4611:
6726:
4604:
3108:
3933:
3919:
6178:
5399:
6164:
3912:
6171:
3114:
647:(NHM OR50100). The casts show some detail that is no longer preserved on the original specimens, which have significantly degraded due to pyrite disease. The discovery of this genus gave North American paleontologists the opportunity to observe an articulated, albeit incomplete, theropod skeleton. During the late 19th century, this genus unfortunately became a wastebasket taxon for the referral of isolated theropod elements from across North America, given that
961:
4618:
140:
1787:
3926:
6739:
5417:
1689:
5405:
884:
5424:
1095:
6733:
504:
6158:
1103:
782:
118:
5412:
1832:
663:
998:
in 1997, in light of the many different theropods discovered since Cope's time. He felt that due to some unusual features it couldn't be placed in any existing family and, like Marsh, he felt that it warranted placement in
Dryptosauridae. This phylogenetic assignment was also supported by the work of
947:
tubercle, which is approximately 1.7 times as wide as the shaft immediately distally, the presence of an ovoid fossa on the medial surface of the femoral shaft immediately proximal to the medial condyle (which is demarcated anteriorly by the mesiodistal crest and demarcated medially by a novel crest)
2199:
Dryptosaurus." In: Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B. The Age of
Dinosaurs. Publications International, LTD. p.
547:
was notified of the discovery and encouraged to visit the marl pit to obtain the specimen. When he arrived, he was thoroughly surprised by the skeleton's completeness and uniqueness, calling it the "finest discovery I have yet made". The skeleton was then deposited at the
948:
the presence of a proximomedially trending ridge on the anterior surface of the fibula immediately proximal to the iliofibularis tubercle, the lip on the lateral surface of the lateral condyle of the astragalus being prominent and overlapping the proximal surface of the
879:
III. The ontological maturity of the holotype individual is supported by the fact that the neurocentral sutures are closed in all of its caudal vertebrae. AMNH FARB 2438 consists of left metatarsal IV, which are likely from the same individual as the holotype.
1805:
may represent the earliest depiction of theropods as highly active and dynamic animals. Knight's artistic hand was guided by E. D. Cope and reflects their progressive opinion about theropod agility, despite their large size, as well as the opinion of
1757:
and its coeval fauna from western North
America, which was dominated by the much larger tyrannosaurids. Although certainly a carnivore, the paucity of known Cretaceous East Coast dinosaurs make ascertaining the specific diet of
614:
possibly belonging to the skeleton was found at the same locale, but this has come into question. Cope presented his description of the specimen to the
Academy on August 21, 1866, naming it a new genus and species of dinosaur,
911:/cm), and recurved. The femur is only 3% longer than the tibia. The longest manual ungual phalanx recovered measured 176 mm (6.9 in) in length. The morphology of the proximal portion of metatarsal IV suggests that
942:
can be distinguished based on the following characteristics: the combination of a reduced humerus (humerus: femur ratio = 0.375) and a large hand (phalanx I-1:femur ratio = 0.200), the strong mediolateral expansion of the
2089:
Chan-gyu Yun (2017). "Teihivenator gen. nov., a new generic name for the tyrannosauroid dinosaur "Laelaps" macropus (Cope, 1868; preoccupied by Koch, 1836)". Journal of
Zoological And Bioscience Research. 4 (2): 7–13.
459:
that could grow up to 7.5 metres (25 ft) long and weigh up to 756–1,500 kilograms (1,667–3,307 lb). Although it is now largely unknown outside of academic circles, the famous 1897 painting of the genus by
1741:
in central New Jersey. Studies suggest that the New Egypt
Formation is a marine unit and it is considered to be the deeper-water equivalent of the Tinton and Red Bank formations. This formation overlies the
2020:
Spamer, E.E., E. Daeschler, and L.G. Vostreys-Shapiro. 1995. A study of fossil vertebrate types in the
Academy of Natural Sciences of Philadelphia. Special Publication 16, Academy of Natural Sciences of
808:. Its hands, which are also relatively large, were once believed to have had three fingers. Brusatte et al. (2011), however, observed an overall similarity in the shape of the available phalanges of
2508:
Olsson, R.K. 1987. Cretaceous stratigraphy of the
Atlantic coastal plain, Atlantic highlands of New Jersey. Geological Society of America Centennial Field Guide–Northeastern Section: 87–90.
903:
had ziphodont dentition. The shape of the alveolus situated on the anterior portion of the fragment suggests that it housed a tooth that was smaller and more circular than the others. This
1778:
were also present, although they were less likely to be hunted due to their armor plating. When hunting, both the skull and hands were important for the capture and processing of prey.
792:
is estimated to have been 7.5 metres (25 ft) long and 756–1,500 kilograms (1,667–3,307 lb), although this is based on partial remains of one individual. Like its relative
2254:
Carr, T.D.; Williamson, T.E. & Schwimmer, D.R. (2005). "A new genus and species of tyrannosauroid from the Late
Cretaceous (middle Campanian) Demopolis Formation of Alabama".
1753:
During the
Maastrichtian stage, the Western Interior Seaway, which stretched in a north–south direction from the present-day Arctic Ocean down to the Gulf of Mexico, separated
6875:
952:, and metatarsal IV being observed with a flat shaft proximally (resulting in a semiovoid cross section that is much wider mediolaterally than it is long anteroposteriorly).
1810:, the curator of vertebrate paleontology at the American Museum of Natural History at the time of the painting's commission. The original painting is now preserved in the
2288:
Holtz, T.R. 2004. Tyrannosauroidea. In D. B. Weishampel, P. Dodson, and H. Osmólska (editors), The Dinosauria, 2nd ed.: 111–136. Berkeley: University of California Press.
2218:
Molnar, R.E. 1990. Problematic Theropoda. In D. B. Weishampel, P. Dodson, and H. Osmólska (editors), The Dinosauria: 306–317. Berkeley: University of California Press.
2007:
1706:
549:
816:
may have had only two functional digits. Each of its fingers were tipped with an eight-inch long, talon-like claw. Its arm morphology suggests that arm reduction in
2209:
Russell, D.A. 1970. Tyrannosauroids from the Late Cretaceous of western Canada. National Museum of Natural Science (Ottawa) Publications in Paleontology 1: 1–34.
2104:
Theropod specimens from the Navesink Formation and their implications for the Diversity and Biogeography of Ornithomimosaurs and Tyrannosauroids on Appalachia
6895:
715:
972:
has been classified in a number of different theropod families. Cope (1866), Leidy (1868), and Lydekker (1888) noted obvious similarities with the genus
766:
and found that only the partial tibia could be definitely classified as that of a tyrannosauroid and that the distal metatarsal could have been from an
1038:. This observation was also supported by the work of Baird and Horner (1979), but did not have wide acceptance until a new discovery was made in 2005.
1864:
Leidy, J. 1856. Notice of remains of extinct reptiles and fishes discovered by F. V. Hayden in the badlands of the Judith River, Nebraska Territory.
643:(2011) noted that well-preserved, historic casts of most of the type material from ANSP 9995 and AMNH FARB 2438 are housed in the collections of the
6803:
4610:
6890:
6885:
4603:
6725:
2052:
2427:"Dinosaurs from the Santonian–Campanian Atlantic coastline substantiate phylogenetic signatures of vicariance in Cretaceous North America"
899:
The fragmentary right maxilla preserves the three alveoli in full and the fourth only partially. The authors were able to ascertain that
827:
The type specimen is a fragmentary skeleton belonging to a single adult individual, ANSP 9995. ANSP 9995 consists of a fragmentary right
6900:
2570:
1811:
1661:
1941:
409:
3932:
5398:
651:
was not recognized as a distinct group of large theropods at the time and numerous theropod species were assigned to it (often as
1845:
1488:
3918:
6163:
516:
3107:
6905:
644:
3911:
6177:
3113:
1966:"Redescription of the holotype of Dryptosaurus aquilunguis (Dinosauria: Theropoda) from the Upper Cretaceous of New Jersey"
6880:
1965:
1716:
6604:
5416:
714:-like characteristics of the teeth are primitive to theropods as a whole and may have been present in other large-bodied
6910:
4617:
139:
6170:
3925:
528:
2490:
Olsson, R.K. (1960). "Foraminifera of latest Cretaceous and earliest Tertiary age in the New Jersey coastal plain".
6738:
2120:
1925:
907:
tooth is common among tyrannosauroids. The disarticulated teeth recovered are transversely narrow, serrated (17–18
512:
5423:
1878:
600:
which are now lost were also noted by Cope in addition to several sacral vertebrae which are now referred to the
2267:
2658:
908:
5404:
627:
who never failed to catch what it was hunting. The genus name was occupied by a mite and replaced by the name
6732:
2326:
2563:
1665:
770:. Brownstein also placed the pedal phalanges in Ornithimimosauria, though he did not create a lectotype for
126:
6758:
1807:
824:
may have used both its arms and its jaws and as weapons when hunting, capturing, and its processing prey.
702:
of Colorado. Five damaged partial tooth crowns from AMNH 5780, mistakenly thought to have belonged to the
1723:. The specimen was collected by quarry workers in marl and sandstone that were deposited during the late
6870:
6829:
5129:
5041:
2300:"The systematics of Late Jurassic tyrannosauroids (Dinosauria: Theropoda) from Europe and North America"
2001:
1771:
1712:
632:
481:
436:
287:
244:
6847:
6157:
2345:
6790:
6690:
4818:
4786:
2438:
2379:
1977:
1921:
1727:
734:
2349:
5142:
4929:
1738:
743:
683:
624:
544:
540:
477:
307:
5411:
1075:
and assigned it as an "intermediate" tyrannosauroid that is more derived than basal taxa, such as
6597:
5118:
4978:
4834:
4774:
4209:
4119:
4002:
2866:
2556:
2341:
2271:
1743:
1015:
979:
916:
725:
699:
316:
134:
6834:
2044:
2032:
960:
6816:
6479:
6396:
6370:
6209:
6196:
6074:
5632:
5584:
4416:
3766:
2845:
2472:
2454:
2407:
2175:
2076:
Holtz, T.R. (2004). "Tyrannosauroidea." Pp. 111-136 in Weishampel, Dodson and Osmolska (eds).
2048:
1937:
1794:
1786:
1536:
1131:
995:
767:
461:
6821:
1042:
was the only large carnivore known in eastern North America until the discovery of the basal
802:
seems to have had relatively long arms when compared with more derived tyrannosaurs, such as
511:
Up until 1866, theropods from the Americas were only known from isolated teeth discovered by
6583:
6340:
6250:
6039:
5971:
5834:
5753:
5725:
5685:
5566:
5536:
5054:
4807:
4671:
4528:
4428:
4277:
4184:
4142:
3709:
3019:
2462:
2446:
2397:
2387:
2357:
2353:
2321:
2311:
2263:
2165:
2157:
1985:
1837:
1311:
1121:
1083:
1064:
1047:
876:
747:
648:
611:
365:
2529:
6646:
6384:
6347:
6279:
6134:
6127:
5955:
5897:
5870:
5860:
5820:
5655:
5514:
5498:
5316:
5288:
5266:
5153:
4998:
4705:
4479:
4362:
4340:
4261:
4090:
4067:
4060:
4038:
4024:
4017:
3978:
3829:
3676:
3603:
3592:
3528:
3416:
3303:
2880:
2789:
2643:
2618:
1820:, in the late 19th century, depicted the animals as large, slow, tail-dragging behemoths.
1731:
1561:
1519:
1152:
1088:
987:
817:
623:", which is derived from the Greek word "hurricane" or "storm wind", was also the name of
605:
577:
444:
426:
226:
30:
2517:
Carr & Williamson (2002). "Evolution of basal Tyrannosauroidea from North America".
2442:
2383:
1981:
1814:
collections. By contrast, the typical illustrations of large carnivorous dinosaurs like
1688:
931:, in which the third metatarsal is "pinched" between the second and fourth metatarsals.
6611:
6574:
6555:
6517:
6506:
6303:
6016:
5982:
5964:
5944:
5841:
5555:
5300:
5162:
4989:
4971:
4799:
4766:
4752:
4713:
4487:
4348:
4305:
4298:
4220:
4196:
4149:
4108:
4045:
4031:
3759:
3744:
3571:
3405:
3269:
3237:
3226:
3203:
3165:
3026:
2996:
2920:
2859:
2758:
2467:
2402:
2365:
2361:
1902:
Cope, E.D. (1866). "Discovery of a gigantic dinosaur in the Cretaceous of New Jersey".
1711:
and was recovered in the West Jersey Marl Company Pit, in what came to be known as the
1673:
1043:
983:
883:
332:
6864:
6653:
6499:
6460:
6439:
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6311:
6238:
6032:
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5774:
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3003:
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2823:
2816:
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1438:
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1028:
1008:
1000:
927:
921:
804:
601:
536:
440:
68:
2275:
2227:
Gilmore, C.W. 1946. A new carnivorous dinosaur from the Lance Formation of Montana.
1094:
6625:
6563:
6533:
6453:
6446:
6424:
6416:
6296:
6288:
6268:
5991:
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5476:
5251:
5183:
5111:
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4236:
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4166:
4135:
4128:
4009:
3836:
3688:
3585:
3487:
3480:
3375:
3359:
3295:
3246:
3178:
2955:
2838:
2831:
2802:
2765:
2734:
2240:
Baird, D., and J. Horner. 1979. Cretaceous dinosaurs (Reptilia) of North Carolina.
2145:
1989:
1815:
1775:
1219:
974:
852:
848:
757:
729:
672:
553:
503:
341:
1102:
2392:
6679:
6660:
6590:
6540:
6526:
6468:
6409:
6354:
6231:
6025:
5917:
5886:
5803:
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5469:
5337:
5259:
5222:
5104:
5076:
5029:
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4827:
4684:
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4355:
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3994:
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3800:
3786:
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3352:
3323:
3253:
3218:
3211:
3011:
2989:
2982:
2941:
2933:
2909:
2894:
2873:
2781:
2773:
2067:
Cope, E.D., On the genus Laelaps, American Journal of Science, 1868; 2: 415-417.
1763:
851:
from the left hand (I-1, II-2, and an ungual), the shafts of the left and right
836:
781:
678:
589:
117:
43:
1993:
6618:
6547:
6492:
6113:
5928:
5746:
5732:
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5603:
5529:
5351:
5344:
5330:
5236:
5169:
5097:
5061:
5014:
4956:
4942:
4909:
4901:
4894:
4875:
4698:
4663:
4451:
4052:
3843:
3793:
3730:
3701:
3656:
3634:
3627:
3619:
3543:
3439:
3398:
3391:
3383:
3367:
3287:
3158:
3034:
2887:
2750:
2609:
2366:"Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans"
1827:
1720:
1359:
1335:
1243:
904:
872:
794:
532:
452:
197:
88:
53:
6781:
2531:- Cretaceous Dinosaurs of the Southeastern United States by David T. King Jr.
2458:
2179:
6639:
6632:
6332:
6325:
5998:
5937:
5591:
5309:
4855:
4759:
4494:
4079:
3964:
3951:
3878:
3814:
3737:
3716:
3473:
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3144:
3131:
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2720:
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2579:
1930:
949:
636:
489:
456:
429:
210:
151:
93:
37:
2476:
2411:
1110:
The following cladogram containing almost all tyrannosauroids is by Loewen
2316:
2299:
1964:
Carpenter, Ken & Russell, Dale A, Donald Baird, and R. Denton (1997).
662:
6775:
6223:
5648:
5641:
5613:
5506:
5006:
4387:
3723:
3550:
3311:
2948:
2627:
2603:
2426:
2370:
1669:
1383:
1186:
1169:
1077:
888:
840:
832:
585:
569:
561:
492:
469:
432:
184:
171:
83:
78:
63:
58:
48:
2450:
698:
was coined by Cope in 1877 for a partial dentary (now missing) from the
6808:
5781:
5543:
5273:
5176:
4501:
3822:
2541:
944:
856:
844:
828:
597:
593:
573:
520:
98:
73:
2150:(Dinosauria: Theropoda) and a Review of its Tyrannosauroid Affinities"
6432:
6120:
3564:
2597:
2170:
1023:
986:. Marsh, however, examined the remains and later assigned to its own
919:
foot, an advanced feature shared by derived tyrannosauroids, such as
868:
581:
473:
161:
6752:
1059:
in both body size and morphology. This discovery made it clear that
2161:
2043:. Bloomington and Indianapolis: Indiana University Press. pp.
1737:
The southern New Jersey Hornerstown Formation is also known as the
1018:
was the first to observe that certain anatomical features may link
1011:, though its exact placement within that group remained uncertain.
6842:
5280:
5021:
4963:
1785:
1687:
1101:
1093:
959:
882:
864:
860:
780:
661:
565:
557:
502:
422:
6795:
6696:
5323:
762:. Earlier that year, Brownstein (2017) analyzed the material of
710:
and probably belong to that genus instead. However, some of the
524:
6756:
6194:
5440:
4634:
3949:
3129:
2705:
2590:
2552:
1936:(Abridged ed.). United Kingdom: Oxford University Press.
1904:
Proceedings of the Academy of Natural Sciences of Philadelphia
1866:
Proceedings of the Academy of Natural Sciences of Philadelphia
2195:
2193:
2191:
2189:
2548:
395:
2268:
10.1671/0272-4634(2005)025[0119:ANGASO]2.0.CO;2
2080:(second edition). University of California Press, Berkeley.
1055:, which is known from more complete remains, is similar to
982:. Based on this line of reasoning, Cope classified it as a
527:
uncovered an incomplete theropod skeleton in a quarry near
439:
approximately 67.6 million years ago during the end of the
398:
386:
1801:
The 1897 watercolor painting by Charles R. Knight titled
978:, which was known at the time from remains discovered in
377:
990:
family, Dryptosauridae. The fossil material assigned to
754:. In 2017, it was informally given the new generic name
1774:, and they may have been a prominent part of its diet.
724:
was also coined by Cope for a partial leg found in the
1087:, but more primitive than members of the more derived
2144:
Brusatte, S.L.; Benson, R.B.J.; Norell, M.A. (2011).
410:
401:
392:
368:
812:
with those of derived tyrannosaurids and noted that
383:
380:
374:
371:
6765:
6678:
6573:
6516:
6478:
6395:
6368:
6278:
6248:
6207:
6105:
6073:
6015:
5981:
5954:
5927:
5896:
5869:
5858:
5818:
5683:
5631:
5602:
5564:
5553:
5453:
5365:
5299:
5221:
5152:
5128:
5040:
4988:
4928:
4884:
4873:
4817:
4785:
4732:
4682:
4647:
4565:
4518:
4465:
4427:
4373:
4324:
4260:
4219:
4195:
4164:
4118:
4088:
4077:
3962:
3877:
3776:
3687:
3666:
3602:
3505:
3449:
3415:
3322:
3267:
3236:
3176:
3142:
3068:
3048:
2972:
2919:
2800:
2718:
2669:
2642:
556:under the catalogue number ANSP 9995 and includes;
389:
1929:
1071:(2013) confirmed the tyrannosauroid affinities of
125:Reconstructed skeletons mounted in fighting pose,
2327:20.500.11820/31f38145-54e7-48f8-819a-262601e93f2b
750:in his contribution to the second edition of the
2139:
2137:
2135:
2133:
2131:
2129:
2039:. In Tanke, Darren; Carpenter, Kenneth (eds.).
1676:, which now includes both "Cryptotyrannus" and
1672:. Brownstein (2021) also reinstated the family
523:. During the summer of 1866, workers from the
2564:
1067:. Detailed phylogenetic analysis by Brusatte
820:may not have proceeded in a uniform fashion.
706:holotype, share many features in common with
8:
2006:: CS1 maint: multiple names: authors list (
1734:period, approximately 67 million years ago.
6876:Late Cretaceous dinosaurs of North America
6753:
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6256:
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6191:
6102:
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5691:
5576:
5572:
5561:
5461:
5450:
5437:
5228:
5218:
5134:
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4881:
4791:
4744:
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4690:
4655:
4644:
4631:
4562:
4471:
4379:
4332:
4201:
4176:
4172:
4100:
4096:
4085:
3970:
3959:
3946:
3874:
3693:
3684:
3513:
3328:
3279:
3275:
3188:
3184:
3150:
3139:
3126:
2925:
2808:
2726:
2715:
2702:
2648:
2639:
2587:
2571:
2557:
2549:
1766:are known from the same time and place as
116:
20:
2466:
2401:
2391:
2325:
2315:
2169:
1664:", an informally named theropod from the
839:, lateral teeth, 11 middle-distal caudal
732:had referred earlier to the ornithomimid
1959:
1957:
1955:
1953:
1003:studies during the 1990s suggested that
999:Russell (1970) and Molnar (1990). Other
2298:Brusatte, S.L.; Benson, R.B.J. (2013).
1857:
2117:The Princeton Field Guide to Dinosaurs
2033:"On the type and referred material of
1999:
435:that lived on the island continent of
2229:Smithsonian Miscellaneous Collections
2098:
2096:
1750:referred material has been reported.
895:with three fingers and lipless snout.
7:
6848:A52387D3-0775-FFC1-FF97-FAB1B85ED2A6
1692:Diagram showing known remains in of
666:An outdated historical depiction of
543:. In August of 1866, paleontologist
6896:Taxa named by Othniel Charles Marsh
1511:
1504:
1480:
1430:
1423:
1399:
1375:
1351:
1327:
1303:
1235:
1211:
1144:
1137:
1127:
1117:
716:Morrison Formation theropod species
2519:Journal of Vertebrate Paleontology
2256:Journal of Vertebrate Paleontology
2037:Cope 1877 (Dinosauria: Theropoda)"
1970:Journal of Vertebrate Paleontology
1715:in Barnsboro, Mantua Township, in
472:of its time, in spite of its poor
14:
968:Since the time of its discovery,
785:Estimated size, compared to human
659:), only to be later reclassified.
6737:
6731:
6724:
6176:
6169:
6162:
6156:
5422:
5415:
5410:
5403:
5397:
4616:
4609:
4602:
3931:
3924:
3917:
3910:
3112:
3106:
2425:Doran Brownstein, Chase (2021).
1846:Timeline of tyrannosaur research
1830:
1489:Appalachiosaurus montgomeriensis
1026:Late Cretaceous tyrannosaurids,
645:Natural History Museum in London
535:with sediments belonging to the
364:
138:
2106:(No. e3105v1). PeerJ Preprints.
1990:10.1080/02724634.1997.10011003
746:listed it as an indeterminate
1:
6891:Fossil taxa described in 1877
6886:Fossil taxa described in 1866
2304:Acta Palaeontologica Polonica
875:, and a midshaft fragment of
507:Elements of the type specimen
480:in 1866 and later renamed by
468:one of the more widely known
2393:10.1371/journal.pone.0079420
1660:to be the sister taxon to "
855:bones, a fragmentary right
584:, and nearly complete left
550:Academy of Natural Sciences
476:record. First described by
6927:
6901:Paleontology in New Jersey
2431:Royal Society Open Science
2121:Princeton University Press
2102:Brownstein, C. D. (2017).
2090:doi:10.24896/jzbr.2017422.
1770:, the island continent of
843:, both the left and right
16:Extinct genus of dinosaurs
6721:
6487:
6404:
6379:
6263:
6259:
6218:
6203:
6190:
6154:
6101:
6069:
5905:
5829:
5698:
5694:
5579:
5575:
5464:
5449:
5436:
5395:
5231:
5217:
5137:
5049:
4937:
4794:
4747:
4743:
4693:
4658:
4643:
4630:
4599:
4561:
4474:
4382:
4335:
4204:
4179:
4175:
4103:
4099:
3973:
3958:
3945:
3907:
3873:
3696:
3516:
3331:
3282:
3278:
3191:
3187:
3153:
3138:
3125:
3103:
2928:
2811:
2729:
2714:
2701:
2651:
2638:
2586:
2154:American Museum Novitates
2031:Chure, Daniel J. (2001).
1559:
1533:
1516:
1509:
1502:
1485:
1478:
1452:
1435:
1428:
1421:
1404:
1397:
1380:
1373:
1356:
1349:
1332:
1325:
1308:
1301:
1257:
1240:
1233:
1216:
1209:
1183:
1166:
1149:
1142:
1135:
1125:
738:, distinguishing it from
322:
315:
293:
286:
135:Scientific classification
133:
124:
115:
23:
2543:Dryptosaurus aquilunguis
2148:Dryptosaurus aquilunguis
2041:Mesozoic Vertebrate Life
1883:www.theropoddatabase.com
1656:Brownstein (2021) found
1456:Xiongguanlong baimoensis
1408:Dryptosaurus aquilunguis
1261:Stokesosaurus clevelandi
1153:Proceratosaurus bradleyi
625:a dog in Greek mythology
525:West Jersey Marl Company
300:Dryptosaurus aquilunguis
2492:Journal of Paleontology
1932:A Greek-English Lexicon
1746:, from which potential
1666:Merchantville Formation
1220:Sinotyrannus kazuoensis
631:"slashing reptile", by
495:ever known to science.
127:New Jersey State Museum
1808:Henry Fairfield Osborn
1798:
1697:
1107:
1099:
965:
934:According to Brusatte
896:
835:, a fragmentary right
831:, a fragmentary right
786:
687:
682:in the background. By
635:, Cope's rival in the
508:
6906:Cretaceous New Jersey
6830:Paleobiology Database
5130:Carcharodontosaurinae
5042:Carcharodontosauridae
2317:10.4202/app.2011.0141
1922:Liddell, Henry George
1789:
1782:Cultural significance
1713:Hornerstown Formation
1700:The type specimen of
1691:
1105:
1097:
963:
886:
784:
670:(center) confronting
665:
506:
499:Discovery and species
482:Othniel Charles Marsh
6881:Maastrichtian genera
6106:Dubious coelurosaurs
4819:Metriacanthosaurinae
4787:Metriacanthosauridae
3069:Dubious neotheropods
1439:Alectrosaurus olseni
1384:Raptorex kriegsteini
1170:Kileskus aristotocus
1106:Limb and other bones
1098:Postcranial elements
980:southeastern England
742:by its longer toes.
639:, in 1877. Brusatte
513:Ferdinand Van Hayden
6911:Symbols of Delaware
5143:Carcharodontosaurus
4930:Carcharodontosauria
2451:10.1098/rsos.210127
2443:2021RSOS....810127D
2384:2013PLoSO...879420L
2115:Paul, G.S. (2010).
2035:Laelaps trihedrodon
1982:1997JVPal..17..561C
1739:New Egypt Formation
1244:Juratyrant langhami
696:Laelaps trihedrodon
691:Misassigned species
684:Edward Drinker Cope
617:Laelaps aquilunguis
545:Edward Drinker Cope
541:New Egypt Formation
488:is among the first
478:Edward Drinker Cope
327:Laelaps aquilunguis
279:D. aquilunguis
6598:Archaeornithomimus
5119:Veterupristisaurus
4979:Veterupristisaurus
4835:Metriacanthosaurus
4775:Piatnitzkysauridae
4210:Eustreptospondylus
4120:Piatnitzkysauridae
4003:Chuandongocoelurus
2867:Notatesseraeraptor
1879:"Tyrannosauroidea"
1799:
1744:Navesink Formation
1698:
1360:Bagaraatan ostromi
1108:
1100:
1016:Charles W. Gilmore
966:
964:Holotype claw bone
917:arctometatarsalian
897:
787:
726:Navesink Formation
700:Morrison Formation
688:
509:
455:, ground-dwelling
6858:
6857:
6817:Open Tree of Life
6759:Taxon identifiers
6750:
6749:
6746:
6745:
6719:
6718:
6715:
6714:
6711:
6710:
6707:
6706:
6674:
6673:
6670:
6669:
6480:Macrocheiriformes
6397:Ornithomimosauria
6371:Maniraptoriformes
6364:
6363:
6210:Maniraptoromorpha
6197:Maniraptoromorpha
6186:
6185:
6152:
6151:
6148:
6147:
6144:
6143:
6097:
6096:
6093:
6092:
6075:Maniraptoromorpha
6065:
6064:
6061:
6060:
6057:
6056:
6053:
6052:
6049:
6048:
5814:
5813:
5679:
5678:
5633:Proceratosauridae
5585:Chingkankousaurus
5432:
5431:
5393:
5392:
5389:
5388:
5385:
5384:
5361:
5360:
5213:
5212:
5209:
5208:
5205:
5204:
5201:
5200:
5197:
5196:
5193:
5192:
4869:
4868:
4865:
4864:
4626:
4625:
4597:
4596:
4593:
4592:
4589:
4588:
4585:
4584:
4557:
4556:
4553:
4552:
4549:
4548:
4545:
4544:
4461:
4460:
4417:Vallibonavenatrix
4320:
4319:
4316:
4315:
4160:
4159:
3941:
3940:
3905:
3904:
3901:
3900:
3897:
3896:
3869:
3868:
3865:
3864:
3861:
3860:
3857:
3856:
3853:
3852:
3501:
3500:
3497:
3496:
3263:
3262:
3121:
3120:
3101:
3100:
3097:
3096:
3093:
3092:
3044:
3043:
2697:
2696:
2693:
2692:
2689:
2688:
2054:978-0-253-33907-2
1795:Charles R. Knight
1717:Gloucester County
1653:
1652:
1644:
1643:
1635:
1634:
1626:
1625:
1617:
1616:
1608:
1607:
1599:
1598:
1590:
1589:
1581:
1580:
1572:
1571:
1548:
1547:
1537:Alioramus remotus
1467:
1466:
1290:
1289:
1281:
1280:
1272:
1271:
1198:
1197:
1132:Proceratosauridae
1063:was also a basal
772:Laelaps macropus.
517:Geological Survey
462:Charles R. Knight
355:
354:
349:
336:
267:
248:
6918:
6851:
6850:
6838:
6837:
6825:
6824:
6812:
6811:
6799:
6798:
6786:
6785:
6784:
6754:
6741:
6735:
6728:
6584:Aepyornithomimus
6485:
6402:
6377:
6341:Sinocalliopteryx
6261:
6257:
6251:Neocoelurosauria
6216:
6205:
6192:
6180:
6173:
6166:
6160:
6103:
6071:
6040:Zhuchengtyrannus
5972:Thanatotheristes
5903:
5867:
5835:Appalachiosaurus
5827:
5696:
5692:
5686:Pantyrannosauria
5577:
5573:
5567:Tyrannosauroidea
5562:
5537:Xinjiangovenator
5462:
5451:
5438:
5426:
5419:
5414:
5407:
5401:
5229:
5219:
5135:
5055:Acrocanthosaurus
5047:
4935:
4882:
4808:Yangchuanosaurus
4792:
4745:
4741:
4691:
4672:Lourinhanosaurus
4656:
4645:
4632:
4620:
4613:
4606:
4563:
4529:Sigilmassasaurus
4472:
4429:Ceratosuchopsini
4380:
4333:
4278:Dubreuillosaurus
4202:
4185:Streptospondylus
4177:
4173:
4143:Piatnitzkysaurus
4101:
4097:
4086:
3971:
3960:
3947:
3935:
3928:
3921:
3914:
3875:
3710:Ekrixinatosaurus
3694:
3685:
3514:
3329:
3280:
3276:
3189:
3185:
3151:
3140:
3127:
3116:
3110:
3020:Procompsognathus
2926:
2809:
2727:
2716:
2703:
2649:
2640:
2633:
2632:
2588:
2573:
2566:
2559:
2550:
2526:
2509:
2506:
2500:
2499:
2487:
2481:
2480:
2470:
2422:
2416:
2415:
2405:
2395:
2338:
2332:
2331:
2329:
2319:
2295:
2289:
2286:
2280:
2279:
2251:
2245:
2238:
2232:
2225:
2219:
2216:
2210:
2207:
2201:
2197:
2184:
2183:
2173:
2146:"The Anatomy of
2141:
2124:
2113:
2107:
2100:
2091:
2087:
2081:
2074:
2068:
2065:
2059:
2058:
2028:
2022:
2018:
2012:
2011:
2005:
1997:
1992:. Archived from
1961:
1948:
1947:
1935:
1918:
1912:
1911:
1899:
1893:
1892:
1890:
1889:
1875:
1869:
1862:
1840:
1838:Dinosaurs portal
1835:
1834:
1833:
1762:very difficult.
1512:
1505:
1481:
1431:
1424:
1400:
1376:
1352:
1336:Eotyrannus lengi
1328:
1312:Dilong paradoxus
1304:
1236:
1212:
1145:
1138:
1128:
1122:Tyrannosauroidea
1118:
1053:Appalachiosaurus
1048:Appalachiosaurus
994:was reviewed by
889:life restoration
764:Laelaps macropus
722:Laelaps macropus
649:Tyrannosauroidea
588:. However, four
576:, an incomplete
570:caudal vertebrae
519:excursions into
427:eutyrannosaurian
417:
413:
408:
407:
404:
403:
400:
397:
394:
391:
388:
385:
382:
379:
376:
373:
370:
347:
330:
302:
298:
265:
258:
243:
238:
225:
143:
142:
120:
110:
40:
29:Temporal range:
21:
6926:
6925:
6921:
6920:
6919:
6917:
6916:
6915:
6861:
6860:
6859:
6854:
6846:
6841:
6833:
6828:
6820:
6815:
6807:
6802:
6794:
6789:
6780:
6779:
6774:
6761:
6751:
6742:
6729:
6703:
6666:
6647:Sinornithomimus
6569:
6512:
6474:
6391:
6385:Compsognathidae
6373:
6360:
6348:Sinosauropteryx
6280:Compsognathidae
6274:
6253:
6244:
6212:
6199:
6182:
6181:
6167:
6140:
6135:Shanyangosaurus
6128:Phaedrolosaurus
6089:
6045:
6011:
5977:
5956:Daspletosaurini
5950:
5923:
5898:Tyrannosaurinae
5892:
5871:Albertosaurinae
5863:
5861:Tyrannosauridae
5854:
5823:
5821:Eutyrannosauria
5810:
5688:
5675:
5656:Proceratosaurus
5627:
5598:
5569:
5558:
5549:
5515:Richardoestesia
5499:Chilantaisaurus
5458:
5445:
5428:
5427:
5408:
5381:
5357:
5317:Australovenator
5295:
5289:Chilantaisaurus
5267:Phuwiangvenator
5189:
5154:Giganotosaurini
5148:
5124:
5036:
4999:Chilantaisaurus
4984:
4924:
4878:
4861:
4813:
4781:
4737:
4728:
4706:Monolophosaurus
4687:
4678:
4652:
4639:
4622:
4621:
4607:
4581:
4541:
4514:
4480:Camarillasaurus
4457:
4423:
4369:
4363:Ostafrikasaurus
4341:Camarillasaurus
4329:
4312:
4262:Afrovenatorinae
4256:
4215:
4191:
4169:
4156:
4114:
4093:
4091:Megalosauroidea
4082:
4073:
4068:Vectaerovenator
4061:Szechuanosaurus
4039:Monolophosaurus
4025:Kaijiangosaurus
4018:Cryolophosaurus
3979:Calamospondylus
3967:
3954:
3937:
3936:
3929:
3922:
3915:
3893:
3849:
3830:Pycnonemosaurus
3772:
3683:
3677:Majungasaurinae
3662:
3604:Majungasaurinae
3598:
3593:Xenotarsosaurus
3529:Dryptosauroides
3510:
3493:
3445:
3417:Elaphrosaurinae
3411:
3318:
3304:Ornithomimoides
3272:
3259:
3232:
3181:
3172:
3147:
3134:
3117:
3111:
3089:
3064:
3040:
2968:
2915:
2881:Shuangbaisaurus
2805:
2796:
2723:
2710:
2685:
2665:
2659:Avemetatarsalia
2644:Avemetatarsalia
2634:
2619:Avemetatarsalia
2593:
2592:
2582:
2577:
2538:
2516:
2513:
2512:
2507:
2503:
2489:
2488:
2484:
2424:
2423:
2419:
2350:Sertich, J.J.W.
2340:
2339:
2335:
2297:
2296:
2292:
2287:
2283:
2253:
2252:
2248:
2239:
2235:
2226:
2222:
2217:
2213:
2208:
2204:
2198:
2187:
2143:
2142:
2127:
2114:
2110:
2101:
2094:
2088:
2084:
2075:
2071:
2066:
2062:
2055:
2030:
2029:
2025:
2019:
2015:
1998:
1963:
1962:
1951:
1944:
1920:
1919:
1915:
1901:
1900:
1896:
1887:
1885:
1877:
1876:
1872:
1863:
1859:
1854:
1836:
1831:
1829:
1826:
1803:Leaping Laelaps
1791:Leaping Laelaps
1784:
1732:Late Cretaceous
1686:
1654:
1645:
1636:
1627:
1618:
1609:
1600:
1591:
1582:
1573:
1562:Tyrannosauridae
1549:
1520:Alioramus altai
1468:
1291:
1282:
1273:
1199:
1187:Guanlong wucaii
1089:Tyrannosauridae
958:
847:, three manual
818:tyrannosauroids
779:
768:ornithomimosaur
744:Thomas R. Holtz
693:
606:Pneumatoarthrus
501:
445:Late Cretaceous
415:
411:
367:
363:
311:
304:
296:
295:
282:
264:
256:
242:
236:
227:Eutyrannosauria
223:
137:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
6924:
6922:
6914:
6913:
6908:
6903:
6898:
6893:
6888:
6883:
6878:
6873:
6863:
6862:
6856:
6855:
6853:
6852:
6839:
6826:
6813:
6800:
6787:
6771:
6769:
6763:
6762:
6757:
6748:
6747:
6744:
6743:
6723:
6722:
6720:
6717:
6716:
6713:
6712:
6709:
6708:
6705:
6704:
6702:
6701:
6700:
6699:
6693:
6684:
6682:
6676:
6675:
6672:
6671:
6668:
6667:
6665:
6664:
6657:
6650:
6643:
6636:
6629:
6622:
6615:
6612:Dromiceiomimus
6608:
6601:
6594:
6587:
6579:
6577:
6575:Ornithomimidae
6571:
6570:
6568:
6567:
6560:
6556:Paraxenisaurus
6552:
6544:
6537:
6530:
6522:
6520:
6518:Deinocheiridae
6514:
6513:
6511:
6510:
6507:Shenzhousaurus
6503:
6496:
6488:
6482:
6476:
6475:
6473:
6472:
6465:
6457:
6450:
6443:
6436:
6429:
6421:
6413:
6405:
6399:
6393:
6392:
6390:
6389:
6380:
6374:
6369:
6366:
6365:
6362:
6361:
6359:
6358:
6351:
6344:
6337:
6329:
6322:
6315:
6308:
6304:Beipiaognathus
6300:
6293:
6284:
6282:
6276:
6275:
6273:
6272:
6264:
6254:
6249:
6246:
6245:
6243:
6242:
6235:
6228:
6219:
6213:
6208:
6201:
6200:
6195:
6188:
6187:
6184:
6183:
6175:
6161:
6155:
6153:
6150:
6149:
6146:
6145:
6142:
6141:
6139:
6138:
6131:
6124:
6117:
6109:
6107:
6099:
6098:
6095:
6094:
6091:
6090:
6088:
6087:
6086:
6085:
6079:
6077:
6067:
6066:
6063:
6062:
6059:
6058:
6055:
6054:
6051:
6050:
6047:
6046:
6044:
6043:
6036:
6029:
6021:
6019:
6017:Tyrannosaurini
6013:
6012:
6010:
6009:
6002:
5995:
5987:
5985:
5983:Teratophoneini
5979:
5978:
5976:
5975:
5968:
5965:Daspletosaurus
5960:
5958:
5952:
5951:
5949:
5948:
5945:Qianzhousaurus
5941:
5933:
5931:
5925:
5924:
5922:
5921:
5914:
5906:
5900:
5894:
5893:
5891:
5890:
5883:
5875:
5873:
5864:
5859:
5856:
5855:
5853:
5852:
5845:
5842:Bistahieversor
5838:
5830:
5824:
5819:
5816:
5815:
5812:
5811:
5809:
5808:
5800:
5793:
5786:
5778:
5771:
5764:
5757:
5750:
5743:
5736:
5729:
5722:
5714:
5707:
5699:
5689:
5684:
5681:
5680:
5677:
5676:
5674:
5673:
5666:
5659:
5652:
5645:
5637:
5635:
5629:
5628:
5626:
5625:
5617:
5609:
5607:
5600:
5599:
5597:
5596:
5588:
5580:
5570:
5565:
5559:
5556:Tyrannoraptora
5554:
5551:
5550:
5548:
5547:
5540:
5533:
5526:
5518:
5511:
5503:
5495:
5488:
5480:
5473:
5465:
5459:
5454:
5447:
5446:
5441:
5434:
5433:
5430:
5429:
5421:
5402:
5396:
5394:
5391:
5390:
5387:
5386:
5383:
5382:
5380:
5379:
5378:
5377:
5371:
5369:
5363:
5362:
5359:
5358:
5356:
5355:
5348:
5341:
5334:
5327:
5320:
5313:
5305:
5303:
5301:Megaraptoridae
5297:
5296:
5294:
5293:
5285:
5277:
5270:
5263:
5256:
5248:
5240:
5232:
5226:
5215:
5214:
5211:
5210:
5207:
5206:
5203:
5202:
5199:
5198:
5195:
5194:
5191:
5190:
5188:
5187:
5180:
5173:
5166:
5163:Giganotosaurus
5158:
5156:
5150:
5149:
5147:
5146:
5138:
5132:
5126:
5125:
5123:
5122:
5115:
5108:
5101:
5094:
5087:
5080:
5073:
5066:
5058:
5050:
5044:
5038:
5037:
5035:
5034:
5026:
5018:
5011:
5003:
4994:
4992:
4990:Neovenatoridae
4986:
4985:
4983:
4982:
4975:
4972:Ulughbegsaurus
4968:
4960:
4953:
4946:
4938:
4932:
4926:
4925:
4923:
4922:
4914:
4906:
4898:
4890:
4888:
4879:
4874:
4871:
4870:
4867:
4866:
4863:
4862:
4860:
4859:
4852:
4845:
4838:
4831:
4823:
4821:
4815:
4814:
4812:
4811:
4804:
4800:Xuanhanosaurus
4795:
4789:
4783:
4782:
4780:
4779:
4771:
4767:Xuanhanosaurus
4763:
4756:
4753:Asfaltovenator
4748:
4738:
4733:
4730:
4729:
4727:
4726:
4718:
4714:Megalosauridae
4710:
4702:
4694:
4688:
4683:
4680:
4679:
4677:
4676:
4668:
4659:
4653:
4648:
4641:
4640:
4635:
4628:
4627:
4624:
4623:
4615:
4601:
4600:
4598:
4595:
4594:
4591:
4590:
4587:
4586:
4583:
4582:
4580:
4579:
4578:
4577:
4571:
4569:
4559:
4558:
4555:
4554:
4551:
4550:
4547:
4546:
4543:
4542:
4540:
4539:
4532:
4524:
4522:
4516:
4515:
4513:
4512:
4505:
4498:
4491:
4488:Ichthyovenator
4484:
4475:
4469:
4463:
4462:
4459:
4458:
4456:
4455:
4448:
4441:
4433:
4431:
4425:
4424:
4422:
4421:
4413:
4405:
4398:
4391:
4383:
4377:
4371:
4370:
4368:
4367:
4359:
4352:
4349:Cristatusaurus
4345:
4336:
4330:
4325:
4322:
4321:
4318:
4317:
4314:
4313:
4311:
4310:
4306:Poekilopleuron
4302:
4299:Piveteausaurus
4295:
4288:
4281:
4274:
4266:
4264:
4258:
4257:
4255:
4254:
4247:
4240:
4233:
4225:
4223:
4221:Megalosaurinae
4217:
4216:
4214:
4213:
4205:
4199:
4197:Megalosauridae
4193:
4192:
4190:
4189:
4180:
4170:
4165:
4162:
4161:
4158:
4157:
4155:
4154:
4150:Xuanhanosaurus
4146:
4139:
4132:
4124:
4122:
4116:
4115:
4113:
4112:
4109:Yunyangosaurus
4104:
4094:
4089:
4083:
4078:
4075:
4074:
4072:
4071:
4064:
4057:
4049:
4046:Pandoravenator
4042:
4035:
4032:Kayentavenator
4028:
4021:
4014:
4006:
3999:
3991:
3983:
3974:
3968:
3963:
3956:
3955:
3950:
3943:
3942:
3939:
3938:
3930:
3923:
3916:
3909:
3908:
3906:
3903:
3902:
3899:
3898:
3895:
3894:
3892:
3891:
3890:
3889:
3883:
3881:
3871:
3870:
3867:
3866:
3863:
3862:
3859:
3858:
3855:
3854:
3851:
3850:
3848:
3847:
3840:
3833:
3826:
3819:
3811:
3804:
3797:
3790:
3782:
3780:
3774:
3773:
3771:
3770:
3763:
3760:Skorpiovenator
3756:
3748:
3741:
3734:
3727:
3720:
3713:
3706:
3697:
3691:
3682:
3681:
3672:
3670:
3664:
3663:
3661:
3660:
3653:
3645:
3638:
3631:
3624:
3616:
3608:
3606:
3600:
3599:
3597:
3596:
3589:
3582:
3575:
3572:Spectrovenator
3568:
3561:
3554:
3547:
3540:
3533:
3525:
3517:
3511:
3506:
3503:
3502:
3499:
3498:
3495:
3494:
3492:
3491:
3484:
3477:
3470:
3463:
3455:
3453:
3447:
3446:
3444:
3443:
3436:
3429:
3421:
3419:
3413:
3412:
3410:
3409:
3406:Spinostropheus
3402:
3395:
3388:
3380:
3372:
3364:
3356:
3349:
3341:
3332:
3326:
3320:
3319:
3317:
3316:
3308:
3300:
3292:
3283:
3273:
3270:Abelisauroidea
3268:
3265:
3264:
3261:
3260:
3258:
3257:
3250:
3242:
3240:
3238:Ceratosauridae
3234:
3233:
3231:
3230:
3227:Saltriovenator
3223:
3215:
3208:
3204:Fosterovenator
3200:
3192:
3182:
3177:
3174:
3173:
3171:
3170:
3166:Lophostropheus
3162:
3154:
3148:
3143:
3136:
3135:
3130:
3123:
3122:
3119:
3118:
3105:
3104:
3102:
3099:
3098:
3095:
3094:
3091:
3090:
3088:
3087:
3080:
3072:
3070:
3066:
3065:
3063:
3062:
3061:
3060:
3054:
3052:
3046:
3045:
3042:
3041:
3039:
3038:
3031:
3027:Pterospondylus
3023:
3016:
3008:
3000:
2997:Lucianovenator
2993:
2986:
2978:
2976:
2970:
2969:
2967:
2966:
2959:
2952:
2945:
2938:
2929:
2923:
2921:Coelophysoidea
2917:
2916:
2914:
2913:
2906:
2898:
2891:
2884:
2877:
2870:
2863:
2860:Lophostropheus
2856:
2849:
2842:
2835:
2828:
2820:
2812:
2806:
2801:
2798:
2797:
2795:
2794:
2786:
2778:
2770:
2762:
2759:Erythrovenator
2755:
2747:
2739:
2730:
2724:
2719:
2712:
2711:
2706:
2699:
2698:
2695:
2694:
2691:
2690:
2687:
2686:
2684:
2683:
2682:
2681:
2675:
2673:
2667:
2666:
2664:
2663:
2662:
2661:
2652:
2646:
2636:
2635:
2631:
2630:
2621:
2612:
2606:
2600:
2591:
2584:
2583:
2578:
2576:
2575:
2568:
2561:
2553:
2547:
2546:
2537:
2536:External links
2534:
2533:
2532:
2527:
2511:
2510:
2501:
2482:
2417:
2378:(11): e79420.
2362:Evans, David C
2358:Sampson, S. D.
2333:
2290:
2281:
2262:(1): 119–143.
2246:
2233:
2220:
2211:
2202:
2185:
2162:10.1206/3717.2
2156:(3717): 1–53.
2125:
2108:
2092:
2082:
2078:The Dinosauria
2069:
2060:
2053:
2023:
2013:
1996:on 2007-03-10.
1976:(3): 561–573.
1949:
1942:
1913:
1894:
1870:
1856:
1855:
1853:
1850:
1849:
1848:
1842:
1841:
1825:
1822:
1783:
1780:
1685:
1682:
1674:Dryptosauridae
1662:Cryptotyrannus
1651:
1650:
1647:
1646:
1642:
1641:
1638:
1637:
1633:
1632:
1629:
1628:
1624:
1623:
1620:
1619:
1615:
1614:
1611:
1610:
1606:
1605:
1602:
1601:
1597:
1596:
1593:
1592:
1588:
1587:
1584:
1583:
1579:
1578:
1575:
1574:
1570:
1569:
1566:
1565:
1558:
1555:
1554:
1551:
1550:
1546:
1545:
1542:
1541:
1532:
1529:
1528:
1525:
1524:
1515:
1510:
1508:
1503:
1501:
1498:
1497:
1494:
1493:
1484:
1479:
1477:
1474:
1473:
1470:
1469:
1465:
1464:
1461:
1460:
1451:
1448:
1447:
1444:
1443:
1434:
1429:
1427:
1422:
1420:
1417:
1416:
1413:
1412:
1403:
1398:
1396:
1393:
1392:
1389:
1388:
1379:
1374:
1372:
1369:
1368:
1365:
1364:
1355:
1350:
1348:
1345:
1344:
1341:
1340:
1331:
1326:
1324:
1321:
1320:
1317:
1316:
1307:
1302:
1300:
1297:
1296:
1293:
1292:
1288:
1287:
1284:
1283:
1279:
1278:
1275:
1274:
1270:
1269:
1266:
1265:
1256:
1253:
1252:
1249:
1248:
1239:
1234:
1232:
1229:
1228:
1225:
1224:
1215:
1210:
1208:
1205:
1204:
1201:
1200:
1196:
1195:
1192:
1191:
1182:
1179:
1178:
1175:
1174:
1165:
1162:
1161:
1158:
1157:
1148:
1143:
1141:
1136:
1134:
1126:
1124:
1116:
1065:tyrannosauroid
1044:tyrannosauroid
957:
956:Classification
954:
778:
775:
748:tyrannosauroid
704:L. trihedrodon
692:
689:
500:
497:
353:
352:
351:
350:
337:
320:
319:
313:
312:
305:
291:
290:
284:
283:
275:
273:
269:
268:
254:
250:
249:
240:Dryptosauridae
234:
230:
229:
221:
214:
213:
208:
201:
200:
195:
188:
187:
182:
175:
174:
169:
165:
164:
159:
155:
154:
149:
145:
144:
131:
130:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
6923:
6912:
6909:
6907:
6904:
6902:
6899:
6897:
6894:
6892:
6889:
6887:
6884:
6882:
6879:
6877:
6874:
6872:
6869:
6868:
6866:
6849:
6844:
6840:
6836:
6831:
6827:
6823:
6818:
6814:
6810:
6805:
6801:
6797:
6792:
6788:
6783:
6777:
6773:
6772:
6770:
6768:
6764:
6760:
6755:
6740:
6736:
6734:
6727:
6698:
6694:
6692:
6688:
6687:
6686:
6685:
6683:
6681:
6677:
6663:
6662:
6658:
6656:
6655:
6654:Struthiomimus
6651:
6649:
6648:
6644:
6642:
6641:
6637:
6635:
6634:
6630:
6628:
6627:
6623:
6621:
6620:
6616:
6614:
6613:
6609:
6607:
6606:
6605:"Coelosaurus"
6602:
6600:
6599:
6595:
6593:
6592:
6588:
6586:
6585:
6581:
6580:
6578:
6576:
6572:
6566:
6565:
6561:
6558:
6557:
6553:
6550:
6549:
6545:
6543:
6542:
6538:
6536:
6535:
6531:
6529:
6528:
6524:
6523:
6521:
6519:
6515:
6509:
6508:
6504:
6502:
6501:
6500:Pelecanimimus
6497:
6495:
6494:
6490:
6489:
6486:
6483:
6481:
6477:
6471:
6470:
6466:
6463:
6462:
6461:Thecocoelurus
6458:
6456:
6455:
6451:
6449:
6448:
6444:
6442:
6441:
6440:Kinnareemimus
6437:
6435:
6434:
6430:
6427:
6426:
6422:
6419:
6418:
6414:
6412:
6411:
6407:
6406:
6403:
6400:
6398:
6394:
6387:
6386:
6382:
6381:
6378:
6375:
6372:
6367:
6357:
6356:
6352:
6350:
6349:
6345:
6343:
6342:
6338:
6335:
6334:
6330:
6328:
6327:
6323:
6321:
6320:
6319:Huaxiagnathus
6316:
6314:
6313:
6312:Compsognathus
6309:
6306:
6305:
6301:
6299:
6298:
6294:
6291:
6290:
6286:
6285:
6283:
6281:
6277:
6271:
6270:
6266:
6265:
6262:
6258:
6255:
6252:
6247:
6241:
6240:
6239:Ornitholestes
6236:
6234:
6233:
6229:
6226:
6225:
6221:
6220:
6217:
6214:
6211:
6206:
6202:
6198:
6193:
6189:
6179:
6174:
6172:
6165:
6159:
6137:
6136:
6132:
6130:
6129:
6125:
6123:
6122:
6118:
6116:
6115:
6111:
6110:
6108:
6104:
6100:
6083:
6082:
6081:
6080:
6078:
6076:
6072:
6068:
6042:
6041:
6037:
6035:
6034:
6033:Tyrannosaurus
6030:
6028:
6027:
6023:
6022:
6020:
6018:
6014:
6008:
6007:
6006:Teratophoneus
6003:
6001:
6000:
5996:
5994:
5993:
5989:
5988:
5986:
5984:
5980:
5974:
5973:
5969:
5967:
5966:
5962:
5961:
5959:
5957:
5953:
5947:
5946:
5942:
5940:
5939:
5935:
5934:
5932:
5930:
5926:
5920:
5919:
5915:
5913:
5912:
5908:
5907:
5904:
5901:
5899:
5895:
5889:
5888:
5884:
5882:
5881:
5880:Albertosaurus
5877:
5876:
5874:
5872:
5868:
5865:
5862:
5857:
5851:
5850:
5846:
5844:
5843:
5839:
5837:
5836:
5832:
5831:
5828:
5825:
5822:
5817:
5806:
5805:
5801:
5799:
5798:
5797:Xiongguanlong
5794:
5792:
5791:
5787:
5784:
5783:
5779:
5777:
5776:
5775:Suskityrannus
5772:
5770:
5769:
5768:Stokesosaurus
5765:
5763:
5762:
5761:Santanaraptor
5758:
5756:
5755:
5751:
5749:
5748:
5744:
5742:
5741:
5737:
5735:
5734:
5730:
5728:
5727:
5723:
5720:
5719:
5715:
5713:
5712:
5708:
5706:
5705:
5704:Alectrosaurus
5701:
5700:
5697:
5693:
5690:
5687:
5682:
5672:
5671:
5667:
5665:
5664:
5660:
5658:
5657:
5653:
5651:
5650:
5646:
5644:
5643:
5639:
5638:
5636:
5634:
5630:
5623:
5622:
5621:Tanycolagreus
5618:
5616:
5615:
5611:
5610:
5608:
5605:
5601:
5594:
5593:
5589:
5587:
5586:
5582:
5581:
5578:
5574:
5571:
5568:
5563:
5560:
5557:
5552:
5546:
5545:
5541:
5539:
5538:
5534:
5532:
5531:
5527:
5524:
5523:
5519:
5517:
5516:
5512:
5509:
5508:
5504:
5501:
5500:
5496:
5494:
5493:
5489:
5486:
5485:
5484:Bahariasaurus
5481:
5479:
5478:
5474:
5472:
5471:
5467:
5466:
5463:
5460:
5457:
5456:Coelurosauria
5452:
5448:
5444:
5443:Coelurosauria
5439:
5435:
5425:
5420:
5418:
5413:
5406:
5400:
5375:
5374:
5373:
5372:
5370:
5368:
5367:Coelurosauria
5364:
5354:
5353:
5349:
5347:
5346:
5342:
5340:
5339:
5335:
5333:
5332:
5328:
5326:
5325:
5321:
5319:
5318:
5314:
5312:
5311:
5307:
5306:
5304:
5302:
5298:
5292:
5290:
5286:
5284:
5282:
5278:
5276:
5275:
5271:
5269:
5268:
5264:
5262:
5261:
5257:
5254:
5253:
5249:
5246:
5245:
5244:Bahariasaurus
5241:
5239:
5238:
5234:
5233:
5230:
5227:
5224:
5220:
5216:
5186:
5185:
5181:
5179:
5178:
5174:
5172:
5171:
5167:
5165:
5164:
5160:
5159:
5157:
5155:
5151:
5145:
5144:
5140:
5139:
5136:
5133:
5131:
5127:
5121:
5120:
5116:
5114:
5113:
5109:
5107:
5106:
5102:
5100:
5099:
5095:
5093:
5092:
5088:
5086:
5085:
5084:Kelmayisaurus
5081:
5079:
5078:
5074:
5072:
5071:
5067:
5064:
5063:
5059:
5057:
5056:
5052:
5051:
5048:
5045:
5043:
5039:
5032:
5031:
5027:
5024:
5023:
5019:
5017:
5016:
5012:
5009:
5008:
5004:
5001:
5000:
4996:
4995:
4993:
4991:
4987:
4981:
4980:
4976:
4974:
4973:
4969:
4966:
4965:
4961:
4959:
4958:
4954:
4952:
4951:
4947:
4945:
4944:
4940:
4939:
4936:
4933:
4931:
4927:
4920:
4919:
4918:Saurophaganax
4915:
4912:
4911:
4907:
4904:
4903:
4899:
4897:
4896:
4892:
4891:
4889:
4887:
4883:
4880:
4877:
4872:
4858:
4857:
4853:
4851:
4850:
4849:Siamotyrannus
4846:
4844:
4843:
4839:
4837:
4836:
4832:
4830:
4829:
4825:
4824:
4822:
4820:
4816:
4810:
4809:
4805:
4802:
4801:
4797:
4796:
4793:
4790:
4788:
4784:
4777:
4776:
4772:
4769:
4768:
4764:
4762:
4761:
4757:
4755:
4754:
4750:
4749:
4746:
4742:
4739:
4736:
4735:Allosauroidea
4731:
4724:
4723:
4722:Spinosauridae
4719:
4716:
4715:
4711:
4708:
4707:
4703:
4701:
4700:
4696:
4695:
4692:
4689:
4686:
4681:
4674:
4673:
4669:
4666:
4665:
4661:
4660:
4657:
4654:
4651:
4646:
4642:
4638:
4633:
4629:
4619:
4614:
4612:
4605:
4575:
4574:
4573:
4572:
4570:
4568:
4564:
4560:
4538:
4537:
4533:
4531:
4530:
4526:
4525:
4523:
4521:
4517:
4511:
4510:
4506:
4504:
4503:
4499:
4497:
4496:
4492:
4490:
4489:
4485:
4482:
4481:
4477:
4476:
4473:
4470:
4468:
4467:Spinosaurinae
4464:
4454:
4453:
4449:
4447:
4446:
4445:Riparovenator
4442:
4440:
4439:
4438:Ceratosuchops
4435:
4434:
4432:
4430:
4426:
4419:
4418:
4414:
4411:
4410:
4406:
4404:
4403:
4402:Riojavenatrix
4399:
4397:
4396:
4392:
4390:
4389:
4385:
4384:
4381:
4378:
4376:
4375:Baryonychinae
4372:
4365:
4364:
4360:
4358:
4357:
4353:
4351:
4350:
4346:
4343:
4342:
4338:
4337:
4334:
4331:
4328:
4327:Spinosauridae
4323:
4308:
4307:
4303:
4301:
4300:
4296:
4294:
4293:
4289:
4287:
4286:
4282:
4280:
4279:
4275:
4273:
4272:
4268:
4267:
4265:
4263:
4259:
4253:
4252:
4251:Wiehenvenator
4248:
4246:
4245:
4241:
4239:
4238:
4234:
4232:
4231:
4227:
4226:
4224:
4222:
4218:
4212:
4211:
4207:
4206:
4203:
4200:
4198:
4194:
4187:
4186:
4182:
4181:
4178:
4174:
4171:
4168:
4163:
4152:
4151:
4147:
4145:
4144:
4140:
4138:
4137:
4133:
4131:
4130:
4126:
4125:
4123:
4121:
4117:
4111:
4110:
4106:
4105:
4102:
4098:
4095:
4092:
4087:
4084:
4081:
4076:
4070:
4069:
4065:
4063:
4062:
4058:
4055:
4054:
4050:
4048:
4047:
4043:
4041:
4040:
4036:
4034:
4033:
4029:
4027:
4026:
4022:
4020:
4019:
4015:
4012:
4011:
4007:
4005:
4004:
4000:
3997:
3996:
3992:
3989:
3988:
3987:Chienkosaurus
3984:
3981:
3980:
3976:
3975:
3972:
3969:
3966:
3961:
3957:
3953:
3948:
3944:
3934:
3927:
3920:
3913:
3887:
3886:
3885:
3884:
3882:
3880:
3876:
3872:
3846:
3845:
3841:
3839:
3838:
3834:
3832:
3831:
3827:
3825:
3824:
3820:
3817:
3816:
3812:
3810:
3809:
3805:
3803:
3802:
3798:
3796:
3795:
3791:
3789:
3788:
3784:
3783:
3781:
3779:
3778:Furileusauria
3775:
3769:
3768:
3764:
3762:
3761:
3757:
3754:
3753:
3752:Rahiolisaurus
3749:
3747:
3746:
3742:
3740:
3739:
3735:
3733:
3732:
3728:
3726:
3725:
3721:
3719:
3718:
3714:
3712:
3711:
3707:
3704:
3703:
3699:
3698:
3695:
3692:
3690:
3686:
3679:
3678:
3674:
3673:
3671:
3669:
3668:Carnotaurinae
3665:
3659:
3658:
3654:
3651:
3650:
3649:Rahiolisaurus
3646:
3644:
3643:
3642:Majungasaurus
3639:
3637:
3636:
3632:
3630:
3629:
3625:
3622:
3621:
3617:
3615:
3614:
3610:
3609:
3607:
3605:
3601:
3595:
3594:
3590:
3588:
3587:
3583:
3581:
3580:
3579:Tarascosaurus
3576:
3574:
3573:
3569:
3567:
3566:
3562:
3560:
3559:
3555:
3553:
3552:
3548:
3546:
3545:
3541:
3539:
3538:
3537:Eoabelisaurus
3534:
3531:
3530:
3526:
3524:
3523:
3522:Chenanisaurus
3519:
3518:
3515:
3512:
3509:
3508:Abelisauridae
3504:
3490:
3489:
3485:
3483:
3482:
3478:
3476:
3475:
3471:
3469:
3468:
3467:Masiakasaurus
3464:
3462:
3461:
3457:
3456:
3454:
3452:
3448:
3442:
3441:
3437:
3435:
3434:
3433:Huinculsaurus
3430:
3428:
3427:
3426:Elaphrosaurus
3423:
3422:
3420:
3418:
3414:
3408:
3407:
3403:
3401:
3400:
3396:
3394:
3393:
3389:
3386:
3385:
3381:
3378:
3377:
3373:
3370:
3369:
3365:
3362:
3361:
3357:
3355:
3354:
3350:
3347:
3346:
3345:Austrocheirus
3342:
3339:
3338:
3334:
3333:
3330:
3327:
3325:
3321:
3314:
3313:
3309:
3306:
3305:
3301:
3298:
3297:
3293:
3290:
3289:
3285:
3284:
3281:
3277:
3274:
3271:
3266:
3256:
3255:
3251:
3249:
3248:
3244:
3243:
3241:
3239:
3235:
3229:
3228:
3224:
3221:
3220:
3216:
3214:
3213:
3209:
3206:
3205:
3201:
3199:
3198:
3197:Berberosaurus
3194:
3193:
3190:
3186:
3183:
3180:
3175:
3168:
3167:
3163:
3161:
3160:
3156:
3155:
3152:
3149:
3146:
3141:
3137:
3133:
3128:
3124:
3115:
3109:
3086:
3085:
3084:Halticosaurus
3081:
3079:
3078:
3077:Teinurosaurus
3074:
3073:
3071:
3067:
3058:
3057:
3056:
3055:
3053:
3051:
3047:
3037:
3036:
3032:
3029:
3028:
3024:
3022:
3021:
3017:
3014:
3013:
3009:
3006:
3005:
3004:Megapnosaurus
3001:
2999:
2998:
2994:
2992:
2991:
2987:
2985:
2984:
2980:
2979:
2977:
2975:
2974:Coelophysidae
2971:
2965:
2964:
2963:Powellvenator
2960:
2958:
2957:
2953:
2951:
2950:
2946:
2944:
2943:
2939:
2936:
2935:
2931:
2930:
2927:
2924:
2922:
2918:
2912:
2911:
2907:
2904:
2903:
2899:
2897:
2896:
2892:
2890:
2889:
2885:
2883:
2882:
2878:
2876:
2875:
2871:
2869:
2868:
2864:
2862:
2861:
2857:
2855:
2854:
2853:Liliensternus
2850:
2848:
2847:
2843:
2841:
2840:
2836:
2834:
2833:
2829:
2826:
2825:
2824:Dolichosuchus
2821:
2819:
2818:
2817:Dilophosaurus
2814:
2813:
2810:
2807:
2804:
2799:
2792:
2791:
2790:Herrerasauria
2787:
2784:
2783:
2779:
2776:
2775:
2771:
2768:
2767:
2763:
2761:
2760:
2756:
2753:
2752:
2748:
2745:
2744:
2743:Daemonosaurus
2740:
2737:
2736:
2732:
2731:
2728:
2725:
2722:
2717:
2713:
2709:
2704:
2700:
2679:
2678:
2677:
2676:
2674:
2672:
2668:
2660:
2656:
2655:
2654:
2653:
2650:
2647:
2645:
2641:
2637:
2629:
2625:
2622:
2620:
2616:
2613:
2611:
2607:
2605:
2601:
2599:
2595:
2594:
2589:
2585:
2581:
2574:
2569:
2567:
2562:
2560:
2555:
2554:
2551:
2545:
2544:
2540:
2539:
2535:
2530:
2528:
2524:
2520:
2515:
2514:
2505:
2502:
2497:
2493:
2486:
2483:
2478:
2474:
2469:
2464:
2460:
2456:
2452:
2448:
2444:
2440:
2437:(8): 210127.
2436:
2432:
2428:
2421:
2418:
2413:
2409:
2404:
2399:
2394:
2389:
2385:
2381:
2377:
2373:
2372:
2367:
2363:
2359:
2355:
2354:Currie, P. J.
2351:
2347:
2343:
2337:
2334:
2328:
2323:
2318:
2313:
2309:
2305:
2301:
2294:
2291:
2285:
2282:
2277:
2273:
2269:
2265:
2261:
2257:
2250:
2247:
2243:
2237:
2234:
2230:
2224:
2221:
2215:
2212:
2206:
2203:
2196:
2194:
2192:
2190:
2186:
2181:
2177:
2172:
2167:
2163:
2159:
2155:
2151:
2149:
2140:
2138:
2136:
2134:
2132:
2130:
2126:
2122:
2118:
2112:
2109:
2105:
2099:
2097:
2093:
2086:
2083:
2079:
2073:
2070:
2064:
2061:
2056:
2050:
2046:
2042:
2038:
2036:
2027:
2024:
2021:Philadelphia.
2017:
2014:
2009:
2003:
1995:
1991:
1987:
1983:
1979:
1975:
1971:
1967:
1960:
1958:
1956:
1954:
1950:
1945:
1943:0-19-910207-4
1939:
1934:
1933:
1927:
1923:
1917:
1914:
1909:
1905:
1898:
1895:
1884:
1880:
1874:
1871:
1867:
1861:
1858:
1851:
1847:
1844:
1843:
1839:
1828:
1823:
1821:
1819:
1818:
1813:
1809:
1804:
1796:
1792:
1788:
1781:
1779:
1777:
1773:
1769:
1765:
1761:
1756:
1751:
1749:
1745:
1740:
1735:
1733:
1729:
1726:
1725:Maastrichtian
1722:
1718:
1714:
1710:
1708:
1703:
1695:
1690:
1683:
1681:
1679:
1675:
1671:
1667:
1663:
1659:
1649:
1648:
1640:
1639:
1631:
1630:
1622:
1621:
1613:
1612:
1604:
1603:
1595:
1594:
1586:
1585:
1577:
1576:
1568:
1567:
1564:
1563:
1557:
1556:
1553:
1552:
1544:
1543:
1540:
1539:
1538:
1531:
1530:
1527:
1526:
1523:
1522:
1521:
1514:
1513:
1507:
1506:
1500:
1499:
1496:
1495:
1492:
1491:
1490:
1483:
1482:
1476:
1475:
1472:
1471:
1463:
1462:
1459:
1458:
1457:
1450:
1449:
1446:
1445:
1442:
1441:
1440:
1433:
1432:
1426:
1425:
1419:
1418:
1415:
1414:
1411:
1410:
1409:
1402:
1401:
1395:
1394:
1391:
1390:
1387:
1386:
1385:
1378:
1377:
1371:
1370:
1367:
1366:
1363:
1362:
1361:
1354:
1353:
1347:
1346:
1343:
1342:
1339:
1338:
1337:
1330:
1329:
1323:
1322:
1319:
1318:
1315:
1314:
1313:
1306:
1305:
1299:
1298:
1295:
1294:
1286:
1285:
1277:
1276:
1268:
1267:
1264:
1263:
1262:
1255:
1254:
1251:
1250:
1247:
1246:
1245:
1238:
1237:
1231:
1230:
1227:
1226:
1223:
1222:
1221:
1214:
1213:
1207:
1206:
1203:
1202:
1194:
1193:
1190:
1189:
1188:
1181:
1180:
1177:
1176:
1173:
1172:
1171:
1164:
1163:
1160:
1159:
1156:
1155:
1154:
1147:
1146:
1140:
1139:
1133:
1130:
1129:
1123:
1120:
1119:
1115:
1113:
1104:
1096:
1092:
1090:
1086:
1085:
1080:
1079:
1074:
1070:
1066:
1062:
1058:
1054:
1050:
1049:
1045:
1041:
1037:
1036:
1035:Tyrannosaurus
1031:
1030:
1029:Albertosaurus
1025:
1021:
1017:
1012:
1010:
1006:
1002:
997:
996:Ken Carpenter
993:
989:
985:
981:
977:
976:
971:
962:
955:
953:
951:
946:
941:
937:
932:
930:
929:
928:Tyrannosaurus
924:
923:
922:Albertosaurus
918:
914:
910:
906:
902:
894:
890:
885:
881:
878:
874:
870:
866:
862:
858:
854:
850:
846:
842:
838:
834:
830:
825:
823:
819:
815:
811:
807:
806:
805:Tyrannosaurus
801:
797:
796:
791:
783:
776:
774:
773:
769:
765:
761:
759:
753:
749:
745:
741:
737:
736:
731:
727:
723:
719:
717:
713:
709:
705:
701:
697:
690:
685:
681:
680:
675:
674:
669:
664:
660:
658:
654:
650:
646:
642:
638:
634:
633:Othniel Marsh
630:
629:Dryptosaurus,
626:
622:
618:
613:
609:
607:
603:
599:
595:
591:
587:
583:
579:
575:
571:
567:
563:
559:
555:
551:
546:
542:
538:
537:Maastrichtian
534:
530:
526:
522:
518:
514:
505:
498:
496:
494:
491:
487:
483:
479:
475:
471:
467:
463:
458:
454:
451:was a large,
450:
446:
442:
441:Maastrichtian
438:
434:
431:
428:
424:
420:
419:
406:
361:
360:
346:
345:
343:
338:
334:
329:
328:
324:
323:
321:
318:
314:
309:
303:
301:
292:
289:
288:Binomial name
285:
281:
280:
274:
271:
270:
263:
262:
255:
252:
251:
246:
241:
235:
232:
231:
228:
222:
219:
216:
215:
212:
209:
206:
203:
202:
199:
196:
193:
190:
189:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
141:
136:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
36:67.6–66
32:
26:
22:
19:
6871:Tyrannosaurs
6767:Dryptosaurus
6766:
6730:
6659:
6652:
6645:
6638:
6631:
6626:Ornithomimus
6624:
6617:
6610:
6603:
6596:
6589:
6582:
6564:Tyrannomimus
6562:
6554:
6546:
6539:
6534:Deinocheirus
6532:
6525:
6505:
6498:
6491:
6467:
6459:
6454:Nqwebasaurus
6452:
6447:Nedcolbertia
6445:
6438:
6431:
6425:Calamosaurus
6423:
6417:Aviatyrannis
6415:
6408:
6383:
6353:
6346:
6339:
6331:
6324:
6317:
6310:
6302:
6297:Aristosuchus
6295:
6289:Aniksosaurus
6287:
6269:Aniksosaurus
6267:
6237:
6230:
6222:
6168:
6133:
6126:
6119:
6112:
6038:
6031:
6024:
6004:
5997:
5992:Dynamoterror
5990:
5970:
5963:
5943:
5936:
5916:
5911:Asiatyrannus
5909:
5885:
5878:
5849:Dryptosaurus
5848:
5847:
5840:
5833:
5802:
5795:
5788:
5780:
5773:
5766:
5759:
5752:
5745:
5740:Jinbeisaurus
5738:
5731:
5724:
5716:
5711:Aviatyrannis
5709:
5702:
5668:
5663:Sinotyrannus
5661:
5654:
5647:
5640:
5619:
5612:
5590:
5583:
5542:
5535:
5528:
5520:
5513:
5505:
5497:
5492:Bicentenaria
5490:
5482:
5477:Asiamericana
5475:
5468:
5409:
5350:
5343:
5336:
5329:
5322:
5315:
5308:
5287:
5279:
5272:
5265:
5258:
5252:Deltadromeus
5250:
5242:
5235:
5184:Tyrannotitan
5182:
5175:
5168:
5161:
5141:
5117:
5112:Taurovenator
5110:
5103:
5096:
5091:Lajasvenator
5089:
5082:
5075:
5070:Concavenator
5068:
5060:
5053:
5028:
5020:
5013:
5005:
4997:
4977:
4970:
4962:
4955:
4948:
4941:
4916:
4908:
4900:
4893:
4886:Allosauridae
4854:
4847:
4842:Shidaisaurus
4840:
4833:
4826:
4806:
4798:
4773:
4765:
4758:
4751:
4720:
4712:
4704:
4697:
4670:
4662:
4650:Avetheropoda
4637:Avetheropoda
4608:
4567:Avetheropoda
4534:
4527:
4520:Spinosaurini
4507:
4500:
4493:
4486:
4478:
4450:
4443:
4436:
4415:
4407:
4400:
4395:Protathlitis
4393:
4386:
4361:
4354:
4347:
4339:
4304:
4297:
4290:
4285:Leshansaurus
4283:
4276:
4269:
4249:
4242:
4237:Megalosaurus
4235:
4230:Duriavenator
4228:
4208:
4183:
4167:Megalosauria
4148:
4141:
4136:Marshosaurus
4134:
4129:Condorraptor
4127:
4107:
4066:
4059:
4051:
4044:
4037:
4030:
4023:
4016:
4010:Cruxicheiros
4008:
4001:
3993:
3985:
3977:
3842:
3837:Quilmesaurus
3835:
3828:
3821:
3813:
3806:
3799:
3792:
3785:
3765:
3758:
3750:
3743:
3736:
3729:
3722:
3715:
3708:
3700:
3689:Brachyrostra
3675:
3655:
3647:
3640:
3633:
3626:
3618:
3611:
3591:
3586:Tralkasaurus
3584:
3577:
3570:
3563:
3556:
3549:
3542:
3535:
3527:
3520:
3488:Vespersaurus
3486:
3481:Velocisaurus
3479:
3472:
3465:
3458:
3438:
3431:
3424:
3404:
3397:
3390:
3382:
3376:Deltadromeus
3374:
3366:
3360:Compsosuchus
3358:
3351:
3343:
3335:
3310:
3302:
3296:Lametasaurus
3294:
3286:
3252:
3247:Ceratosaurus
3245:
3225:
3217:
3210:
3202:
3195:
3179:Ceratosauria
3164:
3157:
3082:
3075:
3033:
3025:
3018:
3010:
3002:
2995:
2988:
2981:
2961:
2956:Podokesaurus
2954:
2947:
2940:
2932:
2908:
2900:
2893:
2886:
2879:
2872:
2865:
2858:
2851:
2844:
2839:Gojirasaurus
2837:
2832:Dracovenator
2830:
2822:
2815:
2803:Neotheropoda
2788:
2780:
2772:
2766:Guaibasaurus
2764:
2757:
2749:
2741:
2735:Chindesaurus
2733:
2623:
2614:
2542:
2522:
2518:
2504:
2495:
2491:
2485:
2434:
2430:
2420:
2375:
2369:
2342:Loewen, M.A.
2336:
2307:
2303:
2293:
2284:
2259:
2255:
2249:
2241:
2236:
2228:
2223:
2214:
2205:
2153:
2147:
2116:
2111:
2103:
2085:
2077:
2072:
2063:
2040:
2034:
2026:
2016:
2002:cite journal
1994:the original
1973:
1969:
1931:
1926:Robert Scott
1916:
1907:
1903:
1897:
1886:. Retrieved
1882:
1873:
1865:
1860:
1817:Megalosaurus
1816:
1802:
1800:
1790:
1768:Dryptosaurus
1767:
1764:Hadrosaurids
1760:Dryptosaurus
1759:
1755:Dryptosaurus
1754:
1752:
1748:Dryptosaurus
1747:
1736:
1705:
1702:Dryptosaurus
1701:
1699:
1694:Dryptosaurus
1693:
1684:Paleoecology
1678:Dryptosaurus
1677:
1658:Dryptosaurus
1657:
1655:
1560:
1535:
1534:
1518:
1517:
1487:
1486:
1454:
1453:
1437:
1436:
1407:
1406:
1405:
1382:
1381:
1358:
1357:
1334:
1333:
1310:
1309:
1259:
1258:
1242:
1241:
1218:
1217:
1185:
1184:
1168:
1167:
1151:
1150:
1111:
1109:
1082:
1076:
1073:Dryptosaurus
1072:
1068:
1061:Dryptosaurus
1060:
1057:Dryptosaurus
1056:
1052:
1046:
1040:Dryptosaurus
1039:
1033:
1027:
1020:Dryptosaurus
1019:
1013:
1005:Dryptosaurus
1004:
1001:phylogenetic
992:Dryptosaurus
991:
984:megalosaurid
975:Megalosaurus
973:
970:Dryptosaurus
969:
967:
940:Dryptosaurus
939:
935:
933:
926:
920:
913:Dryptosaurus
912:
901:Dryptosaurus
900:
898:
893:Dryptosaurus
892:
887:An outdated
826:
822:Dryptosaurus
821:
814:Dryptosaurus
813:
810:Dryptosaurus
809:
803:
800:Dryptosaurus
799:
793:
790:Dryptosaurus
789:
788:
771:
763:
758:Teihivenator
755:
751:
740:Dryptosaurus
739:
733:
730:Joseph Leidy
721:
720:
711:
707:
703:
695:
694:
677:
673:Elasmosaurus
671:
668:Dryptosaurus
667:
656:
652:
640:
628:
620:
616:
610:An isolated
604:
580:, a partial
554:Philadelphia
510:
486:Dryptosaurus
485:
466:Dryptosaurus
465:
449:Dryptosaurus
448:
359:Dryptosaurus
358:
357:
356:
348:(Cope, 1866)
342:Megalosaurus
340:
339:
331:Cope, 1866 (
326:
325:
299:
294:
278:
277:
261:Dryptosaurus
260:
259:
239:
217:
204:
191:
178:
25:Dryptosaurus
24:
18:
6691:Maniraptora
6680:Maniraptora
6661:Tototlmimus
6591:Anserimimus
6541:Garudimimus
6527:Beishanlong
6469:Valdoraptor
6410:Arkansaurus
6355:Xunmenglong
6232:Juravenator
6026:Tarbosaurus
5918:Nanuqsaurus
5887:Gorgosaurus
5804:Megaraptora
5790:Timurlengia
5522:Sciurumimus
5470:Aratasaurus
5338:Murusraptor
5260:Fukuiraptor
5223:Megaraptora
5105:Shaochilong
5077:Eocarcharia
5030:Megaraptora
4950:Lusovenator
4828:Alpkarakush
4685:Carnosauria
4536:Spinosaurus
4509:Siamosaurus
4409:Suchosaurus
4356:Iberospinus
4292:Magnosaurus
4271:Afrovenator
4244:Torvosaurus
3995:Chilesaurus
3808:Carnotaurus
3801:Caletodraco
3787:Abelisaurus
3613:Arcovenator
3460:Laevisuchus
3451:Noasaurinae
3353:Berthasaura
3324:Noasauridae
3254:Genyodectes
3219:Lukousaurus
3212:Jubbulpuria
3012:Panguraptor
2990:Coelophysis
2983:Camposaurus
2942:Panguraptor
2934:Dracoraptor
2910:Zupaysaurus
2895:Tachiraptor
2874:Sarcosaurus
2774:Nhandumirim
2346:Irmis, R.B.
1009:coelurosaur
871:, the left
867:, the left
863:, the left
859:, the left
777:Description
735:Coelosaurus
679:Hadrosaurus
676:, with two
602:protostegid
564:fragments,
443:age of the
344:aquilunguis
333:preoccupied
266:Marsh, 1877
6865:Categories
6619:Gallimimus
6548:Harpymimus
6493:Harpymimus
6114:Iliosuchus
6084:see below↓
5929:Alioramini
5747:Juratyrant
5733:Eotyrannus
5718:Bagaraatan
5670:Yutyrannus
5604:Coeluridae
5530:Vayuraptor
5376:see below↓
5352:Tratayenia
5345:Orkoraptor
5331:Megaraptor
5237:Aoniraptor
5170:Mapusaurus
5098:Sauroniops
5062:Altispinax
5015:Neovenator
4957:Siamraptor
4943:Datanglong
4910:Epanterias
4902:Antrodemus
4895:Allosaurus
4876:Allosauria
4699:Altispinax
4664:Gasosaurus
4576:see below↓
4452:Suchomimus
4053:Sinosaurus
3888:see below↓
3844:Viavenator
3794:Aucasaurus
3731:Ilokelesia
3702:Dahalokely
3657:Rajasaurus
3635:Indosaurus
3628:Genusaurus
3620:Dahalokely
3544:Indosuchus
3440:Limusaurus
3399:Ligabueino
3392:Kiyacursor
3384:Genusaurus
3368:Dahalokely
3288:Betasuchus
3159:Dornraptor
3059:see below↓
3035:Segisaurus
2888:Sinosaurus
2751:Eodromaeus
2680:see below↓
2628:Dinosauria
2610:Sauropsida
2242:Brimleyana
2231:106: 1–19.
1910:: 275–279.
1888:2024-08-08
1852:References
1772:Appalachia
1721:New Jersey
905:incisiform
877:metatarsal
873:astragalus
837:surangular
795:Eotyrannus
752:Dinosauria
712:Allosaurus
708:Allosaurus
612:metatarsal
533:New Jersey
437:Appalachia
198:Saurischia
185:Dinosauria
6695:includes
6640:Rativates
6633:Qiupalong
6333:Scipionyx
6326:Mirischia
5999:Lythronax
5938:Alioramus
5592:Labocania
5310:Aerosteon
4856:Sinraptor
4760:Erectopus
4495:Irritator
4080:Orionides
3965:Tetanurae
3952:Tetanurae
3879:Tetanurae
3815:Elemgasem
3738:Llukalkan
3717:Elemgasem
3474:Noasaurus
3337:Afromimus
3145:Averostra
3132:Averostra
3050:Averostra
2902:Velocipes
2721:Theropoda
2708:Theropoda
2671:Theropoda
2596:Kingdom:
2580:Theropoda
2525:(3): 41A.
2459:2054-5703
2310:: 47–54.
2180:0003-0082
2171:2246/6117
2123:, p. 100.
1868:8: 72–73.
1776:Nodosaurs
1051:in 2005.
1014:In 1946,
988:monotypic
950:calcaneum
909:denticles
849:phalanges
841:vertebrae
637:Bone Wars
529:Barnsboro
493:dinosaurs
484:in 1877,
470:dinosaurs
457:carnivore
425:of basal
272:Species:
211:Theropoda
158:Kingdom:
152:Eukaryota
6796:52570689
6776:Wikidata
6224:Coelurus
5649:Kileskus
5642:Guanlong
5614:Coelurus
5507:Gualicho
5007:Gualicho
4388:Baryonyx
3724:Guemesia
3551:Kryptops
3312:Ozraptor
2949:Pendraig
2604:Chordata
2602:Phylum:
2598:Animalia
2477:34457333
2412:24223179
2371:PLoS ONE
2360:(2013).
2276:86243316
2244:2: 1–18.
1928:(1980).
1824:See also
1696:in brown
1670:Delaware
1114:(2013).
1078:Guanlong
938:(2011),
596:, and a
590:chevrons
586:hindlimb
490:theropod
447:period.
433:dinosaur
430:theropod
317:Synonyms
310:, 1866 )
233:Family:
172:Chordata
168:Phylum:
162:Animalia
148:Domain:
6822:4946657
6809:3238991
6782:Q131634
5782:Timimus
5544:Zuolong
5274:Rapator
5177:Meraxes
4502:Oxalaia
3823:Koleken
2846:Lepidus
2608:Class:
2498:: 1–58.
2468:8385347
2439:Bibcode
2403:3819173
2380:Bibcode
2364:(ed.).
2200:112-113
1978:Bibcode
1730:of the
945:ischial
915:had an
857:ischium
833:dentary
829:maxilla
657:Laelaps
621:Laelaps
598:scapula
594:sternum
562:dentary
521:Montana
515:during
453:bipedal
421:) is a
297:†
276:†
253:Genus:
105:↓
6433:Hexing
6121:Kakuru
5726:Dilong
3767:Thanos
3745:Niebla
3565:Rugops
3558:Kurupi
2475:
2465:
2457:
2410:
2400:
2274:
2178:
2051:
1940:
1924:&
1797:, 1897
1112:et al.
1084:Dilong
1069:et al.
1032:, and
1024:coeval
1007:was a
936:et al.
869:fibula
845:humeri
686:, 1869
653:Lælaps
641:et al.
582:pelvis
574:humeri
539:-aged
474:fossil
247:, 1890
6843:Plazi
6835:38605
6697:birds
5754:Moros
5281:Siats
5022:Siats
4964:Siats
2624:Clade
2615:Clade
2272:S2CID
2045:10–18
1728:stage
1022:with
865:tibia
861:femur
853:pubic
728:that
578:manus
566:teeth
558:skull
464:made
423:genus
414:-toh-
245:Marsh
218:Clade
205:Clade
192:Clade
179:Clade
6804:GBIF
6689:see
5324:Maip
2782:Tawa
2657:see
2473:PMID
2455:ISSN
2408:PMID
2176:ISSN
2049:ISBN
2008:link
1938:ISBN
1812:AMNH
1709:9995
1707:ANSP
1081:and
925:and
592:, a
560:and
412:DRIP
308:Cope
44:PreꞒ
6791:EoL
2463:PMC
2447:doi
2398:PMC
2388:doi
2322:hdl
2312:doi
2264:doi
2166:hdl
2158:doi
1986:doi
1793:by
1704:is
1668:of
891:of
655:or
619:. "
552:in
418:-əs
416:SOR
396:ɔːr
6867::
6845::
6832::
6819::
6806::
6793::
6778::
2626::
2617::
2523:22
2521:.
2496:34
2494:.
2471:.
2461:.
2453:.
2445:.
2433:.
2429:.
2406:.
2396:.
2386:.
2374:.
2368:.
2356:;
2352:;
2348:;
2344:;
2320:.
2308:58
2306:.
2302:.
2270:.
2260:25
2258:.
2188:^
2174:.
2164:.
2152:.
2128:^
2119:,
2095:^
2047:.
2004:}}
2000:{{
1984:.
1974:17
1972:.
1968:.
1952:^
1908:18
1906:.
1881:.
1719:,
1680:.
1091:.
798:,
718:.
572:,
568:,
531:,
387:oʊ
220::
207::
194::
181::
129:.
94:Pg
38:Ma
33:,
6559:?
6551:?
6464:?
6428:?
6420:?
6388:?
6336:?
6307:?
6292:?
6227:?
5807:?
5785:?
5721:?
5624:?
5606:?
5595:?
5525:?
5510:?
5502:?
5487:?
5291:?
5283:?
5255:?
5247:?
5225:?
5065:?
5033:?
5025:?
5010:?
5002:?
4967:?
4921:?
4913:?
4905:?
4803:?
4778:?
4770:?
4725:?
4717:?
4709:?
4675:?
4667:?
4483:?
4420:?
4412:?
4366:?
4344:?
4309:?
4188:?
4153:?
4056:?
4013:?
3998:?
3990:?
3982:?
3818:?
3755:?
3705:?
3680:?
3652:?
3623:?
3532:?
3387:?
3379:?
3371:?
3363:?
3348:?
3340:?
3315:?
3307:?
3299:?
3291:?
3222:?
3207:?
3169:?
3030:?
3015:?
3007:?
2937:?
2905:?
2827:?
2793:?
2785:?
2777:?
2769:?
2754:?
2746:?
2738:?
2572:e
2565:t
2558:v
2479:.
2449::
2441::
2435:8
2414:.
2390::
2382::
2376:8
2330:.
2324::
2314::
2278:.
2266::
2182:.
2168::
2160::
2057:.
2010:)
1988::
1980::
1946:.
1891:.
760:"
756:"
608:.
405:/
402:s
399:ə
393:s
390:ˈ
384:t
381:p
378:ɪ
375:r
372:d
369:ˌ
366:/
362:(
335:)
306:(
257:†
237:†
224:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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