:
1370:
1356:
1348:
503:
410:
81:
1634:
1582:
646:
1716:
1665:
1495:
848:
465:
1418:
766:
1260:
1060:
1052:
969:
328:
1457:
1100:
1011:
1002:
961:
1706:
1426:
1228:
1220:
1212:
1164:
953:
417:
304:
176:
1144:
1108:
1530:
1504:
1284:
1268:
1236:
722:
406:
394:
215:
1652:
1292:
1244:
1204:
1184:
1068:
46:
1441:
1328:
1116:
1044:
608:
604:
361:
346:
described the
Douglas squirrel as "by far the most interesting and influential of the California
76:
1574:
1509:
373:
Mating can occur as early as
February. Gestation is about four weeks, and the young (which are
353:
1673:
1517:
1252:
1076:
748:
740:
689:
666:
656:
402:
308:
268:
37:
1678:
1308:
992:
804:
730:
488:
441:
405:
or mature second-growth forests, and some authors regard them as dependent on its presence.
276:
231:
1613:
1092:
1084:
835:
628:
288:
726:
1136:
476:
461:
449:
1695:
1608:
1393:
585:
457:
397:(mountains) of California, northwards to the southwestern coast of British Columbia.
272:
264:
260:
153:
66:
61:
498:
416:
Throughout most their range, Douglas squirrels essentially replace the niche of the
1522:
1300:
1276:
928:
917:
193:
621:
532:
1647:
1626:
1556:
1489:
735:
710:
433:
312:
987:
576:
553:
386:
332:
284:
1480:
744:
876:
825:
795:
in commercially thinned and unthinned stands of coastal coniferous forest".
670:
374:
347:
343:
93:
20:
1587:
752:
464:, and some fruit including strawberries and plums. Douglas squirrels are
409:
is a distinctive subspecies of the
Douglas squirrel that instead inhabits
1474:
899:
880:
872:
787:
Ransome, D.B.; Sullivan, T.P. (2002). "Short-term population dynamics of
143:
113:
460:
cones, as well as acorns, berries, mushrooms, the eggs of birds such as
1548:
1025:
468:
hoarders, storing their food in a single location or 'larder' called a
390:
655:(3rd ed.). The Johns Hopkins University Press. pp. 754–818.
1561:
911:
905:
893:
480:
469:
292:
280:
133:
123:
103:
1451:
808:
497:
352:
652:
Mammal
Species of the World: a taxonomic and geographic reference
331:
called it the "Pillillooeet", in imitation of its characteristic
1535:
1455:
844:
650:
502:
Pacific silver fir cone debris from
Douglas squirrels feeding,
432:
Douglas squirrels mostly eat seeds of coniferous trees such as
631:. Description of several new Species of American quadrupeds.
492:
484:
633:
Journal of the
Academy of Natural Sciences of Philadelphia, 8
511:
413:
pine forests in a small portion of Baja
California.
19:"Chickaree" redirects here. Not to be confused with
1464:
1391:
1368:
1023:
1000:
926:
271:(including the northwestern coastal states of the
856:
645:Thorington, R.W. Jr.; Hoffmann, R.S. (2005).
303:, although these names are also used for the
8:
456:). They also eat the fleshy scales of green
831:John Muir's account of the Douglas Squirrel
684:Muir, John (1989). "The Douglas Squirrel".
1722:Fauna of the Sierra Nevada (United States)
1452:
863:
849:
841:
575:Linzey, A. V. & Hammerson, G. (2008).
287:, with an isolated subspecies in northern
230:
55:
36:
27:
734:
826:Tracks and calls of the Douglas Squirrel
1732:Fauna of the Northwestern United States
649:. In Wilson, D.E.; Reeder, D.M (eds.).
567:
551:
767:"Gray squirrels and scatter hoarding"
715:Molecular Phylogenetics and Evolution
275:as well as the southwestern coast of
7:
540:Alarm call of the Douglas squirrel (
1702:IUCN Red List least concern species
836:Smithsonian: "The Douglas Squirrel"
797:Canadian Journal of Forest Research
586:IUCN Red List of Threatened Species
1727:Fauna of the Western United States
393:along the Pacific Coast, from the
14:
991:
986:
981:
946:Central American dwarf squirrel
688:. The Sierra Club. p. 173.
552:Problems playing this file? See
530:
358:Anacortes Community Forest Lands
80:
295:. It is sometimes known as the
1:
1357:Southern Amazon red squirrel
1349:Northern Amazon red squirrel
972:(Microsciurus santanderensis)
1712:Rodents of the United States
504:North Cascades National Park
736:10.1016/j.ympev.2016.04.014
686:The Mountains of California
1758:
1742:Taxa named by John Bachman
1419:Southwestern red squirrel
1380:Bangs's mountain squirrel
1053:South Yungas red squirrel
1014:(Rheithrosciurus macrotis)
956:(Microsciurus flaviventer)
385:Douglas squirrels live in
18:
1737:Mammals described in 1839
1438:
1429:(Tamiasciurus hudsonicus)
1261:Calabrian black squirrel
1085:Atlantic Forest squirrel
1061:Yellow-throated squirrel
979:
970:Santander dwarf squirrel
888:
871:Extant species of family
238:
229:
206:
201:
182:
175:
77:Scientific classification
75:
53:
44:
35:
30:
1413:(Tamiasciurus douglasii)
1382:(Syntheosciurus brochus)
627:January 7, 2009, at the
475:Their predators include
239:Range (does not include
1421:(Tamiasciurus fremonti)
1295:(Sciurus variegatoides)
1012:Tufted ground squirrel
962:Western dwarf squirrel
1510:Tamiasciurus_douglasii
1496:Tamiasciurus douglasii
1466:Tamiasciurus douglasii
1427:American red squirrel
1311:(Sciurus yucatanensis)
1271:(Sciurus nayaritensis)
1263:(Sciurus meridionalis)
1231:(Sciurus carolinensis)
1229:Eastern gray squirrel
1221:Mexican gray squirrel
1213:Arizona gray squirrel
1165:Western gray squirrel
1063:(Sciurus gilvigularis)
964:(Microsciurus mimulus)
954:Amazon dwarf squirrel
793:Tamiasciurus douglasii
579:Tamiasciurus douglasii
542:Tamiasciurus douglasii
524:Tamiasciurus douglasii
516:
506:
399:Tamiasciurus douglasii
365:
256:Tamiasciurus douglasii
186:Tamiasciurus douglasii
1351:(Sciurus igniventris)
1269:Mexican fox squirrel
1223:(Sciurus aureogaster)
1215:(Sciurus arizonensis)
1071:(Sciurus granatensis)
1055:(Sciurus argentinius)
948:(Microsciurus alfari)
515:
501:
438:Pseudotsuga menziesii
418:American red squirrel
356:
305:American red squirrel
1293:Variegated squirrel
1119:(Sciurus stramineus)
1101:Richmond's squirrel
1095:(Sciurus pucheranii)
1069:Red-tailed squirrel
1359:(Sciurus spadiceus)
1329:Caucasian squirrel
1147:(Sciurus pyrrhinus)
1145:Junín red squirrel
1139:(Sciurus flammifer)
1117:Guayaquil squirrel
1109:Sanborn's squirrel
1103:(Sciurus richmondi)
1045:Brazilian squirrel
769:. 24 September 2013
727:2016MolPE.100..170H
47:Conservation status
1331:(Sciurus anomalus)
1303:(Sciurus vulgaris)
1287:(Sciurus oculatus)
1285:Peters's squirrel
1253:Japanese squirrel
1239:(Sciurus colliaei)
1237:Collie's squirrel
1111:(Sciurus sanborni)
1077:Bolivian squirrel
1047:(Sciurus aestuans)
789:Glaucomys sabrinus
647:"Family Sciuridae"
622:Bachman, J. (1839)
607:}}: old-form url (
517:
507:
403:old-growth forests
366:
321:Douglas's squirrel
222:T. d. mollipilosus
1717:Rodents of Canada
1689:
1688:
1674:Open Tree of Life
1458:Taxon identifiers
1449:
1448:
1411:Douglas squirrel
1309:Yucatan squirrel
1245:Deppe's squirrel
1205:Allen's squirrel
1185:Abert's squirrel
1167:(Sciurus griseus)
1087:(Sciurus ingrami)
1079:(Sciurus ignitus)
936:(Dwarf squirrels)
803:(11): 2043–2050.
535:
489:northern goshawks
407:Mearns's squirrel
317:Douglas' squirrel
309:Variant spellings
269:Pacific Northwest
263:found in western
247:
246:
168:T. douglasii
70:
31:Douglas squirrel
16:Species of rodent
1749:
1682:
1681:
1669:
1668:
1656:
1655:
1643:
1642:
1630:
1629:
1617:
1616:
1604:
1603:
1591:
1590:
1578:
1577:
1565:
1564:
1552:
1551:
1539:
1538:
1526:
1525:
1513:
1512:
1500:
1499:
1498:
1485:
1484:
1483:
1453:
1404:
1403:
1402:
1401:(Pine squirrels)
1345:
1325:
1247:(Sciurus deppei)
1207:(Sciurus alleni)
1201:
1187:(Sciurus aberti)
1181:
1161:
1133:
1093:Andean squirrel
1041:
995:
990:
985:
939:
938:
937:
865:
858:
851:
842:
813:
812:
784:
778:
777:
775:
774:
763:
757:
756:
738:
706:
700:
699:
681:
675:
674:
642:
636:
619:
613:
612:
601:
599:
597:
572:
537:
536:
514:
446:Picea sitchensis
325:Native Americans
277:British Columbia
251:Douglas squirrel
234:
188:
85:
84:
64:
59:
58:
40:
28:
1757:
1756:
1752:
1751:
1750:
1748:
1747:
1746:
1692:
1691:
1690:
1685:
1677:
1672:
1664:
1659:
1651:
1646:
1638:
1633:
1625:
1620:
1612:
1607:
1599:
1594:
1586:
1581:
1573:
1568:
1560:
1555:
1547:
1542:
1534:
1529:
1521:
1516:
1508:
1503:
1494:
1493:
1488:
1479:
1478:
1473:
1460:
1450:
1445:
1434:
1400:
1399:
1398:
1397:
1387:
1364:
1338:
1318:
1279:(Sciurus niger)
1194:
1174:
1154:
1137:Fiery squirrel
1126:
1034:
1019:
1003:Rheithrosciurus
996:
977:
935:
934:
933:
932:
922:
884:
869:
822:
817:
816:
809:10.1139/x02-126
786:
785:
781:
772:
770:
765:
764:
760:
708:
707:
703:
696:
683:
682:
678:
663:
644:
643:
639:
629:Wayback Machine
620:
616:
602:
595:
593:
574:
573:
569:
564:
559:
558:
550:
548:
547:
546:
545:
538:
531:
528:
518:
512:
477:Pacific martens
462:yellow warblers
430:
383:
371:
341:
289:Baja California
210:T. d. douglasii
197:
190:
184:
171:
79:
71:
60:
56:
49:
24:
17:
12:
11:
5:
1755:
1753:
1745:
1744:
1739:
1734:
1729:
1724:
1719:
1714:
1709:
1704:
1694:
1693:
1687:
1686:
1684:
1683:
1670:
1657:
1644:
1631:
1618:
1605:
1592:
1579:
1566:
1553:
1540:
1527:
1514:
1501:
1486:
1470:
1468:
1462:
1461:
1456:
1447:
1446:
1439:
1436:
1435:
1433:
1432:
1424:
1416:
1407:
1405:
1389:
1388:
1386:
1385:
1376:
1374:
1371:Syntheosciurus
1366:
1365:
1363:
1362:
1354:
1346:
1335:
1334:
1326:
1315:
1314:
1306:
1298:
1290:
1282:
1274:
1266:
1258:
1250:
1242:
1234:
1226:
1218:
1210:
1202:
1191:
1190:
1182:
1171:
1170:
1162:
1158:Hesperosciurus
1151:
1150:
1142:
1134:
1123:
1122:
1114:
1106:
1098:
1090:
1082:
1074:
1066:
1058:
1050:
1042:
1031:
1029:
1021:
1020:
1018:
1017:
1008:
1006:
998:
997:
980:
978:
976:
975:
967:
959:
951:
942:
940:
924:
923:
921:
920:
914:
908:
902:
896:
889:
886:
885:
870:
868:
867:
860:
853:
845:
839:
838:
833:
828:
821:
820:External links
818:
815:
814:
779:
758:
701:
694:
676:
661:
637:
614:
566:
565:
563:
560:
549:
539:
529:
521:
520:
519:
510:
509:
508:
454:Pinus contorta
429:
426:
382:
379:
370:
367:
340:
337:
245:
244:
236:
235:
227:
226:
225:
224:
219:
212:
204:
203:
199:
198:
191:
180:
179:
173:
172:
165:
163:
159:
158:
151:
147:
146:
141:
137:
136:
131:
127:
126:
121:
117:
116:
111:
107:
106:
101:
97:
96:
91:
87:
86:
73:
72:
54:
51:
50:
45:
42:
41:
33:
32:
15:
13:
10:
9:
6:
4:
3:
2:
1754:
1743:
1740:
1738:
1735:
1733:
1730:
1728:
1725:
1723:
1720:
1718:
1715:
1713:
1710:
1708:
1705:
1703:
1700:
1699:
1697:
1680:
1675:
1671:
1667:
1662:
1658:
1654:
1649:
1645:
1641:
1636:
1632:
1628:
1623:
1619:
1615:
1610:
1606:
1602:
1597:
1593:
1589:
1584:
1580:
1576:
1571:
1567:
1563:
1558:
1554:
1550:
1545:
1541:
1537:
1532:
1528:
1524:
1519:
1515:
1511:
1506:
1502:
1497:
1491:
1487:
1482:
1476:
1472:
1471:
1469:
1467:
1463:
1459:
1454:
1444:
1443:
1437:
1431:
1430:
1425:
1423:
1422:
1417:
1415:
1414:
1409:
1408:
1406:
1396:
1395:
1390:
1384:
1383:
1378:
1377:
1375:
1373:
1372:
1367:
1361:
1360:
1355:
1353:
1352:
1347:
1344:
1343:
1337:
1336:
1333:
1332:
1327:
1324:
1323:
1317:
1316:
1313:
1312:
1307:
1305:
1304:
1301:Red squirrel
1299:
1297:
1296:
1291:
1289:
1288:
1283:
1281:
1280:
1277:Fox squirrel
1275:
1273:
1272:
1267:
1265:
1264:
1259:
1257:
1256:
1255:(Sciurus lis)
1251:
1249:
1248:
1243:
1241:
1240:
1235:
1233:
1232:
1227:
1225:
1224:
1219:
1217:
1216:
1211:
1209:
1208:
1203:
1200:
1199:
1193:
1192:
1189:
1188:
1183:
1180:
1179:
1173:
1172:
1169:
1168:
1163:
1160:
1159:
1153:
1152:
1149:
1148:
1143:
1141:
1140:
1135:
1132:
1131:
1125:
1124:
1121:
1120:
1115:
1113:
1112:
1107:
1105:
1104:
1099:
1097:
1096:
1091:
1089:
1088:
1083:
1081:
1080:
1075:
1073:
1072:
1067:
1065:
1064:
1059:
1057:
1056:
1051:
1049:
1048:
1043:
1040:
1039:
1038:Guerlinguetus
1033:
1032:
1030:
1028:
1027:
1022:
1016:
1015:
1010:
1009:
1007:
1005:
1004:
999:
994:
989:
984:
974:
973:
968:
966:
965:
960:
958:
957:
952:
950:
949:
944:
943:
941:
931:
930:
925:
919:
915:
913:
909:
907:
903:
901:
897:
895:
891:
890:
887:
882:
878:
874:
866:
861:
859:
854:
852:
847:
846:
843:
837:
834:
832:
829:
827:
824:
823:
819:
810:
806:
802:
798:
794:
790:
783:
780:
768:
762:
759:
754:
750:
746:
742:
737:
732:
728:
724:
720:
716:
712:
705:
702:
697:
695:0-87156-663-X
691:
687:
680:
677:
672:
668:
664:
662:0-8018-8221-4
658:
654:
653:
648:
641:
638:
634:
630:
626:
623:
618:
615:
610:
606:
592:
588:
587:
582:
580:
571:
568:
561:
557:
555:
543:
527:
525:
505:
500:
496:
494:
490:
486:
485:domestic cats
482:
478:
473:
471:
467:
463:
459:
458:giant sequoia
455:
451:
447:
443:
439:
435:
427:
425:
421:
419:
414:
412:
408:
404:
400:
396:
395:Sierra Nevada
392:
388:
380:
378:
376:
368:
363:
359:
355:
351:
349:
345:
338:
336:
334:
330:
326:
322:
318:
314:
310:
306:
302:
301:pine squirrel
298:
294:
290:
286:
283:) to central
282:
278:
274:
273:United States
270:
266:
265:North America
262:
261:pine squirrel
258:
257:
252:
242:
241:T. d. mearnsi
237:
233:
228:
223:
220:
218:
217:
216:T. d. mearnsi
213:
211:
208:
207:
205:
200:
195:
189:
187:
181:
178:
177:Binomial name
174:
170:
169:
164:
161:
160:
157:
156:
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
125:
122:
119:
118:
115:
112:
109:
108:
105:
102:
99:
98:
95:
92:
89:
88:
83:
78:
74:
68:
63:
62:Least Concern
52:
48:
43:
39:
34:
29:
26:
22:
1707:Tamiasciurus
1465:
1440:
1428:
1420:
1412:
1410:
1394:Tamiasciurus
1392:
1381:
1369:
1358:
1350:
1341:
1339:
1330:
1321:
1319:
1310:
1302:
1294:
1286:
1278:
1270:
1262:
1254:
1246:
1238:
1230:
1222:
1214:
1206:
1197:
1195:
1186:
1177:
1175:
1166:
1157:
1155:
1146:
1138:
1130:Hadrosciurus
1129:
1127:
1118:
1110:
1102:
1094:
1086:
1078:
1070:
1062:
1054:
1046:
1037:
1035:
1024:
1013:
1001:
971:
963:
955:
947:
929:Microsciurus
927:
918:Sciuromorpha
800:
796:
792:
788:
782:
771:. Retrieved
761:
718:
714:
704:
685:
679:
651:
640:
632:
617:
594:. Retrieved
590:
584:
578:
570:
541:
523:
474:
453:
445:
442:Sitka spruce
437:
431:
422:
415:
398:
384:
372:
342:
320:
316:
300:
296:
255:
254:
250:
248:
240:
221:
214:
209:
185:
183:
167:
166:
155:Tamiasciurus
154:
25:
1648:NatureServe
1557:iNaturalist
1490:Wikispecies
875:(subfamily
721:: 170–182.
434:Douglas fir
339:Description
329:Kings River
313:common name
267:, from the
202:Subspecies
1696:Categories
1342:Urosciurus
1178:Otosciurus
916:Suborder:
773:2015-11-20
562:References
554:media help
526:alarm call
450:shore pine
387:coniferous
362:Washington
333:alarm call
285:California
1653:2.1156423
1340:Subgenus
1320:Subgenus
1196:Subgenus
1176:Subgenus
1156:Subgenus
1128:Subgenus
1036:Subgenus
892:Kingdom:
877:Sciurinae
873:Sciuridae
745:1055-7903
605:cite iucn
596:8 January
375:altricial
344:John Muir
297:chickaree
162:Species:
144:Sciuridae
100:Kingdom:
94:Eukaryota
21:Chickadee
1640:12400250
1575:11334808
1475:Wikidata
1442:Category
912:Rodentia
906:Mammalia
900:Chordata
898:Phylum:
894:Animalia
881:Sciurini
753:27083861
671:26158608
635:, 57-74.
625:Archived
391:habitats
369:Breeding
348:Sciuridæ
140:Family:
134:Rodentia
124:Mammalia
114:Chordata
110:Phylum:
104:Animalia
90:Domain:
67:IUCN 3.1
1627:1001704
1549:2437281
1481:Q386410
1198:Sciurus
1026:Sciurus
910:Order:
904:Class:
723:Bibcode
481:bobcats
401:prefer
389:forest
381:Habitat
311:of the
259:) is a
196:, 1839)
194:Bachman
150:Genus:
130:Order:
120:Class:
65: (
1679:345257
1666:144198
1601:180167
1536:347429
883:tribe)
751:
743:
692:
669:
659:
491:, and
470:midden
466:larder
448:) and
323:. The
293:Mexico
281:Canada
1614:42586
1588:62769
1570:IRMNG
1562:46259
1523:54MTJ
1322:Tenes
411:xeric
319:and
1661:NCBI
1609:IUCN
1596:ITIS
1544:GBIF
791:and
749:PMID
741:ISSN
690:ISBN
667:OCLC
657:ISBN
609:help
598:2009
591:2008
493:owls
428:Diet
315:are
249:The
1635:MSW
1622:MDD
1583:ISC
1531:EoL
1518:CoL
1505:ADW
805:doi
731:doi
719:100
440:),
327:of
307:.
299:or
279:in
1698::
1676::
1663::
1650::
1637::
1624::
1611::
1598::
1585::
1572::
1559::
1546::
1533::
1520::
1507::
1492::
1477::
879:,
801:32
799:.
747:.
739:.
729:.
717:.
713:.
665:.
603:{{
589:.
583:.
487:,
483:,
479:,
335:.
291:,
243:)
864:e
857:t
850:v
811:.
807::
776:.
755:.
733::
725::
698:.
673:.
611:)
600:.
581:"
577:"
556:.
544:)
452:(
444:(
436:(
364:)
360:(
253:(
192:(
69:)
23:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.